treatments-xml/data/74/27/E3/7427E30DF6B85974B2937820FFBC61A3.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1026.61554" ID-GBIF-Dataset="0a2d723c-d292-4f0c-9d00-6cdd176c49a6" ID-PMC="PMC8016817" ID-Pensoft-Pub="1313-2970-1026-45" ID-Pensoft-UUID="2EFE20EFF6705E34BFA20CF497608C4D" ID-PubMed="33850418" ID-ZooBank="A3212DAACEF24A1F93CC1823940C0108" ModsDocID="1313-2970-1026-45" checkinTime="1616697129133" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob &amp; Balke, Michael" docDate="2021" docId="7427E30DF6B85974B2937820FFBC61A3" docLanguage="en" docName="ZooKeys 1026: 45-67" docOrigin="ZooKeys 1026" docPubDate="2021-03-25" docSource="http://dx.doi.org/10.3897/zookeys.1026.61554" docTitle="Exocelina apistefti Shaverdo, Surbakti &amp; Balke 2021, sp. nov." docType="treatment" docUuid="7862C435-10FA-4ED3-B73E-BACCE3D10EBB" docUuidSource="ZooBank" docVersion="5" id="2EFE20EFF6705E34BFA20CF497608C4D" lastPageNumber="45" masterDocId="2EFE20EFF6705E34BFA20CF497608C4D" masterDocTitle="Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="67" masterPageNumber="45" pageNumber="45" updateTime="1668149974149" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Seven new species of the Exocelina ekari group from New Guinea central and coastal mountains (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shaverdo, Helena</mods:namePart>
<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Surbakti, Suriani</mods:namePart>
<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sumoked, Bob</mods:namePart>
<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Balke, Michael</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1026</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>45</mods:start>
<mods:end>67</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1026.61554</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1026.61554</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1026-45</mods:identifier>
<mods:identifier type="ZooBank">A3212DAACEF24A1F93CC1823940C0108</mods:identifier>
<mods:identifier type="Pensoft-UUID">2EFE20EFF6705E34BFA20CF497608C4D</mods:identifier>
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<treatment ID-GBIF-Taxon="180680995" LSID="urn:lsid:zoobank.org:act:7862C435-10FA-4ED3-B73E-BACCE3D10EBB" httpUri="http://treatment.plazi.org/id/7427E30DF6B85974B2937820FFBC61A3" lastPageNumber="45" pageId="0" pageNumber="45">
<subSubSection pageId="0" pageNumber="45" type="nomenclature">
<paragraph pageId="0" pageNumber="45">
<taxonomicName LSID="http://zoobank.org/7862C435-10FA-4ED3-B73E-BACCE3D10EBB" authority="Shaverdo, Surbakti &amp; Balke" authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti" status="sp. nov.">Exocelina apistefti Shaverdo, Surbakti &amp; Balke</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="45">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">Figures 4</figureCitation>
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">, 8</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="reference_group">
<paragraph pageId="0" pageNumber="45">
<taxonomicName authority="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke, 2021" authorityName="Shaverdo &amp; Surbakti &amp; Sumoked &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis" status="sp. nov.">Exocelina nr. brahminensis</taxonomicName>
#7287: 
<bibRefCitation DOI="https://doi.org/10.1186/s12862-021-01764-2" author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" refId="B10" refString="Toussaint, EFA, White, LT, Shaverdo, H, Lam, A, Surbakti, S, Panjaitan, R, Sumoked, B, von Rintelen, T, Sagata, K, Balke, M, 2021. [in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology. https://doi.org/10.1186/s12862-021-01764-2" title="[in press] New Guinean orogenic dynamics and biota evolution revealed using a custom geospatial analyses pipeline. BMS Evolutionary Biology." url="https://doi.org/10.1186/s12862-021-01764-2" year="2021">Toussaint et al. 2021</bibRefCitation>
: figs 3-6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type locality">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
locality.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138321" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" elevation="1700" latitude="-2.5718334" location="Foja Mts" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
, 
<collectingMunicipality>Sarmi Regency</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:262B8B0DC67DE37EBE707C5FE03779D0" country="Indonesia" latitude="-2.5718334" longLatPrecision="1" longitude="138.71724" municipality="Sarmi Regency" name="Foja Mts" stateProvince="Papua Barat">Foja Mts</location>
, 
<geoCoordinate degrees="02" direction="south" minutes="34" orientation="latitude" precision="1" seconds="18.6" value="-2.5718334">02°34'18.6&quot;S</geoCoordinate>
, 
<geoCoordinate degrees="138" direction="east" minutes="43" orientation="longitude" precision="1" seconds="02.1" value="138.71724">138°43'02.1&quot;E</geoCoordinate>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="type material">
<paragraph pageId="0" pageNumber="45">
<typeStatus>Type</typeStatus>
material.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3062138339" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB" country="Indonesia" elevation="1700" latitude="-2.571839" location="Foja Mountains" longLatPrecision="1" longitude="138.71725" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
male &quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:8D62B64180210C7DCA41E6DD99853E8D" country="Indonesia" latitude="-2.571839" longLatPrecision="1" longitude="138.71725" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, bog camp, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700m</elevation>
</quantity>
, 
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
, 
<geoCoordinate degrees="2.571839" direction="south" orientation="latitude" precision="1" value="-2.571839">-2.571839</geoCoordinate>
<geoCoordinate degrees="138.717250" direction="east" orientation="longitude" precision="1" value="138.71725">138.717250</geoCoordinate>
, Sumoked (Pap058)&quot;, 
<normalizedToken originalValue="“7287”">&quot;7287&quot;</normalizedToken>
[green text] (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138334" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="KSP" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount type="female">2 females</specimenCount>
with the same label as the 
<typeStatus>holotype</typeStatus>
(
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3062138320" collectingDate="2016-05-23" collectingDateMax="2016-06-03" collectingDateMin="2016-05-23" collectionCode="MZB, KSP" country="Indonesia" elevation="1600" latitude="-2.561006" location="Foja Mountains" longLatPrecision="1" longitude="138.71149" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount type="male">1 male</specimenCount>
, 
<specimenCount type="female">1 female</specimenCount>
&quot; 
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:C5F1BEF221719927C587AB1FC529DB04" country="Indonesia" latitude="-2.561006" longLatPrecision="1" longitude="138.71149" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, river camp, 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="m" value="1600.0">1600m</elevation>
</quantity>
, 
<date value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">
<collectingDate value="2016-05-23" valueMax="2016-06-03" valueMin="2016-05-23">23.v.-3.vi.2016</collectingDate>
</date>
, 
<geoCoordinate degrees="2.561006" direction="south" orientation="latitude" precision="1" value="-2.561006">-2.561006</geoCoordinate>
<geoCoordinate degrees="138.711487" direction="east" orientation="longitude" precision="1" value="138.71149">138.711487</geoCoordinate>
, Sumoked (Pap059)&quot; (
<collectionCode country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" type="Museum">MZB</collectionCode>
, 
<collectionCode country="USA" name="Pittsburg State University" type="Herbarium">KSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="description">
<paragraph pageId="0" pageNumber="45">Description.</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Body size and form</emphasis>
:
</emphasis>
Beetle small: TL-H 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.575" metricValueMax="3.7" metricValueMin="3.45" unit="mm" value="3.575" valueMax="3.7" valueMin="3.45">3.45-3.7 mm</quantity>
, TL 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.925" metricValueMax="4.05" metricValueMin="3.8" unit="mm" value="3.925" valueMax="4.05" valueMin="3.8">3.8-4.05 mm</quantity>
, MW 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.925" metricValueMax="2.0" metricValueMin="1.85" unit="mm" value="1.925" valueMax="2.0" valueMin="1.85">1.85-2 mm</quantity>
(
<typeStatus>holotype</typeStatus>
: TL-H 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
, TL 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.9" unit="mm" value="3.9">3.9 mm</quantity>
, 
<collectionCode country="Netherlands" name="Museum Wasmann">MW</collectionCode>
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.9" unit="mm" value="1.9">1.9 mm</quantity>
), with oblong-oval habitus (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">4</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Colouration</emphasis>
:
</emphasis>
Dorsally piceous, with paler lateral sides of pronotum (Fig. 
