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<document id="D163100BEEF853C7BAC6C96CADF82C7F" ID-DOI="10.11646/zootaxa.3980.1.8" ID-GBIF-Dataset="a5e69751-ca89-4d63-b089-9b91161d134f" ID-ISSN="1175-5326" ID-Zenodo-Dep="234472" ID-ZooBank="F6681D7E-0C35-4366-AEC5-3D92F0B0D772" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461227405397" checkinUser="plazi" docAuthor="Ashman, Lauren G., Oberprieler, Rolf G. &amp; Ślipiński, Adam" docDate="2015" docId="8D7087C6FFBF1042CD930E308AF8FE45" docLanguage="en" docName="zt03980p142.pdf" docOrigin="Zootaxa 3980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhopalomma Ashman, Oberprieler &amp; Slipinski, gen. n." docType="treatment" docVersion="7" lastPageNumber="138" masterDocId="7149FFBEFFBE1040CD040A2A8842FFCD" masterDocTitle="Rhopalomma stefaniae gen. et sp. n., the first ommatid beetle from the Upper Jurassic in Australia (Coleoptera: Archostemata: Ommatidae)" masterLastPageNumber="142" masterPageNumber="136" pageNumber="137" updateTime="1698604906761" updateUser="plazi">
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<mods:title id="E47BF6D765A5EDCFA1B4E5117CCB12CB">Rhopalomma stefaniae gen. et sp. n., the first ommatid beetle from the Upper Jurassic in Australia (Coleoptera: Archostemata: Ommatidae)</mods:title>
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<mods:namePart id="88440BA1AEB457A7BCBD2FDD3D62D35F">Ashman, Lauren G.</mods:namePart>
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<mods:namePart id="A24CF02784559CE2A3A38F83FC718455">Oberprieler, Rolf G.</mods:namePart>
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<mods:namePart id="B61D7E932AF2AA458B72B45ECB47FDC3">Ślipiński, Adam</mods:namePart>
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<mods:title id="86F82137BB8A899F2FBE38AEB3497259">Zootaxa</mods:title>
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<mods:date id="490843C5CA674B18E7524DE543CCA605">2015</mods:date>
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<mods:number id="7C02346CDCEA5B7D0F69FE9888C91229">3980</mods:number>
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<mods:identifier id="17A43C53A595274FC75B0DAC5B2ABDFA" type="DOI">10.11646/zootaxa.3980.1.8</mods:identifier>
<mods:identifier id="069AD2610CA76BE474C4125EE86D8EB8" type="GBIF-Dataset">a5e69751-ca89-4d63-b089-9b91161d134f</mods:identifier>
<mods:identifier id="228CFEB006F064805136AC8812D65930" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7BE390429B95464299E020A7B32609E0" type="Zenodo-Dep">234472</mods:identifier>
<mods:identifier id="522E410CFDDDD97039A2B205BCFDF0F9" type="ZooBank">F6681D7E-0C35-4366-AEC5-3D92F0B0D772</mods:identifier>
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<treatment id="8D7087C6FFBF1042CD930E308AF8FE45" ID-DOI="http://doi.org/10.5281/zenodo.6096489" ID-GBIF-Taxon="119632858" ID-Zenodo-Dep="6096489" LSID="urn:lsid:plazi:treatment:8D7087C6FFBF1042CD930E308AF8FE45" httpUri="http://treatment.plazi.org/id/8D7087C6FFBF1042CD930E308AF8FE45" lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137">
<subSubSection id="4DC3655BFFBF1041CD930E308B79FBF8" box="[151,827,1050,1077]" pageId="1" pageNumber="137" type="nomenclature">
<paragraph id="056636D0FFBF1041CD930E308B79FBF8" blockId="1.[151,827,1050,1077]" box="[151,827,1050,1077]" pageId="1" pageNumber="137">
<heading id="5E2E81BCFFBF1041CD930E308B79FBF8" bold="true" box="[151,827,1050,1077]" fontSize="11" level="1" pageId="1" pageNumber="137" reason="1">
<emphasis id="37ADEAC2FFBF1041CD930E308B79FBF8" bold="true" box="[151,827,1050,1077]" pageId="1" pageNumber="137">
