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<document id="FB0DD032653EAEE7D603CBEA4D0A7204" ID-CLB-Dataset="22062" ID-DOI="10.5852/ejt.2016.170" ID-GBIF-Dataset="f0106ae9-b2bc-4e28-905e-2653f8f0cbd3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3832608" ID-ZooBank="5203E7A5-1592-4F1D-9F3C-24E6791D9429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589831302177" checkinUser="valdenar" docAuthor="Hsiao, Yun, Okushima, Yûichi, Yang, Ping-Shih &amp; Ko, Chium-Cheng" docDate="2016" docId="2D34EF515F5FFFD2FDE5E1BF5ABDFE7F" docLanguage="en" docName="ejt-2016-170.pdf.imf" docOrigin="European Journal of Taxonomy 170" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lycocerus kintaroi Hsiao &amp; Okushima &amp; Yang 2016, sp. nov." docType="treatment" docUuid="FE9AC62A-4B2E-49A0-BD5E-58CD46500D58" docUuidSource="ZooBank" docVersion="11" lastPageNumber="30" masterDocId="D10D97295F44FFCFFFE6E7145B24FFBB" masterDocTitle="Taxonomic revision of the Lycocerus hanatanii species group (Coleoptera, Cantharidae), with the description of new species from Taiwan" masterLastPageNumber="33" masterPageNumber="1" pageNumber="28" updateTime="1698840141833" updateUser="ExternalLinkService">
<mods:mods id="044360BA04B4C883557B583658AD0678" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="872E073E9B0A8E01B1F033BEC2B2067C">
<mods:title id="A5A09CAB94F1D2A3DE0F5AB2EB9CB47B">Taxonomic revision of the Lycocerus hanatanii species group (Coleoptera, Cantharidae), with the description of new species from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="8E0306960DC736BD09708BB6DD5098A5" type="personal">
<mods:role id="1B2C02CFA8B571CDC22DA7005C2BA7B7">
<mods:roleTerm id="0660D4171F168E34D8470B5901D0DB01">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FFFA94BABDD2C5DF3B6565841C23DC4">Hsiao, Yun</mods:namePart>
<mods:nameIdentifier id="69B1430A54E00627BA8809DD4C3923C7" type="LSID">6E5C734A-1809-46A1-A595-5A0CB7B39BA0</mods:nameIdentifier>
<mods:affiliation id="3C061A109408739BE7C0FB4D18F8096D">Department of Entomology, National Taiwan University, No. 27, Lane 113, Sec. 4, Roosevelt Rd., Taipei 10617, Taiwan. Kurashiki Museum of Natural History, Chûô 2 - 6 - 1, Kurashiki-shi, Okayama Pref., 710 - 0046 Japan.</mods:affiliation>
<mods:nameIdentifier id="18ED8ECE4BE202515EDEDB1AA10DB1A5" type="email">yunhsiao@outlook.com</mods:nameIdentifier>
</mods:name>
<mods:name id="AB7A78F9440C49041B3A957B73CC727A" type="personal">
<mods:role id="3CC8C423A36E0135D594C78C6067452B">
<mods:roleTerm id="D50DA487FF4918E35D930612E2EF1AFD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="571855A19F84DEB6F00E3DB160AF2C3D">Okushima, Yûichi</mods:namePart>
<mods:nameIdentifier id="43CD424589C5A326464E064B051F7323" type="LSID">BB590861-E5AE-438B-AC0E-A8724F8FAAB5</mods:nameIdentifier>
<mods:nameIdentifier id="3A70EC43BE3A2972B5BB5DD03DE44884" type="email">musoku@city.kurashiki.okayama.jp</mods:nameIdentifier>
</mods:name>
<mods:name id="59B4E7CBD2230B7D7A8CE0C0699C248A" type="personal">
<mods:role id="5CEFC525BD7DC3E0337841204A73D9E6">
<mods:roleTerm id="AFF1D16D426969BCA7866EA632AC3E1E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB432D61CE224A332A669D57AEAD1604">Yang, Ping-Shih</mods:namePart>
<mods:nameIdentifier id="59E8D78314A3737C570319EBA535496F" type="LSID">0CD84731-DCC1-4A68-BE78-E543D35FA5A2&amp;290CF231-2518-4C91-A2EA-C184C9728C7F</mods:nameIdentifier>
