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<document id="C312531CF0A4C9EC6C8F062AC0DD35AE" ID-DOI="10.5281/zenodo.186086" ID-GBIF-Dataset="34491f5d-e23b-40d7-9d4c-41714503582e" ID-ISSN="1175-5326" ID-Zenodo-Dep="186086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045940587" checkinUser="plazi" docAuthor="Campos, Ernesto" docDate="2009" docId="743587B5FFC6FFF419BF239EFB30FC02" docLanguage="en" docName="zt02022p044.pdf" docOrigin="Zootaxa 2022" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enigmatheres canfieldi Rathbun 1918, new combination" docType="treatment" docVersion="9" lastPageNumber="33" masterDocId="880CFFCDFFC5FFF019282309FF92FF92" masterDocTitle="Pacific Ocean, with a reappraisal of the subfamily Pinnotherinae de Haan, 1833 (Crustacea: Brachyura: Pinnotheridae)" masterLastPageNumber="44" masterPageNumber="29" pageNumber="32" updateTime="1698576862496" updateUser="plazi">
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<mods:title id="F82DD6145AFCCA94E67FCBB0D5B44760">Pacific Ocean, with a reappraisal of the subfamily Pinnotherinae de Haan, 1833 (Crustacea: Brachyura: Pinnotheridae)</mods:title>
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<mods:namePart id="0A5D474EF96CF6A0F673CDACF75BAA6A">Campos, Ernesto</mods:namePart>
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<mods:date id="7DB9194601F55730C01ECCC810FAC58E">2009</mods:date>
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<subSubSection id="B4866528FFC6FFF319BF239EFEBEFF46" pageId="3" pageNumber="32" type="nomenclature">
<paragraph id="FC2336A3FFC6FFF319BF239EFCFBFF23" blockId="3.[151,873,151,212]" box="[151,873,151,177]" pageId="3" pageNumber="32">
<heading id="A76B81CFFFC6FFF319BF239EFCFBFF23" bold="true" box="[151,873,151,177]" fontSize="11" level="1" pageId="3" pageNumber="32" reason="1">
<emphasis id="CEE8EAB1FFC6FFF319BF239EFCFBFF23" bold="true" box="[151,873,151,177]" pageId="3" pageNumber="32">
<taxonomicName id="3B9C4D20FFC6FFF319BF239EFD17FF23" ID-CoL="39V5H" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" box="[151,645,151,177]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="canfieldi" status="comb. nov.">
<emphasis id="CEE8EAB1FFC6FFF319BF239EFE23FF23" bold="true" box="[151,433,151,177]" italics="true" pageId="3" pageNumber="32">Enigmatheres canfieldi</emphasis>
(
<bibRefCitation id="980D4B52FFC6FFF318E9239EFDEFFF23" author="Rathbun" box="[449,637,151,177]" pageId="3" pageNumber="44" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="D5DB57CAFFC6FFF31BBB239EFCFBFF23" box="[659,873,151,177]" pageId="3" pageNumber="32">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FC2336A3FFC6FFF319B623B3FEBEFF46" blockId="3.[151,873,151,212]" box="[158,300,186,212]" pageId="3" pageNumber="32">
(
<figureCitation id="64A72A26FFC6FFF3198F23B3FF7AFF46" box="[167,232,186,212]" captionStart="FIGURE 1" captionStartId="3.[151,257,1824,1848]" captionTargetBox="[179,1421,817,1806]" captionTargetId="figure@3.[168,1433,804,1818]" captionTargetPageId="3" captionText="FIGURE 1. Enigmatheres canfieldi (Rathbun, 1918) new combination, Monterey, California (USNM 8445). Adult female: A B, left outer and right inner view of MXP 3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; C D not at scale; E = 1 mm." httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="3" pageNumber="32">Fig. 1</figureCitation>
