treatments-xml/data/3B/42/DD/3B42DDB70A215D02894745755B5830D5.xml

<document ID-CLB-Dataset="9589" ID-DOI="http://dx.doi.org/10.3897/phytokeys.166.58831" ID-GBIF-Dataset="5e97bad8-e3d8-4cc1-b0f9-18eaad6eb913" ID-PMC="PMC7679347" ID-Pensoft-Pub="1314-2003-166-117" ID-Pensoft-UUID="561609DBFBAA57C79592B7CB04A5CF97" ID-PubMed="33239959" ModsDocID="1314-2003-166-117" checkinTime="1605365356784" checkinUser="pensoft" docAuthor="Chen, Jun-Tong, Huang, Xian-Han, Lv, Zhen-Yu, Kuang, Tian-Hui, Luo, Jian, Deng, Yun-Fei &amp; Deng, Tao" docDate="2020" docId="3B42DDB70A215D02894745755B5830D5" docLanguage="en" docName="PhytoKeys 166: 117-127" docOrigin="PhytoKeys 166" docSource="http://dx.doi.org/10.3897/phytokeys.166.58831" docTitle="Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng 2020, sp. nov." docType="treatment" docVersion="6" id="561609DBFBAA57C79592B7CB04A5CF97" lastPageNumber="117" masterDocId="561609DBFBAA57C79592B7CB04A5CF97" masterDocTitle="Strobilanthes sunhangii (Acanthaceae), a new species from Tibet, China" masterLastPageNumber="127" masterPageNumber="117" pageNumber="117" updateTime="1732940323979" updateUser="ExternalLinkService">
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<mods:title id="05B60069D3699318CF09A01658700B90">Strobilanthes sunhangii (Acanthaceae), a new species from Tibet, China</mods:title>
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<mods:namePart id="535EA908F4DA3AAE9C019063C5F893BC">Chen, Jun-Tong</mods:namePart>
<mods:affiliation id="B1F0B8534FE55E4870A9F610762E79FE">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:namePart id="E26EF5B75195336B9761D1629EDCD4AE">Huang, Xian-Han</mods:namePart>
<mods:affiliation id="644C192A8BE5D4A0B861EB73B5B75575">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China</mods:affiliation>
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<mods:namePart id="56E385064F5E29AA510BBBEC1C74802F">Lv, Zhen-Yu</mods:namePart>
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<mods:namePart id="FC93D70536F85B7756BE1B19D36C1E44">Kuang, Tian-Hui</mods:namePart>
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<mods:namePart id="3BD344469154435172DD5309415A0BBF">Luo, Jian</mods:namePart>
<mods:affiliation id="3C51783ED1EF482872846A05C99CC418">Institute of Plateau Ecology, Tibet Agricultural and Animal Husbandry University, Nyingchi 860000, Tibet, China</mods:affiliation>
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<mods:namePart id="461C40FAB2260931B195D5BAFDDA6B07">Deng, Yun-Fei</mods:namePart>
<mods:affiliation id="E341D005F6B675F741F4105B9F5B40A3">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, Guangdong, China</mods:affiliation>
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<mods:namePart id="98E6FF91651F45A3F05E90045C27D064">Deng, Tao</mods:namePart>
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<mods:date id="F39874254E6E41C74F3A39B72E29D9A3">2020</mods:date>
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<paragraph id="B5C495F874C9F1D0A7231306110ACBF0" pageId="0" pageNumber="117">
<taxonomicName id="50C95F5B029E62761CC9C8FC59F02DE2" ID-CoL="7Z4YQ" LSID="3B42DDB7-0A21-5D02-8947-45755B5830D5" authority="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii" status="sp. nov.">Strobilanthes sunhangii T. Deng, J.T. Chen &amp; Y.F. Deng</taxonomicName>
<taxonomicNameLabel id="CCD3A2B86DE8C988FC3A8C28BF240513" pageId="0" pageNumber="117">sp. nov.</taxonomicNameLabel>