<mods:affiliation id="C0DDED09F151F6026335A59A83C6D5B6">Department of Entomology, National Taiwan University, No. 27, Lane 113, Sec. 4, Roosevelt Rd., Taipei 10617, Taiwan. Kurashiki Museum of Natural History, Chûô 2 - 6 - 1, Kurashiki-shi, Okayama Pref., 710 - 0046 Japan. &amp; Department of Entomology, National Taiwan University, No. 27, Lane 113, Sec. 4, Roosevelt Rd., Taipei 10617, Taiwan. Kurashiki Museum of Natural History, Chûô 2 - 6 - 1, Kurashiki-shi, Okayama Pref., 710 - 0046 Japan.</mods:affiliation>
<mods:nameIdentifier id="1FE09361484C180602B9E53C126DB299" type="email">psyang@ntu.edu.tw&amp;kocc2501@ntu.edu.tw</mods:nameIdentifier>
</mods:name>
<mods:name id="EA5BB251BD38F8ADAAE449D3F770E190" type="personal">
<mods:role id="0665E4E81598277C0BABBAEAC8962BA9">
<mods:roleTerm id="172CCDD306218595BDF83568C503A66C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B76349C3FB8D5E517A7183B4F7DDF481">Ko, Chium-Cheng</mods:namePart>
</mods:name>
<mods:typeOfResource id="BF5AD0FE28D40504A71B2E5A91415AF2">text</mods:typeOfResource>
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<mods:titleInfo id="8297DFA5F34A28EB563EB5C6F5A8DF27">
<mods:title id="E5D9E7E927F2DBD45A51D1CB9D73B174">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C573C5B368B9DC5B75290AC94859DF05">
<mods:date id="5FBA4D7BC8F6CF8D86A3B0618631E503">2016</mods:date>
<mods:detail id="51CD2201DDF646C2BE78F0CB6D940110" type="pubDate">
<mods:number id="6B0418E0DFC329795E816976931EC6B3">2016-01-05</mods:number>
</mods:detail>
<mods:detail id="2F7537582B245BEC5C19DA283FCA29F1" type="volume">
<mods:number id="2E890884B9E27A6A0C5E4E0C73230662">170</mods:number>
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<mods:start id="88D1EE6CCBA2B5B6F819540571B9C814">1</mods:start>
<mods:end id="B8D404B525EF95153B82F2095A3F0A56">33</mods:end>
</mods:extent>
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<mods:classification id="F11D1EF02601B68411F5C035F443A779">journal article</mods:classification>
<mods:identifier id="3E6AF8BB8241955FDC3AC1A6CEF1BC97" type="CLB-Dataset">22062</mods:identifier>
<mods:identifier id="E222390E6F64F15B0E88A044538BE7F5" type="DOI">10.5852/ejt.2016.170</mods:identifier>
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<mods:identifier id="A370F96DBC24EAF8B89CD5843E49FA33" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="14DD3CC21FBE12D659D5BAD2EB067C17" type="Zenodo-Dep">3832608</mods:identifier>
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</mods:mods>
<treatment id="2D34EF515F5FFFD2FDE5E1BF5ABDFE7F" ID-DOI="http://doi.org/10.5281/zenodo.3850470" ID-GBIF-Taxon="164167252" ID-Zenodo-Dep="3850470" LSID="urn:lsid:zoobank.org:act:FE9AC62A-4B2E-49A0-BD5E-58CD46500D58" httpUri="http://treatment.plazi.org/id/2D34EF515F5FFFD2FDE5E1BF5ABDFE7F" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="ED870DCC5F5FFFD4FDE5E1BF5F15F97E" box="[515,1073,1706,1733]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="A5225E475F5FFFD4FDE5E1BF5F15F97E" blockId="27.[358,1230,1706,1803]" box="[515,1073,1706,1733]" pageId="27" pageNumber="28">
<heading id="FE6AE92B5F5FFFD4FDE5E1BF5F15F97E" box="[515,1073,1706,1733]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
<taxonomicName id="629D25C45F5FFFD4FDE5E1BF58EBF97F" authority="Hsiao &amp; Okushima" authorityName="Hsiao &amp; Okushima &amp; Yang" authorityYear="2016" box="[515,975,1706,1733]" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kintaroi" status="sp. nov.">
<emphasis id="97E982555F5FFFD4FDE5E1BF59C1F97E" bold="true" box="[515,741,1706,1733]" italics="true" pageId="27" pageNumber="28">Lycocerus kintaroi</emphasis>
Hsiao &amp; Okushima
