treatments-xml/data/7F/3D/87/7F3D87954A30FF8DFF5AFF196CF896BC.xml

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<mods:title id="490762A68AE8E691B710C2A91F647886">A revision of the genus Morphostenophanes Pic, 1925 (Coleoptera, Tenebrionidae, Stenochiinae, Cnodalonini)</mods:title>
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<mods:namePart id="12FBB877F74DC32C56CAE5F193410F7D">Zhou, De-Yao</mods:namePart>
<mods:affiliation id="4D53954D317B1C5C7528870ADA47D2FB">Room 901, No. 126, Lane 1331, Chenggu Road, Shanghai, 201800 China. E-mail: scydmaeninae @ 163. com</mods:affiliation>
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<taxonomicName id="30944D004A30FF8FFF5AFF196CD094FB" authority="(Kaszab, 1960)" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[151,677,153,180]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFF5AFF196F9194FB" bold="true" box="[151,484,153,180]" italics="true" pageId="41" pageNumber="42">Morphostenophanes atavus</emphasis>
(
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)
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ÑẎȐAEƤ
</emphasis>
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<paragraph id="F72B36834A30FF8FFF5AFF3E6CC79497" blockId="41.[151,832,153,216]" box="[151,690,189,216]" pageId="41" pageNumber="42">
(
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A–G; 31A, E, K, P; 32A, G–I; 33A, G)
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A30FF8FFF5AFE826C86953A" pageId="41" pageNumber="42" type="reference_group">
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<treatmentCitation id="763510924A30FF8FFF5AFE826CE19555" author="Kaszab, Z." box="[151,660,257,282]" page="278" pageId="41" pageNumber="42" year="1960">
<taxonomicName id="30944D004A30FF8FFF5AFE826CE19555" authority="Kaszab, 1960: 278" authorityName="Kaszab" authorityPageNumber="278" authorityYear="1960" box="[151,660,257,282]" class="Insecta" family="Tenebrionidae" genus="Promorphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFF5AFE826FA49555" box="[151,465,258,282]" italics="true" pageId="41" pageNumber="42">Promorphostenophanes atavus</emphasis>
<bibRefCitation id="93054B724A30FF8FFE1AFE816CE19555" author="Kaszab, Z." box="[471,660,257,282]" pageId="41" pageNumber="42" pagination="265 - 294" refId="ref52710" refString="Kaszab, Z. (1960) Neue orientalische Misolampinen (Coleoptera, Tenebrionidae). Annales historico-naturales Musei nationalis hungarici, 52, 265 - 294." type="journal article" year="1960">Kaszab, 1960: 278</bibRefCitation>
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(
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locality: 
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), fig. 9; 
<treatmentCitation id="763510924A30FF8FFC1CFE816AC09555" author="Ando, K. &amp; Ren, G. D." box="[977,1205,257,282]" page="90" pageId="41" pageNumber="42" year="2006">
<bibRefCitation id="93054B724A30FF8FFC1CFE816AC09555" author="Ando, K. &amp; Ren, G. D." box="[977,1205,257,282]" pageId="41" pageNumber="42" pagination="81 - 94" refId="ref52400" refString="Ando, K. &amp; Ren, G. D. (2006) Contribution to the knowledge of Chinese Tenebrionidae I (Coleoptera). Entomological Review of Japan, 61 (1), 81 - 94." type="journal article" year="2006">Ando &amp; Ren 2006: 90</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="763510924A30FF8FFB0DFE816BED9555" author="Lobl, I. &amp; Merkl, K. &amp; Ando, K. &amp; Bouchard, P. &amp; Lillig, M. &amp; Masumoto, K. &amp; Schawaller, W." box="[1216,1432,257,282]" page="347" pageId="41" pageNumber="42" year="2008">
<bibRefCitation id="93054B724A30FF8FFB0DFE816BED9555" author="Lobl, I. &amp; Merkl, K. &amp; Ando, K. &amp; Bouchard, P. &amp; Lillig, M. &amp; Masumoto, K. &amp; Schawaller, W." box="[1216,1432,257,282]" pageId="41" pageNumber="42" pagination="105 - 352" refId="ref52846" refString="Lobl, I., Merkl, K., Ando, K., Bouchard, P., Lillig, M., Masumoto, K. &amp; Schawaller, W. (2008) Tenebrionidae Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. 5. Tenebrionoidea. Apollo Books, Stenstrup, pp. 105 - 352." type="book chapter" year="2008">
Löbl 
<emphasis id="C5E0EA914A30FF8FFB3BFE836B569555" box="[1270,1315,258,282]" italics="true" pageId="41" pageNumber="42">et al</emphasis>
. 2008: 347
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; 
<treatmentCitation id="763510924A30FF8FFF0AFEA16FDC9576" author="Gao, Z. H. &amp; Ren, G. D." box="[199,425,289,313]" page="311" pageId="41" pageNumber="42" year="2009">
<bibRefCitation id="93054B724A30FF8FFF0AFEA16FDC9576" author="Gao, Z. H. &amp; Ren, G. D." box="[199,425,289,313]" pageId="41" pageNumber="42" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren 2009: 311</bibRefCitation>
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.
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</paragraph>
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<treatmentCitationGroup id="D78411AD4A30FF8FFF5AFEBF6C86953A" pageId="41" pageNumber="42">
<taxonomicName id="30944D004A30FF8FFF5AFEBF6C2E9517" authority="(Kaszab, 1960)" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[151,603,319,344]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFF5AFEBF6FC49518" box="[151,433,319,343]" italics="true" pageId="41" pageNumber="42">Morphostenophanes atavus</emphasis>
(
<bibRefCitation id="93054B724A30FF8FFE0CFEBF6C269517" author="Kaszab, Z." box="[449,595,319,344]" pageId="41" pageNumber="42" pagination="265 - 294" refId="ref52710" refString="Kaszab, Z. (1960) Neue orientalische Misolampinen (Coleoptera, Tenebrionidae). Annales historico-naturales Musei nationalis hungarici, 52, 265 - 294." type="journal article" year="1960">Kaszab, 1960</bibRefCitation>
)
</taxonomicName>
: 
<treatmentCitation id="763510924A30FF8FFDAAFEC06DC69518" author="Masumoto, K. &amp; Becvar, S." box="[615,947,319,343]" page="209" pageId="41" pageNumber="42" year="2008">
<bibRefCitation id="93054B724A30FF8FFDAAFEC06DC69518" author="Masumoto, K. &amp; Becvar, S." box="[615,947,319,343]" pageId="41" pageNumber="42" pagination="205 - 211" refId="ref53052" refString="Masumoto, K. &amp; Becvar, S. (2008) A study of genera Morphostenophanes Pic and Promorphostenophanes Kaszab (Coleoptera: Tenebrionidae). Entomological Review of Japan, 62 (2), 205 - 211." type="journal article" year="2008">Masumoto &amp; Bečvář 2008: 209</bibRefCitation>
</treatmentCitation>
, fig. 4; 
<treatmentCitation id="763510924A30FF8FFBC5FEBF6A879518" author="Gao, Z. H. &amp; Ren, G. D." box="[1032,1266,319,343]" page="311" pageId="41" pageNumber="42" year="2009">
<bibRefCitation id="93054B724A30FF8FFBC5FEBF6A879518" author="Gao, Z. H. &amp; Ren, G. D." box="[1032,1266,319,343]" pageId="41" pageNumber="42" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren 2009: 311</bibRefCitation>
</treatmentCitation>
, figs. 16–22, 43 [misidentification of 
<taxonomicName id="30944D004A30FF8FFE50FEDE6C919539" box="[413,740,350,374]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A30FF8FFE50FEDE6C919539" box="[413,740,350,374]" italics="true" pageId="41" pageNumber="42">Morphostenophanes brevigaster</emphasis>
</taxonomicName>
].