<taxonomicName id="C2D94D53FFBF1041CD930E308AA0FBF8" authority="Ashman, Oberprieler &amp; Slipinski" authorityName="Ashman, Oberprieler &amp; Slipinski" box="[151,738,1050,1077]" class="Insecta" family="Ommatidae" genus="Rhopalomma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="37ADEAC2FFBF1041CD930E308978FBF9" bold="true" box="[151,314,1050,1076]" italics="true" pageId="1" pageNumber="137">Rhopalomma</emphasis>
Ashman, Oberprieler &amp; Ślipiński
</taxonomicName>
<taxonomicNameLabel id="2C9E57B9FFBF1041CFED0E318B79FBF8" box="[745,827,1051,1077]" pageId="1" pageNumber="137" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4DC3655BFFBF1041CD930E4B8BE5FBBA" box="[151,935,1121,1143]" pageId="1" pageNumber="137" type="materials_examined">
<paragraph id="056636D0FFBF1041CD930E4B8BE5FBBA" blockId="1.[151,935,1121,1143]" box="[151,935,1121,1143]" pageId="1" pageNumber="137">
<emphasis id="37ADEAC2FFBF1041CD930E4B8964FBBA" bold="true" box="[151,294,1121,1143]" pageId="1" pageNumber="137">
<typeStatus id="DA628872FFBF1041CD930E4B888EFBBA" box="[151,204,1121,1143]" pageId="1" pageNumber="137">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="C2D94D53FFBF1041CC2F0E4B8B28FBBA" authority="Ashman, Oberprieler &amp; Slipinski" authorityName="Ashman, Oberprieler &amp; Slipinski" box="[299,874,1121,1143]" class="Insecta" family="Ommatidae" genus="Rhopalomma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="species" species="stefaniae" status="sp. nov.">
<emphasis id="37ADEAC2FFBF1041CC2F0E4B8A57FBBA" box="[299,533,1121,1143]" italics="true" pageId="1" pageNumber="137">Rhopalomma stefaniae</emphasis>
Ashman, Oberprieler &amp; Ślipiński
</taxonomicName>
<taxonomicNameLabel id="2C9E57B9FFBF1041CE750E488BE5FBBA" box="[881,935,1122,1143]" pageId="1" pageNumber="137" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="4DC3655BFFBF1041CD930E8E8B84F9CD" pageId="1" pageNumber="137" type="description">
<paragraph id="056636D0FFBF1041CD930E8E8B84F9CD" blockId="1.[151,1436,1188,2005]" pageId="1" pageNumber="137">
<emphasis id="37ADEAC2FFBF1041CD930E8E896BFB70" bold="true" box="[151,297,1188,1213]" pageId="1" pageNumber="137">Description.</emphasis>
Body elongate-oval, 2.5 times longer than wide (
<figureCitation id="9DE22A55FFBF1041CE6D0E8E8B98FB71" box="[873,986,1188,1213]" captionStart="FIGURES 1 – 2" captionStartId="2.[151,264,1555,1577]" captionTargetBox="[151,1436,447,1531]" captionTargetId="figure@2.[151,1436,447,1531]" captionTargetPageId="2" captionText="FIGURES 1 – 2. Rhopalomma stefaniae gen. et sp. n. 1. part (AM F. 139992); 2. counterpart (AM F. 139993). Scale bars are 1 mm." httpUri="https://zenodo.org/record/234473/files/figure.png" pageId="1" pageNumber="137">Figs. 1, 2</figureCitation>
,
<figureCitation id="9DE22A55FFBF1041CEED0E8E8C5AFB70" box="[1001,1048,1188,1213]" captionStart="FIGURES 5 – 6" captionStartId="4.[151,264,1611,1633]" captionTargetBox="[211,1371,429,1569]" captionTargetId="figure@4.[199,1388,411,1590]" captionTargetPageId="4" captionText="FIGURES 5 – 6. Rhopalomma stefaniae gen. et sp. n. line drawings. 5. part (AM F. 139992); 6. counterpart (AM F. 139993)." httpUri="https://zenodo.org/record/234475/files/figure.png" pageId="1" pageNumber="137">5, 6</figureCitation>
). Head slightly longer than wide; clypeus not distinctly separated from frons; short labrum indicated; antennae laterally inserted, 11-segmented, slightly shorter than head, last three antennomeres enlarged to form a slight club (
<figureCitation id="9DE22A55FFBF1041C93E0EC68CC3FAC9" box="[1082,1153,1260,1285]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1528,1550]" captionTargetBox="[151,1436,772,1506]" captionTargetId="figure@3.[151,1436,772,1507]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Rhopalomma stefaniae gen. et sp. n., details of head and prothorax. 3. part (AM F. 139992); 4. counterpart (AM F. 139993), after further exposure of structures, with antennal club and pronotum labelled. Scale bars are 1 mm." httpUri="https://zenodo.org/record/234474/files/figure.png" pageId="1" pageNumber="137">Fig. 4</figureCitation>