</taxonomicName>
<taxonomicNameLabel id="8CDA3F2E5F5FFFD4FC30E1BF5F15F97E" box="[982,1073,1707,1733]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F5FFFD4FE80E1D9585AF8B0" pageId="27" pageNumber="28" type="description">
<paragraph id="A5225E475F5FFFD4FE80E1D95FEAF953" blockId="27.[358,1230,1706,1803]" box="[358,1230,1741,1768]" pageId="27" pageNumber="28">
<uri id="D10C52455F5FFFD4FE80E1D95FEAF953" box="[358,1230,1741,1768]" pageId="27" pageNumber="28">
urn:lsid:zoobank.org:act:
<uuid id="D13B64925F5FFFD4FD65E1D95FEAF953" box="[643,1230,1741,1768]" pageId="27" pageNumber="28">FE9AC62A-4B2E-49A0-BD5E-58CD46500D58</uuid>
</uri>
</paragraph>
<paragraph id="A5225E475F5FFFD4FD53E1E5585AF8B0" blockId="27.[358,1230,1706,1803]" box="[693,894,1776,1803]" pageId="27" pageNumber="28">
<figureCitation id="3DA642C25F5FFFD4FD53E1E55830F8B0" box="[693,788,1777,1803]" captionStart="Fig" captionStartId="8.[189,232,1822,1848]" captionTargetBox="[188,1394,422,1777]" captionTargetId="figure@8.[149,1432,382,1818]" captionTargetPageId="8" captionText="Fig. 2. Habitus, dorsal view. A–B. Lycocerus pictus (Wittmer, 1983). C–E. L. yitingi Hsiao &amp; Okushima sp. nov. F–G. L. araticollis (Fairmaire, 1897). H–I. L. griseopubens (Pic, 1928). J–M. L. nigripennis (Pic, 1938). N–O. L. aurantiacus Hsiao &amp; Okushima sp. nov. P–Q. L. evangelium Hsiao &amp; Okushima sp. nov. R. L. kintaroi Hsiao &amp; Okushima sp. nov. C, F, H, J, N, P, R. Name-bearing types. D, O, Q. Paratypes. A–B, E, G, I, K–M. Non-type specimens. A, C, F–G, I, K–L, N, P, R. Male. B, D–E, H, J, M, O, Q. Female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3832612" httpUri="https://zenodo.org/record/3832612/files/figure.png" pageId="27" pageNumber="28">Figs 2R</figureCitation>
,
<figureCitation id="3DA642C25F5FFFD4FCC6E1E45809F8B0" box="[800,813,1776,1803]" captionStart="Fig" captionStartId="16.[190,233,1956,1982]" captionTargetBox="[194,1497,270,1908]" captionTargetId="figure@16.[154,1506,264,1921]" captionTargetPageId="16" captionText="Fig. 5. Aedeagus. A–C. Lycocerus evangelium Hsiao &amp; Okushima sp. nov. D–F. L. kintaroi Hsiao &amp; Okushima sp. nov. A, D. Ventral view. B, E. Lateral view. C, F. Dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3832618" httpUri="https://zenodo.org/record/3832618/files/figure.png" pageId="27" pageNumber="28">5</figureCitation>
D–F, 8
</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F5FFFD4FF5BE0265A8FF85F" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="A5225E475F5FFFD4FF5BE0265A12F8F7" blockId="27.[189,310,1842,1868]" box="[189,310,1842,1868]" pageId="27" pageNumber="28">
<heading id="FE6AE92B5F5FFFD4FF5BE0265A12F8F7" bold="true" box="[189,310,1842,1868]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="97E982555F5FFFD4FF5BE0265A12F8F7" bold="true" box="[189,310,1842,1868]" pageId="27" pageNumber="28">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A5225E475F5FFFD4FF5BE0755A8FF85F" blockId="27.[189,1399,1889,2020]" pageId="27" pageNumber="28">This species can easily be distinguished from other members of this species group by the black pronotum, with yellowish fringed margins; black elytra, with very slender yellowish stripe on each humeral side; aedeagus with dorsal plate broad and subtruncated; apex of laterophyse far from apical margin of dorsal plate in lateral view.</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F58FFD3FF5BE61F5AFDFECD" pageId="28" pageNumber="29" type="etymology">