<figureCitation id="27E5CE25DA9A123265FDF65BC65B03A9" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Photograph of the holotype of Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng (KUN barcode 1345286!)." figureDoi="10.3897/phytokeys.166.58831.figure1" httpUri="https://binary.pensoft.net/fig/474755" pageId="0" pageNumber="117">Figs 1</figureCitation>
<figureCitation id="6519258CF1DA006A5BAF6400DB85A29C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A plant showing flowering branch and leaves B flower (view from side) C pistil, ovary and style D calyx E opened corolla showing androecium (Drawn by Xiaoshuang Zhang based on the holotype of Sunhang 19964)." figureDoi="10.3897/phytokeys.166.58831.figure2" httpUri="https://binary.pensoft.net/fig/474756" pageId="0" pageNumber="117">, 2</figureCitation>
<figureCitation id="3C139F4B6D8DEDD52CEB382D7D2C7E34" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A habitat and flowering branch B, C flowering branch and flower (view from front) D, E flower (view from side) F calyx G style H leaf adaxial surfaces I leaf abaxial surfaces." figureDoi="10.3897/phytokeys.166.58831.figure3" httpUri="https://binary.pensoft.net/fig/474757" pageId="0" pageNumber="117">, 3</figureCitation>
<figureCitation id="397D870B4325C33792152A8CC6A19CC5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pollen grains of Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A equatorial view B polar view C ornamentation." figureDoi="10.3897/phytokeys.166.58831.figure4" httpUri="https://binary.pensoft.net/fig/474758" pageId="0" pageNumber="117">, 4</figureCitation>
</paragraph>
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<paragraph id="AC078ADEF5C4B29B234AC928F219E9BE" pageId="0" pageNumber="117">Type.</paragraph>
<paragraph id="CEE57974048C5BB8816CB91E1992CE4D" pageId="0" pageNumber="117">
China. Tibet: 
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County, Beibeng Town, 
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, 
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, elev. ca. 1,470 m, 5 Oct 2018, 
<emphasis id="D2297B9287D5B903E5E5D76B14254952" italics="true" pageId="0" pageNumber="117">H. Sun</emphasis>
, 
<emphasis id="D3B8D0E90A01BF0E75B35575AF10B544" italics="true" pageId="0" pageNumber="117">T. Deng</emphasis>
&amp; 
<emphasis id="54D6A8BE919AA327CA41DC88F1FF1B41" italics="true" pageId="0" pageNumber="117">Z.M. Li Sunhang19964</emphasis>
(
<emphasis id="98A088298DC067B64E3E9284F424DB71" bold="true" italics="true" pageId="0" pageNumber="117">holotype</emphasis>
: KUN1345286!; 
<emphasis id="2182124064C69E97F8653F3BC1953F98" bold="true" italics="true" pageId="0" pageNumber="117">isotypes</emphasis>
: IBSC!, KUN1345287!, KUN1345288!)
</paragraph>
<caption id="B8E846160EF6B120BC12118622E4DE88" doi="10.3897/phytokeys.166.58831.figure1" httpUri="https://binary.pensoft.net/fig/474755" pageId="0" pageNumber="117" start="Figure 1" startId="F1">
<paragraph id="41059DE18858E7F51205F62A7DD9C0EE" pageId="0" pageNumber="117">
<emphasis id="30CE41EBB605844F203C55E936FC3427" bold="true" pageId="0" pageNumber="117">Figure 1.</emphasis>
Photograph of the holotype of 
<taxonomicName id="FA770D5C14054841682F3F90FA6C2876" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="0EBA99CE848BF460390DB5592FFF1E4D" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
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T. Deng, J.T. Chen &amp; Y.F. Deng (KUN barcode 1345286!).