<figureCitation captionStart="Figures 1–4" captionStartId="F1" captionText="Figures 1 - 4. Habitus and colouration of holotype 1 Exocelina foja sp. nov. 2 E. waaf sp. nov. 3 E. riberai sp. nov. 4 E. apistefti sp. nov." figureDoi="10.3897/zookeys.1026.61554.figures1-4" httpUri="https://binary.pensoft.net/fig/524445" pageId="0" pageNumber="45">4</figureCitation>
). Head piceous, with slightly paler, dark brown, anterior margin; pronotum piceous, slightly paler towards lateral sides, lateral sides brown to dark brown, yellowish to reddish brown at anterior angles; elytra uniformly piceous; head appendages and proximal part of legs yellowish brown, legs distally brown.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Surface sculpture</emphasis>
:
</emphasis>
Shiny dorsally, with fine microreticulation and almost invisible punctation on elytra. Head with dense and coarse punctation (spaces between punctures 0-3 times size of punctures), distinctly finer and sparser anteriorly and posteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with distinctly finer and sparser punctation than on head. Elytra with very sparse and fine punctation, almost invisible. Elytra with weakly impressed microreticulation; pronotum and especially head with stronger microreticulation. Metaventrite and metacoxa distinctly but weakly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with weak microreticulation, strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Structures</emphasis>
:
</emphasis>
Pronotum without lateral bead. Base of prosternum and neck of prosternal process with ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex medially, with distinct bead and few setae. Abdominal ventrite 6 broadly rounded apically.
</paragraph>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Male</emphasis>
:
</emphasis>
Antenna simple. Pro- and mesotarsomeres 1-3 not dilated, narrow. Protarsomere 4 cylindrical, narrow, with medium-sized, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 13 and posterior row of six short setae (Fig. 
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8A</figureCitation>
). Median lobe with distinctly discontinuous outline; in lateral view, almost straight, with apex broad, curved downwards, and pointed at tip; in ventral view, with distinct submedian constriction, distal part narrower than proximal one, apex deeply and narrowly concave (Fig. 
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8B, C</figureCitation>
). Paramere with strong notch on dorsal side, with median notch tip sharply pointed, and subdistal part large and elongate; subdistal setae long and dense, of two kind: more numerous upper ones thin and lower setae shorter, thicker and flattened; proximal setae hair-like, distinctly more inconspicuous than subdistal ones (Fig. 
<figureCitation captionStart="Figures 7, 8" captionStartId="F3" captionText="Figures 7, 8. 7 Exocelina riberai sp. nov., holotype 8 E. apistefti sp. nov., holotype A right protarsomeres 4 and 5 in ventral view B median lobe in ventral view C median lobe in lateral view D right paramere in external view." figureDoi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45">8D</figureCitation>
). Abdominal ventrite 6 broadly rounded, with 9-11 lateral striae on each side.