<paragraph id="A5225E475F58FFD3FF5BE61F5A60FE9E" blockId="28.[189,324,267,293]" box="[189,324,267,293]" pageId="28" pageNumber="29">
<heading id="FE6AE92B5F58FFD3FF5BE61F5A60FE9E" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="97E982555F58FFD3FF5BE61F5A60FE9E" bold="true" box="[189,324,267,293]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE62D5AFDFECD" blockId="28.[189,1399,313,375]" pageId="28" pageNumber="29">The specific epithet is dedicated to its collector, the late Dr. Kintaro Baba who was one of the great Japanese entomologists.</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F58FFD3FF5BE68A5837FDD4" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="A5225E475F58FFD3FF5BE68A5A48FE03" blockId="28.[189,364,414,440]" box="[189,364,414,440]" pageId="28" pageNumber="29">
<heading id="FE6AE92B5F58FFD3FF5BE68A5A48FE03" bold="true" box="[189,364,414,440]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="97E982555F58FFD3FF5BE68A5A48FE03" bold="true" box="[189,364,414,440]" pageId="28" pageNumber="29">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="15F5541A5F58FFD3FF0CE6D95FFDFDB1" ID-GBIF-Occurrence="2620522301" collectingDate="1986-05-05" collectionCode="NMNS" collectorName="K. Baba" country="Taiwan" location="Shi Nan Shan" municipality="Liu Kui" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="A5225E475F58FFD3FF0CE6D95A78FE5C" blockId="28.[189,1241,461,523]" box="[234,348,461,487]" pageId="28" pageNumber="29">
<emphasis id="97E982555F58FFD3FF0CE6D95A78FE5C" bold="true" box="[234,348,461,487]" pageId="28" pageNumber="29">
<typeStatus id="7A26E0E55F58FFD3FF0CE6D95A78FE5C" box="[234,348,461,487]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE6E45FFDFDB1" blockId="28.[189,1241,461,523]" box="[189,1241,496,523]" pageId="28" pageNumber="29">
<collectingCountry id="DD8A1ED75F58FFD3FF5BE6E45A17FDB1" box="[189,307,496,522]" name="Taiwan" pageId="28" pageNumber="29">TAIWAN</collectingCountry>
:
<specimenCount id="B39B95CE5F58FFD3FED8E6E45A71FDB1" box="[318,341,496,522]" pageId="28" pageNumber="29" type="male">♂</specimenCount>
,
<location id="A042089C5F58FFD3FE84E6E45923FDB1" LSID="urn:lsid:plazi:treatment:2D34EF515F5FFFD2FDE5E1BF5ABDFE7F:A042089C5F58FFD3FE84E6E45923FDB1" box="[354,519,496,522]" country="Taiwan" municipality="Liu Kui" name="Shi Nan Shan" pageId="28" pageNumber="29">Shi Nan Shan</location>
,
<location id="A042089C5F58FFD3FDF4E6E4598FFDB1" LSID="urn:lsid:plazi:treatment:2D34EF515F5FFFD2FDE5E1BF5ABDFE7F:A042089C5F58FFD3FDF4E6E4598FFDB1" box="[530,683,496,522]" country="Taiwan" municipality="Liu Kui" name="near Liu Kui" pageId="28" pageNumber="29">near Liu Kui</location>
,
<location id="A042089C5F58FFD3FD51E6E45803FDB1" LSID="urn:lsid:plazi:treatment:2D34EF515F5FFFD2FDE5E1BF5ABDFE7F:A042089C5F58FFD3FD51E6E45803FDB1" box="[695,807,496,522]" country="Taiwan" municipality="Liu Kui" name="S Taiwan" pageId="28" pageNumber="29">S Taiwan</location>
,
<date id="D12378875F58FFD3FCD5E6E458E7FDB1" box="[819,963,496,523]" pageId="28" pageNumber="29" value="1986-05-05">
<collectingDate id="C167816F5F58FFD3FCD5E6E458E7FDB1" box="[819,963,496,523]" pageId="28" pageNumber="29" value="1986-05-05">5 May 1986</collectingDate>
</date>
,
<collectorName id="08683B915F58FFD3FC29E6E45F0BFDB1" box="[975,1071,496,522]" pageId="28" pageNumber="29">K. Baba</collectorName>
leg. (
<collectionCode id="C38CC6825F58FFD3FB97E6E45FE8FDB1" box="[1137,1228,496,522]" collectionName="Taiwan [Republic of China], Taichung, National Museum of Natural Science" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="28" pageNumber="29">NMNS</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="A5225E475F58FFD3FF0CE5255A78FDF0" blockId="28.[189,787,561,623]" box="[234,348,561,587]" pageId="28" pageNumber="29">
