312 lines
40 KiB
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312 lines
40 KiB
XML
<document id="B5F1CDCC0CCD98CEF85ECC0596EE11B8" ID-CLB-Dataset="38738" ID-DOI="10.11646/phytotaxa.238.1.1" ID-GBIF-Dataset="30c1c7b1-db98-407b-9694-239c1d9dd243" ID-ISSN="1179-3163" ID-Zenodo-Dep="399630" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="admin" checkinTime="1458887842333" checkinUser="plazi" docAuthor="Thiago E. C. Meneguzzo, José F. A. Baumgratz & Cássio Van Den Berg" docDate="2015" docId="F10187B5F1463277C5BCCCB39CFE1C97" docLanguage="en" docName="Phytotaxa238.1.001-039.pdf.imf" docOrigin="Phytotaxa 238 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Aganisia Lindley" docType="treatment" docVersion="12" lastPageNumber="5" masterDocId="0D38FFCDF1453273C534CD1B99551F06" masterDocTitle="Taxonomic studies in the Aganisia complex (Orchidaceae, Zygopetalinae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="4" updateTime="1698186280166" updateUser="plazi">
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<mods:mods id="A6CE55130D71372A07DAB7D2FB2732EF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CAC9D1790E8165B375AA75679A2C747C">
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<mods:title id="214A242CD4DFAAFE151650997D403E27">Taxonomic studies in the Aganisia complex (Orchidaceae, Zygopetalinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A54A8B766B6E00727DC4CF4F38154DED" type="personal">
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<mods:roleTerm id="937E9AAC2EC71E92543FD0CD02F7882F">Author</mods:roleTerm>
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<mods:namePart id="E28CD3AF0CF2DD3B97557893C2F7D178">Thiago E. C. Meneguzzo</mods:namePart>
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<mods:affiliation id="E528D641B7CF1A588C1D25C984B234D5">Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rua Pacheco Leão, 915, Jardim Botânico, 22460 - 030, Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:nameIdentifier id="ED30977711212FFD17FEE37B7E0B10EA" type="email">meneguzzotec@gmail.com</mods:nameIdentifier>
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<mods:name id="18F1596B162FDC1B271F5F270AEAD8B9" type="personal">
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<mods:roleTerm id="F2E95FFD61B60C025FA2BBF631FBA27B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C9B39D88E6A9F86C4FED1BFF0C05678E">José F. A. Baumgratz</mods:namePart>
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<mods:affiliation id="7963ACC7D29C6D28EA12D49F43937CA3">Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rua Pacheco Leão, 915, Jardim Botânico, 22460 - 030, Rio de Janeiro, Rio de Janeiro, Brazil</mods:affiliation>
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</mods:name>
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<mods:name id="1A4C4ECD56C8082C605C64D68DCA71BD" type="personal">
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<mods:roleTerm id="5451E3EEA42FFFD1D7103CA532607636">Author</mods:roleTerm>
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<mods:namePart id="51B1F968E3ECE2D89E8613199968EE16">Cássio Van Den Berg</mods:namePart>
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<mods:affiliation id="BE12C709346EE72B46444CD6EB529365">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Avenida Transnordestina, s. n., 44036 - 900, Feira de Santana, Bahia, Brazil</mods:affiliation>
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<mods:typeOfResource id="FB8B776F1B2DCFF12EE1FD5D1F264022">text</mods:typeOfResource>
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<mods:title id="DC4827BED6268EE2379022AC3B7C7856">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="74706EDF414AD8ABC70A191AE011AAB5">
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<mods:date id="65FFA3EE159BBDBB0EF90C392D841F88">2015</mods:date>
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<mods:number id="40D6A55F5D01B26F2C9AA0C5BA764482">2015-12-09</mods:number>
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<mods:detail id="B679D9D8A65A65D79BAE24F4294E092C" type="volume">
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<mods:number id="DA5660E2BA39014156CFED3F686AFCE5">238</mods:number>
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</mods:detail>
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<mods:detail id="F6FB36F280E51C15C1C9A8B5DEE8CDA2" type="issue">
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<mods:number id="7A7AA2B6B5D9D7F41EE6D1B255B0C628">1</mods:number>
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<mods:start id="F30468888D1C0405BEEEB47E66011E32">1</mods:start>
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<mods:end id="0F618C8B19A9CA4E0A91DFAAABFC096C">39</mods:end>
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<mods:identifier id="15F1ACC1A5C53A448351562F373FF3ED" type="DOI">10.11646/phytotaxa.238.1.1</mods:identifier>
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<mods:identifier id="1C48713CC5584266530E9B96E22528E7" type="ISSN">1179-3163</mods:identifier>
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<treatment id="F10187B5F1463277C5BCCCB39CFE1C97" ID-DOI="http://doi.org/10.5281/zenodo.4779851" ID-GBIF-Taxon="121065265" ID-Zenodo-Dep="4779851" LSID="urn:lsid:plazi:treatment:F10187B5F1463277C5BCCCB39CFE1C97" httpUri="http://treatment.plazi.org/id/F10187B5F1463277C5BCCCB39CFE1C97" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<subSubSection id="31B26528F1463270C5BCCCB39B781EC4" box="[136,557,423,450]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="791736A3F1463270C5BCCCB39B781EC4" blockId="3.[136,1452,423,487]" box="[136,557,423,450]" pageId="3" pageNumber="4">
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1.