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</paragraph>
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<paragraph id="F72B36834A30FF8FFF5AFE1F6F69966A" blockId="41.[151,1436,414,874]" pageId="41" pageNumber="42">
<emphasis id="C5E0EA914A30FF8FFF5AFE1F6FC895F6" bold="true" box="[151,445,414,441]" pageId="41" pageNumber="42">Type material examined.</emphasis>
<materialsCitation id="47FC3CDE4A30FF8FFE0EFE1F6F539592" ID-GBIF-Occurrence="2610567379" location="Tientshun" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="C5E0EA914A30FF8FFE0EFE1F6C4595F6" bold="true" box="[451,560,415,441]" pageId="41" pageNumber="42">
<typeStatus id="282F88214A30FF8FFE0EFE1F6C4595F6" box="[451,560,415,441]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
of 
<taxonomicName id="30944D004A30FF8FFD9EFE1F6DD995F6" authorityName="Kaszab" authorityYear="1960" box="[595,940,415,441]" class="Insecta" family="Tenebrionidae" genus="Promorphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFD9EFE1F6DD995F6" box="[595,940,415,441]" italics="true" pageId="41" pageNumber="42">Promorphostenophanes atavus</emphasis>
</taxonomicName>
labelled: ‘ 
<collectingRegion id="3550F8614A30FF8FFBD1FE1F6A0295F6" box="[1052,1143,415,441]" country="China" name="Yunnan" pageId="41" pageNumber="42">Yunnan</collectingRegion>
<location id="F24B60584A30FF8FFBB0FE1F6A9995F6" LSID="urn:lsid:plazi:treatment:7F3D87954A30FF8DFF5AFF196CF896BC:F24B60584A30FF8FFBB0FE1F6A9995F6" box="[1149,1260,415,441]" name="Tientshun" pageId="41" pageNumber="42" stateProvince="Yunnan">Tientshun</location>
[handwritten in pink label] //
</materialsCitation>
<materialsCitation id="47FC3CDE4A30FF8FFEE1FE436F6D966A" ID-GBIF-Occurrence="2610567484" collectionCode="HNHM" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="282F88214A30FF8FFEE1FE436FD49592" box="[300,417,451,477]" pageId="41" pageNumber="42" type="holotype">Holotypus</typeStatus>
1960 
<taxonomicName id="30944D004A30FF8FFE2BFE436DED9592" authority="Kaszab" authorityName="Kaszab" authorityYear="1960" box="[486,920,451,477]" class="Insecta" family="Tenebrionidae" genus="Promorphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFE2BFE436D359592" box="[486,832,451,477]" italics="true" pageId="41" pageNumber="42">Promorphostenophanes atavus</emphasis>
Kaszab
</taxonomicName>
[handwritten on a pink label with red border]’ (female, HMHM, 
<figureCitation id="6FAF2A064A30FF8FFEAEFE676FC7964E" box="[355,434,486,513]" captionStart="FIGURE 25" captionStartId="41.[151,250,1937,1962]" captionTargetBox="[201,1386,905,1912]" captionTargetId="figure@41.[201,1386,895,1912]" captionTargetPageId="41" captionText="FIGURE 25. Habitus of Morphostenophanes atavus (Kaszab). Male in dorsal (A) and lateral view (B). Holotype female in dorsal (C), ventral (E) and lateral view (G), with an enlargement of ventral apical portion of left mesotibiae (D), and labels (F). Scale bars of A, B = 5 mm; C–G not to scale." figureDoi="http://doi.org/10.5281/zenodo.3797181" httpUri="https://zenodo.org/record/3797181/files/figure.png" pageId="41" pageNumber="42">Fig. 25</figureCitation>
C–G). Examined through five photographs taken by Tamás Németh (
<collectionCode id="9185AE464A30FF8FFB78FE676B7B964E" box="[1205,1294,487,513]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="41" pageNumber="42">HNHM</collectionCode>
) and sent by Ottó Merkl
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.
</paragraph>
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<emphasis id="C5E0EA914A30FF8FFF0AFDAE6C4C9606" bold="true" box="[199,569,558,585]" pageId="41" pageNumber="42">Additional material examined</emphasis>
(
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). 