<materialsCitation id="15F5541A5F58FFD3FF0CE5255A78FDF0" ID-GBIF-Occurrence="3763220416" box="[234,348,561,587]" collectionCode="EUMJ" country="Taiwan" location="TAIWAN" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="97E982555F58FFD3FF0CE5255A78FDF0" bold="true" box="[234,348,561,587]" pageId="28" pageNumber="29">
<typeStatus id="7A26E0E55F58FFD3FF0CE5255A78FDF0" box="[234,348,561,587]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE5415837FDD4" blockId="28.[189,787,561,623]" box="[189,787,596,623]" pageId="28" pageNumber="29">
<collectingCountry id="DD8A1ED75F58FFD3FF5BE5415A17FDD4" box="[189,307,597,623]" name="Taiwan" pageId="28" pageNumber="29">TAIWAN</collectingCountry>
:
<specimenCount id="B39B95CE5F58FFD3FED8E5415A4FFDD4" box="[318,363,597,623]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
, same data as for
<typeStatus id="7A26E0E55F58FFD3FDD8E5405981FDD5" box="[574,677,596,622]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(
<collectionCode id="C38CC6825F58FFD3FD52E5415823FDD4" box="[692,775,597,623]" collectionName="Japan, Matsuyama, Ehime University" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="28" pageNumber="29">EUMJ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F58FFD3FF5BE5825A1DF814" pageId="28" pageNumber="29" type="description">
<paragraph id="A5225E475F58FFD3FF5BE5825A6AFD0B" blockId="28.[189,334,662,688]" box="[189,334,662,688]" pageId="28" pageNumber="29">
<heading id="FE6AE92B5F58FFD3FF5BE5825A6AFD0B" bold="true" box="[189,334,662,688]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="97E982555F58FFD3FF5BE5825A6AFD0B" bold="true" box="[189,334,662,688]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A5225E475F58FFD3FF0CE5D05A55FD65" blockId="28.[189,1399,708,911]" box="[234,369,708,734]" pageId="28" pageNumber="29">
<emphasis id="97E982555F58FFD3FF0CE5D05A55FD65" bold="true" box="[234,369,708,734]" pageId="28" pageNumber="29">Coloration</emphasis>
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE5FC5837FC35" blockId="28.[189,1399,708,911]" pageId="28" pageNumber="29">Eyes and head black. Antennae dark brown to black. Mandibles yellowish brown. Pronotum black, fringed with yellowish margins. Elytra black, with very slender yellowish stripe on each humeral side. Legs black. Pro-, meso- and metasterna, as well as abdomen blackish brown, with margins of abdominal ventrites yellowish brown. Body closely covered with pale pubescence; anterior margin of clypeus, antennae, elytra and legs fringed with pale bristles.</paragraph>
<paragraph id="A5225E475F58FFD3FF0CE4A25AB8FC6B" blockId="28.[189,1399,950,1223]" box="[234,412,950,976]" pageId="28" pageNumber="29">
<emphasis id="97E982555F58FFD3FF0CE4A25A0EFC6B" bold="true" box="[234,298,950,976]" pageId="28" pageNumber="29">Male</emphasis>
(
<figureCitation id="3DA642C25F58FFD3FEDCE4A25AB0FC6B" box="[314,404,950,976]" captionStart="Fig" captionStartId="8.[189,232,1822,1848]" captionTargetBox="[188,1394,422,1777]" captionTargetId="figure@8.[149,1432,382,1818]" captionTargetPageId="8" captionText="Fig. 2. Habitus, dorsal view. A–B. Lycocerus pictus (Wittmer, 1983). C–E. L. yitingi Hsiao &amp; Okushima sp. nov. F–G. L. araticollis (Fairmaire, 1897). H–I. L. griseopubens (Pic, 1928). J–M. L. nigripennis (Pic, 1938). N–O. L. aurantiacus Hsiao &amp; Okushima sp. nov. P–Q. L. evangelium Hsiao &amp; Okushima sp. nov. R. L. kintaroi Hsiao &amp; Okushima sp. nov. C, F, H, J, N, P, R. Name-bearing types. D, O, Q. Paratypes. A–B, E, G, I, K–M. Non-type specimens. A, C, F–G, I, K–L, N, P, R. Male. B, D–E, H, J, M, O, Q. Female. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3832612" httpUri="https://zenodo.org/record/3832612/files/figure.png" pageId="28" pageNumber="29">Fig. 2R</figureCitation>
