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<mods:title id="76CF8365418AE5D2C1C5D6FAE2BAA7AA">A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini)</mods:title>
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<heading id="AC6381EF4A5FFFE0FF5AFA3A6DB1919B" bold="true" box="[151,964,1466,1493]" fontSize="11" level="1" pageId="70" pageNumber="71" reason="1">
<emphasis id="C5E0EA914A5FFFE0FF5AFA3A6DB1919B" bold="true" box="[151,964,1466,1493]" pageId="70" pageNumber="71">
<taxonomicName id="30944D004A5FFFE0FF5AFA3A6D78919B" authority="Gao &amp; Ren, 2009" authorityName="Gao &amp; Ren" authorityYear="2009" box="[151,781,1466,1493]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FF5AFA3A6C5B919A" bold="true" box="[151,558,1466,1493]" italics="true" pageId="70" pageNumber="71">Morphostenophanes tuberculatus</emphasis>
<bibRefCitation id="93054B724A5FFFE0FDF8FA3A6D78919B" author="Gao, Z. H. &amp; Ren, G. D." box="[565,781,1466,1493]" pageId="70" pageNumber="71" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren, 2009</bibRefCitation>
</taxonomicName>
ṬǾẎȐAEƤ
</emphasis>
</heading>
</paragraph>
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<subSubSection id="BF8E65084A5FFFE0FF5AFA5E6F5D91B6" box="[151,296,1502,1529]" pageId="70" pageNumber="71" type="description">
<paragraph id="F72B36834A5FFFE0FF5AFA5E6F5D91B6" blockId="70.[151,964,1466,1529]" box="[151,296,1502,1529]" pageId="70" pageNumber="71">
(
<figureCitation id="6FAF2A064A5FFFE0FF6DFA5E6F5591B6" box="[160,288,1502,1529]" captionStart="FIGURE 46" captionStartId="71.[151,250,819,844]" captionTargetBox="[291,1298,181,794]" captionTargetId="figure@71.[290,1298,181,794]" captionTargetPageId="71" captionText="FIGURE 46. Habitus of Morphostenophanes species in the jendeki-group: M. tanikadoi Masumoto, male (A, C), and female (B, D); in dorsal (A, B), and lateral view (C, D). Holotype male of M. tuberculatus in dorsal view (E) with labels (F), not to scale. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3797267" httpUri="https://zenodo.org/record/3797267/files/figure.png" pageId="70" pageNumber="71">Fig. 46E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A5FFFE0FF5AF99C6F41921C" pageId="70" pageNumber="71" type="reference_group">
<paragraph id="F72B36834A5FFFE0FF5AF99C6F41921C" blockId="70.[151,1437,1563,1619]" pageId="70" pageNumber="71">
<treatmentCitationGroup id="D78411AD4A5FFFE0FF5AF99C6F41921C" pageId="70" pageNumber="71">
<treatmentCitation id="763510924A5FFFE0FF5AF99C6C95927B" author="Gao, Z. H. &amp; Ren, G. D." box="[151,736,1563,1588]" page="315" pageId="70" pageNumber="71" year="2009">
<taxonomicName id="30944D004A5FFFE0FF5AF99C6C95927B" authority="Gao &amp; Ren, 2009: 315" authorityName="Gao &amp; Ren" authorityPageNumber="315" authorityYear="2009" box="[151,736,1563,1588]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FF5AF99C6F99927B" box="[151,492,1564,1588]" italics="true" pageId="70" pageNumber="71">Morphostenophanes tuberculatus</emphasis>
<materialsCitation id="47FC3CDE4A5FFFE0FE3EF99C6C95927B" ID-GBIF-Occurrence="2610567422" box="[499,736,1563,1588]" country="China" county="Lushui County" location="Yakou" municipality="Pianma" pageId="70" pageNumber="71" specimenCount="1" stateProvince="Yunnan">
<bibRefCitation id="93054B724A5FFFE0FE3EF99C6C95927B" author="Gao, Z. H. &amp; Ren, G. D." box="[499,736,1563,1588]" pageId="70" pageNumber="71" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren, 2009: 315</bibRefCitation>
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(
<typeStatus id="282F88214A5FFFE0FD23F99C6D6E927B" box="[750,795,1564,1588]" pageId="70" pageNumber="71">type</typeStatus>
locality:
<location id="F24B60584A5FFFE0FCB0F99B6DB4927B" LSID="urn:lsid:plazi:treatment:7F3D87954A5FFFE0FF5AFA3A6CF893B2:F24B60584A5FFFE0FCB0F99B6DB4927B" box="[893,961,1563,1588]" country="China" county="Lushui County" municipality="Pianma" name="Yakou" pageId="70" pageNumber="71" stateProvince="Yunnan">Yakou</location>