AE)
</paragraph>
</subSubSection>
<subSubSection id="B4866528FFC6FFF319BF2201FE5CFE0D" pageId="3" pageNumber="32" type="reference_group">
<paragraph id="FC2336A3FFC6FFF319BF2201FDFDFEAD" blockId="3.[151,1436,264,415]" pageId="3" pageNumber="32">
<treatmentCitationGroup id="DC8C118DFFC6FFF319BF2201FDFDFEAD" pageId="3" pageNumber="32">
<taxonomicName id="3B9C4D20FFC6FFF319BF2201FD06FE8D" authority="Rathbun, 1904: 106" authorityName="Rathbun" authorityPageNumber="106" authorityYear="1904" box="[151,660,264,287]" class="Malacostraca" family="Pinnotheridae" genus="Raphonotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="subquadratus">
<emphasis id="CEE8EAB1FFC6FFF319BF2201FE23FE8D" box="[151,433,264,287]" italics="true" pageId="3" pageNumber="32">Raphonotus subquadratus</emphasis>
<treatmentCitation id="7D3D10B2FFC6FFF318932201FD06FE8D" author="Rathbun" box="[443,660,264,287]" page="106" pageId="3" pageNumber="32" year="1904">
<bibRefCitation id="980D4B52FFC6FFF318932201FDCCFE8D" author="Rathbun" box="[443,606,264,287]" pageId="3" pageNumber="44" refString="Rathbun, M. J. (1904) Decapod crustaceans of the northwest coast of North America. Harriman Alaska Expedition 10, 1 - 190." type="journal article" year="1904">Rathbun, 1904</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
(part: specimen from Monterey, California,
<emphasis id="CEE8EAB1FFC6FFF31DA82201FB3AFE8D" box="[1152,1192,264,287]" italics="true" pageId="3" pageNumber="32">fide</emphasis>
<bibRefCitation id="980D4B52FFC6FFF31D9A2201FAC7FE8D" author="Rathbun" box="[1202,1365,264,287]" pageId="3" pageNumber="44" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
). (Not
<taxonomicName id="3B9C4D20FFC6FFF319EF2221FDF1FEAD" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" box="[199,611,296,319]" class="Malacostraca" family="Pinnotheridae" genus="Raphonotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="subquadratus">
<emphasis id="CEE8EAB1FFC6FFF319EF2221FE4CFEAD" box="[199,478,296,319]" italics="true" pageId="3" pageNumber="32">Raphonotus subquadratus</emphasis>
<bibRefCitation id="980D4B52FFC6FFF318CC2221FDF1FEAD" author="Dana" box="[484,611,296,319]" pageId="3" pageNumber="44" refString="Dana, J. D. (1851) On the classification of the crustacean Grapsoidea. American Journal of Science and Arts, series 2 (12), 283 - 291." type="journal article" year="1851">Dana, 1851</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="FC2336A3FFC6FFF319BF2241FE5CFE0D" blockId="3.[151,1436,264,415]" pageId="3" pageNumber="32">
<treatmentCitationGroup id="DC8C118DFFC6FFF319BF2241FE5CFE0D" pageId="3" pageNumber="32">
<taxonomicName id="3B9C4D20FFC6FFF319BF2241FD9BFECD" ID-CoL="3DQY7" authority="Rathbun, 1918: 12" authorityName="Rathbun" authorityPageNumber="12" authorityYear="1918" box="[151,521,328,351]" class="Malacostraca" family="Pinnotheridae" genus="Fabia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="canfieldi">