)
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE4CE5F37FB7C" blockId="28.[189,1399,950,1223]" pageId="28" pageNumber="29">HEAD. Nearly as long as wide; vertex faintly hollowed, depressed along apical margin of clypeus and in lateral areas before eyes; surface densely and finely punctate and lustrous; anterior margin of clypeus arcuate and faintly indented in middle; eyes not so large, globular and slightly prominent, ratio of eye diameter to interocular space 1:5.0; terminal labial palpomeres rounded ax-shaped; terminal maxillary palpomeres rounded ax-shaped; antennae filiform, extending to elytral midlength; antennomere I clavate, II short and a little expanded apicad, III to XI subcylindrical, all antennomeres without groove, ratio of antennomere lengths: 16: 10: 13.5:15.5: 15.5: 15: 14.5: 15: 14:13: 14.5.</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE3FA5E51FAC9" blockId="28.[189,1399,1261,1394]" pageId="28" pageNumber="29">PRONOTUM. Subquadrate, slightly narrower than head, about 0.95 times as long as wide; anterior and posterior margins moderately arcuate; lateral margins feebly sinuate; anterior angles rounded; posterior angles obtuse; dorsum convex in postero-lateral and hollowed in antero-lateral areas; medio-longitudinal groove distinct in posterior half; surface smooth and lustrous. Scutellum triangular, with rounded apex.</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE28D587AFA6D" blockId="28.[189,1399,1432,1494]" pageId="28" pageNumber="29">ELYTRA. Conjointly about 1.3 times as wide as pronotum, about 2.5 times as long as wide, sides subparallel; surface densely and finely punctate and lustrous.</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE2EA5A79F981" blockId="28.[189,1398,1533,1594]" pageId="28" pageNumber="29">LEGS. Moderately slender; femora mostly straight; tibiae mostly straight, with basal part feebly arcuate; claws simple.</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE1765908F8B2" blockId="28.[189,1399,1634,1801]" pageId="28" pageNumber="29">
AEDEAGUS (
<figureCitation id="3DA642C25F58FFD3FEA7E1765AA6F9C6" box="[321,386,1634,1661]" captionStart="Fig" captionStartId="16.[190,233,1956,1982]" captionTargetBox="[194,1497,270,1908]" captionTargetId="figure@16.[154,1506,264,1921]" captionTargetPageId="16" captionText="Fig. 5. Aedeagus. A–C. Lycocerus evangelium Hsiao &amp; Okushima sp. nov. D–F. L. kintaroi Hsiao &amp; Okushima sp. nov. A, D. Ventral view. B, E. Lateral view. C, F. Dorsal view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3832618" httpUri="https://zenodo.org/record/3832618/files/figure.png" pageId="28" pageNumber="29">Fig. 5</figureCitation>
D–F). Ventral process moderately expanded and rounded apically, apex curved inwards, basal part broad and extending inwards onto ventral side; dorsal plate of each paramere as long as ventral process, broad and subtruncated, apical margin with some hairs on it, concave on inner margin. Laterophyse very short with obtuse apex, invisible in lateral view. Inner sac swollen, lengthened behind, somewhat shorter than tegmen.