</paragraph>
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<paragraph id="CBF1ACEFDB1F3F5DC0BB11E021311E87" pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph id="64C0252A6E931400BE227BEF050A203F" pageId="0" pageNumber="117">
<taxonomicName id="D7C8D5C3A14E5155C41BA0E02623A21E" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="949157CFDEF72FAE4319C250F5C75A13" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
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resembles 
<taxonomicName id="025F2DB92A4C590A7BFE7104283F8B20" lsidName="S. medogensis" pageId="0" pageNumber="117" rank="species" species="medogensis">
<emphasis id="B8B203A9BA9154CB056015771706A1E0" italics="true" pageId="0" pageNumber="117">S. medogensis</emphasis>
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, but differs by its stems subterete, glabrous (vs. slightly sulcate, bifariously puberulent); spikes (7-)11-22 cm long (vs. 3-6 cm long), rachis glabrous (vs. bifariously pubescent); corolla outside and lobes pinkish-white, inside purplish-pink (vs. corolla yellowish-white, but dull purple on lobes), the tube bent to ca. 90° twice (vs. straight), lobes apices emarginate (vs. rounded); calyx 7-8 mm long, glabrous, 5-lobed to middle (vs. ca. 12 mm long, gland-tipped pubescent, 5-lobed to base), lobes ovate, equal (vs. narrowly oblong-elliptic, subequal); stamens included (vs. exserted), filaments of shorter pair ca. 2 mm long (vs. 7-8 mm long), longer pair ca. 4 mm long (vs. 9-10 mm long), anther thecae ca. 4 mm (vs. ca.1 mm); style with sparse gland-tipped trichomes (vs. glabrous).
</paragraph>
<caption id="D3A774EADA6DB252C15633999619D047" doi="10.3897/phytokeys.166.58831.figure2" httpUri="https://binary.pensoft.net/fig/474756" pageId="0" pageNumber="117" start="Figure 2" startId="F2">
<paragraph id="4E8D322F69F7C03AB67296B6749E613C" pageId="0" pageNumber="117">
<emphasis id="5BFD5787ECEC8038270D5639FD119EAC" bold="true" pageId="0" pageNumber="117">Figure 2.</emphasis>
<taxonomicName id="BD0ED137C1AB21F5DA1E87055664A485" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="92BBD70F05AEEBBD6806181D0E4D09A2" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
</taxonomicName>
T. Deng, J.T. Chen &amp; Y.F. Deng 
<emphasis id="36A723AB1175E57AA76D748CC3E3A533" bold="true" pageId="0" pageNumber="117">A</emphasis>
plant showing flowering branch and leaves 
<emphasis id="F3DFC246907567D6F505D377DFA08A79" bold="true" pageId="0" pageNumber="117">B</emphasis>
flower (view from side) 
<emphasis id="71D9BDD8DA43D35F4C098578F88E9068" bold="true" pageId="0" pageNumber="117">C</emphasis>
pistil, ovary and style 
<emphasis id="090390387AC97C33905245BB96FA8FAE" bold="true" pageId="0" pageNumber="117">D</emphasis>
calyx 
<emphasis id="BA0ED3A4978B3D29DC25B14EE3C7762C" bold="true" pageId="0" pageNumber="117">E</emphasis>
opened corolla showing androecium (Drawn by Xiaoshuang Zhang based on the holotype of Sunhang19964).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="875C2C9989ABC7B235A43DE01B6D95ED" pageId="0" pageNumber="117" type="description">
<paragraph id="AED2CB37A81472F74735C5BA0892E7DF" pageId="0" pageNumber="117">Description.</paragraph>
<paragraph id="01507E6E4F451689B93CBDEEA9D7FA4E" pageId="0" pageNumber="117">
Undershrubs 1-2 m high, gregarious. Stems subterete, glabrous, slightly swollen at node. Leaves slightly anisophyllous, smaller one ca. 2/3 of the larger one in size; petiole 3-7 mm long or petiole of smaller leaf in each pair shorter, sulcate, glabrous; blades ovate to lanceolate-ovate, smaller ones 4.2-6.9 
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1.2-2.0 cm, larger ones 8.2-10.1 
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2.2-3.2 cm, both surfaces glabrous, densely covered with numerous linear cystoliths, secondary veins 3-5 on each side of midvein and prominent on both surfaces, base rounded to broadly cuneate, slightly oblique, margin serrulate, apex acuminate and narrowly caudate. Inflorescences of axillary spikes, simple or rarely 2-furcate, slender, (7-) 11-22 cm long; rachis glabrous; bracts and bracteoles not seen, caducous. Flowers (0.6-) 1.0-2.5 (-3.0) cm apart on rachis. Calyx 7-8 mm long, glabrous, 5-lobed to middle; lobes ovate, equal, margins narrowly scarious, 1-nerved abaxially, conspicuous after drying, apices acuminate. Corolla 2.8-3.3 cm long, campanulate, outside and lobes pinkish-white, inside purplish-pink, outside glabrous, inside glabrous except for bifarious hirsute retaining the style (Fig. 