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<taxonomicName id="BEA84D20F1463270C597CCB398E11EC4" ID-CoL="8VTF5" ID-ENA="310109" authority="Lindley" authorityName="Lindley" box="[163,436,424,450]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="4BDCEAB1F1463270C597CCB3985F1EC4" bold="true" box="[163,266,424,450]" italics="true" pageId="3" pageNumber="4">Aganisia</emphasis>
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<bibRefCitation id="1D394B52F1463270C425CCB398E11EC4" author="Lindley, J." box="[273,436,424,450]" journalOrPublisher="Edwards's Botanical Register" pageId="3" pageNumber="4" pagination="45 - 46" part="25" refId="ref28982" refString="Lindley, J. (1839) Aganisia pulchella. Edwards's Botanical Register 25 (misc.): 45 - 46." title="Aganisia pulchella" type="journal article" year="1839">Lindley (1839</bibRefCitation>
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</taxonomicName>
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: misc. 45).
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</paragraph>
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</subSubSection>
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<subSubSection id="31B26528F1463270C707CCB298141D9F" pageId="3" pageNumber="4" type="reference_group">
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<paragraph id="791736A3F1463270C707CCB29B811EE0" blockId="3.[136,1452,423,487]" pageId="3" pageNumber="4">
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<taxonomicName id="BEA84D20F1463270C707CCB29AE31EC5" authority="Kuntze" authorityName="Kuntze" box="[563,950,425,451]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="section" section="Typaganisia">
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<emphasis id="4BDCEAB1F1463270C707CCB29BC21EC5" box="[563,663,425,451]" italics="true" pageId="3" pageNumber="4">Aganisia</emphasis>
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sect.
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<emphasis id="4BDCEAB1F1463270C7E2CCB29A0B1EC4" box="[726,862,425,450]" italics="true" pageId="3" pageNumber="4">Typaganisia</emphasis>
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Kuntze
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</taxonomicName>
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<emphasis id="4BDCEAB1F1463270C688CCB29A871EC4" box="[956,978,425,450]" italics="true" pageId="3" pageNumber="4">in</emphasis>
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<bibRefCitation id="1D394B52F1463270C6ECCCB29C7C1EC4" author="von Post, T. & Kuntze, O." bookContentInfo="714 pp." box="[984,1321,424,451]" journalOrPublisher="Wretmanianis, Upsalla" pageId="3" pageNumber="4" publicationUrl="http://dx.doi.org/10.5962/bhl.title.58320" refId="ref30014" refString="von Post, T. & Kuntze, O. (1903) Lexicon generum phanerogamarum inde ab anno MDCXXXVII cum nomenclatura legitima internationali simul scientifica. Wretmanianis, Upsalla, 714 pp. http: // dx. doi. org / 10.5962 / bhl. title. 58320" title="Lexicon generum phanerogamarum inde ab anno MDCXXXVII cum nomenclatura legitima internationali simul scientifica" type="book" year="1903">von Post & Kuntze (1903: 13)</bibRefCitation>
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,
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<emphasis id="4BDCEAB1F1463270C001CCB29CF91EC4" box="[1333,1452,424,450]" italics="true" pageId="3" pageNumber="4">nom. illeg.</emphasis>
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Type:—
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<taxonomicName id="BEA84D20F1463270C5D7CCD69B341EE0" ID-CoL="65LN9" authority="Lindley" authorityName="Lindley" box="[227,609,460,487]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="pulchella">
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<emphasis id="4BDCEAB1F1463270C5D7CCD698E31EE0" box="[227,438,460,487]" italics="true" pageId="3" pageNumber="4">Aganisia pulchella</emphasis>
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<bibRefCitation id="1D394B52F1463270C489CCD79B341EE0" author="Lindley, J." box="[445,609,460,486]" journalOrPublisher="Edwards's Botanical Register" pageId="3" pageNumber="4" pagination="45 - 46" part="25" refId="ref28982" refString="Lindley, J. (1839) Aganisia pulchella. Edwards's Botanical Register 25 (misc.): 45 - 46." title="Aganisia pulchella" type="journal article" year="1839">Lindley (1839</bibRefCitation>
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</taxonomicName>
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: misc. 45)
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</paragraph>
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<paragraph id="791736A3F1463270C5BCCF0C98731D54" blockId="3.[136,1452,535,665]" pageId="3" pageNumber="4">
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<taxonomicName id="BEA84D20F1463270C5BCCF0C98CE1D28" ID-CoL="8VT5X" authority="Lindley (1853 a: 1)" authorityName="Lindley" authorityPageNumber="1" authorityYear="1853" box="[136,411,535,558]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="4BDCEAB1F1463270C5BCCF0C99B41D2A" box="[136,225,535,556]" italics="true" pageId="3" pageNumber="4">Acacallis</emphasis>
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<bibRefCitation id="1D394B52F1463270C5D3CF0C98CE1D28" author="Lindley, J." box="[231,411,535,558]" journalOrPublisher="Matthews, London" pageId="3" pageNumber="4" pagination="1" part="4" publicationUrl="http://dx.doi.org/10.5962/bhl.title.51709" refId="ref29107" refString="Lindley, J. (1853 a) Acacallis. In: Folia orchidacea 4 (Acacallis). Matthews, London, p. 1. http: // dx. doi. org / 10.5962 / bhl. title. 51709" title="Acacallis" type="journal article" volumeTitle="Folia orchidacea" year="1853">Lindley (1853a: 1)</bibRefCitation>
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</taxonomicName>
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.
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<taxonomicName id="BEA84D20F1463270C491CF039A761D2B" authority="(Lindl.) Kuntze" authorityName="Kuntze" baseAuthorityName="Lindl." box="[421,803,535,557]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="section" section="Acacallis">
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<emphasis id="4BDCEAB1F1463270C491CF0398AF1D2B" box="[421,506,536,557]" italics="true" pageId="3" pageNumber="4">Aganisia</emphasis>
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sect.
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<emphasis id="4BDCEAB1F1463270C704CF0C9BDC1D2A" box="[560,649,535,556]" italics="true" pageId="3" pageNumber="4">Acacallis</emphasis>
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(Lindl.) Kuntze
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</taxonomicName>
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<emphasis id="4BDCEAB1F1463270C61DCF039A691D2B" box="[809,828,536,557]" italics="true" pageId="3" pageNumber="4">in</emphasis>
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<bibRefCitation id="1D394B52F1463270C675CF039D371D28" author="von Post, T. & Kuntze, O." bookContentInfo="714 pp." box="[833,1122,536,558]" journalOrPublisher="Wretmanianis, Upsalla" pageId="3" pageNumber="4" publicationUrl="http://dx.doi.org/10.5962/bhl.title.58320" refId="ref30014" refString="von Post, T. & Kuntze, O. (1903) Lexicon generum phanerogamarum inde ab anno MDCXXXVII cum nomenclatura legitima internationali simul scientifica. Wretmanianis, Upsalla, 714 pp. http: // dx. doi. org / 10.5962 / bhl. title. 58320" title="Lexicon generum phanerogamarum inde ab anno MDCXXXVII cum nomenclatura legitima internationali simul scientifica" type="book" year="1903">von Post & Kuntze (1903: 13)</bibRefCitation>
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. Type:—
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<taxonomicName id="BEA84D20F1463270C18DCF0C98771D54" ID-CoL="8NP4" authority="Lindley (1853 a: 1)" authorityName="Lindley" authorityPageNumber="1" authorityYear="1853" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="cyanea">
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<emphasis id="4BDCEAB1F1463270C18DCF0C9C0E1D2B" box="[1209,1371,535,557]" italics="true" pageId="3" pageNumber="4">Acacallis cyanea</emphasis>
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<bibRefCitation id="1D394B52F1463270C055CF0C98771D54" author="Lindley, J." journalOrPublisher="Matthews, London" pageId="3" pageNumber="4" pagination="1" part="4" publicationUrl="http://dx.doi.org/10.5962/bhl.title.51709" refId="ref29107" refString="Lindley, J. (1853 a) Acacallis. In: Folia orchidacea 4 (Acacallis). Matthews, London, p. 1. http: // dx. doi. org / 10.5962 / bhl. title. 51709" title="Acacallis" type="journal article" volumeTitle="Folia orchidacea" year="1853">Lindley (1853a: 1)</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="791736A3F1463270C5BCCF4498141D9F" blockId="3.[136,1452,535,665]" pageId="3" pageNumber="4">
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<taxonomicName id="BEA84D20F1463270C5BCCF4498AB1D73" ID-CoL="8WGWK" authority="Schlechter ex Cogniaux" authorityName="Schlechter ex Cogniaux" box="[136,510,607,629]" class="Liliopsida" family="Orchidaceae" genus="Kochiophyton" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
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<emphasis id="4BDCEAB1F1463270C5BCCF44985A1D72" box="[136,271,607,628]" italics="true" pageId="3" pageNumber="4">Kochiophyton</emphasis>
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Schlechter ex Cogniaux
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</taxonomicName>
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<emphasis id="4BDCEAB1F1463270C731CF7B9B4D1D73" box="[517,536,608,629]" italics="true" pageId="3" pageNumber="4">in</emphasis>
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<bibRefCitation id="1D394B52F1463270C72BCF449A501D73" author="Cogniaux, A." box="[543,773,607,629]" editor="Martius, C. F. P." journalOrPublisher="Typographia Regia, Munich" pageId="3" pageNumber="4" pagination="573 - 576" part="3" refId="ref27278" refString="Cogniaux, A. (1906 b) Orchidaceae: Kochiophyton. In: Martius, C. F. P., Eicher, A. G. & Urban, I. (Eds.) Flora Brasiliensis 3 (6). Typographia Regia, Munich. pp. 573 - 576, t. 119." title="Orchidaceae: Kochiophyton" type="journal article" volumeTitle="Flora Brasiliensis" year="1906">Cogniaux (1906b: 574)</bibRefCitation>
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.
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<emphasis id="4BDCEAB1F1463270C63BCF449A0E1D72" box="[783,859,607,628]" italics="true" pageId="3" pageNumber="4">Type:—</emphasis>
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<taxonomicName id="BEA84D20F1463270C66FCF449C791D73" ID-CoL="6NN8V" authority="Schlechter ex Cogniaux" authorityName="Schlechter ex Cogniaux" box="[859,1324,607,629]" class="Liliopsida" family="Orchidaceae" genus="Kochiophyton" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="negrense">
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<emphasis id="4BDCEAB1F1463270C66FCF449D6B1D73" box="[859,1086,607,629]" italics="true" pageId="3" pageNumber="4">Kochiophyton negrense</emphasis>
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Schlechter ex Cogniaux
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</taxonomicName>
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<emphasis id="4BDCEAB1F1463270C000CF7B9C121D73" box="[1332,1351,608,629]" italics="true" pageId="3" pageNumber="4">in</emphasis>
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<bibRefCitation id="1D394B52F1463270C07ACF4498681D9F" author="Cogniaux, A." editor="Martius, C. F. P." journalOrPublisher="Typographia Regia, Munich" pageId="3" pageNumber="4" pagination="573 - 576" part="3" refId="ref27278" refString="Cogniaux, A. (1906 b) Orchidaceae: Kochiophyton. In: Martius, C. F. P., Eicher, A. G. & Urban, I. (Eds.) Flora Brasiliensis 3 (6). Typographia Regia, Munich. pp. 573 - 576, t. 119." title="Orchidaceae: Kochiophyton" type="journal article" volumeTitle="Flora Brasiliensis" year="1906">Cogniaux (1906b: 574)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="31B26528F1463270C5BCCFD39DFB1C2C" pageId="3" pageNumber="4" type="etymology">
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<paragraph id="791736A3F1463270C5BCCFD39DFB1C2C" blockId="3.[136,1452,711,2071]" pageId="3" pageNumber="4">
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<emphasis id="4BDCEAB1F1463270C5BCCFD3987B1DE4" bold="true" box="[136,302,712,738]" pageId="3" pageNumber="4">Etymology:—</emphasis>
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Lindley did not explain the source of the name. It probably originates from Greek,
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<emphasis id="4BDCEAB1F1463270C1FDCFDC9C5C1DE4" box="[1225,1289,711,738]" italics="true" pageId="3" pageNumber="4">γάνος</emphasis>
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, splendid, due to the author being delighted with the beauty of this plant, as cited in the protologue—“A very pretty new genus”—or an obscure nymph from Alpine mythology, Veneto, northeastern Italy, also known as Anguana.
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</paragraph>
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</subSubSection>
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<subSubSection id="31B26528F1463270C589CE2F9A851AD1" pageId="3" pageNumber="4" type="description">
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<paragraph id="791736A3F1463270C589CE2F9A851AD1" blockId="3.[136,1452,711,2071]" pageId="3" pageNumber="4">Herbs epiphytic or hemiepiphytic, sympodial, rhizomatous, with thick, spongy roots. Flowering pseudobulbs inconspicuous, homoblastic, fusiform, orbicular in transverse section; cataphylls on pseudobulbs non-leafy, scarious, non-articulate. Leaves 1 per pseudobulb; petiole conduplicate, distinct from blade, margin non-hyaline; blade articulate, plicate, elliptic to narrow-elliptic, base cuneate, apex obtuse to acute, abaxial face with 5 more prominent nerves. Inflorescences racemose, axillary from an inner cataphyll, shorter than plant, rarely longer, erect, pedunculate, rachis lax, blooming acropetally; bracts lanceolate, adpressed; bracteole 1, lanceolate, shorter than the ovary. Flowers resupinate, pedicellate. Sepals with the dorsal one free, and laterals adnate to column feet and oblique, narrow-ovate or ovate. Petals free, narrowly elliptic, elliptic or orbiculate. Lip free, trilobed, unguiculate; lateral lobes highly reduced, deltoid or semi-elliptic; midlobe lateral lobes strongly reduced, margin entire; midlobe flabellate, elliptic or ovate, margin entire, undulate, minutely crenate or fimbriate; callus composed of an outer pair of lamellae with erose margins, fimbriate in distal part between lamellae, or apex with erose laminar margins, antrorse, or longitudinal laminae irregularly denticulate, the external pair higher than the inner one. Column semi-terete, foot shorter than the column, apex with oblong and conspicuous stigmatic wings; anther incumbent, clinandrium not projecting over the anther, margin entire, external face of anther cap bilobed, apex obtuse-denticulate or digitiform, inner face bilocular; pollinia 4, in 2 pairs, juxtaposed, oblong-ovoid, laterally compressed, unequal in size and shape, tegula oblong to elliptic, caudicle amorphous, viscidium transverse-oblong; stigma transverse; rostellum trilobed, midlobe narrowly triangular, lateral lobes reduced with concave margin, not exceeding the rostellum, rostellum remnant trilobed, midlobe acute and unequal to the lateral ones, lateral lobes truncate or trilobed, in the latter case the midlobe longer than the truncate and lateral ones. Capsules fusiform, 6-carinate, smooth surface; seeds not seen.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="31B26528F1463270C589C8C49CC61711" pageId="3" pageNumber="4" type="distribution">
|
|
<paragraph id="791736A3F1463270C589C8C49DC319D5" blockId="3.[136,1452,711,2071]" pageId="3" pageNumber="4">
|
|
<emphasis id="4BDCEAB1F1463270C589C8C49B5A1AFC" bold="true" box="[189,527,1503,1530]" pageId="3" pageNumber="4">Distribution and habitat:—</emphasis>
|
|
The genus occurs in northern and western South America, in the Amazon and Solimies Basins, Guiana and Guaporé Shields. It grows in the Amazon rainforests, in shady places close to water, from sea level to 200 m, rarely higher.
|
|
<taxonomicName id="BEA84D20F1463270C703CB329A5C1944" authorityName="Lindley" box="[567,777,1576,1603]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="pulchella">
|
|
<emphasis id="4BDCEAB1F1463270C703CB329A5C1944" box="[567,777,1576,1603]" italics="true" pageId="3" pageNumber="4">Aganisia pulchella</emphasis>
|
|
</taxonomicName>
|
|
is also recorded for Trinidad and Suriname in riparian forest surrounded by savannahs and in a disjunct area in southern Bahia, eastern Brazil. In Brazil, the species grows in biome Amazônia and vegetation type
|
|
<emphasis id="4BDCEAB1F1463270C4C4CB6B9BE6198D" box="[496,691,1648,1675]" italics="true" pageId="3" pageNumber="4">floresta de igapó</emphasis>
|
|
(black water seasonal floodplain forest),
|
|
<emphasis id="4BDCEAB1F1463270C1A0CB6B9C37198D" box="[1172,1378,1648,1675]" italics="true" pageId="3" pageNumber="4">floresta de várzea</emphasis>
|
|
(clear water seasonal floodplain forest) and
|
|
<emphasis id="4BDCEAB1F1463270C719CB8E9B9719A8" box="[557,706,1685,1710]" italics="true" pageId="3" pageNumber="4">campinarana</emphasis>
|
|
(white sand forest); in biome Cerrado in
|
|
<emphasis id="4BDCEAB1F1463270C1A6CB8E9C1119A8" box="[1170,1348,1684,1710]" italics="true" pageId="3" pageNumber="4">mata de galeria</emphasis>
|
|
(riparian forest); and biome Floresta Atlântica in
|
|
<emphasis id="4BDCEAB1F1463270C772CBA29BF419D4" box="[582,673,1721,1746]" italics="true" pageId="3" pageNumber="4">restinga</emphasis>
|
|
(sedimentary coastal shrub field and forest).
|
|
</paragraph>
|
|
<paragraph id="791736A3F1463270C589CBC69CC61711" blockId="3.[136,1452,711,2071]" pageId="3" pageNumber="4">
|
|
<taxonomicName id="BEA84D20F1463270C589CBC698DA19F0" authorityName="Lindley" box="[189,399,1756,1783]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="pulchella">
|
|
<emphasis id="4BDCEAB1F1463270C589CBC698DA19F0" box="[189,399,1756,1783]" italics="true" pageId="3" pageNumber="4">Aganisia pulchella</emphasis>
|
|
</taxonomicName>
|
|
has a disjunct distribution in northern South America and eastern Brazil. There is evidence that Floresta Atlântica and Amazônia once were a single biome connected in northeastern Brazil, which, due to increased aridity during the Pleistocene, became separated by Cerrado and Caatinga (
|
|
<bibRefCitation id="1D394B52F1463270C6FDCA3F9DE51838" author="Bigarella, J. J. & Andrade-Lima, D. & Riehns, P. J." box="[969,1200,1827,1854]" journalOrPublisher="Anais da Academia Brasileira de Ciencias" pageId="3" pageNumber="4" pagination="412 - 462" part="47" refId="ref26487" refString="Bigarella, J. J., Andrade-Lima, D. & Riehns, P. J. (1975) Consideracies a respeito das mudancas paleoambientais na distribuic " o de algumas especies vegetais e animais no Brasil. Anais da Academia Brasileira de Ciencias 47 (supl.): 412 - 462." title="Consideracies a respeito das mudancas paleoambientais na distribuic " o de algumas especies vegetais e animais no Brasil" type="journal article" year="1975">
|
|
Bigarella
|
|
<emphasis id="4BDCEAB1F1463270C10CCA3E9D3A1838" box="[1080,1135,1828,1854]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
|
1975
|
|
</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="1D394B52F1463270C18ECA3F9C271838" author="Mori, S. A. & Boom, B. M. & Prance, G. T." box="[1210,1394,1828,1854]" journalOrPublisher="Brittonia" pageId="3" pageNumber="4" pagination="233 - 245" part="33" publicationUrl="http://dx.doi.org/10.2307/2806330" refId="ref29646" refString="Mori, S. A., Boom, B. M. & Prance, G. T. (1981) Distribution patterns and conservation of eastern Brazilian coastal forest tree species. Brittonia 33: 233 - 245. http: // dx. doi. org / 10.2307 / 2806330" title="Distribution patterns and conservation of eastern Brazilian coastal forest tree species" type="journal article" year="1981">
|
|
Mori
|
|
<emphasis id="4BDCEAB1F1463270C1CDCA3E9C651838" box="[1273,1328,1828,1854]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
|
|
1981
|
|
</bibRefCitation>
|
|
). For the same species, occurrence in Trinidad could at first also sound like a disjunction because it is politically included in the Caribbean, but it is a continental island on the South American platform near the coast of Venezuela and not on the Caribbean platform like the rest of the Antilles (
|
|
<bibRefCitation id="1D394B52F1463270C7F5CA8B9A3118AD" author="Schultes, R. E." bookContentInfo="275 pp." box="[705,868,1936,1963]" journalOrPublisher="Pergamon, London" pageId="3" pageNumber="4" refId="ref31519" refString="Schultes, R. E. (1960) Native orchids of Trinidad and Tobago. Pergamon, London, 275 pp." title="Native orchids of Trinidad and Tobago" type="book" year="1960">Schultes 1960</bibRefCitation>
|
|
). Recent field observations in southern Bahia by one of us (TECM) suggest that its life form can be classified tentatively as a hemiepiphyte because most individuals grow on plant litter and later climb up the bases of tree trunks. TDWG code: 81 TRT-OO 82 FRG-OO GUY-OO SUR-OO VEN-OO 83 COL-OO PER-OO 84 BZC-MT BZE-BA BZE-MA BZN-AC BZN-AM BZN-PA BZN-RO BZN-RR.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="31B26528F1413277C589CD849B881E4C" pageId="4" pageNumber="5" type="diagnosis">
|
|
<paragraph id="791736A3F1413277C589CD849B881E4C" blockId="4.[136,1452,159,619]" pageId="4" pageNumber="5">
|
|
<emphasis id="4BDCEAB1F1413277C589CD8498B01FBC" bold="true" box="[189,485,159,186]" pageId="4" pageNumber="5">Diagnostic characters:—</emphasis>
|
|
The genus differs from the others in this study by its medium-sized plants (20 to 50 cm long), rhizomatous, inconspicous pseudobulbs covered by non-leafy, non-articulate and scarious cataphylls, articulate leaves with hyaline margins and plicate blades, inflorescences shorter than the leaves, rarely longer, lax rachis, big flowers (sepals up to 3.8 cm long), lateral sepals adnate to the column foot, lip with a hippocrepiform and ornamented callus and column with conspicuous stigmatic wings.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="31B26528F1413277C589CC489B0D1D6D" pageId="4" pageNumber="5" type="discussion">
|
|
<paragraph id="791736A3F1413277C589CC489B0D1D6D" blockId="4.[136,1452,159,619]" pageId="4" pageNumber="5">
|
|
<emphasis id="4BDCEAB1F1413277C589CC4898891E68" bold="true" box="[189,476,339,366]" pageId="4" pageNumber="5">Generic delimitation:—</emphasis>
|
|
The segregation of
|
|
<taxonomicName id="BEA84D20F1413277C78ECC4F9A771E68" authorityName="Lindley" authorityYear="1853" box="[698,802,340,366]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C78ECC4F9A771E68" box="[698,802,340,366]" italics="true" pageId="4" pageNumber="5">Acacallis</emphasis>
|
|
</taxonomicName>
|
|
from
|
|
<taxonomicName id="BEA84D20F1413277C65FCC4F9A9A1E68" box="[875,975,340,366]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C65FCC4F9A9A1E68" box="[875,975,340,366]" italics="true" pageId="4" pageNumber="5">Aganisia</emphasis>
|
|
</taxonomicName>
|
|
was accepted without critical analysis by some authors (
|
|
<bibRefCitation id="1D394B52F1413277C41ECC6398A51E94" author="Schlechter, R." box="[298,496,376,402]" journalOrPublisher="Orchis" pageId="4" pageNumber="5" pagination="6 - 16" part="12" refId="ref31347" refString="Schlechter, R. (1918 a) Die Gattung Aganisia Ldl. und ihre Verwandten. Orchis 12: 6 - 16." title="Die Gattung Aganisia Ldl. und ihre Verwandten" type="journal article" year="1918">Schlechter 1918a</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="1D394B52F1413277C4CFCC639BC11E94" author="Hoehne, F. C." box="[507,660,375,402]" editor="Hoehne, F. C." journalOrPublisher="Secretaria de Agricultura do Estado de S " o Paulo, S " o Paulo" pageId="4" pageNumber="5" pagination="69 - 192" part="12" refId="ref28602" refString="Hoehne, F. C. (1953) Orchidaceas: 121 - Promenaea a 138 - Chaubardia. In: Hoehne, F. C. (Ed.) Flora Brasilica 12 (2). Secretaria de Agricultura do Estado de S " o Paulo, S " o Paulo, pp. 69 - 192, t. 36 - 84." title="Orchidaceas: 121 - Promenaea a 138 - Chaubardia" type="journal article" volumeTitle="Flora Brasilica" year="1953">Hoehne 1953</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="1D394B52F1413277C7ABCC639A921E94" author="Senghas, K. & Gerlach, G." box="[671,967,376,402]" editor="Brieger, F. G." journalOrPublisher="Paul Parey, Berlin" pageId="4" pageNumber="5" pagination="1713 - 1727" part="3" refId="ref31682" refString="Senghas, K. & Gerlach, G. (1993 c) Subtribus Zygopetalinae: 708. Batemania - 713. Eriopsis. In: Brieger, F. G., Maatsch, R. & Senghas, K. (Eds.) Rudolf Schlechter Die Orchideen I / B, 28 3 ed. Paul Parey, Berlin, pp. 1713 - 1727." title="Subtribus Zygopetalinae: 708. Batemania - 713. Eriopsis" type="journal article" volumeTitle="Rudolf Schlechter Die Orchideen I / B, 28" year="1993">Senghas & Gerlach 1993c</bibRefCitation>
|
|
).
|
|
<taxonomicName id="BEA84D20F1413277C6EDCC639D141E94" authorityName="Lindley" authorityYear="1853" box="[985,1089,376,402]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C6EDCC639D141E94" box="[985,1089,376,402]" italics="true" pageId="4" pageNumber="5">Acacallis</emphasis>
|
|
</taxonomicName>
|
|
would include
|
|
<taxonomicName id="BEA84D20F1413277C1D9CC639CFE1E94" authorityName="Lindley" authorityYear="1853" box="[1261,1451,376,402]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="cyanea">
|
|
<emphasis id="4BDCEAB1F1413277C1D9CC639CFE1E94" box="[1261,1451,376,402]" italics="true" pageId="4" pageNumber="5">Acacallis cyanea</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="BEA84D20F1413277C582CC8798C21EB0" box="[182,407,412,438]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="frimbriata">
|
|
<emphasis id="4BDCEAB1F1413277C582CC8798C21EB0" box="[182,407,412,438]" italics="true" pageId="4" pageNumber="5">Acacallis frimbriata</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="BEA84D20F1413277C4E6CC879B631EB0" box="[466,566,412,438]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C4E6CC879B631EB0" box="[466,566,412,438]" italics="true" pageId="4" pageNumber="5">Aganisia</emphasis>
|
|
</taxonomicName>
|
|
only
|
|
<taxonomicName id="BEA84D20F1413277C747CC879A101EB0" authorityName="Lindley" box="[627,837,412,438]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="pulchella">
|
|
<emphasis id="4BDCEAB1F1413277C747CC879A101EB0" box="[627,837,412,438]" italics="true" pageId="4" pageNumber="5">Aganisia pulchella</emphasis>
|
|
</taxonomicName>
|
|
. These authors referred to the lip of
|
|
<taxonomicName id="BEA84D20F1413277C1E1CC879C6C1EB0" box="[1237,1337,412,438]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C1E1CC879C6C1EB0" box="[1237,1337,412,438]" italics="true" pageId="4" pageNumber="5">Aganisia</emphasis>
|
|
</taxonomicName>
|
|
as sessile.
|
|
<bibRefCitation id="1D394B52F1413277C5BCCCDB986E1EDC" author="Schultes, R. E." box="[136,315,447,474]" journalOrPublisher="Lloydia" pageId="4" pageNumber="5" pagination="88 - 99" part="21" refId="ref31495" refString="Schultes, R. E. (1958) Orchidaceae Neotropicales V: generis Aganisiae Synopsis. Lloydia 21: 88 - 99." title="Orchidaceae Neotropicales V: generis Aganisiae Synopsis" type="journal article" year="1958">Schultes (1958)</bibRefCitation>
|
|
discussed this, and we agree with his views on the reliability of this character to distinguish these genera. We concluded that in fact the lip is unguiculate, and we also found that the column foot is shorter than column. Apart from that, we did not find any other diagnosable characters.
|
|
<bibRefCitation id="1D394B52F1413277C6A4CF139DD11D24" author="Whitten, M. W. & Williams, N. H. & Dressler, R. L. & Gerlach, G. & Pupulin, F." box="[912,1156,519,546]" journalOrPublisher="Lankesteriana" pageId="4" pageNumber="5" pagination="87 - 107" part="5" refId="ref32053" refString="Whitten, M. W., Williams, N. H., Dressler, R. L., Gerlach, G. & Pupulin, F. (2005) Generic relationships of Zygopetalinae (Orchidaceae: Cymbidieae): combined molecular evidence. Lankesteriana 5: 87 - 107." title="Generic relationships of Zygopetalinae (Orchidaceae: Cymbidieae): combined molecular evidence" type="journal article" year="2005">
|
|
Whitten
|
|
<emphasis id="4BDCEAB1F1413277C6C0CF129D7A1D24" box="[1012,1071,520,546]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
|
|
(2005)
|
|
</bibRefCitation>
|
|
in a combined molecular analysis showed that
|
|
<taxonomicName id="BEA84D20F1413277C441CF379B121D40" authorityName="Lindley" box="[373,583,556,582]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="pulchella">
|
|
<emphasis id="4BDCEAB1F1413277C441CF379B121D40" box="[373,583,556,582]" italics="true" pageId="4" pageNumber="5">Aganisia pulchella</emphasis>
|
|
</taxonomicName>
|
|
is sister to
|
|
<taxonomicName id="BEA84D20F1413277C7F0CF379AD71D40" authorityName="Lindley" authorityYear="1853" box="[708,898,556,582]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="cyanea">
|
|
<emphasis id="4BDCEAB1F1413277C7F0CF379AD71D40" box="[708,898,556,582]" italics="true" pageId="4" pageNumber="5">Acacallis cyanea</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="BEA84D20F1413277C68CCF379DDB1D40" box="[952,1166,556,582]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="fimbriata">
|
|
<emphasis id="4BDCEAB1F1413277C68CCF379DDB1D40" box="[952,1166,556,582]" italics="true" pageId="4" pageNumber="5">Acacallis fimbriata</emphasis>
|
|
</taxonomicName>
|
|
. Thus, we conclude there is no reason to separate these two genera.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="31B26528F1413277C5BCCFB49BD41DCE" box="[136,641,686,713]" pageId="4" pageNumber="5" type="multiple">
|
|
<paragraph id="791736A3F1413277C5BCCFB49BD41DCE" blockId="4.[136,641,686,713]" box="[136,641,686,713]" pageId="4" pageNumber="5">
|
|
<heading id="225F81CFF1413277C5BCCFB49BD41DCE" bold="true" box="[136,641,686,713]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="0">
|
|
<emphasis id="4BDCEAB1F1413277C5BCCFB49BD41DCE" bold="true" box="[136,641,686,713]" pageId="4" pageNumber="5">
|
|
Key to the species and forms of
|
|
<taxonomicName id="BEA84D20F1413277C721CFB59BD41DCE" box="[533,641,686,712]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="4BDCEAB1F1413277C721CFB59BD41DCE" bold="true" box="[533,641,686,712]" italics="true" pageId="4" pageNumber="5">Aganisia</emphasis>
|
|
</taxonomicName>
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="31B26528F1413277C5BCCFF09CFE1C97" pageId="4" pageNumber="5" type="key">
|
|
<key id="0D396192F1413277C5BCCFF09CFE1C97" pageId="4" pageNumber="5">
|
|
<keyStep id="C25C2E06F1413277C5BCCFF09CFE1C1B" pageId="4" pageNumber="5">
|
|
<paragraph id="791736A3F1413277C5BCCFF09CFF1C06" blockId="4.[136,1451,747,913]" box="[136,1450,747,769]" pageId="4" pageNumber="5">
|
|
<keyLead id="C2599596F1413277C5BCCFF09CFF1C06" box="[136,1450,747,769]" pageId="4" pageNumber="5">
|
|
1 Lip with lateral lobes unconstricted, margin of midlobe entire........................................................................................
|
|
<taxonomicName id="BEA84D20F1413277C003CFF79CFF1C06" ID-CoL="65LN9" authorityName="Lindley" box="[1335,1450,747,769]" class="Liliopsida" family="Orchidaceae" genus="Aganisia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="pulchella">
|
|
<emphasis id="4BDCEAB1F1413277C003CFF79CFF1C06" box="[1335,1450,747,769]" italics="true" pageId="4" pageNumber="5">A. pulchella</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="791736A3F1413277C5BCCE139CFE1C1B" blockId="4.[136,1451,747,913]" box="[136,1451,776,798]" pageId="4" pageNumber="5">
|
|
<keyLead id="C2599596F1413277C5BCCE139CFE1C1B" box="[136,1451,776,798]" pageId="4" pageNumber="5">- Lip with lateral lobes strongly constricted distally, midlobe margin undulate and inconspicuously crenate or fimbriate ................2</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="C25C2E06F1413277C5BCCE3E9CFE1C51" pageId="4" pageNumber="5">
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<paragraph id="791736A3F1413277C5BCCE3E9CFF1C3C" blockId="4.[136,1451,747,913]" box="[136,1450,805,827]" pageId="4" pageNumber="5">
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<keyLead id="C2599596F1413277C5BCCE3E9CFF1C3C" box="[136,1450,805,827]" pageId="4" pageNumber="5">
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2 Lip with midlobe elliptic to ovate, strongly concave, margin fimbriate ..........................................................................
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|
<taxonomicName id="BEA84D20F1413277C003CE3E9CFF1C3C" ID-CoL="8NP5" box="[1335,1450,805,826]" class="Liliopsida" family="Orchidaceae" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="fimbriata">
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<emphasis id="4BDCEAB1F1413277C003CE3E9CFF1C3C" box="[1335,1450,805,826]" italics="true" pageId="4" pageNumber="5">A. fimbriata</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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|
<paragraph id="791736A3F1413277C5BCCE599CFE1C51" blockId="4.[136,1451,747,913]" box="[136,1451,834,856]" pageId="4" pageNumber="5">
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<keyLead id="C2599596F1413277C5BCCE599CFE1C51" box="[136,1451,834,856]" pageId="4" pageNumber="5">- Lip with midlobe flabellate, slightly concave, margin undulate or inconspicuously crenate ............................................................3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="C25C2E06F1413277C5BCCE449CFE1C97" pageId="4" pageNumber="5">
|
|
<paragraph id="791736A3F1413277C5BCCE449CFE1C75" blockId="4.[136,1451,747,913]" box="[136,1451,862,884]" pageId="4" pageNumber="5">
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|
<keyLead id="C2599596F1413277C5BCCE449CFE1C75" box="[136,1451,862,884]" pageId="4" pageNumber="5">
|
|
3 Sepals, petals and lip white, rarely lip yellowish .....................................................................................................
|
|
<taxonomicName id="BEA84D20F1413277C033CE449CFE1C75" ID-CoL="5S6WV" box="[1287,1451,862,884]" class="Liliopsida" family="Orchidaceae" form="alba" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="form" species="cyanea">
|
|
<emphasis id="4BDCEAB1F1413277C033CE449C331C72" box="[1287,1382,863,884]" italics="true" pageId="4" pageNumber="5">A. cyanea</emphasis>
|
|
f.
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|
<emphasis id="4BDCEAB1F1413277C0B4CE459CFE1C75" box="[1408,1451,862,883]" italics="true" pageId="4" pageNumber="5">alba</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="791736A3F1413277C5BCCE679CFE1C97" blockId="4.[136,1451,747,913]" box="[136,1451,891,913]" pageId="4" pageNumber="5">
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|
<keyLead id="C2599596F1413277C5BCCE679CFE1C97" box="[136,1451,891,913]" pageId="4" pageNumber="5">
|
|
- Sepals, petals and lip lilac, rarely midlobe of lip yellowish.................................................................................
|
|
<taxonomicName id="BEA84D20F1413277C1DACE679CFE1C97" ID-CoL="8NP4" authority="f. cyanea" box="[1262,1451,891,913]" class="Liliopsida" family="Orchidaceae" form="cyanea" genus="Acacallis" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="form" species="cyanea">
|
|
<emphasis id="4BDCEAB1F1413277C1DACE679C181C97" box="[1262,1357,892,913]" italics="true" pageId="4" pageNumber="5">A. cyanea</emphasis>
|
|
f.
|
|
<emphasis id="4BDCEAB1F1413277C053CE679CFE1C97" box="[1383,1451,892,913]" italics="true" pageId="4" pageNumber="5">cyanea</emphasis>
|
|
</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</key>
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</subSubSection>
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</treatment>
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</document> |