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<collectingCountry id="8F8376134A30FF8FFD42FDAE6C849606" box="[655,753,558,585]" name="China" pageId="41" pageNumber="42">CHINA</collectingCountry>
: 
<collectingRegion id="3550F8614A30FF8FFCCFFDAF6D119606" box="[770,868,559,585]" country="China" name="Yunnan" pageId="41" pageNumber="42">Yunnan</collectingRegion>
</materialsCitation>
:
</emphasis>
<materialsCitation id="47FC3CDE4A30FF8FFCBBFDAF6AE29621" ID-GBIF-Occurrence="2610567477" collectingDate="2014-04-11" collectionCode="MYNU" collectorName="Xuan-Kong Jiang &amp; Tian Lu" country="China" elevation="2402" latitude="25.296759" location="Mount Gaoligong" longLatPrecision="1" longitude="98.76472" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
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(
<collectionCode id="9185AE464A30FF8FFC63FDAF6A7C9606" box="[942,1033,559,585]" pageId="41" pageNumber="42">MYNU</collectionCode>
), east slope of 
<location id="F24B60584A30FF8FFB0AFDB06BED9606" LSID="urn:lsid:plazi:treatment:7F3D87954A30FF8DFF5AFF196CF896BC:F24B60584A30FF8FFB0AFDB06BED9606" box="[1223,1432,559,586]" country="China" latitude="25.296759" longLatPrecision="1" longitude="98.76472" name="Mount Gaoligong" pageId="41" pageNumber="42" stateProvince="Yunnan">Mount Gaoligong</location>
, 
<geoCoordinate id="92A050444A30FF8FFF5AFDD26F339622" box="[151,326,594,621]" degrees="25" direction="north" minutes="17" orientation="latitude" pageId="41" pageNumber="42" precision="1" seconds="48.33" value="25.296759">25°17’48.33”N</geoCoordinate>
, 
<geoCoordinate id="92A050444A30FF8FFE9CFDD26F889622" box="[337,509,594,621]" degrees="98" direction="east" minutes="45" orientation="longitude" pageId="41" pageNumber="42" precision="1" seconds="52.98" value="98.76472">98°45’52.98”E</geoCoordinate>
, 
<quantity id="306C9B664A30FF8FFDC5FDD36C2A9621" box="[520,607,595,622]" metricMagnitude="3" metricUnit="m" metricValue="2.402" pageId="41" pageNumber="42" unit="m" value="2402.0">
<elevation id="7CB9D1B04A30FF8FFDC5FDD36C2A9621" box="[520,607,595,622]" metricMagnitude="3" metricUnit="m" metricValue="2.402" pageId="41" pageNumber="42" unit="m" value="2402.0">2402 m</elevation>
</quantity>
, 
<date id="832A10434A30FF8FFDA6FDD36C979622" box="[619,738,595,621]" pageId="41" pageNumber="42" value="2014-04-11">
<collectingDate id="936EE9AB4A30FF8FFDA6FDD36C979622" box="[619,738,595,621]" pageId="41" pageNumber="42" value="2014-04-11">2014.iv.11</collectingDate>
</date>
, at night, 
<collectorName id="5A6153554A30FF8FFC9CFDD46A629622" box="[849,1047,595,622]" pageId="41" pageNumber="42">Xuan-Kong Jiang</collectorName>
&amp; 
<collectorName id="5A6153554A30FF8FFBF7FDD36AE29621" box="[1082,1175,595,622]" pageId="41" pageNumber="42">Tian Lu</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A30FF8FFF0AFDF66D4896B2" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="F72B36834A30FF8FFF0AFDF66D4896B2" blockId="41.[151,1436,414,874]" pageId="41" pageNumber="42">
<emphasis id="C5E0EA914A30FF8FFF0AFDF66F3796DE" bold="true" box="[199,322,630,657]" pageId="41" pageNumber="42">Diagnosis.</emphasis>
Body large and elongate, black. Antennae short, reaching basal 2/7 of elytra; Pronotum widest in anterior fourth; elytra striate, with aspect ratio exceeding 
<quantity id="306C9B664A30FF8FFCD6FD1B6D2E96FA" box="[795,859,667,693]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="41" pageNumber="42" unit="in" value="1.8">1.8 in</quantity>
male, and 
<quantity id="306C9B664A30FF8FFC1BFD1B6A6396FA" box="[982,1046,667,693]" metricMagnitude="-2" metricUnit="m" metricValue="4.318" pageId="41" pageNumber="42" unit="in" value="1.7">1.7 in</quantity>
female; apical spurs of mesotibiae exposed and visible in ventral view; metafemora reaching anterior half of sternite VII; metatibiae nearly straight; apical lobes of sternite VIII narrowly hooked in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A30FF8CFF0AFC866A18921C" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="F72B36834A30FF8FFF0AFC866FF79726" blockId="41.[151,1436,414,874]" pageId="41" pageNumber="42">
<emphasis id="C5E0EA914A30FF8FFF0AFC866FCB976E" bold="true" box="[199,446,774,801]" pageId="41" pageNumber="42">Description of male.</emphasis>
Male (
<figureCitation id="6FAF2A064A30FF8FFDD8FC876CD6976D" box="[533,675,774,802]" captionStart="FIGURE 25" captionStartId="41.[151,250,1937,1962]" captionTargetBox="[201,1386,905,1912]" captionTargetId="figure@41.[201,1386,895,1912]" captionTargetPageId="41" captionText="FIGURE 25. Habitus of Morphostenophanes atavus (Kaszab). Male in dorsal (A) and lateral view (B). Holotype female in dorsal (C), ventral (E) and lateral view (G), with an enlargement of ventral apical portion of left mesotibiae (D), and labels (F). Scale bars of A, B = 5 mm; C–G not to scale." figureDoi="http://doi.org/10.5281/zenodo.3797181" httpUri="https://zenodo.org/record/3797181/files/figure.png" pageId="41" pageNumber="42">Fig. 25A, B</figureCitation>
). Color black, antennae, all tibiae, mouthparts, and claws reddish brown, shagreened. Body elongate, length 
<quantity id="306C9B664A30FF8FFDBDFCAB6CA19709" box="[624,724,811,838]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="41" pageNumber="42" unit="mm" value="23.7">23.7 mm</quantity>
, width 
<quantity id="306C9B664A30FF8FFCE9FCAB6D0E9709" box="[804,891,811,838]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="41" pageNumber="42" unit="mm" value="8.4">8.4 mm</quantity>
, strongly convex, noticeably constricted between pronotum and elytra.
</paragraph>
<caption id="A3EB660B4A30FF8FFF5AF8116C1793BD" ID-DOI="http://doi.org/10.5281/zenodo.3797181" ID-Zenodo-Dep="3797181" httpUri="https://zenodo.org/record/3797181/files/figure.png" pageId="41" pageNumber="42" startId="41.[151,250,1937,1962]" targetBox="[201,1386,905,1912]" targetPageId="41">
<paragraph id="F72B36834A30FF8FFF5AF8116C1793BD" blockId="41.[151,1437,1937,2034]" pageId="41" pageNumber="42">
<emphasis id="C5E0EA914A30FF8FFF5AF8116F5793E5" bold="true" box="[151,290,1937,1962]" pageId="41" pageNumber="42">FIGURE 25.</emphasis>
Habitus of 
<taxonomicName id="30944D004A30FF8FFE52F8116D6A93E5" authority="(Kaszab)" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[415,799,1937,1962]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A30FF8FFE52F8116CCD93E6" box="[415,696,1937,1961]" italics="true" pageId="41" pageNumber="42">Morphostenophanes atavus</emphasis>
(Kaszab)
</taxonomicName>
. Male in dorsal (
<emphasis id="C5E0EA914A30FF8FFC1FF8116D9193E5" bold="true" box="[978,996,1937,1962]" pageId="41" pageNumber="42">A</emphasis>
) and lateral view (
<emphasis id="C5E0EA914A30FF8FFB61F8126AC893E5" bold="true" box="[1196,1213,1938,1962]" pageId="41" pageNumber="42">B</emphasis>
). Holotype female in dorsal (
<emphasis id="C5E0EA914A30FF8FFF29F8356E839381" bold="true" box="[228,246,1973,1998]" pageId="41" pageNumber="42">C</emphasis>
), ventral (
<emphasis id="C5E0EA914A30FF8FFE92F8366F059381" bold="true" box="[351,368,1974,1998]" pageId="41" pageNumber="42">E</emphasis>
) and lateral view (
<emphasis id="C5E0EA914A30FF8FFDE3F8356C379381" bold="true" box="[558,578,1973,1998]" pageId="41" pageNumber="42">G</emphasis>
), with an enlargement of ventral apical portion of left mesotibiae (
<emphasis id="C5E0EA914A30FF8FFB2EF8366A809381" bold="true" box="[1251,1269,1974,1998]" pageId="41" pageNumber="42">D</emphasis>
), and labels (
<emphasis id="C5E0EA914A30FF8FFAB3F8366BF89381" bold="true" box="[1406,1421,1974,1998]" pageId="41" pageNumber="42">F</emphasis>
). Scale bars of A, B = 5 mm; C–G not to scale.