<figureCitation id="492E929AA283EE5083B21EDD38CD8849" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A habitat and flowering branch B, C flowering branch and flower (view from front) D, E flower (view from side) F calyx G style H leaf adaxial surfaces I leaf abaxial surfaces." figureDoi="10.3897/phytokeys.166.58831.figure3" httpUri="https://binary.pensoft.net/fig/474757" pageId="0" pageNumber="117">3E</figureCitation>
); tube basally cylindrical, ca. 3 mm wide and ca. 6 mm long, then bent to ca. 90° and gradually widened to 9-12 mm wide at middle and 1.6-1.8 cm long, then secondly bent to ca. 90° and tube upper cylindrical 9-12 mm wide and 1.6-1.8 cm long; lobes broadly elliptic, 8-9 
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7-8 mm, apices emarginate (Fig. 
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). Stamens 4, didynamous, included; the united part of filaments glabrous at base, with dense villous on middle and margin, with slightly sparse villous on upper (Fig. 
<figureCitation id="43FCDEA96344AF7B3F1E3F31A166776D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A habitat and flowering branch B, C flowering branch and flower (view from front) D, E flower (view from side) F calyx G style H leaf adaxial surfaces I leaf abaxial surfaces." figureDoi="10.3897/phytokeys.166.58831.figure3" httpUri="https://binary.pensoft.net/fig/474757" pageId="0" pageNumber="117">3E</figureCitation>
), shorter pair ca. 2 mm long, longer pair ca. 4 mm long; anther thecae oblong, ca. 4 mm long. Pollen grains prolate in outline, 74.1 (72.2-76.1) 
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49.7 (48.1-50.4) 
<normalizedToken id="C01C32B71953A59F42E253D23DCB5A43" originalValue="μm">μm</normalizedToken>
, P/E = 1.49, tricolporate, ribbed pseudocolpi 12, exine reticulate (Fig. 
<figureCitation id="8FED36F1CB4603D9A48C3099DD3C0D30" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Pollen grains of Strobilanthes sunhangii T. Deng, J. T. Chen &amp; Y. F. Deng A equatorial view B polar view C ornamentation." figureDoi="10.3897/phytokeys.166.58831.figure4" httpUri="https://binary.pensoft.net/fig/474758" pageId="0" pageNumber="117">4</figureCitation>
). Ovary ca. 5 mm long, glabrous; style ca. 1.8 cm long, with sparse gland-tipped trichomes. Capsule not seen.
</paragraph>
<caption id="9BD26DD03BEE643640CD5ABA17A34A80" doi="10.3897/phytokeys.166.58831.figure3" httpUri="https://binary.pensoft.net/fig/474757" pageId="0" pageNumber="117" start="Figure 3" startId="F3">
<paragraph id="80424FBDA014D9A4C63F74679275966C" pageId="0" pageNumber="117">
<emphasis id="6E875105B22EEADCBEA3AE3BC43CD083" bold="true" pageId="0" pageNumber="117">Figure 3.</emphasis>
<taxonomicName id="C639C79A881F2CE68A6FEC371A0AD615" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="F6BE7E83E86C22ED0DAE02B0111730C5" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
</taxonomicName>
T. Deng, J.T. Chen &amp; Y.F. Deng 
<emphasis id="3203D5243FABEB46BB8DD89D4D62F2FF" bold="true" pageId="0" pageNumber="117">A</emphasis>
habitat and flowering branch 
<emphasis id="E863BAE2ACB0BFB9DA31DD24E0C107E1" bold="true" pageId="0" pageNumber="117">B, C</emphasis>
flowering branch and flower (view from front) 
<emphasis id="7FDA619E8ABE70A69CAEB7B87E1C276D" bold="true" pageId="0" pageNumber="117">D, E</emphasis>
flower (view from side) 
<emphasis id="AEBD4ABB0256B57F570F9F797FFB833B" bold="true" pageId="0" pageNumber="117">F</emphasis>
calyx 
<emphasis id="78CD92982C0885B55622B16E0F06CF90" bold="true" pageId="0" pageNumber="117">G</emphasis>
style 
<emphasis id="77245237B2FEE580189050F526DF4CFD" bold="true" pageId="0" pageNumber="117">H</emphasis>
leaf adaxial surfaces 
<emphasis id="7A0B53B4150B6A5C65C78EA0C05A5711" bold="true" pageId="0" pageNumber="117">I</emphasis>
leaf abaxial surfaces.