). Prothorax subquadrate; pronotum with anterior margin straight, posterior margin arcuate with distinct ridge (
<figureCitation id="9DE22A55FFBF1041C94D0F3A8CCFFAE5" box="[1097,1165,1296,1321]" captionStart="FIGURES 3 – 4" captionStartId="3.[151,264,1528,1550]" captionTargetBox="[151,1436,772,1506]" captionTargetId="figure@3.[151,1436,772,1507]" captionTargetPageId="3" captionText="FIGURES 3 – 4. Rhopalomma stefaniae gen. et sp. n., details of head and prothorax. 3. part (AM F. 139992); 4. counterpart (AM F. 139993), after further exposure of structures, with antennal club and pronotum labelled. Scale bars are 1 mm." httpUri="https://zenodo.org/record/234474/files/figure.png" pageId="1" pageNumber="137">Fig. 3</figureCitation>
). Prothoracic legs short, stout; femora broad, tibiae narrowly cylindrical. Scutellar shield small, subtriangular. Elytra flat dorsally, fairly abruptly sloping laterally, rounded apically, sides expanded into narrow, complete flanges of approximately uniform width; epipleura complete. Elytra clathrate, each with 10 complete striae of deep cell-like punctures extending from base to apex, and a scutellary striole with 3 punctures; striae 1–6 on disc, with approximately 30– 34 cells each, striae 7–10 on lateral slope; odd intervals 1, 3, 5 and 7 raised, almost as broad as strial punctures and twice as broad as even intervals; intervals 3 and 5 confluent before apex.
</paragraph>
</subSubSection>
<subSubSection id="4DC3655BFFBF1041CDC30C268C2AF984" pageId="1" pageNumber="137" type="etymology">
<paragraph id="056636D0FFBF1041CDC30C268C2AF984" blockId="1.[151,1436,1188,2005]" pageId="1" pageNumber="137">
<emphasis id="37ADEAC2FFBF1041CDC30C26890CF9E8" bold="true" box="[199,334,1548,1573]" pageId="1" pageNumber="137">Etymology.</emphasis>
The generic name is a combination of the Greek noun
<emphasis id="37ADEAC2FFBF1041CEDC0C268C79F9E8" box="[984,1083,1548,1573]" italics="true" pageId="1" pageNumber="137">rhopalos</emphasis>
(club, referring to the slightly clubbed antennae) and
<taxonomicName id="C2D94D53FFBF1041CC9E0C1B89A7F985" box="[410,485,1585,1608]" class="Insecta" family="Ommatidae" genus="Omma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis id="37ADEAC2FFBF1041CC9E0C1B89A7F985" box="[410,485,1585,1608]" italics="true" pageId="1" pageNumber="137">Omma</emphasis>
</taxonomicName>
(the
<typeStatus id="DA628872FFBF1041CF1A0C1B8A0CF984" box="[542,590,1585,1609]" pageId="1" pageNumber="137">type</typeStatus>
genus of the family) and its gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="4DC3655BFFBF1042CDC30C7E8AF8FE45" lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137" type="discussion">
<paragraph id="056636D0FFBF1041CDC30C7E8C66F931" blockId="1.[151,1436,1188,2005]" pageId="1" pageNumber="137">
<emphasis id="37ADEAC2FFBF1041CDC30C7E8979F9A0" bold="true" box="[199,315,1620,1645]" pageId="1" pageNumber="137">Remarks.</emphasis>
We assign
<emphasis id="37ADEAC2FFBF1041CCC70C7E8A1AF9A0" box="[451,600,1620,1645]" italics="true" pageId="1" pageNumber="137">Rhopalomma</emphasis>
to Archostemata based on several characters: the closely placed convex tubercles on the cuticular pieces of the pronotum, head and elytral margins; the complete epipleura; the ten striae of cell-like punctures on the elytra and the strongly developed scutellary strioles. Of the families of Archostemata, we place