</paragraph>
</caption>
<paragraph id="F72B36834A33FF8CFF0AFF196D229654" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Head (
<figureCitation id="6FAF2A064A33FF8CFED9FF196FEE94FC" box="[276,411,153,179]" captionStart="FIGURE 31" captionStartId="51.[151,250,1867,1892]" captionTargetBox="[189,1398,489,1842]" captionTargetId="figure@51.[189,1398,489,1842]" captionTargetPageId="51" captionText="FIGURE 31. External characters and anatomic structure of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, E, K, P), M. bannaensis Zhou new species (B, G, L, Q), M. birmanicus (Kaszab) (C, H, M, R), M. brevigaster Zhou new species (D, I, N, S), M. lincangensis Zhou new species (E, J, O, T, Y), and M. vietnamicus (Kaszab) (U–X, Z). Male heads in dorsal (A–E, U) and ventral view (F–J, W); male prothoraces in dorsal (K–O, V) and ventral view (P–T, X); female reproductive organs (Y); and male mesotibiae in dorsal view (Z). Scale bars of A–X = 1 mm; Y = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797203" httpUri="https://zenodo.org/record/3797203/files/figure.png" pageId="42" pageNumber="43">Fig. 31A, F</figureCitation>
) transversely quadrate, sparsely and finely punctate, with outer margin distinctly notched between genae and clypeus; clypeus transversely hexagonal, slightly convex in middle, gently bent downwards in front, anterior margin nearly straight, weakly emarginate; frontoclypeal suture finely depressed, widely U-shaped, weakly protruding backwards in middle; genae strongly raised, depressed before eyes, strongly and roundly produced anterolaterally; frons broad, anterior part gradually sloping forwards, flattened in middle; eyes transversely reniform, strongly convex laterally; inner ocular sulci finely depressed, moderately grooved along inner margins, becoming broader posteriorly; tempora moderately convex, more coarsely punctate than frons. OI = 46.4. Antennae (
<figureCitation id="6FAF2A064A33FF8CFF6DFE156F7395E0" box="[160,262,405,431]" captionStart="FIGURE 32" captionStartId="52.[151,250,1249,1274]" captionTargetBox="[151,1437,561,1224]" captionTargetId="figure@52.[151,1437,561,1224]" captionTargetPageId="52" captionText="FIGURE 32. External characters of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G–I), M. bannaensis Zhou new species (B, J–L), M. birmanicus (Kaszab) (C, M–O), M. brevigaster Zhou new species (D, P–R), M. lincangensis Zhou new species (E, S–U), and M. vietnamicus (Kaszab) (F, V–X). Male antennae in dorsal view (A–F). Male protibiae in dorsal view (G, J, M, P, S, V), mesotibiae in ventral view (H, K, N, Q, T, W) and metatibiae in dorsal view (I, L, O, R, U, S). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797207" httpUri="https://zenodo.org/record/3797207/files/figure.png" pageId="42" pageNumber="43">Fig. 32A</figureCitation>
) slender, reaching basal 2/7 of elytra, with antennomeres weakly thickened to apices; relative lengths of antennomeres: 0.71: 0.34: 1.19: 1.14: 1.29: 1.31: 1.29: 1.16: 1.14: 1.04: 1.19. 
<collectingRegion id="3550F8614A33FF8CFBD9FE396A06959C" box="[1044,1139,441,467]" country="Myanmar" name="Chin" pageId="42" pageNumber="43">Mentum</collectingRegion>
(
<figureCitation id="6FAF2A064A33FF8CFB49FE396A92959C" box="[1156,1255,441,467]" captionStart="FIGURE 31" captionStartId="51.[151,250,1867,1892]" captionTargetBox="[189,1398,489,1842]" captionTargetId="figure@51.[189,1398,489,1842]" captionTargetPageId="51" captionText="FIGURE 31. External characters and anatomic structure of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, E, K, P), M. bannaensis Zhou new species (B, G, L, Q), M. birmanicus (Kaszab) (C, H, M, R), M. brevigaster Zhou new species (D, I, N, S), M. lincangensis Zhou new species (E, J, O, T, Y), and M. vietnamicus (Kaszab) (U–X, Z). Male heads in dorsal (A–E, U) and ventral view (F–J, W); male prothoraces in dorsal (K–O, V) and ventral view (P–T, X); female reproductive organs (Y); and male mesotibiae in dorsal view (Z). Scale bars of A–X = 1 mm; Y = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797203" httpUri="https://zenodo.org/record/3797203/files/figure.png" pageId="42" pageNumber="43">Fig. 31F</figureCitation>
) inversely trapezoidal, lateral margins weakly rounded; medial surface finely punctate, with several large pores bearing long setae, gradually rising from base to apex, depressed along both sides.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFDA56FEF96BC" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Pronotum (
<figureCitation id="6FAF2A064A33FF8CFE84FDA56FC79670" box="[329,434,549,575]" captionStart="FIGURE 31" captionStartId="51.[151,250,1867,1892]" captionTargetBox="[189,1398,489,1842]" captionTargetId="figure@51.[189,1398,489,1842]" captionTargetPageId="51" captionText="FIGURE 31. External characters and anatomic structure of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, E, K, P), M. bannaensis Zhou new species (B, G, L, Q), M. birmanicus (Kaszab) (C, H, M, R), M. brevigaster Zhou new species (D, I, N, S), M. lincangensis Zhou new species (E, J, O, T, Y), and M. vietnamicus (Kaszab) (U–X, Z). Male heads in dorsal (A–E, U) and ventral view (F–J, W); male prothoraces in dorsal (K–O, V) and ventral view (P–T, X); female reproductive organs (Y); and male mesotibiae in dorsal view (Z). Scale bars of A–X = 1 mm; Y = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797203" httpUri="https://zenodo.org/record/3797203/files/figure.png" pageId="42" pageNumber="43">Fig. 31K</figureCitation>
) inversely trapezoidal, PW/PL = 1.10, widest in anterior fourth, anterior margin nearly straight, anterior marginal border fine, interrupted in middle; lateral margins weakly rounded, lateral marginal borders thin, visible in dorsal view slightly before anterior half; posterior margin weakly rounded, emarginate in middle, posterior marginal border marked; anterior angles rounded; posterior angles obtuse; disc strongly convex, shagreened, finely and sparsely punctate, with a pair of vague impressions on middle. Scutellum widely triangular, glossy, finely punctate.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFD7D6F3397CC" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Elytra elongate oval, widest near middle, EL/EW = 1.84; strongly convex, highest in basal 2/5; with 9 continuous furrowed striae; 9 