<emphasis id="CEE8EAB1FFC6FFF319BF2241FEA8FECD" box="[151,314,328,351]" italics="true" pageId="3" pageNumber="32">Fabia canfieldi</emphasis>
<treatmentCitation id="7D3D10B2FFC6FFF318692241FD9BFECD" author="Rathbun" box="[321,521,328,351]" page="12" pageId="3" pageNumber="32" year="1918">
<bibRefCitation id="980D4B52FFC6FFF318692241FE70FECD" author="Rathbun" box="[321,482,328,351]" pageId="3" pageNumber="44" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
: 12
</treatmentCitation>
</taxonomicName>
, 101, 102, 106107, pl. 24, fig. 57, text-fig. 57;
<treatmentCitation id="7D3D10B2FFC6FFF31D302241FF52FEED" author="Schmitt" page="254" pageId="3" pageNumber="32" year="1921">
<bibRefCitation id="980D4B52FFC6FFF31D302241FB23FECD" author="Schmitt" box="[1048,1201,328,351]" pageId="3" pageNumber="44" refString="Schmitt, W. L. (1921) The marine decapod Crustacea of California. University of California Publications in Zoology, 23, 1 - 470." type="journal article" year="1921">Schmitt, 1921</bibRefCitation>
: 254
</treatmentCitation>
255, pl.39, fig. 56:
<treatmentCitation id="7D3D10B2FFC6FFF3188D2261FDFAFEED" author="Balss" box="[421,616,360,383]" page="1420" pageId="3" pageNumber="32" year="1957">
<bibRefCitation id="980D4B52FFC6FFF3188D2261FDB7FEED" author="Balss" box="[421,549,360,383]" pageId="3" pageNumber="43" refString="Balss, H. (1957) Decapoda, VIII: Systematik, In: H. G. Bronn. Klassen und Ordungen des Tierreichs, Band 5, Abteilung I, 7 (12), 1505 - 1672." type="journal article" year="1957">Balss, 1957</bibRefCitation>
: 1420
</treatmentCitation>
;
<treatmentCitation id="7D3D10B2FFC6FFF31B5B2261FCF3FEED" author="Schmitt" box="[627,865,360,383]" page="7" pageId="3" pageNumber="32" year="1973">
<bibRefCitation id="980D4B52FFC6FFF31B5B2261FCDAFEED" author="Schmitt" box="[627,840,360,383]" pageId="3" pageNumber="44" refString="Schmitt, W. L., McCain, J. C. &amp; Davidson, E. S. (1973) Fam. Pinnotheridae, Brachyura I: Decapoda I. In, H. - E. Gruner &amp; L. B. Holthuis, eds. Crustaceorum Catalogus. W. Junk, Den Haag 3, 1 - 160." type="journal article" year="1973">
Schmitt
<emphasis id="CEE8EAB1FFC6FFF31BE42261FC90FEED" box="[716,770,360,383]" italics="true" pageId="3" pageNumber="32">et al.</emphasis>
, 1973
</bibRefCitation>
: 7
</treatmentCitation>
, 22:
<treatmentCitation id="7D3D10B2FFC6FFF31ABD2261FBA5FEED" author="Dai" box="[917,1079,360,383]" page="136" pageId="3" pageNumber="32" year="1980">
<bibRefCitation id="980D4B52FFC6FFF31ABD2261FB90FEED" author="Dai" box="[917,1026,360,383]" lastPageId="3" lastPageNumber="44" pageId="3" pageNumber="43" refString="Dai, Ai-Yun (1980) Fabia obtusidentata Dai, sp. nov. In: Dai, Ai-Yun, Zhong-qi, Feng, Yu-zhi, Song &amp; Guo-xiao Chen. New species and new records of family Pinnotheridae from Hainan Island. Acta Zootaxonomica Sinica, 5, 129 - 142 [in Chinese, with English summary]." type="journal article" year="1980">Dai, 1980</bibRefCitation>
: 136
</treatmentCitation>
;
<treatmentCitation id="7D3D10B2FFC6FFF31D6A2261FABFFEED" author="Campos" box="[1090,1325,360,383]" page="1174" pageId="3" pageNumber="32" year="1996">