<emphasis id="37ADEAC2FFBF1041CDDD0CEA892CF914" box="[217,366,1728,1753]" italics="true" pageId="1" pageNumber="137">Rhopalomma</emphasis>
in
<taxonomicName id="C2D94D53FFBF1041CC970CEA8A54F914" box="[403,534,1728,1753]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="family">Ommatidae</taxonomicName>
because of the relatively short antennae (not longer than the head and prothorax together) and their lateral insertion (
<bibRefCitation id="61484B21FFBF1041CF2E0CCE8A91F930" author="Crowson" box="[554,723,1764,1789]" pageId="1" pageNumber="137" refString="Crowson, R. A. (1962) Observations on the beetle family Cupedidae, with descriptions of two new fossil forms and a key to the recent genera. Annals and Magazine of Natural History, Series 13, 5, 147 - 157. http: // dx. doi. org / 10.1080 / 00222936208651227" type="journal article" year="1962">Crowson 1962</bibRefCitation>
;
<bibRefCitation id="61484B21FFBF1041CFDB0CCF8C5BF931" author="Lawrence" box="[735,1049,1764,1789]" pageId="1" pageNumber="137" refString="Lawrence, J. F. &amp; Slipinski, A. (2013) Australian Beetles: Morphology, Classification and Keys. CSIRO publishing, Collingwood, Australia, 561 pp." type="book" year="2013">Lawrence &amp; Ślipiński 2013</bibRefCitation>
).
</paragraph>
<paragraph id="056636D0FFBF1042CDC30D238D3CFF18" blockId="1.[151,1436,1188,2005]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="138" pageId="1" pageNumber="137">
Due to the poor preservation and incompleteness of the specimen, we cannot confidently assign
<emphasis id="37ADEAC2FFBF1041C8020D228DD9F8EC" box="[1286,1435,1800,1825]" italics="true" pageId="1" pageNumber="137">Rhopalomma</emphasis>
to a subfamily or tribe of
<taxonomicName id="C2D94D53FFBF1041CCB20D068A7EF888" box="[438,572,1836,1861]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="family">Ommatidae</taxonomicName>
, and therefore we classify it as
<taxonomicName id="C2D94D53FFBF1041CE990D068C62F888" box="[925,1056,1836,1861]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="family">Ommatidae</taxonomicName>
<emphasis id="37ADEAC2FFBF1041C9230D078C80F888" box="[1063,1218,1836,1861]" italics="true" pageId="1" pageNumber="137">incertae sedis</emphasis>
. One key character distinguishing
<taxonomicName id="C2D94D53FFBF1041CC3A0D7A8983F8A4" box="[318,449,1872,1897]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="subFamily" subFamily="Ommatinae">Ommatinae</taxonomicName>
from the other two subfamilies (
<taxonomicName id="C2D94D53FFBF1041CE3D0D7A8BABF8A4" box="[825,1001,1872,1897]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="subFamily" subFamily="Tetraphalerinae">Tetraphalerinae</taxonomicName>
and Notocupedinae) is the absence of lateral carinae on the pronotum (
<bibRefCitation id="61484B21FFBF1041CF0E0D5E8AF7F840" author="Crowson" box="[522,693,1908,1933]" pageId="1" pageNumber="137" refString="Crowson, R. A. (1962) Observations on the beetle family Cupedidae, with descriptions of two new fossil forms and a key to the recent genera. Annals and Magazine of Natural History, Series 13, 5, 147 - 157. http: // dx. doi. org / 10.1080 / 00222936208651227" type="journal article" year="1962">Crowson 1962</bibRefCitation>
;
<bibRefCitation id="61484B21FFBF1041CFC50D5F8B33F841" author="Tan" box="[705,881,1908,1933]" pageId="1" pageNumber="137" refString="Tan, J., Wang, Y., Ren, D. &amp; Yang, X. (2012) New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae. BioMed Central Evolutionary Biology, 12, 113. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 12 - 113" type="journal article" year="2012">
Tan
<emphasis id="37ADEAC2FFBF1041CFF70D5F8B6EF840" box="[755,812,1908,1933]" italics="true" pageId="1" pageNumber="137">et al.</emphasis>
2012
</bibRefCitation>