<superScript id="00E19BCB4A33FF8CFE43FCA06FEE9761" attach="left" box="[398,411,800,814]" fontSize="6" pageId="42" pageNumber="43">th</superScript>
stria branching before apex, 1 
<superScript id="00E19BCB4A33FF8CFD3CFCA06C899761" attach="left" box="[753,764,800,814]" fontSize="6" pageId="42" pageNumber="43">st</superScript>
and 9 
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, 2 
<superScript id="00E19BCB4A33FF8CFCA6FCA06D0E9761" attach="left" box="[875,891,800,814]" fontSize="6" pageId="42" pageNumber="43">nd</superScript>
and 7 
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, 3 
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and 6 
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, and 4 
<superScript id="00E19BCB4A33FF8CFB5BFCA06AD69761" attach="left" box="[1174,1187,800,814]" fontSize="6" pageId="42" pageNumber="43">th</superScript>
and 5 
<superScript id="00E19BCB4A33FF8CFB25FCA06A809761" attach="none" box="[1256,1269,800,814]" fontSize="6" pageId="42" pageNumber="43">th</superScript>
, converging at ends, 2 
<superScript id="00E19BCB4A33FF8CFF24FCC46E8C971D" attach="left" box="[233,249,836,850]" fontSize="6" pageId="42" pageNumber="43">nd</superScript>
and 3 
<superScript id="00E19BCB4A33FF8CFE8CFCC46F3A971D" attach="none" box="[321,335,836,850]" fontSize="6" pageId="42" pageNumber="43">rd</superScript>
, and 5 
<superScript id="00E19BCB4A33FF8CFE52FCC46FD9971D" attach="left" box="[415,428,836,850]" fontSize="6" pageId="42" pageNumber="43">th</superScript>
and 6 
<superScript id="00E19BCB4A33FF8CFE39FCC46C74971D" attach="left" box="[500,513,836,850]" fontSize="6" pageId="42" pageNumber="43">th</superScript>
converging at anterior ends; intervals strongly convex, shagreened, sparsely and finely punctate.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFC0D6F21905C" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Prosternum (
<figureCitation id="6FAF2A064A33FF8CFE9AFC0D6FCD97E8" box="[343,440,909,935]" captionStart="FIGURE 31" captionStartId="51.[151,250,1867,1892]" captionTargetBox="[189,1398,489,1842]" captionTargetId="figure@51.[189,1398,489,1842]" captionTargetPageId="51" captionText="FIGURE 31. External characters and anatomic structure of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, E, K, P), M. bannaensis Zhou new species (B, G, L, Q), M. birmanicus (Kaszab) (C, H, M, R), M. brevigaster Zhou new species (D, I, N, S), M. lincangensis Zhou new species (E, J, O, T, Y), and M. vietnamicus (Kaszab) (U–X, Z). Male heads in dorsal (A–E, U) and ventral view (F–J, W); male prothoraces in dorsal (K–O, V) and ventral view (P–T, X); female reproductive organs (Y); and male mesotibiae in dorsal view (Z). Scale bars of A–X = 1 mm; Y = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797203" httpUri="https://zenodo.org/record/3797203/files/figure.png" pageId="42" pageNumber="43">Fig. 31P</figureCitation>
) shagreened, finely and sparsely punctate; prosternal process declivous, truncate at apex; hypomeron weakly rugulose, shagreened. Metasternum glossy, metaventral anterior process weakly wrinkled. Abdomen depressed, somewhat rough and finely punctate, sternites III and IV sulcate on both sides, sternite V depressed laterally.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFB9D6C7C90EC" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Legs slender. Protibiae (
<figureCitation id="6FAF2A064A33FF8CFE19FB9D6C4F9078" box="[468,570,1053,1079]" captionStart="FIGURE 32" captionStartId="52.[151,250,1249,1274]" captionTargetBox="[151,1437,561,1224]" captionTargetId="figure@52.[151,1437,561,1224]" captionTargetPageId="52" captionText="FIGURE 32. External characters of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G–I), M. bannaensis Zhou new species (B, J–L), M. birmanicus (Kaszab) (C, M–O), M. brevigaster Zhou new species (D, P–R), M. lincangensis Zhou new species (E, S–U), and M. vietnamicus (Kaszab) (F, V–X). Male antennae in dorsal view (A–F). Male protibiae in dorsal view (G, J, M, P, S, V), mesotibiae in ventral view (H, K, N, Q, T, W) and metatibiae in dorsal view (I, L, O, R, U, S). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797207" httpUri="https://zenodo.org/record/3797207/files/figure.png" pageId="42" pageNumber="43">Fig. 32G</figureCitation>
) weakly curved, apical 3/5 of inner margins pubescent; mesotibiae (
<figureCitation id="6FAF2A064A33FF8CFAE2FB9D6BE19078" box="[1327,1428,1053,1079]" captionStart="FIGURE 32" captionStartId="52.[151,250,1249,1274]" captionTargetBox="[151,1437,561,1224]" captionTargetId="figure@52.[151,1437,561,1224]" captionTargetPageId="52" captionText="FIGURE 32. External characters of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G–I), M. bannaensis Zhou new species (B, J–L), M. birmanicus (Kaszab) (C, M–O), M. brevigaster Zhou new species (D, P–R), M. lincangensis Zhou new species (E, S–U), and M. vietnamicus (Kaszab) (F, V–X). Male antennae in dorsal view (A–F). Male protibiae in dorsal view (G, J, M, P, S, V), mesotibiae in ventral view (H, K, N, Q, T, W) and metatibiae in dorsal view (I, L, O, R, U, S). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797207" httpUri="https://zenodo.org/record/3797207/files/figure.png" pageId="42" pageNumber="43">Fig. 32H</figureCitation>
) weakly curved near apical third, apical 3/5 of inner margins pubescent, ventral apical tuft interrupted, mesotibial spur uncovered (marked in 
<figureCitation id="6FAF2A064A33FF8CFE1FFBE56C4E9030" box="[466,571,1125,1151]" captionStart="FIGURE 32" captionStartId="52.[151,250,1249,1274]" captionTargetBox="[151,1437,561,1224]" captionTargetId="figure@52.[151,1437,561,1224]" captionTargetPageId="52" captionText="FIGURE 32. External characters of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G–I), M. bannaensis Zhou new species (B, J–L), M. birmanicus (Kaszab) (C, M–O), M. brevigaster Zhou new species (D, P–R), M. lincangensis Zhou new species (E, S–U), and M. vietnamicus (Kaszab) (F, V–X). Male antennae in dorsal view (A–F). Male protibiae in dorsal view (G, J, M, P, S, V), mesotibiae in ventral view (H, K, N, Q, T, W) and metatibiae in dorsal view (I, L, O, R, U, S). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797207" httpUri="https://zenodo.org/record/3797207/files/figure.png" pageId="42" pageNumber="43">Fig. 32H</figureCitation>
