261 lines
21 KiB
XML
261 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.145.1713" ID-GBIF-Dataset="cc9664ec-527b-4315-b69c-7392f72f096a" ID-PMC="PMC3267456" ID-Pensoft-Pub="1313-2970-145-129" ID-PubMed="22287886" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-145-129" ModsDocOrigin="ZooKeys 145" ModsDocTitle="Two new species of Harpactea (Araneae, Dysderidae) from Turkey" checkinTime="1451249699236" checkinUser="pensoft" docAuthor="Kunt, Kadir Bogac, Elverici, Mert, Oezkuetuek, Recep Sulhi & Yagmur, Ersen Aydin" docDate="2011" docId="967F490DAF29022A0D357D6847FECB20" docLanguage="en" docName="ZooKeys 145: 129-141" docOrigin="ZooKeys 145" docSource="http://dx.doi.org/10.3897/zookeys.145.1713" docTitle="Harpactea arnedoi Kunt, Elverici, Oezkuetuek & Yagmur, 2011, sp. n." docType="treatment" docVersion="4" lastPageNumber="131" masterDocId="E226FF82FFCA6557FF847A37FFF8F833" masterDocTitle="Two new species of Harpactea (Araneae, Dysderidae) from Turkey" masterLastPageNumber="141" masterPageNumber="129" pageNumber="130" updateTime="1668152571754" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of Harpactea (Araneae, Dysderidae) from Turkey</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kunt, Kadir Bogac</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Elverici, Mert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oezkuetuek, Recep Sulhi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yagmur, Ersen Aydin</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>145</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>129</mods:start>
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<mods:end>141</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.145.1713</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.145.1713</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-145-129</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031866" LSID="urn:lsid:zoobank.org:act:8BC9D2B7-4E3D-4A17-8017-6DAB08F82D8F" httpUri="http://treatment.plazi.org/id/967F490DAF29022A0D357D6847FECB20" lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130">
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<subSubSection pageId="1" pageNumber="130" type="nomenclature">
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<paragraph pageId="1" pageNumber="130">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:8BC9D2B7-4E3D-4A17-8017-6DAB08F82D8F" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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<taxonomicNameLabel pageId="1" pageNumber="130">sp. n.</taxonomicNameLabel>
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Figs 28
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="130" type="material examined">
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<paragraph pageId="1" pageNumber="130">Material examined.</paragraph>
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<paragraph pageId="1" pageNumber="130">
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Holotype ♂ (AUZM), Turkey, Gaziantep Province,
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<normalizedToken originalValue="Kuşçubeli">Kuscubeli</normalizedToken>
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Pass [
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="37.11394">37°6'50.20"N</geoCoordinate>
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;
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="36.609497">36°36'34.20"E</geoCoordinate>
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], 13.XI.2010, under leaf litter, leg. E.A.
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<normalizedToken originalValue="Yağmur">Yagmur</normalizedToken>
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. Paratypes 1 ♀, 1 ♂ (AUZM); 2 ♀♀, 2 ♂♂, 9 juveniles (cKBK & SMF), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="130" type="derivatio nominis">
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<paragraph pageId="1" pageNumber="130">Derivatio nominis.</paragraph>
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<paragraph pageId="1" pageNumber="130">
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The new species is dedicated to the Spanish arachnologist, Dr. Miquel Arnedo (Barcelona University, Barcelona, Spain), who has made important contributions to the taxonomy of the family
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<taxonomicName family="Dysderidae" lsidName="" pageId="1" pageNumber="130" rank="family">Dysderidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="130" type="diagnosis">
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<paragraph pageId="1" pageNumber="130">Diagnosis.</paragraph>
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<paragraph pageId="1" pageNumber="130">
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. differs from all other
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Harpactea</taxonomicName>
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species in the structure of the copulatory organs. However, the general morphology of the male palp resembles that of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea zoiai" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="zoiai">Harpactea zoiai</taxonomicName>
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Gasparo, 1999 known from Greece (see
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<bibRefCitation author="Gasparo, F" journalOrPublisher="Bollettino Museo Regionale di Scienze Naturali Torino" pageId="5" pageNumber="134" pagination="303 - 308" title="Descrizione di Harpactea zoiai n. sp. del Peloponneso meridionale (Araneae, Dysderidae)." volume="16" year="1999">Gasparo 1999</bibRefCitation>
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). The male of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. differs in having an uneven spherical shape of the palpal bulb; the region between the bulb and the distal continuation is elongated and has a funnel-like appearance; the embolus is shorter, hook-like and bends downwards towards the tip. The vulva is apparent and characterized by a peripherally sclerotized posterior diverticulumand folded distal expansion.