); the lateral margins of the specimen’s pronotum are unclear, and hence we cannot use this character to undertake a further taxonomic placement. Furthermore, the crucial characters distinguishing
<taxonomicName id="C2D94D53FFBF1041CF0A0D968AD3F818" box="[526,657,1980,2005]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="subFamily" subFamily="Ommatinae">Ommatinae</taxonomicName>
from
<taxonomicName id="C2D94D53FFBF1041CFDD0D968BC8F818" box="[729,906,1980,2005]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="137" phylum="Arthropoda" rank="subFamily" subFamily="Tetraphalerinae">Tetraphalerinae</taxonomicName>
, the presence of antennal grooves on the head and of protibial grooves on the propleura (
<bibRefCitation id="61484B21FFBC1042CF6B0AB28B5AFF7D" author="Crowson" box="[623,792,151,176]" pageId="2" pageNumber="138" refString="Crowson, R. A. (1962) Observations on the beetle family Cupedidae, with descriptions of two new fossil forms and a key to the recent genera. Annals and Magazine of Natural History, Series 13, 5, 147 - 157. http: // dx. doi. org / 10.1080 / 00222936208651227" type="journal article" year="1962">Crowson 1962</bibRefCitation>
), are not visible on the specimen. However,
<emphasis id="37ADEAC2FFBC1042C8020ABD8DD9FF7D" box="[1286,1435,151,176]" italics="true" pageId="2" pageNumber="138">Rhopalomma</emphasis>
is more likely to belong in
<taxonomicName id="C2D94D53FFBC1042CCC10A968A0AFF18" box="[453,584,188,213]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="subFamily" subFamily="Ommatinae">Ommatinae</taxonomicName>
than
<taxonomicName id="C2D94D53FFBC1042CF820A968B75FF18" box="[646,823,188,213]" class="Insecta" family="Ommatidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="subFamily" subFamily="Tetraphalerinae">Tetraphalerinae</taxonomicName>
, as its head does not have raised tubercles or ridges.
</paragraph>
<paragraph id="056636D0FFBC1042CDC30AF58AF8FE45" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="138">
The pattern of the elytral striae of
<emphasis id="37ADEAC2FFBC1042CF4A0AF58AA1FF35" box="[590,739,223,248]" italics="true" pageId="2" pageNumber="138">Rhopalomma</emphasis>
is very similar to that of
<taxonomicName id="C2D94D53FFBC1042C9020ACB8D63FF35" authority="Ponomarenko" authorityName="Ponomarenko" box="[1030,1313,223,248]" class="Insecta" family="Ommatidae" genus="Notocupes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="37ADEAC2FFBC1042C9020ACB8C3EFF35" box="[1030,1148,225,248]" italics="true" pageId="2" pageNumber="138">Notocupes</emphasis>
Ponomarenko
</taxonomicName>
(and other Notocupedini), in that there are four prominent intervals on each elytron, with two striae of punctures in between, and the second and third prominent intervals are confluent before the elytral apex (
<bibRefCitation id="61484B21FFBC1042C9650B0D8DD3FE8D" author="Jarzembowski" box="[1121,1425,295,320]" pageId="2" pageNumber="138" refString="Jarzembowski, E. A., Wang, B., Zhang, H. &amp; Fang, Y. (2014) Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana. Cretaceous Research, 52, 431 - 439. http: // dx. doi. org / 10.1016 / j. cretres. 2014.03.006" type="journal article" year="2014">
Jarzembowski
<emphasis id="37ADEAC2FFBC1042C80A0B038D08FE8D" box="[1294,1354,295,320]" italics="true" pageId="2" pageNumber="138">et al.</emphasis>
2014
</bibRefCitation>
). However, the clubbed antennae and the lack of a ‘neck’ region on the head are characters unique to
<emphasis id="37ADEAC2FFBC1042C8020B668DD9FEA8" box="[1286,1435,332,357]" italics="true" pageId="2" pageNumber="138">Rhopalomma</emphasis>
and merit placing the specimen in its own genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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