); metatibiae (
<figureCitation id="6FAF2A064A33FF8CFD1BFBE56D479030" box="[726,818,1125,1151]" captionStart="FIGURE 32" captionStartId="52.[151,250,1249,1274]" captionTargetBox="[151,1437,561,1224]" captionTargetId="figure@52.[151,1437,561,1224]" captionTargetPageId="52" captionText="FIGURE 32. External characters of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G–I), M. bannaensis Zhou new species (B, J–L), M. birmanicus (Kaszab) (C, M–O), M. brevigaster Zhou new species (D, P–R), M. lincangensis Zhou new species (E, S–U), and M. vietnamicus (Kaszab) (F, V–X). Male antennae in dorsal view (A–F). Male protibiae in dorsal view (G, J, M, P, S, V), mesotibiae in ventral view (H, K, N, Q, T, W) and metatibiae in dorsal view (I, L, O, R, U, S). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3797207" httpUri="https://zenodo.org/record/3797207/files/figure.png" pageId="42" pageNumber="43">Fig. 32I</figureCitation>
) straight, apical 3/5 of inner margins pubescent, outer margins depressed before apices.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFB2D6A6990A4" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
Aedeagus (
<figureCitation id="6FAF2A064A33FF8CFE89FB2D6FDD9088" box="[324,424,1197,1223]" captionStart="FIGURE 33" captionStartId="53.[151,250,1235,1260]" captionTargetBox="[165,1421,183,1207]" captionTargetId="figure@53.[164,1424,181,1209]" captionTargetPageId="53" captionText="FIGURE 33. Anatomic structures of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G), M. bannaensis Zhou new species (B, H), M. birmanicus (Kaszab) (C, I), M. brevigaster Zhou new species (D, J), M. lincangensis Zhou new species (E, K), and M. vietnamicus (Kaszab) (F, L). Aedeagi (A–F) and male sternites VIII (G–L); in dorsal view (d), lateral view (l), and enlargements of parameres apices (p). Scale bars = 1 mm; all parameres apices not to scale." figureDoi="http://doi.org/10.5281/zenodo.3797213" httpUri="https://zenodo.org/record/3797213/files/figure.png" pageId="42" pageNumber="43">Fig. 33A</figureCitation>
) elongate, curved in lateral view; parameres slender, 0.22 as long as total length, with ovoid apex. Sternite VIII (
<figureCitation id="6FAF2A064A33FF8CFEBAFB516FA890A4" box="[375,477,1233,1259]" captionStart="FIGURE 33" captionStartId="53.[151,250,1235,1260]" captionTargetBox="[165,1421,183,1207]" captionTargetId="figure@53.[164,1424,181,1209]" captionTargetPageId="53" captionText="FIGURE 33. Anatomic structures of Morphostenophanes species in the atavus-group: M. atavus (Kaszab) (A, G), M. bannaensis Zhou new species (B, H), M. birmanicus (Kaszab) (C, I), M. brevigaster Zhou new species (D, J), M. lincangensis Zhou new species (E, K), and M. vietnamicus (Kaszab) (F, L). Aedeagi (A–F) and male sternites VIII (G–L); in dorsal view (d), lateral view (l), and enlargements of parameres apices (p). Scale bars = 1 mm; all parameres apices not to scale." figureDoi="http://doi.org/10.5281/zenodo.3797213" httpUri="https://zenodo.org/record/3797213/files/figure.png" pageId="42" pageNumber="43">Fig. 33G</figureCitation>
) with apical lobes narrowly hooked in lateral view.
</paragraph>
<paragraph id="F72B36834A33FF8CFF0AFB746A18921C" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
<emphasis id="C5E0EA914A33FF8CFF0AFB746FC69140" bold="true" box="[199,435,1268,1295]" pageId="42" pageNumber="43">Comparative notes.</emphasis>
<taxonomicName id="30944D004A33FF8CFE71FB756C869140" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[444,755,1269,1295]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFE71FB756C869140" box="[444,755,1269,1295]" italics="true" pageId="42" pageNumber="43">Morphostenophanes atavus</emphasis>
</taxonomicName>
most resembles 
<taxonomicName id="30944D004A33FF8CFC78FB766A2E9140" box="[949,1115,1269,1295]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFC78FB766DB99140" box="[949,972,1270,1295]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFC10FB756A2E9140" box="[989,1115,1269,1295]" italics="true" pageId="42" pageNumber="43">brevigaster</emphasis>
</taxonomicName>
. They share similar appearance and indistinguishable aedeagi. However, 
<taxonomicName id="30944D004A33FF8CFD5DFA9A6C89917C" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[656,764,1305,1331]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFD5DFA9A6CD2917C" box="[656,679,1306,1331]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFD79FA9A6C89917C" box="[692,764,1306,1331]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
can be easily distinguished from 
<taxonomicName id="30944D004A33FF8CFBA5FA9A6B7C917C" box="[1128,1289,1305,1331]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFBA5FA9A6A0A917C" box="[1128,1151,1306,1331]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFB40FA996B7C917C" box="[1165,1289,1305,1331]" italics="true" pageId="42" pageNumber="43">brevigaster</emphasis>
</taxonomicName>
by its shorter antennae which reaches 2/7 of elytra, compared to those of 
<taxonomicName id="30944D004A33FF8CFCE9FABE6DB09118" box="[804,965,1341,1367]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFCE9FABE6DB09118" box="[804,965,1341,1367]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
reaching third of elytra; each antennomere less sharply dilated towards apex; pronotum widest in apical fourth, compared to that of 
<taxonomicName id="30944D004A33FF8CFB9EFAE26A879134" box="[1107,1266,1377,1403]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFB9EFAE26A879134" box="[1107,1266,1377,1403]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
widest near the middle; elytra more elongate, highest posterior to the midpoint, compared to those of 
<taxonomicName id="30944D004A33FF8CFB85FA066A9F91D0" box="[1096,1258,1413,1439]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFB85FA066A9F91D0" box="[1096,1258,1413,1439]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
highest anterior to the midpoint; elytral striae less depressed than those of the latter; all legs shorter, metafemora reaching anterior half of sternite VII, compared to those 
<taxonomicName id="30944D004A33FF8CFDC3FA4E6CDB91A8" box="[526,686,1485,1511]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFDC3FA4E6CDB91A8" box="[526,686,1485,1511]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
reaching apex of sternite VII; protibiae in male evenly curved, compared to that of 
<taxonomicName id="30944D004A33FF8CFE89FA726F909244" box="[324,485,1521,1547]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFE89FA726F909244" box="[324,485,1521,1547]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
curved at apical third; mesotibiae with apical spur exposed and visible in ventral view, compared to those of 
<taxonomicName id="30944D004A33FF8CFE44F9966C5E9260" box="[393,555,1557,1583]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFE44F9966FD59260" box="[393,416,1558,1583]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFE62F9956C5E9260" box="[431,555,1557,1583]" italics="true" pageId="42" pageNumber="43">brevigaster</emphasis>
</taxonomicName>
covered by tuft and obscured; metatibiae slightly more straight; apical lobes of male sternite VIII narrowly hooked, compared to those of 
<taxonomicName id="30944D004A33FF8CFCD9F9BA6DC6921C" box="[788,947,1593,1619]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A33FF8CFCD9F9BA6DC6921C" box="[788,947,1593,1619]" italics="true" pageId="42" pageNumber="43">M. brevigaster</emphasis>
</taxonomicName>
broadly hooked.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A33FF8DFF0AF9DC6CFA9680" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="discussion">
<paragraph id="F72B36834A33FF8CFF0AF9DC6A7A92AC" blockId="42.[151,1437,153,2015]" pageId="42" pageNumber="43">
<emphasis id="C5E0EA914A33FF8CFF0AF9DC6F3A9238" bold="true" box="[199,335,1628,1655]" pageId="42" pageNumber="43">Comments.</emphasis>
<taxonomicName id="30944D004A33FF8CFE95F9DD6CFA9238" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[344,655,1629,1655]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFE95F9DD6CFA9238" box="[344,655,1629,1655]" italics="true" pageId="42" pageNumber="43">Morphostenophanes atavus</emphasis>
</taxonomicName>
was described from a single female from ‘Yunnan’ without detailed location (‘ohne nähere Angabe des Fundortes’ in original description). The male was then illustrated by 
<bibRefCitation id="93054B724A33FF8CFAE9F9016F2D92F0" author="Masumoto, K. &amp; Becvar, S." pageId="42" pageNumber="43" pagination="205 - 211" refId="ref53052" refString="Masumoto, K. &amp; Becvar, S. (2008) A study of genera Morphostenophanes Pic and Promorphostenophanes Kaszab (Coleoptera: Tenebrionidae). Entomological Review of Japan, 62 (2), 205 - 211." type="journal article" year="2008">Masumoto &amp; Bečvář (2008)</bibRefCitation>
, identified as 
<taxonomicName id="30944D004A33FF8CFE3BF9266C1192F0" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[502,612,1701,1727]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFE3BF9266C7892F0" box="[502,525,1702,1727]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFDD6F9266C1192F0" box="[539,612,1702,1727]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
, however, no description or locality was given. Later, 
<bibRefCitation id="93054B724A33FF8CFB07F9256BE992F0" author="Gao, Z. H. &amp; Ren, G. D." box="[1226,1436,1701,1727]" pageId="42" pageNumber="43" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren (2009)</bibRefCitation>
provided detiled locality for this species, as well as a photograph of the female.
</paragraph>
<paragraph id="F72B36834A33FF8DFF0AF96D6D3B95C4" blockId="42.[151,1437,153,2015]" lastBlockId="43.[151,1437,153,755]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
In the present study, the author examined the 
<typeStatus id="282F88214A33FF8CFD73F96D6D559348" box="[702,800,1773,1799]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="30944D004A33FF8CFC89F96E6DC49348" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[836,945,1773,1799]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFC89F96E6D2E9348" box="[836,859,1774,1799]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFCA5F96E6DC49348" box="[872,945,1774,1799]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
through photos. Specimens with similar appearances from three localities were studied, with one male from the eastern slope of Gaoligong Mountains, and two series containing both males and females from Yingjiang County and an adjacent area and Lincang City. According to the original description, the elytral aspect ratio of the 
<taxonomicName id="30944D004A33FF8CFCD8F8DA6DF0933C" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[789,901,1881,1907]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFCD8F8DA6D59933C" box="[789,812,1882,1907]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFCF1F8DA6DF0933C" box="[828,901,1882,1907]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
<typeStatus id="282F88214A33FF8CFC40F8D96D9A933C" box="[909,1007,1881,1907]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
female is 1.7, which is different from those from around Yingjiang County with an average ratio of 1.59 (n = 10) and maximum of 1.65. Moreover, the elytral striae of the 
<taxonomicName id="30944D004A33FF8CFEA1F8226FAC93F4" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[364,473,1953,1979]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFEA1F8226FF693F4" box="[364,387,1954,1979]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFE5DF8226FAC93F4" box="[400,473,1954,1979]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
<typeStatus id="282F88214A33FF8CFE12F8216C3493F4" box="[479,577,1953,1979]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
are distinctly less depressed than those of the latter. The elytral aspect ratio of those from Lincang ranges from 1.7 to over 1.9, which matches that of the 
<taxonomicName id="30944D004A33FF8CFC06F8466A4D9390" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[971,1080,1989,2015]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A33FF8CFC06F8466D979390" box="[971,994,1990,2015]" italics="true" pageId="42" pageNumber="43">M</emphasis>
. 
<emphasis id="C5E0EA914A33FF8CFC22F8466A4D9390" box="[1007,1080,1990,2015]" italics="true" pageId="42" pageNumber="43">atavus</emphasis>
</taxonomicName>
<typeStatus id="282F88214A33FF8CFBF3F8456AD69390" box="[1086,1187,1989,2015]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
, but they are distinctly different by the latter having much more convex pronotum and elytra. The elytral aspect ratio of the male from the western slope of Gaoligong Mountains is 1.84, which is comprable to the 
<taxonomicName id="30944D004A32FF8DFC29FF3E6A219498" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[996,1108,189,215]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A32FF8DFC29FF3E6D8E9498" box="[996,1019,190,215]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFBC6FF3E6A219498" box="[1035,1108,190,215]" italics="true" pageId="43" pageNumber="44">atavus</emphasis>
</taxonomicName>
female. Moreover, this male has antennomeres less thickened anteriorly, elytral intervals less convex, and ventral apical tuft is interrupted, which are identical with those of the female 
<typeStatus id="282F88214A32FF8DFDF0FE856CD69550" box="[573,675,261,287]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
. Therefore, this male from the Gaoligong Mountains is identified as 
<taxonomicName id="30944D004A32FF8DFF5AFEAA6F73950C" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[151,262,297,323]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A32FF8DFF5AFEAA6EDB950C" box="[151,174,298,323]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFF70FEAA6F73950C" box="[189,262,298,323]" italics="true" pageId="43" pageNumber="44">atavus</emphasis>
</taxonomicName>
. The female 
<typeStatus id="282F88214A32FF8DFE50FEA96F8A950C" box="[413,511,297,323]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
possesses bulges at each side of the base of the pronotum, which are indistinct in the male. This is probably due to interspecific variation. Specimens from Yingjiang County and Lincang City are identified as belonging to 
<taxonomicName id="30944D004A32FF8DFE71FEF26C1595C4" box="[444,608,369,395]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A32FF8DFE71FEF26FA695C4" box="[444,467,370,395]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFE2CFEF16C1595C4" box="[481,608,369,395]" italics="true" pageId="43" pageNumber="44">brevigaster</emphasis>
</taxonomicName>
and 
<taxonomicName id="30944D004A32FF8DFD5BFEF26D3295C4" box="[662,839,369,395]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="lincangensis">
<emphasis id="C5E0EA914A32FF8DFD5BFEF26CD895C4" box="[662,685,370,395]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFD76FEF16D3295C4" box="[699,839,369,395]" italics="true" pageId="43" pageNumber="44">lincangensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="F72B36834A32FF8DFF0AFE156B2C9654" blockId="43.[151,1437,153,755]" pageId="43" pageNumber="44">
On the label of 
<taxonomicName id="30944D004A32FF8DFEB1FE166F9895E0" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[380,493,405,431]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A32FF8DFEB1FE166FE695E0" box="[380,403,406,431]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFE69FE166F9895E0" box="[420,493,406,431]" italics="true" pageId="43" pageNumber="44">atavus</emphasis>
</taxonomicName>
<typeStatus id="282F88214A32FF8DFE38FE156C2F95E0" box="[501,602,405,431]" pageId="43" pageNumber="44" type="holotype">holotype</typeStatus>
, the author found ‘Tientshun’ after the locality ‘Yunnan’, which refers to ‘Tengchong’, a city reaching the western slope of the Gaoligong Mountains. Such detailed locality was not mentioned in the original publication. The type locality associated with data of here examined male suggest a potential distribution of 
<taxonomicName id="30944D004A32FF8DFEF3FD826FD99654" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[318,428,513,539]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A32FF8DFEF3FD826F209654" box="[318,341,514,539]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFEAEFD826FD99654" box="[355,428,514,539]" italics="true" pageId="43" pageNumber="44">atavus</emphasis>
</taxonomicName>
along the southern Gaoligong Mountains, spreading over both slopes of mountains.
</paragraph>
<paragraph id="F72B36834A32FF8DFF0AFDA56CFA9680" blockId="43.[151,1437,153,755]" pageId="43" pageNumber="44">
The male specimen in Masumoto &amp; Bečvář’s photograph (2008, 
<figureCitation id="6FAF2A064A32FF8DFC0DFDA56A739670" box="[960,1030,549,575]" captionStart="FIGURE 4" captionStartId="11.[151,250,1975,2000]" captionTargetBox="[162,1427,931,1942]" captionTargetId="figure@11.[151,1436,921,1950]" captionTargetPageId="11" captionText="FIGURE 4. Habitus of Morphostenophanes aenescens Pic. Male from Xishan District, Qujin City (A, C) and female from Mianmian Mountain (B, D); in dorsal (A, B) and lateral view (C, D). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3797100" httpUri="https://zenodo.org/record/3797100/files/figure.png" pageId="43" pageNumber="44">Fig. 4</figureCitation>
) was in dorsal view, so the author measured its elytral aspect ratio, the result is 1.86. Moreover, its pronotum is constricted before the base, and its protibiae are only slightly curved, based on the revised diagnostic characters, it is undoubtedly a true 
<taxonomicName id="30944D004A32FF8DFAE8FDEE6BE096C8" baseAuthorityName="Kaszab" baseAuthorityYear="1960" box="[1317,1429,621,647]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="C5E0EA914A32FF8DFAE8FDEE6B4996C8" box="[1317,1340,622,647]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFA81FDEE6BE096C8" box="[1356,1429,622,647]" italics="true" pageId="43" pageNumber="44">atavus</emphasis>
</taxonomicName>
. Based on the known distribution, the female specimen mentioned by 
<bibRefCitation id="93054B724A32FF8DFC6AFD116A0F96E4" author="Gao, Z. H. &amp; Ren, G. D." box="[935,1146,657,683]" pageId="43" pageNumber="44" pagination="307 - 319" refId="ref52584" refString="Gao, Z. H. &amp; Ren, G. D. (2009) Taxonomy of the genus Morphostenophanes Pic from China, with two new species (Coleoptera, Tenebrionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 55 (4), 307 - 319." type="journal article" year="2009">Gao &amp; Ren (2009)</bibRefCitation>
from Yingjiang county is considered a misidentified of 
<taxonomicName id="30944D004A32FF8DFE29FD366CF29680" box="[484,647,693,719]" class="Insecta" family="Tenebrionidae" genus="Morphostenophanes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="brevigaster">
<emphasis id="C5E0EA914A32FF8DFE29FD366F8E9680" box="[484,507,694,719]" italics="true" pageId="43" pageNumber="44">M</emphasis>
. 
<emphasis id="C5E0EA914A32FF8DFDC4FD356CF29680" box="[521,647,693,719]" italics="true" pageId="43" pageNumber="44">brevigaster</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BF8E65084A32FF8DFF0AFD586CF896BC" box="[199,653,728,755]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="F72B36834A32FF8DFF0AFD586CF896BC" blockId="43.[151,1437,153,755]" box="[199,653,728,755]" pageId="43" pageNumber="44">
<emphasis id="C5E0EA914A32FF8DFF0AFD586F1496BC" bold="true" box="[199,353,728,755]" pageId="43" pageNumber="44">Distribution.</emphasis>
(
<figureCitation id="6FAF2A064A32FF8DFEA2FD596FC296BC" box="[367,439,729,755]" captionStart="Map 1" captionStartId="8.[151,202,1286,1310]" captionTargetBox="[151,1436,705,1261]" captionTargetId="figure@8.[151,1436,705,1261]" captionText="Map 1. Distribution of the Morphostenophanes species. A. Enlargement of Central and Western Yunnan Province." figureDoi="http://doi.org/10.5281/zenodo.3797090" httpUri="https://zenodo.org/record/3797090/files/figure.png" pageId="43" pageNumber="44">Map 1</figureCitation>
) 
<collectingCountry id="8F8376134A32FF8DFE04FD596C5396BC" box="[457,550,729,755]" name="China" pageId="43" pageNumber="44">CHINA</collectingCountry>
: 
<collectingRegion id="3550F8614A32FF8DFDFDFD596CFC96BC" box="[560,649,729,755]" country="China" name="Yunnan" pageId="43" pageNumber="44">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>