</paragraph>
</caption>
<caption id="DE6E0FF3766039F46D0BDEF6ADBA9D20" doi="10.3897/phytokeys.166.58831.figure4" httpUri="https://binary.pensoft.net/fig/474758" pageId="0" pageNumber="117" start="Figure 4" startId="F4">
<paragraph id="3A9EFD899483633E25C17017F776CBDA" pageId="0" pageNumber="117">
<emphasis id="63A82EF235703438888817CFDBDB41A7" bold="true" pageId="0" pageNumber="117">Figure 4.</emphasis>
Pollen grains of 
<taxonomicName id="A3B027BE404F7A87154B99F2D65B90EF" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="A8513AC0E9943457F4D3ADB27FC089E5" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
</taxonomicName>
T. Deng, J.T. Chen &amp; Y.F. Deng 
<emphasis id="024863218EB1FB22AF669EB2ACF513C0" bold="true" pageId="0" pageNumber="117">A</emphasis>
equatorial view 
<emphasis id="3526E0B51C584103D98603905ADAAEDF" bold="true" pageId="0" pageNumber="117">B</emphasis>
polar view 
<emphasis id="507F0970332A8E0352E9F97439933F0B" bold="true" pageId="0" pageNumber="117">C</emphasis>
ornamentation.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="77B6B9624D494D7FC8C49E7B4BFF93B9" pageId="0" pageNumber="117" type="phenology">
<paragraph id="A7CBC33805567812D181197507886E62" pageId="0" pageNumber="117">Phenology.</paragraph>
<paragraph id="89985F5B34D227F521ABF53EF4075530" pageId="0" pageNumber="117">Flowering from June-October.</paragraph>
</subSubSection>
<subSubSection id="4BE26A1EE454D844B067F7810F2B6495" pageId="0" pageNumber="117" type="etymology">
<paragraph id="7EBB1285D176F4ABAA0A6320F1394529" pageId="0" pageNumber="117">Etymology.</paragraph>
<paragraph id="E5E8827E4454B8638FA9B190CB4C182E" pageId="0" pageNumber="117">
<taxonomicName id="6DE901DC876DBF6F0B6F67CB0D7E7AEF" authorityName="T. Deng, J. T. Chen &amp; Y. F. Deng" authorityYear="2020" class="Magnoliopsida" family="Acanthaceae" genus="Strobilanthes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Strobilanthes sunhangii" order="Lamiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis id="7C34987C6D049C932A9E11F7797C4280" italics="true" pageId="0" pageNumber="117">Strobilanthes sunhangii</emphasis>
</taxonomicName>
is named after Prof. Hang Sun (1963-) for his outstanding contributions to the flora of Himalayan regions and collecting this new species for the first time. Verucular name: The Chinese name is given as 
<normalizedToken id="776912A2E6F9B5F858575B1620F3F1EA" originalValue="“双曲马蓝”">&quot;双曲马蓝&quot;</normalizedToken>
(
<normalizedToken id="0DD0C1C77C283CF17B9223672C892AB4" originalValue="shuāng">shuang</normalizedToken>
<normalizedToken id="3CCB3DA3FD1420EF98AD8A8BD95BD12E" originalValue="qū">qū</normalizedToken>
<normalizedToken id="96A1E5F9367021146E6317E12BC03133" originalValue="mǎ">mǎ</normalizedToken>
<normalizedToken id="B366E65AE08F4C3B640E21EE06F912A7" originalValue="lán">lan</normalizedToken>
), referring to the double-curved corolla of the new species.