,
<collectingMunicipality id="174FACF94A5FFFE0FC01F99B6A6E927B" box="[972,1051,1563,1588]" pageId="70" pageNumber="71">Pianma</collectingMunicipality>
,
<collectingCounty id="1E4A4E0F4A5FFFE0FBEAF99B6AB4927B" box="[1063,1217,1563,1588]" pageId="70" pageNumber="71">Lushui County</collectingCounty>
,
<collectingRegion id="3550F8614A5FFFE0FB06F99C6B6E927C" box="[1227,1307,1564,1587]" country="China" name="Yunnan" pageId="70" pageNumber="71">Yunnan</collectingRegion>
,
<collectingCountry id="8F8376134A5FFFE0FAEAF99B6B12927B" box="[1319,1383,1563,1588]" name="China" pageId="70" pageNumber="71">China</collectingCountry>
), fig. 1214, 40.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="BF8E65084A5FFFE0FF5AF9FB6C74936B" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="F72B36834A5FFFE0FF5AF9FB6C74936B" blockId="70.[151,1436,1658,2045]" pageId="70" pageNumber="71">
<emphasis id="C5E0EA914A5FFFE0FF5AF9FB6FC992DA" bold="true" box="[151,444,1658,1685]" pageId="70" pageNumber="71">Type material examined.</emphasis>
<materialsCitation id="47FC3CDE4A5FFFE0FE0CF9FB6CED9292" ID-GBIF-Occurrence="2610567392" collectingDate="2005-05-19" county="Lushui Pianma Yakou Xiao-Hong Ou" location="University" municipality="Pianma Yakou Xiao-Hong Ou Museum" pageId="70" pageNumber="71" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="C5E0EA914A5FFFE0FE0CF9FB6C5B92DA" bold="true" box="[449,558,1659,1685]" pageId="70" pageNumber="71">
<typeStatus id="282F88214A5FFFE0FE0CF9FB6C5B92DA" box="[449,558,1659,1685]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
</emphasis>
of
<taxonomicName id="30944D004A5FFFE0FD9CF9FA6DB392DB" authorityName="Gao &amp; Ren" authorityYear="2009" box="[593,966,1658,1684]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FD9CF9FA6DB392DB" box="[593,966,1658,1684]" italics="true" pageId="70" pageNumber="71">Morphostenophanes tuberculatus</emphasis>
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labelled (translated from Chinese):
<date id="832A10434A5FFFE0FA9BF9FA6EBF92F6" pageId="70" pageNumber="71" value="2005-05-19">2005- 5·19</date>
Lushui Pianma Yakou [handwritten in white label] //
<date id="832A10434A5FFFE0FCEDF91E6DED92F6" box="[800,920,1694,1721]" pageId="70" pageNumber="71" value="2005-05-19">
<collectingDate id="936EE9AB4A5FFFE0FCEDF91E6DED92F6" box="[800,920,1694,1721]" pageId="70" pageNumber="71" value="2005-05-19">2005-V-19</collectingDate>
</date>
<collectingRegion id="3550F8614A5FFFE0FC50F91F6D8692F6" box="[925,1011,1695,1721]" country="China" name="Yunnan" pageId="70" pageNumber="71">Yunnan</collectingRegion>
<collectingCounty id="1E4A4E0F4A5FFFE0FC34F91E6BE992F6" box="[1017,1436,1694,1721]" pageId="70" pageNumber="71">
Lushui
<collectingMunicipality id="174FACF94A5FFFE0FB83F91E6BE992F6" box="[1102,1436,1694,1721]" pageId="70" pageNumber="71">Pianma Yakou Xiao-Hong Ou</collectingMunicipality>
</collectingCounty>
Museum of
<collectingRegion id="3550F8614A5FFFE0FED0F9426F159293" box="[285,352,1730,1756]" country="China" name="Hebei" pageId="70" pageNumber="71">Hebei</collectingRegion>
<location id="F24B60584A5FFFE0FEABF9426FA99293" LSID="urn:lsid:plazi:treatment:7F3D87954A5FFFE0FF5AFA3A6CF893B2:F24B60584A5FFFE0FEABF9426FA99293" box="[358,476,1730,1756]" county="Lushui Pianma Yakou Xiao-Hong Ou" municipality="Pianma Yakou Xiao-Hong Ou Museum" name="University" pageId="70" pageNumber="71" stateProvince="Yunnan">University</location>
[printed label] //
</materialsCitation>
<typeStatus id="282F88214A5FFFE0FD53F9426D739293" box="[670,774,1730,1756]" pageId="70" pageNumber="71" type="holotype">Holotype</typeStatus>
[red label] //
<emphasis id="C5E0EA914A5FFFE0FC51F9426A039293" box="[924,1142,1730,1756]" italics="true" pageId="70" pageNumber="71">Morphostenophane</emphasis>