</paragraph>
<caption doi="10.3897/zookeys.1026.61554.figures7-8" httpUri="https://binary.pensoft.net/fig/524447" pageId="0" pageNumber="45" start="Figures 7, 8" startId="F3">
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">Figures 7, 8.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="45">7</emphasis>
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina riberai" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="riberai">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina riberai</emphasis>
</taxonomicName>
sp. nov., 
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">8</emphasis>
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis italics="true" pageId="0" pageNumber="45">E. apistefti</emphasis>
</taxonomicName>
sp. nov., 
<typeStatus>holotype</typeStatus>
<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
right protarsomeres 4 and 
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.27" unit="in" value="5.0">5 in</quantity>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
median lobe in ventral view 
<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
median lobe in lateral view 
<emphasis bold="true" pageId="0" pageNumber="45">D</emphasis>
right paramere in external view.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="45">
<emphasis bold="true" pageId="0" pageNumber="45">
<emphasis italics="true" pageId="0" pageNumber="45">Female</emphasis>
:
</emphasis>
Pro- and mesotarsi not modified. Abdominal ventrite 6 without striae.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="affinities">
<paragraph pageId="0" pageNumber="45">Affinities.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species evidently belongs to the 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. ekari" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="ekari">
<emphasis italics="true" pageId="0" pageNumber="45">E. ekari</emphasis>
</taxonomicName>
group due to the discontinuous outline of its median lobe. The species is very similar to 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis">
<emphasis italics="true" pageId="0" pageNumber="45">E. brahminensis</emphasis>
</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.250.3715" author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="0" pageNumber="45" pagination="1 - 76" refId="B3" refString="Shaverdo, HV, Surbakti, S, Hendrich, LM, Balke, M, 2012. Introduction of the Exocelina ekari -group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 250: 1 - 76, DOI: https://doi.org/10.3897/zookeys.250.3715" title="Introduction of the Exocelina ekari - group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">
Shaverdo et al., 
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</bibRefCitation>
general appearance and structure of male genitalia, especially in the sharply pointed median notch tip of paramere, but differs from it in lager size (TL-H 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="3.2249999999999996" metricValueMax="3.3" metricValueMin="3.15" unit="mm" value="3.2249999999999996" valueMax="3.3" valueMin="3.15">3.15-3.3 mm</quantity>
for 
<taxonomicName class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="E. brahminensis" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="brahminensis">
<emphasis italics="true" pageId="0" pageNumber="45">E. brahminensis</emphasis>
</taxonomicName>
), darker dorsal colouration, shape of the median lobe, and paramere setation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="distribution">
<paragraph pageId="0" pageNumber="45">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="45">
<materialsCitation ID-GBIF-Occurrence="3062138322" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Indonesia" location="Foja Mountains" municipality="This" specimenCount="1" stateProvince="Papua Barat">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
: 
<collectingRegion country="Indonesia" name="Papua Barat">Papua Province</collectingRegion>
. 
<collectingMunicipality>This</collectingMunicipality>
species is known only from 
<location LSID="urn:lsid:plazi:treatment:7427E30DF6B85974B2937820FFBC61A3:A185B40B3578D71336FE84E7FDA4F5D5" country="Indonesia" municipality="This" name="Foja Mountains" stateProvince="Papua Barat">Foja Mountains</location>
, from and near the 
<typeStatus>type</typeStatus>
locality (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F7" captionText="Figure 15. Map of the western part of New Guinea showing the species distribution." figureDoi="10.3897/zookeys.1026.61554.figure15" httpUri="https://binary.pensoft.net/fig/524451" pageId="0" pageNumber="45">15</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="habitat">
<paragraph pageId="0" pageNumber="45">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="45">The specimens were collected from small, shallow forest creeks.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="45" type="etymology">
<paragraph pageId="0" pageNumber="45">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="45">
The species name 
<taxonomicName authorityName="Shaverdo, Surbakti &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina apistefti" order="Coleoptera" pageId="0" pageNumber="45" phylum="Arthropoda" rank="species" species="apistefti">
<emphasis italics="true" pageId="0" pageNumber="45">Exocelina apistefti</emphasis>
</taxonomicName>
derives from Greek 
<emphasis italics="true" pageId="0" pageNumber="45">
<normalizedToken originalValue="απίστευτος">απίστευτος</normalizedToken>
</emphasis>
(feminine 
<emphasis italics="true" pageId="0" pageNumber="45">
<normalizedToken originalValue="απίστευτη">απίστευτη</normalizedToken>
</emphasis>
) meaning unbelievable. The name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
</treatment>
</document>