</paragraph>
</subSubSection>
<subSubSection id="9AEDCA0849CE0D61E2450D51DC008580" pageId="0" pageNumber="117" type="distribution">
<paragraph id="0FE0665FD5DC30D88F5AFB8F359414FA" pageId="0" pageNumber="117">Distribution and habitat.</paragraph>
<paragraph id="DF98CF208D57B454D150A21A5ADAF66D" pageId="0" pageNumber="117">
The new species is currently known only from 
<normalizedToken id="50C77D35472BEBA67CD8699FB792AFC7" originalValue="Mêdog">Medog</normalizedToken>
County, Tibet, China. It grows in the moist evergreen forest at an elevation of 1,200-1,800 m. The associations include 
<taxonomicName id="99B373612529A48B1BD69E65CF2342ED" authorityName="R. W. Brown" authorityYear="1946" baseAuthorityName="R. Br. ex Griff." class="Magnoliopsida" family="Hamamelidaceae" genus="Exbucklandia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Exbucklandia populnea" order="Saxifragales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="populnea">
<emphasis id="86C37701A6799929F0222FCDEA6D6552" italics="true" pageId="0" pageNumber="117">Exbucklandia populnea</emphasis>
</taxonomicName>
(R. Br.) R. W. Brown (
<taxonomicName id="E014E5DFD7E41D7836A438C221497724" authorityName="R.W.Brown" authorityYear="1946" class="Magnoliopsida" family="Hamamelidaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Saxifragales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Hamamelidaceae</taxonomicName>
), 
<taxonomicName id="D53781AB7F8AD6B85C54D61430CD2F49" class="Magnoliopsida" family="Viburnaceae" genus="Viburnum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Viburnum" order="Dipsacales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis id="82ECAAC33C0677EA8111AD899393508D" italics="true" pageId="0" pageNumber="117">Viburnum</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="7F2C3EE97D00EA038D1ACC2ECC51F523" class="Magnoliopsida" family="Viburnaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Dipsacales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Viburnaceae</taxonomicName>
), 
<taxonomicName id="F2CD4389C1E56528037CE8E449D111F4" class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis id="5697ADA835C108E11F113473ABE0E0AE" italics="true" pageId="0" pageNumber="117">Rubus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="750BF87A7E5BB463DCB3BAD6B67697F8" class="Magnoliopsida" family="Rosaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Rosales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
), 
<taxonomicName id="1C02C0E566999BCDF5B7AC24BA84D62B" authorityName="C. B. Clarke" authorityYear="1881" baseAuthorityName="Hooker f." class="Magnoliopsida" family="Campanulaceae" genus="Campanumoea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Campanumoea inflata" order="Asterales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="inflata">
<emphasis id="E475796E40E1548015F263428EFE03D7" italics="true" pageId="0" pageNumber="117">Campanumoea inflata</emphasis>
</taxonomicName>
(Hook. f. &amp; Thomson) C.B. Clarke (
<taxonomicName id="ACDCF9FC7FBC1F6C4222D27E5E1C2A5D" class="Magnoliopsida" family="Campanulaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Asterales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Campanulaceae</taxonomicName>
), 
<taxonomicName id="B0A17946F73CA125DCA4193FD933B9AA" class="Magnoliopsida" family="Gentianaceae" genus="Tripterospermum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Tripterospermum volubile" order="Gentianales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="volubile">
<emphasis id="22897B4216DF3A6F7DAD0D1C3FAFC3FD" italics="true" pageId="0" pageNumber="117">Tripterospermum volubile</emphasis>
</taxonomicName>
(D. Don) H. Hara (
<taxonomicName id="F0C04459899BAA769E06E1C5E3545293" class="Magnoliopsida" family="Gentianaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Gentianales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Gentianaceae</taxonomicName>
) and 
<taxonomicName id="C3E42AD4419D58F417C97AEADB814ED0" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Impatiens" order="Ericales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis id="AB186A35A939DD02DC3474DAB33F65F3" italics="true" pageId="0" pageNumber="117">Impatiens</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="B2FE24F9A335B9938C5F9B5927162D17" class="Magnoliopsida" family="Balsaminaceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="family">Balsaminaceae</taxonomicName>
), amongst others.