s
<taxonomicName id="30944D004A5FFFE0FB4AF9426EA6934E" authority="Gao et Ren, 2009" authorityName="Gao et Ren" authorityYear="2009" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FB4AF9426B679293" box="[1159,1298,1730,1756]" italics="true" pageId="70" pageNumber="71">tuberculatus</emphasis>
<bibRefCitation id="93054B724A5FFFE0FAD4F9436EA6934E" author="Gao, Z. H. &amp; Ren, G. D." pageId="70" pageNumber="71" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">
Gao
<emphasis id="C5E0EA914A5FFFE0FA83F9436B179293" box="[1358,1378,1731,1756]" italics="true" pageId="70" pageNumber="71">et</emphasis>
Ren, 2009
</bibRefCitation>
</taxonomicName>
:
<emphasis id="C5E0EA914A5FFFE0FF12F9676F38934F" box="[223,333,1766,1793]" italics="true" pageId="70" pageNumber="71">Acta Zool</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FE91F9676FEF934F" box="[348,410,1767,1792]" italics="true" pageId="70" pageNumber="71">Hung</emphasis>
., 55(4): 315-318, figs. 31-38, 40 (male, MHBU,
<figureCitation id="6FAF2A064A5FFFE0FC02F9666A40934E" box="[975,1077,1766,1793]" captionStart="FIGURE 46" captionStartId="71.[151,250,819,844]" captionTargetBox="[291,1298,181,794]" captionTargetId="figure@71.[290,1298,181,794]" captionTargetPageId="71" captionText="FIGURE 46. Habitus of Morphostenophanes species in the jendeki-group: M. tanikadoi Masumoto, male (A, C), and female (B, D); in dorsal (A, B), and lateral view (C, D). Holotype male of M. tuberculatus in dorsal view (E) with labels (F), not to scale. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.3797267" httpUri="https://zenodo.org/record/3797267/files/figure.png" pageId="70" pageNumber="71">Fig. 46E</figureCitation>
). Examined through two photographs taken by Xing-Long Bai.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A5FFFE0FF0AF8AE6B029397" pageId="70" pageNumber="71" type="discussion">
<paragraph id="F72B36834A5FFFE0FF0AF8AE6B029397" blockId="70.[151,1436,1658,2045]" pageId="70" pageNumber="71">
<emphasis id="C5E0EA914A5FFFE0FF0AF8AE6F3A9306" bold="true" box="[199,335,1838,1865]" pageId="70" pageNumber="71">Comments.</emphasis>
<taxonomicName id="30944D004A5FFFE0FE95F8AE6CA59307" authorityName="Gao et Ren" authorityYear="2009" box="[344,720,1838,1864]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FE95F8AE6CA59307" box="[344,720,1838,1864]" italics="true" pageId="70" pageNumber="71">Morphostenophanes tuberculatus</emphasis>
</taxonomicName>
has the elytra with densely scattered unevenly sized tubercles, which resemble those of
<taxonomicName id="30944D004A5FFFE0FE66F8D36C0B9323" authority="chongli" box="[427,638,1874,1901]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="chongli" subSpecies="chongli">
<emphasis id="C5E0EA914A5FFFE0FE66F8D36FB79323" box="[427,450,1875,1900]" italics="true" pageId="70" pageNumber="71">M</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FE1DF8D26C0B9323" box="[464,638,1874,1900]" italics="true" pageId="70" pageNumber="71">chongli chongli</emphasis>
</taxonomicName>
and
<taxonomicName id="30944D004A5FFFE0FD7FF8D36D0F9323" box="[690,890,1874,1901]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="chongli" subSpecies="glaber">
<emphasis id="C5E0EA914A5FFFE0FD7FF8D36CBC9323" box="[690,713,1875,1900]" italics="true" pageId="70" pageNumber="71">M</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FD1AF8D26D0F9323" box="[727,890,1874,1900]" italics="true" pageId="70" pageNumber="71">chongli glaber</emphasis>
</taxonomicName>
. However, these two species are very different in many ways (see comparative notes of
<taxonomicName id="30944D004A5FFFE0FD88F8F76CCA93DF" box="[581,703,1910,1937]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="chongli">