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</paragraph>
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<caption pageId="1" pageNumber="130">
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<paragraph pageId="1" pageNumber="130">
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Figures 2-6.
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. 2 Cheliceral teeth 3 left palp, retrolateral view 4 ditto, nearly anterior view 5 ditto, nearly retrolateral view 6 ditto, distal view. Scale line: 0.5 mm.
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</paragraph>
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</caption>
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<caption pageId="1" pageNumber="130">
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<paragraph pageId="1" pageNumber="130">
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Figures 7-8. Vulva of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. 7 dorsal view 8 ventral view. Scale lines: 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="130" type="notes">
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<paragraph pageId="1" pageNumber="130">Notes.</paragraph>
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<paragraph pageId="1" pageNumber="130">
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While describing
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n., it was difficult to decide whether it would be best assigned to
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Harpactea</taxonomicName>
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or to
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Stalagtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stalagtia" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Stalagtia</taxonomicName>
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. By the characteristic structure of the male copulatory organ (globular tegulum, long embolus, poorly developed conductor), the new species is similar to members of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Stalagtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stalagtia" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Stalagtia</taxonomicName>
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. However, the male palp possesses a poorly developed conductor (Fig. 4), and the female copulatory organs are similar to those of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Harpactea</taxonomicName>
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species in having a short posterior diverticulum and anterior spermathecae. Thus, we feel the new species is correctly placed in
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Harpactea</taxonomicName>
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. Further evidence in support of this generic placement is the absence of ventral spines on the anterior tibiae and metatarsi in the new species, particularly as
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<bibRefCitation author="Deeleman-Reinhold, CL" journalOrPublisher="Revue Arachnologique" pageId="5" pageNumber="134" pagination="105 - 135" title="The genus Rhode and the harpacteine genera Stalagtia, Folkia, Minotauria, and Kaemis (Araneae, Dysderidae) of Yugoslavia nad Crete, with remark on the genus Harpactea." volume="10" year="1993">Deeleman-Reinhold (1993)</bibRefCitation>
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stated that their presence was diagnostic of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Stalagtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stalagtia" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Stalagtia</taxonomicName>
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. However, the only known Turkish species of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Stalagtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stalagtia" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="genus">Stalagtia</taxonomicName>
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,
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Stalagtia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stalagtia thaleriana" order="Araneae" pageId="1" pageNumber="130" phylum="Arthropoda" rank="species" species="thaleriana">Stalagtia thaleriana</taxonomicName>
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Chatzaki & Arnedo, 2006, does not possess ventral spines on the anterior tibiae and metatarsi, as was mentioned earlier by
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<bibRefCitation author="Chatzaki, M" journalOrPublisher="Zootaxa" pageId="5" pageNumber="134" pagination="1 - 32" title="Taxonomic revision of the epigean representatives of the spider subfamily Harpacteinae (Araneae: Dysderidae) on the island of Crete." volume="1169" year="2006">Chatzaki and Arnedo (2006)</bibRefCitation>
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and
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<bibRefCitation author="Kunt, KB" journalOrPublisher="Biological Diversity and conservation" pageId="5" pageNumber="134" pagination="45 - 49" title="A new record for spider fauna of Turkey, Stalagtia thaleriana Chatzaki & Arnedo, 2006 (Araneae; Dysderidae)." volume="2" year="2009">Kunt et al. (2009)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130" type="description">
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<paragraph pageId="1" pageNumber="130">Description.</paragraph>
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<paragraph lastPageId="2" lastPageNumber="131" pageId="1" pageNumber="130">
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Measurements: [Holotype ♂ / Paratype ♀]: AL 2.26 / 2.50; CL 1.75 / 1.73; CWmax 1.45 / 1.35; CWmin 0.63 / 0.65; AMEd 0.10 / 0.11; PLEd 0.09 /
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<pageBreakToken pageId="2" pageNumber="131" start="start">0.09</pageBreakToken>
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; PMEd 0.08 / 0.08; ChF 0.37 / 0.38; ChG 0.18 / 0.19; ChL 0.62 / 0.64 mm. Leg measurements are given in Table 1.