</paragraph>
</subSubSection>
<subSubSection id="74247055D9FB37B19C93F8BCE64B0C9C" pageId="0" pageNumber="117" type="conservation status">
<paragraph id="ED9DB329F633E3EB71D9187AB1B30F4D" pageId="0" pageNumber="117">Conservation status.</paragraph>
<paragraph id="E8BA3F15FB7E022866B94BF451EF7CEE" pageId="0" pageNumber="117">
The new species was only discovered in Beibeng Town, 
<normalizedToken id="4F92958E27274680D041A6D3183801CA" originalValue="Mêdog">Medog</normalizedToken>
County. About 1,000 individuals were observed and the extent of occurrence was ca. 5 km2. Further explorations to obtain the precise population status are needed to assess its conservation status. Based on available data, the new species is assigned to the category 'Data 
<normalizedToken id="5330135692C8E7561D740372F019E164" originalValue="Deficient’">Deficient'</normalizedToken>
(DD) of the International Union for Conservation of Nature (
<bibRefCitation id="FD8DF3EE21172399C58832BA01E2350C" author="IUCN" journalOrPublisher="19. Beijing &amp; Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="117" publicationUrl="http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" refId="B15" refString="IUCN, 2019. Guidelines for Using the IUCN Red List Categories and Criteria. version 13. Prepared by the Standards and Petitions Subcommittee of the IUCN Species Survival Commission, 113 pp. http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. version 13. Prepared by the Standards and Petitions Subcommittee of the IUCN Species Survival Commission, 113 pp." url="http://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B2F57383C9EC4BCA777C1E245C0234E3" pageId="0" pageNumber="117" type="specimens of s. medogensis examined">
<paragraph id="F39FAB458084C9AFA0481401F91470BE" pageId="0" pageNumber="117">
Specimens of 
<taxonomicName id="8D33CD3B54CBCBD2E6975E83AF769350" lsidName="S. medogensis" pageId="0" pageNumber="117" rank="species" species="medogensis">
<emphasis id="CFC3266432F42C02A93632C2C0A86A6F" italics="true" pageId="0" pageNumber="117">S. medogensis</emphasis>
</taxonomicName>
examined.
</paragraph>
<paragraph id="5C426327F7F38A776552460217D04383" pageId="0" pageNumber="117">
- China. 
<emphasis id="15BCE6E04D42E1108E6C178FE27E3A06" bold="true" pageId="0" pageNumber="117">Tibet</emphasis>
: 
<normalizedToken id="98B7B1972E95CB9FAAAAA24C78D21AD3" originalValue="Mêdog">Medog</normalizedToken>
County, Gedang Town, elev. ca. 2,400 m, 27 Aug1974, 
<emphasis id="65240037B5C4CBC93EA86758BC4BFE7E" italics="true" pageId="0" pageNumber="117">QTP Expedition 74-4911</emphasis>
(holotype: KUN barcode 1219209!); 
<normalizedToken id="4ACD629EC8BC23ACF8061423E0BCAF7A" originalValue="Mêdog">Medog</normalizedToken>
County, Gedang Town, 2,450 m elev., 11 Sept 1982, 
<emphasis id="5D6150096C679ABEE26EC87FFF0F8CA6" italics="true" pageId="0" pageNumber="117">B.S. Li &amp; S.Z.Cheng 814</emphasis>
(PE); 
<normalizedToken id="CABECE8B5D4C17409FE798F066197E9F" originalValue="Mêdog">Medog</normalizedToken>
County, Gedang Town, 1,940 m elev., 13 Oct 1982, 
<emphasis id="B4F8F8C33126D4ED6969C15BBE6B503D" italics="true" pageId="0" pageNumber="117">S.Z. Cheng &amp; B.S. Li 1457</emphasis>
(PE); 
<normalizedToken id="5AC6E65B828C8503069BC5C895CCE53F" originalValue="Mêdog">Medog</normalizedToken>
County, 1,100 m elev., 1 Dec 1982, 
<emphasis id="9428EA68B174A0C1B204A0846FE732BA" italics="true" pageId="0" pageNumber="117">S.Z. Cheng &amp; B.S. Li 3238</emphasis>
(PE); Nyingchi County, Pailong Town, 2,175 m elev., 8 Sept 2006, 
<emphasis id="FFF0D5391E05CCB503D8CFCC336AF315" italics="true" pageId="0" pageNumber="117">Tiber-MacArthur 261</emphasis>
(A); Lung County, Chayne chu, 2,750 m elev., 10 Jul 1936, 
<emphasis id="361F884B6A78476ABE6034EF3E97F2A5" italics="true" pageId="0" pageNumber="117">Ludlow &amp; Sherriff 2336</emphasis>
(BM, E).