<emphasis id="C5E0EA914A5FFFE0FD88F8F76C2993DF" box="[581,604,1911,1936]" italics="true" pageId="70" pageNumber="71">M</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FDA6F8F66CCA93DF" box="[619,703,1910,1936]" italics="true" pageId="70" pageNumber="71">chongli</emphasis>
</taxonomicName>
) so that they clearly not belong to the same species group. Small body (body length
<quantity id="306C9B664A5FFFE0FEA7F81A6FBB93FA" box="[362,462,1946,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="70" pageNumber="71" unit="mm" value="15.5">15.5 mm</quantity>
as mentioned in original publication), carinate apical margin of parameres (
<bibRefCitation id="93054B724A5FFFE0FAD1F81B6EA79396" author="Gao, Z. H. &amp; Ren, G. D." pageId="70" pageNumber="71" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren 2009</bibRefCitation>
,
<figureCitation id="6FAF2A064A5FFFE0FF10F83E6F5B9396" box="[221,302,1982,2009]" captionStart="FIGURE 37" captionStartId="59.[151,250,1970,1995]" captionTargetBox="[206,1380,999,1943]" captionTargetId="figure@59.[201,1386,993,1945]" captionTargetPageId="59" captionText="FIGURE 37. Habitus of Morphostenophanes furvus Zhou new species. Male (A, C), and female (B, D); in dorsal (A, B), and lateral view (C, D). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3797229" httpUri="https://zenodo.org/record/3797229/files/figure.png" pageId="70" pageNumber="71">Fig. 37</figureCitation>
) in
<taxonomicName id="30944D004A5FFFE0FE97F83F6C7F9397" authorityName="Gao et Ren" authorityYear="2009" box="[346,522,1982,2009]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FE97F83F6F049397" box="[346,369,1983,2008]" italics="true" pageId="70" pageNumber="71">M</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FEB2F83E6C7F9397" box="[383,522,1982,2008]" italics="true" pageId="70" pageNumber="71">tuberculatus</emphasis>
</taxonomicName>
suggest potential relation between
<taxonomicName id="30944D004A5FFFE0FC5BF83F6A339397" authorityName="Gao et Ren" authorityYear="2009" box="[918,1094,1982,2009]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="C5E0EA914A5FFFE0FC5BF83F6DD89397" box="[918,941,1983,2008]" italics="true" pageId="70" pageNumber="71">M</emphasis>
.
<emphasis id="C5E0EA914A5FFFE0FC76F83E6A339397" box="[955,1094,1982,2008]" italics="true" pageId="70" pageNumber="71">tuberculatus</emphasis>
</taxonomicName>
and
<taxonomicName id="30944D004A5FFFE0FBB1F83E6AB89397" box="[1148,1229,1982,2008]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="jendeki">
<emphasis id="C5E0EA914A5FFFE0FBB1F83E6AB89397" box="[1148,1229,1982,2008]" italics="true" pageId="70" pageNumber="71">jendeki</emphasis>
</taxonomicName>
-group species.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A5FFFE0FF0AF8626CF893B2" box="[199,653,2018,2045]" pageId="70" pageNumber="71" type="distribution">
<paragraph id="F72B36834A5FFFE0FF0AF8626CF893B2" blockId="70.[151,1436,1658,2045]" box="[199,653,2018,2045]" pageId="70" pageNumber="71">
<emphasis id="C5E0EA914A5FFFE0FF0AF8626F1493B2" bold="true" box="[199,353,2018,2045]" pageId="70" pageNumber="71">Distribution.</emphasis>
(
<figureCitation id="6FAF2A064A5FFFE0FEA2F8636FC293B2" box="[367,439,2019,2045]" captionStart="Map 1" captionStartId="8.[151,202,1286,1310]" captionTargetBox="[151,1436,705,1261]" captionTargetId="figure@8.[151,1436,705,1261]" captionText="Map 1. Distribution of the Morphostenophanes species. A. Enlargement of Central and Western Yunnan Province." figureDoi="http://doi.org/10.5281/zenodo.3797090" httpUri="https://zenodo.org/record/3797090/files/figure.png" pageId="70" pageNumber="71">Map 1</figureCitation>
)
<collectingCountry id="8F8376134A5FFFE0FE04F8636C5393B2" box="[457,550,2019,2045]" name="China" pageId="70" pageNumber="71">CHINA</collectingCountry>
:
<collectingRegion id="3550F8614A5FFFE0FDFDF8636CFC93B2" box="[560,649,2019,2045]" country="China" name="Yunnan" pageId="70" pageNumber="71">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>