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE0245FA3F8F0" blockId="28.[189,1159,1840,1867]" box="[189,1159,1840,1867]" pageId="28" pageNumber="29">
Body length 5.50–6.00 mm (
<typeStatus id="7A26E0E55F58FFD3FDE8E024595FF8F1" box="[526,635,1840,1866]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
:
<quantity id="6265F3A25F58FFD3FD60E02459D0F8F0" box="[646,756,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="28" pageNumber="29" unit="mm" value="5.5">5.50 mm</quantity>
); width
<quantity id="6265F3A25F58FFD3FCB4E02458D8F8F0" box="[850,1020,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="28" pageNumber="29" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.50–1.70 mm</quantity>
(
<quantity id="6265F3A25F58FFD3FBEAE0245F5FF8F0" box="[1036,1147,1840,1867]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="mm" value="1.5">1.50 mm</quantity>
).
</paragraph>
<paragraph id="A5225E475F58FFD3FF0CE0665A61F837" blockId="28.[189,325,1906,1967]" box="[234,325,1906,1932]" pageId="28" pageNumber="29">
<emphasis id="97E982555F58FFD3FF0CE0665A61F837" bold="true" box="[234,325,1906,1932]" pageId="28" pageNumber="29">Female</emphasis>
</paragraph>
<paragraph id="A5225E475F58FFD3FF5BE0815A1DF814" blockId="28.[189,325,1906,1967]" box="[189,313,1941,1967]" pageId="28" pageNumber="29">Unknown.</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F59FFD2FF5BE61F59EEFEE8" box="[189,714,267,340]" pageId="29" pageNumber="30" type="discussion">
<paragraph id="A5225E475F59FFD2FF5BE61F5A0BFE9E" blockId="29.[189,303,267,293]" box="[189,303,267,293]" pageId="29" pageNumber="30">
<heading id="FE6AE92B5F59FFD2FF5BE61F5A0BFE9E" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="97E982555F59FFD2FF5BE61F5A0BFE9E" bold="true" box="[189,303,267,293]" pageId="29" pageNumber="30">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="A5225E475F59FFD2FF5BE62D59EEFEE8" blockId="29.[189,714,313,340]" box="[189,714,313,340]" pageId="29" pageNumber="30">
The right antenna of the
<typeStatus id="7A26E0E55F59FFD2FE38E62D5961FEE8" box="[478,581,313,339]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
is missing.
</paragraph>
</subSubSection>
<subSubSection id="ED870DCC5F59FFD2FF5BE66F5ABDFE7F" pageId="29" pageNumber="30" type="distribution">
<paragraph id="A5225E475F59FFD2FF5BE66F5A92FE2E" blockId="29.[189,438,379,405]" box="[189,438,379,405]" pageId="29" pageNumber="30">
<emphasis id="97E982555F59FFD2FF5BE66F5A72FE2E" bold="true" box="[189,342,379,405]" pageId="29" pageNumber="30">Distribution</emphasis>
(
<figureCitation id="3DA642C25F59FFD2FE80E66F5A88FE2E" box="[358,428,379,405]" captionStart="Fig" captionStartId="22.[189,232,1964,1990]" captionTargetBox="[229,1359,527,1924]" captionTargetId="figure@22.[229,1359,527,1924]" captionTargetPageId="22" captionText="Fig. 8. Distribution map of L. nigripennis (Pic, 1938), L. evangelium Hsiao &amp; Okushima sp. nov., L. aurantiacus Hsiao &amp; Okushima sp. nov. and L. kintaroi Hsiao &amp; Okushima sp. nov. from Taiwan." figureDoi="http://doi.org/10.5281/zenodo.3832624" httpUri="https://zenodo.org/record/3832624/files/figure.png" pageId="29" pageNumber="30">Fig. 8</figureCitation>
)
</paragraph>
<paragraph id="A5225E475F59FFD2FF5BE6BD5ABDFE7F" blockId="29.[189,409,425,452]" box="[189,409,425,452]" pageId="29" pageNumber="30">
<collectingCountry id="DD8A1ED75F59FFD2FF5BE6BD5A37FE78" box="[189,275,425,451]" name="Taiwan" pageId="29" pageNumber="30">Taiwan</collectingCountry>
(endemic).
</paragraph>
</subSubSection>
</treatment>
</document>
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