</paragraph>
</subSubSection>
<subSubSection id="E2EF208B018EA7B9F98A8344E00E4751" pageId="0" pageNumber="117" type="specimens of s. divaricata examined">
<paragraph id="F4D8B4058F121EBA35E9DCBE89A0F083" pageId="0" pageNumber="117">
Specimens of 
<taxonomicName id="4B55DB6EB47F0B986F752C366F7C71FE" lsidName="S. divaricata" pageId="0" pageNumber="117" rank="species" species="divaricata">
<emphasis id="F78D1477DE90CEBC62A3C3F952D681A0" italics="true" pageId="0" pageNumber="117">S. divaricata</emphasis>
</taxonomicName>
examined.
</paragraph>
<paragraph id="831517D1C9139E1D5E613E26B8B5271A" pageId="0" pageNumber="117">
- India. 
<emphasis id="590983E2313255A245C2FE697920AA3F" bold="true" pageId="0" pageNumber="117">Meghalaya</emphasis>
: Khasia, 1,524 m elev., 5 Oct 1886, 
<emphasis id="33713549D89F885DA6911E34B5829659" italics="true" pageId="0" pageNumber="117">C. B. Clarke 45219</emphasis>
(BM, CAL, LE); Khasia, 1,219 m elev., 12 Nov 1871, 
<emphasis id="9A0A5ED5FB8EF4993BFB2EE1C42594AA" italics="true" pageId="0" pageNumber="117">C. B. Clarke 15479</emphasis>
(CAL, K, LE); Khasia, Oct 1850, Simons 482 (CAL). 
<emphasis id="37881EF1C0F48606E8850FD56E776832" bold="true" pageId="0" pageNumber="117">Sikkim</emphasis>
: Khuping, 1,676 m elev., 26 Sept1884, 
<emphasis id="7CDB60BD6B07E36CB9F594DC8839EEB0" italics="true" pageId="0" pageNumber="117">C. B. Clarke 35904</emphasis>
(LE); Khuping, 1,676 m elev., 26 Sept 1884, 
<emphasis id="8ECCA9A1B4F53526857600F6C89A6329" italics="true" pageId="0" pageNumber="117">C. B. Clarke 35920</emphasis>
(LE). - Nepal. Koshi Zone, Sankhuwasabha District, 2,020 m elev., 16 Aug 1998, 
<emphasis id="85AA061179C9DD2EACAEAAF87F5D0E67" italics="true" pageId="0" pageNumber="117">S. Noshiroet al. 9840046</emphasis>
(A); Dhawalagiri Zone, Myagdi District, 1,820-2,360 m elev., 10 Sept 1996, 
<emphasis id="8B01178E7ED15AEB24F7A4A92277E742" italics="true" pageId="0" pageNumber="117">M. Mikage et al. 9681329</emphasis>
(A).
</paragraph>
</subSubSection>
</treatment>
</document>