<bibRefCitation id="980D4B52FFC6FFF31D6A2261FB7FFEED" author="Campos" box="[1090,1261,360,383]" pageId="3" pageNumber="43" refString="Campos, E. (1996 b) Partial revision of the genus Fabia Dana, 1851 (Crustacea: Brachyura: Pinnotheridae). Journal of Natural History, 30, 1157 - 1178." type="journal article" year="1996" yearSuffix="b">Campos, 1996b</bibRefCitation>
:
<date id="88221063FFC6FFF31DDF2261FABFFEED" box="[1271,1325,360,383]" pageId="3" pageNumber="32" value="1174" valueMax="1175">1174</date>
</treatmentCitation>
1175; Ng
<emphasis id="CEE8EAB1FFC6FFF319EF2281FF6FFE0D" box="[199,253,392,415]" italics="true" pageId="3" pageNumber="32">et al.</emphasis>
, 2008: 249 (listed).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="B4866528FFC6FFF319BF22C7FD26FD9D" pageId="3" pageNumber="32" type="distribution">
<paragraph id="FC2336A3FFC6FFF319BF22C7FD26FD9D" blockId="3.[151,1437,462,759]" pageId="3" pageNumber="32">
<emphasis id="CEE8EAB1FFC6FFF319BF22C7FE3FFE7A" bold="true" box="[151,429,462,488]" pageId="3" pageNumber="32">Distribution and host</emphasis>
. Known from only Monterey, California, the
<typeStatus id="23278801FFC6FFF31AF522C7FB83FE7A" box="[989,1041,462,488]" pageId="3" pageNumber="32">type</typeStatus>
locality; in folds of the keyhole limpet
<taxonomicName id="3B9C4D20FFC6FFF319C122FCFD22FD9D" authority="Sowerby, 1825" authorityName="Sowerby" authorityYear="1825" box="[233,688,501,527]" class="Gastropoda" family="Fissurellidae" genus="Megathura" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="3" pageNumber="32" phylum="Mollusca" rank="species" species="crenulata">
<emphasis id="CEE8EAB1FFC6FFF319C122FCFE70FD9C" box="[233,482,501,526]" italics="true" pageId="3" pageNumber="32">Megathura crenulata</emphasis>
(Sowerby, 1825)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B4866528FFC6FFF419ED2112FC8BFE52" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32" type="materials_examined">
<paragraph id="FC2336A3FFC6FFF319ED2112FAC9FDA4" blockId="3.[151,1437,462,759]" box="[197,1371,539,566]" pageId="3" pageNumber="32">
<emphasis id="CEE8EAB1FFC6FFF319ED2112FE23FDA7" bold="true" box="[197,433,539,565]" pageId="3" pageNumber="32">Material examined</emphasis>
.
<typeStatus id="23278801FFC6FFF318E82115FDBFFDA4" box="[448,557,540,566]" pageId="3" pageNumber="32" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="3B9C4D20FFC6FFF31B7C2115FC48FDA4" authority="Rathbun, 1918: 1" authorityName="Rathbun" authorityPageNumber="1" authorityYear="1918" box="[596,986,540,566]" class="Malacostraca" family="Pinnotheridae" genus="Fabia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="canfieldi">
<emphasis id="CEE8EAB1FFC6FFF31B7C2115FC9AFDA7" box="[596,776,540,565]" italics="true" pageId="3" pageNumber="32">Fabia canfieldi</emphasis>
<bibRefCitation id="980D4B52FFC6FFF31A272115FC52FDA4" author="Rathbun" box="[783,960,540,566]" pageId="3" pageNumber="44" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461." type="journal article" year="1918">Rathbun, 1918</bibRefCitation>
: 1
</taxonomicName>
damaged female (
<collectionCode id="9A8DAE66FFC6FFF31D992115FA9AFDA4" LSID="urn:lsid:biocol.org:col:34871" box="[1201,1288,540,566]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="32">USNM</collectionCode>
8445).