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</paragraph>
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<caption pageId="2" pageNumber="131">
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<paragraph pageId="2" pageNumber="131">
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Table 1. Leg measurements of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n.<br/>
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="131">
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<table pageId="2" pageNumber="131">
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">(Holotype ♂ / Paratype ♀)</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Fe</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Pa</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Ti</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Me</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Ta</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg I</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg II</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg III</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg IV</td>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="2" pageNumber="131">
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MALE: Small sized spider. Carapace greenish - light brown, with smooth surface and distinct fovea. AME, PLE and PME closely grouped; AME separated. Sternum, labium, gnathocoxae and chelicerae light brown. Sternum dark brown laterally, with thin, long hair near margins. Cheliceral groove with four teeth: retromargin with two tiny teeth; promargin with two strong teeth, largest tooth closest to base of the cheliceral groove (Fig. 2). Top of the labium and gnathocoxae with short, strong hair, sparsely distributed. Abdomen greyish-light brown, with short, thin blackish hair over the entire surface. Legs yellowish-light brown with sparse blackish setae. Ventral surface of coxae with long, thin, black sparse hair. Leg IV> Leg I> Leg II> Leg III. Tarsi with three claws. Tarsi III and IV with fine scopulae. Legs III and IV with fine metatarsal scopulae covering slightly less than the distal half of the segment (ventral surface only). Prolateral part of coxae III and IV with 1 spine. Dorsal parts of femora, tibiae and metatarsi with spines. Number of spines can vary among individuals. Detailed leg spination of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. is given in Table 2.
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</paragraph>
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<caption pageId="2" pageNumber="131">
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<paragraph pageId="2" pageNumber="131">
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Table 2. Leg spination of
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n.<br/>
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="131">
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<table pageId="2" pageNumber="131">
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">(Holotype)</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg I</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg II</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg III</td>
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Leg IV</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">C</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Fe</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Pa</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Ti</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Me</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">(Paratype)</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">C</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Fe</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Pa</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Ti</td>
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</tr>
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<tr pageId="2" pageNumber="131">
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<td colspan="1" pageId="2" pageNumber="131" rowspan="1">Me</td>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="2" pageNumber="131">Palpal tarsus of the male covered with thin and elongated setae. Tegulum yellow, lighter than the legs. Bulb almost spherical. The anteroventral region of the bulb has a chitinized edge. Between bulb and distal appendages there is a neck-shaped transition region (Figs 3-5). This region is lightly chitinized and dusky in patches. From the base of the embolus, conductor and accessory apophysis have dark brown tips. Embolus is slender and cylindrical up to its hook-like tip and it is not so heavily sclerotized. At the periphery it is membranous and projects downwards, parallel to the palpal tibia. Conductor and accessory apophysis are developed at the end of the neck-shaped transition region, on the opposite tips of an ear-shaped structure. Both are triangular in shape, whereas the accessory apophysis is longer and jagged (Fig. 6). The relative positions of the distal appendages are most easily seen at a 90 degree angle in ventral view. Embolus separated from conductor and accessory apophysis by a broad base, at first it follows the course of the conductor, but later bends with a sharp curve and accompanies the accessory apophysis.</paragraph>
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<paragraph pageId="2" pageNumber="131">FEMALE: No differences found between male and female, in terms of body colour and morphology. Vulva sclerotized almost uniformly. Distal crest long and butt-ended at the tip. Distal expansion of spermatheca convoluted. Rod-shaped part of the anterior spermatheca short and cylindrical. Basal transverse part of the anterior spermatheca separates from rod-shaped part laterally at an acute angle and forms a triangular shape. Anterior basal arc lies linearly through the centre and periphery, but widens at the edges. Transverse bar short and crescent-shaped with tips turned downwards. Posterior diverticulum shaped as a membranous sac and is sclerotized at the periphery (Figs 7, 8).</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="131" type="ecology">
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<paragraph pageId="2" pageNumber="131">Ecology.</paragraph>
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<paragraph pageId="2" pageNumber="131">
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Specimens were collected in
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<normalizedToken originalValue="Kuşçubeli">Kuscubeli</normalizedToken>
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Pass, located in the Amanos Mountains, from habitats covered by Turkish Pine (
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<taxonomicName class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus brutia" order="Pinales" pageId="2" pageNumber="131" phylum="Tracheophyta" rank="species" species="brutia">Pinus brutia</taxonomicName>
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) and scrub type oak forests (
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<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus infectoria" order="Fagales" pageId="2" pageNumber="131" phylum="Tracheophyta" rank="species" species="infectoria">Quercus infectoria</taxonomicName>
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). A variety of herbaceous plants and low shrubs such as
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<taxonomicName class="Liliopsida" family="Asparagaceae" genus="Ruscus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ruscus aculeatus" order="Asparagales" pageId="2" pageNumber="131" phylum="Tracheophyta" rank="species" species="aculeatus">Ruscus aculeatus</taxonomicName>
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are also widely represented in those forests. Sampling was done by sifting leaf litter during the early winter.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="131" type="distribution">
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<paragraph pageId="2" pageNumber="131">Distribution.</paragraph>
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<paragraph pageId="2" pageNumber="131">
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<taxonomicName class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea arnedoi" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="arnedoi">Harpactea arnedoi</taxonomicName>
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sp. n. is known from thetype locality only.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |