286 lines
35 KiB
XML
286 lines
35 KiB
XML
<document ID-DOI="10.5252/zoosystema2022v44a20" ID-ISSN="1638-9387" ID-Zenodo-Dep="7304128" ID-ZooBank="urn:lsid:zoobank.org:pub:26622CAD-FF77-45CD-A95E-F0BD31BA4FBF" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1667910815094" checkinUser="felipe" docAuthor="Cutrim, Marcelo, M, Alberto, Silva-Neto, oreira da, Rafael, José Albertino, García, Alfonso Nery & Aldrete" docDate="2022" docId="D875A26CFFD6BB1C2914F99E5C860BDB" docLanguage="en" docName="Zoosystema.44.20.493-501.pdf" docOrigin="Zoosystema 44 (20)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.3:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="3" docTitle="Ptiloneura leonardoi Cutrim & M & Silva-Neto & Rafael & García & Aldrete 2022, n. sp." docType="treatment" docVersion="2" lastPageNumber="498" masterDocId="244CDA14FFD3BB1B2820FF8C5E190A55" masterDocTitle="The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species" masterLastPageNumber="501" masterPageNumber="493" pageNumber="496" updateTime="1667911224788" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Ptiloneura Enderlein, 1901 (Psocodea, ‘ Psocoptera’, Ptiloneuridae) in the Brazilian Amazon Forest and Atlantic Forest: new species, variations in forewings and a key to the species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cutrim, Marcelo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>M, Alberto</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Silva-Neto, oreira da</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rafael, José Albertino</mods:namePart>
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<mods:affiliation>Instituto Nacional de Pesquisas da Amazônia, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas (Brazil) marcelocutrim @ gmail. com (corresponding author).</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>García, Alfonso Nery</mods:namePart>
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<mods:affiliation>Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma Apdo. Postal 70 - 153, Ciudad de</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aldrete</mods:namePart>
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<mods:affiliation>México, México (Mexico)</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-11-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>44</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>20</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>493</mods:start>
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<mods:end>501</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5252/zoosystema2022v44a20</mods:identifier>
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<mods:identifier type="ISSN">1638-9387</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7304128</mods:identifier>
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<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:26622CAD-FF77-45CD-A95E-F0BD31BA4FBF</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7304144" ID-Zenodo-Dep="7304144" LSID="urn:lsid:plazi:treatment:D875A26CFFD6BB1C2914F99E5C860BDB" httpUri="http://treatment.plazi.org/id/D875A26CFFD6BB1C2914F99E5C860BDB" lastPageId="7" lastPageNumber="498" pageId="5" pageNumber="496">
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<subSubSection box="[308,599,1554,1581]" pageId="5" pageNumber="496" type="nomenclature">
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<paragraph blockId="5.[308,599,1554,1613]" box="[308,599,1554,1581]" pageId="5" pageNumber="496">
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<heading box="[308,599,1554,1581]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="496" reason="2">
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<taxonomicName authority="Cutrim & M & Silva-Neto & Rafael & García & Aldrete, 2022" authorityName="Cutrim & M & Silva-Neto & Rafael & García & Aldrete" authorityYear="2022" box="[308,533,1554,1581]" class="Insecta" family="Ptiloneuridae" genus="Ptiloneura" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="496" phylum="Arthropoda" rank="species" species="leonardoi" status="n. sp.">
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<emphasis bold="true" box="[308,533,1554,1581]" italics="true" pageId="5" pageNumber="496">Ptiloneura leonardoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[540,599,1555,1581]" pageId="5" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="496" type="description">
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<paragraph blockId="5.[308,599,1554,1613]" box="[414,493,1586,1613]" pageId="5" pageNumber="496">
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(
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<figureCitation box="[421,485,1586,1613]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2</figureCitation>
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)
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</paragraph>
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<paragraph blockId="5.[155,752,1654,1672]" box="[155,752,1654,1672]" pageId="5" pageNumber="496">
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<uri box="[155,752,1654,1672]" pageId="5" pageNumber="496">
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||
urn:lsid:zoobank.org:act:
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<uuid box="[355,752,1654,1672]" pageId="5" pageNumber="496">13EFDBFB-79B6-4C5A-AF4A-2E35DD9B941F</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="496" type="materials_examined">
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<paragraph blockId="5.[132,776,1704,1754]" pageId="5" pageNumber="496">
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<materialsCitation collectingDate="1972-06" collectionCode="TYPE, MATERIAL, INPA" collectorName="Fritz Plaumann" country="Brazil" latitude="-27.150278" location="Nova Teutonia" longLatPrecision="20" longitude="-52.417778" pageId="5" pageNumber="496" specimenCount="1" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="holotype">
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<collectionCode box="[132,280,1704,1727]" pageId="5" pageNumber="496">TYPE MATERIAL</collectionCode>
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. —
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<emphasis bold="true" box="[325,420,1704,1728]" pageId="5" pageNumber="496">
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<typeStatus box="[325,420,1704,1728]" pageId="5" pageNumber="496">Holotype</typeStatus>
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</emphasis>
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.