</paragraph>
<paragraph id="FC2336A3FFC6FFF319ED214BFC08FD11" blockId="3.[151,1437,462,759]" pageId="3" pageNumber="32">
<emphasis id="CEE8EAB1FFC6FFF319ED214BFD99FDCE" bold="true" box="[197,523,578,604]" pageId="3" pageNumber="32">
Redescription of
<typeStatus id="23278801FFC6FFF318B6214BFD99FDCE" box="[414,523,578,604]" pageId="3" pageNumber="32" type="holotype">holotype</typeStatus>
</emphasis>
. Carapace damaged (crushed as to be for the most part beyond description). Front devoid of setae, with short, longitudinal median depression.
</paragraph>
<paragraph id="FC2336A3FFC6FFF319ED2199FBE5FD65" blockId="3.[151,1437,462,759]" pageId="3" pageNumber="32">
Third maxilliped obliquely placed in buccal cavity; ischium, merus indistinguishably fused; palp 3- segmented; carpus larger than conical propodus; dactylus small, narrow, inserted on middle third of medioventral margin of propodus. Exopod with 2-segmented flagellum (
<figureCitation id="64A72A26FFC6FFF31AC121D4FBB8FD65" box="[1001,1066,733,759]" captionStart="FIGURE 1" captionStartId="3.[151,257,1824,1848]" captionTargetBox="[179,1421,817,1806]" captionTargetId="figure@3.[168,1433,804,1818]" captionTargetPageId="3" captionText="FIGURE 1. Enigmatheres canfieldi (Rathbun, 1918) new combination, Monterey, California (USNM 8445). Adult female: A B, left outer and right inner view of MXP 3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; C D not at scale; E = 1 mm." httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="3" pageNumber="32">Fig. 1</figureCitation>
AB).
</paragraph>
<caption id="A8E3662BFFC6FFF319BF2429FB4AF8EE" httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="3" pageNumber="32" targetBox="[179,1421,817,1806]" targetPageId="3">
<paragraph id="FC2336A3FFC6FFF319BF2429FB4AF8EE" blockId="3.[151,1436,1824,1916]" pageId="3" pageNumber="32">
<emphasis id="CEE8EAB1FFC6FFF319BF2429FE8FF8AA" bold="true" box="[151,285,1824,1848]" pageId="3" pageNumber="32">FIGURE 1.</emphasis>
<taxonomicName id="3B9C4D20FFC6FFF3180D2429FD73F8A5" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" box="[293,737,1824,1847]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="canfieldi" status="comb. nov.">
<emphasis id="CEE8EAB1FFC6FFF3180D2429FDB7F8A5" box="[293,549,1824,1847]" italics="true" pageId="3" pageNumber="32">Enigmatheres canfieldi</emphasis>
(Rathbun, 1918)
</taxonomicName>
<emphasis id="CEE8EAB1FFC6FFF31BC12429FC23F8AA" bold="true" box="[745,945,1824,1848]" pageId="3" pageNumber="32">
<taxonomicNameLabel id="D5DB57CAFFC6FFF31BC12429FC23F8AA" box="[745,945,1824,1848]" pageId="3" pageNumber="32">new combination</taxonomicNameLabel>
</emphasis>
, Monterey, California (USNM 8445). Adult female: AB, left outer and right inner view of MXP3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; CD not at scale; E = 1 mm.
</paragraph>
</caption>
<paragraph id="FC2336A3FFC6FFF419ED24A6FC8AFF4A" blockId="3.[151,1437,1967,2031]" lastBlockId="4.[151,1437,152,912]" lastPageId="4" lastPageNumber="33" pageId="3" pageNumber="32">
Chelipeds elongated (
<figureCitation id="64A72A26FFC6FFF318FB24A6FD88F85B" box="[467,538,1967,1993]" captionStart="FIGURE 1" captionStartId="3.[151,257,1824,1848]" captionTargetBox="[179,1421,817,1806]" captionTargetId="figure@3.[168,1433,804,1818]" captionTargetPageId="3" captionText="FIGURE 1. Enigmatheres canfieldi (Rathbun, 1918) new combination, Monterey, California (USNM 8445). Adult female: A B, left outer and right inner view of MXP 3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; C D not at scale; E = 1 mm." httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="3" pageNumber="32">Fig. 1</figureCitation>
C), palm slightly increasing in width distally, fingers shorter than palm, slightly deflexed, ventral margin of propodus slightly concave; fingers not gaping when closed, setose along inner surface; with prehensile tooth at proximal third of cutting edge of dactylus, smaller one at base of pollex; cutting edge of both fingers with small teeth (
<figureCitation id="64A72A26FFC1FFF41B8523B7FD62FF4A" box="[685,752,190,216]" captionStart="FIGURE 1" captionStartId="3.[151,257,1824,1848]" captionTargetBox="[179,1421,817,1806]" captionTargetId="figure@3.[168,1433,804,1818]" captionTargetPageId="3" captionText="FIGURE 1. Enigmatheres canfieldi (Rathbun, 1918) new combination, Monterey, California (USNM 8445). Adult female: A B, left outer and right inner view of MXP 3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; C D not at scale; E = 1 mm." httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="4" pageNumber="33">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="FC2336A3FFC1FFF419ED23ECFE9AFE08" blockId="4.[151,1437,152,912]" pageId="4" pageNumber="33">
Walking legs subcylindrical, naked, relative length 2&gt; 3&gt; 1&gt; 4; second leg asymmetrical, right 1/3 longer than left; WL1 stouter than others; propodi slightly curved, slightly stouter at distal end than at proximal, except right WL2 which tapers to distal end; longest right WL2 dactylus differs from others dactili in being nearly straight to tip; other dactyli shorter, slender, conical, more curved toward hooked, hard tip (
<figureCitation id="64A72A26FFC1FFF419B72289FF73FE08" box="[159,225,384,410]" captionStart="FIGURE 1" captionStartId="3.[151,257,1824,1848]" captionTargetBox="[179,1421,817,1806]" captionTargetId="figure@3.[168,1433,804,1818]" captionTargetPageId="3" captionText="FIGURE 1. Enigmatheres canfieldi (Rathbun, 1918) new combination, Monterey, California (USNM 8445). Adult female: A B, left outer and right inner view of MXP 3; C, left cheliped; D, pollex and dactylus of left cheliped; E, left and right walking legs, respectively. Scales: A = 0.57 mm; B = 0.47 mm; C D not at scale; E = 1 mm." httpUri="https://zenodo.org/record/186087/files/figure.png" pageId="4" pageNumber="33">Fig. 1</figureCitation>
E).
</paragraph>
<paragraph id="FC2336A3FFC1FFF419ED22AFFC8BFE52" blockId="4.[151,1437,152,912]" box="[197,793,422,448]" pageId="4" pageNumber="33">Abdomen of six somites and telson well separated.</paragraph>
</subSubSection>
<subSubSection id="B4866528FFC1FFF419ED22C4FB30FC02" pageId="4" pageNumber="33" type="discussion">
<paragraph id="FC2336A3FFC1FFF419ED22C4FB30FC02" blockId="4.[151,1437,152,912]" pageId="4" pageNumber="33">
<emphasis id="CEE8EAB1FFC1FFF419ED22C4FED2FE75" bold="true" box="[197,320,461,487]" pageId="4" pageNumber="33">Remarks</emphasis>
. The presence of a thin, translucent and soft carapace, enlarged carpus of MXP3 and asymmetrical WL2 are characters that suggest
<taxonomicName id="3B9C4D20FFC1FFF41BE222FDFCFCFD9F" box="[714,878,500,525]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41BE222FDFCFCFD9F" box="[714,878,500,525]" italics="true" pageId="4" pageNumber="33">Enigmatheres</emphasis>
</taxonomicName>
is related to the genus
<taxonomicName id="3B9C4D20FFC1FFF41DAF22FDFF4AFDA6" authority="Campos, 2002" authorityName="Campos" authorityYear="2002" class="Malacostraca" family="Pinnotheridae" genus="Austinotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41DAF22FDFAB8FD9F" box="[1159,1322,500,525]" italics="true" pageId="4" pageNumber="33">Austinotheres</emphasis>
<bibRefCitation id="980D4B52FFC1FFF41C1B22FDFF4AFDA6" author="Campos" pageId="4" pageNumber="43" refString="Campos, E. (2002) Two new genera of pinnotherid crabs from the tropical eastern Pacific (Brachyura: Pinnotheridae). Journal of Crustacean Biology, 16 (3), 556 - 563." type="journal article" year="2002">Campos, 2002</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="3B9C4D20FFC1FFF419CC2113FE1FFDA1" box="[228,397,538,563]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF419CC2113FE1FFDA1" box="[228,397,538,563]" italics="true" pageId="4" pageNumber="33">Enigmatheres</emphasis>
</taxonomicName>
differs from
<taxonomicName id="3B9C4D20FFC1FFF41B1A2113FD48FDA1" box="[562,730,538,563]" class="Malacostraca" family="Pinnotheridae" genus="Austinotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41B1A2113FD48FDA1" box="[562,730,538,563]" italics="true" pageId="4" pageNumber="33">Austinotheres</emphasis>
</taxonomicName>
in the gently curved outer margin of the MXP3 ischiummerus, and by the insertion of the dactylus on the middle third of the ventral margin of the conical propodus.
<taxonomicName id="3B9C4D20FFC1FFF419BF2161FEABFD13" box="[151,313,616,641]" class="Malacostraca" family="Pinnotheridae" genus="Austinotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF419BF2161FEABFD13" box="[151,313,616,641]" italics="true" pageId="4" pageNumber="33">Austinotheres</emphasis>
</taxonomicName>
has an obtuse angle on the MXP3 outer margin and the dactylus of this appendage is inserted further distally on the propodus.
<taxonomicName id="3B9C4D20FFC1FFF41B3D2186FD27FD3A" box="[533,693,655,680]" class="Malacostraca" family="Pinnotheridae" genus="Austinotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41B3D2186FD27FD3A" box="[533,693,655,680]" italics="true" pageId="4" pageNumber="33">Austinotheres</emphasis>
</taxonomicName>
can be also distinguished from
<taxonomicName id="3B9C4D20FFC1FFF41D0F2186FB5BFD3A" box="[1063,1225,655,680]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41D0F2186FB5BFD3A" box="[1063,1225,655,680]" italics="true" pageId="4" pageNumber="33">Enigmatheres</emphasis>
</taxonomicName>
by its slender and long walking legs, with blade-shaped and slightly curved dactyli. Both
<taxonomicName id="3B9C4D20FFC1FFF41AC321BCFB1CFD5C" box="[1003,1166,693,718]" class="Malacostraca" family="Pinnotheridae" genus="Enigmatheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41AC321BCFB1CFD5C" box="[1003,1166,693,718]" italics="true" pageId="4" pageNumber="33">Enigmatheres</emphasis>
</taxonomicName>
and
<taxonomicName id="3B9C4D20FFC1FFF41DE121BCFAF9FD5C" box="[1225,1387,693,718]" class="Malacostraca" family="Pinnotheridae" genus="Austinotheres" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="CEE8EAB1FFC1FFF41DE121BCFAF9FD5C" box="[1225,1387,693,718]" italics="true" pageId="4" pageNumber="33">Austinotheres</emphasis>
</taxonomicName>
can be distinguished from other pinnotherids by the asymmetrical WL2 and the shape and relative length of the MXP3 palp articles, particularly the carpus being larger than the propodus, as well as the small and narrow dactylus being inserted medially or distally on the medioventral margin of the propodus. Others genera currently in the
<taxonomicName id="3B9C4D20FFC1FFF4187B2059FE64FCF8" box="[339,502,848,874]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
have symmetrical walking legs and/or the dactylus of MXP3 is inserted on the middle or proximal ventral margin of a propodus which is always larger than the carpus.
</paragraph>
</subSubSection>
</treatment>
</document>