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<emphasis bold="true" box="[432,490,1705,1729]" pageId="5" pageNumber="496">
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<collectingCountry box="[432,490,1705,1729]" name="Brazil" pageId="5" pageNumber="496">Brazil</collectingCountry>
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</emphasis>
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•
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<specimenCount box="[514,553,1704,1728]" pageId="5" pageNumber="496" type="male">1 ♂</specimenCount>
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;
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<collectingRegion box="[566,713,1704,1728]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="496">Santa Catarina</collectingRegion>
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,
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<location LSID="urn:lsid:plazi:treatment:D875A26CFFD6BB1C2914F99E5C860BDB:550345A1FFD6BB1E2AF3F9245EC10C8F" country="Brazil" latitude="-27.150278" longLatPrecision="20" longitude="-52.417778" name="Nova Teutonia" pageId="5" pageNumber="496" stateProvince="Santa Catarina">Nova Teutônia</location>
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;
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<geoCoordinate box="[224,320,1730,1754]" degrees="27" direction="south" minutes="09" orientation="latitude" pageId="5" pageNumber="496" precision="15" seconds="1" value="-27.150278">27°09’1”S</geoCoordinate>
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,
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<geoCoordinate box="[327,431,1730,1754]" degrees="52" direction="west" minutes="25" orientation="longitude" pageId="5" pageNumber="496" precision="15" seconds="4" value="-52.417778">52°25’4”W</geoCoordinate>
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;
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<date box="[441,521,1730,1754]" pageId="5" pageNumber="496" value="1972-06">
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<collectingDate box="[441,521,1730,1754]" pageId="5" pageNumber="496" value="1972-06">VI.1972</collectingDate>
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</date>
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;
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<collectorName box="[528,669,1730,1754]" pageId="5" pageNumber="496">Fritz Plaumann</collectorName>
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leg.;
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<collectionCode box="[714,772,1730,1754]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="496">INPA</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph blockId="5.[131,775,1776,1853]" pageId="5" pageNumber="496">
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<typeStatus box="[132,237,1777,1800]" pageId="5" pageNumber="496">HOLOTYPE</typeStatus>
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CONDITION. — Head, fore and hind wings, phallosome, hypandrium, epiproct, and paraproct mounted on microslide. The remaining body parts are preserved in 70% ethanol.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="496" type="etymology">
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<paragraph blockId="5.[132,775,1875,1926]" pageId="5" pageNumber="496">ETYMOLOGY. — The species honors Leonardo de Assis Castro, son of Marcelo Cutrim.</paragraph>
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<paragraph blockId="5.[132,775,1948,2025]" pageId="5" pageNumber="496">
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GEOGRAPHICAL RECORDS. — Known only from the
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<typeStatus box="[651,693,1948,1972]" pageId="5" pageNumber="496">type</typeStatus>
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locality,
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<collectingCountry box="[132,191,1974,1998]" name="Brazil" pageId="5" pageNumber="496">Brazil</collectingCountry>
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,
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<collectingRegion box="[200,397,1974,1999]" country="Brazil" name="Santa Catarina" pageId="5" pageNumber="496">Santa Catarina state</collectingRegion>
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, Nova Teutônia city, on Atlantic Forest in southeastern
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<collectingCountry box="[287,346,2001,2025]" name="Brazil" pageId="5" pageNumber="496">Brazil</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="496" type="diagnosis">
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<paragraph blockId="5.[811,1456,215,399]" pageId="5" pageNumber="496">
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DIAGNOSIS. — Central sclerite of the hypandrium with two curved, postero-lateral processes, distally acuminate, curved outward resulting in a U-shaped posterior concavity; side sclerites short, almost rectangular (
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<figureCitation box="[933,1008,295,319]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2G</figureCitation>
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); forewings with a slender, marginal pigmented band from R4+5 to areola postica (
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<figureCitation box="[1143,1216,322,346]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2B</figureCitation>
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); mesal endophallic sclerite of phallosome transverse, with large, median, pointed process, and long, slender, postero-lateral processes (
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<figureCitation box="[1238,1312,375,399]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2F</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="7" lastPageNumber="498" pageId="5" pageNumber="496" type="description">
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<paragraph blockId="5.[813,1458,471,752]" box="[813,957,471,497]" pageId="5" pageNumber="496">DESCRIPTION</paragraph>
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<paragraph blockId="5.[813,1458,471,752]" box="[813,974,502,528]" pageId="5" pageNumber="496">
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<emphasis box="[813,974,502,528]" italics="true" pageId="5" pageNumber="496">
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Male (
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<typeStatus box="[880,967,502,528]" pageId="5" pageNumber="496">holotype</typeStatus>
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)
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</emphasis>
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</paragraph>
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<paragraph blockId="5.[813,1458,471,752]" pageId="5" pageNumber="496">
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<emphasis bold="true" box="[813,883,534,560]" pageId="5" pageNumber="496">Color.</emphasis>
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Compound eyes black, ocelli hyaline; head pattern (
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<figureCitation box="[822,908,566,592]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2A</figureCitation>
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). Scape and pedicel pale yellow, f1–f3 pale brown. Mx4 brown. Femora pale yellow; tibiae pale yellow with a brown band distally; tarsomeres 1-3 pale yellow. Forewing (
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<figureCitation box="[822,907,662,688]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2B</figureCitation>
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) veins brown, pterostigma brown with a median hyaline area, a brown spot in R
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<subScript attach="left" box="[1148,1178,706,723]" fontSize="7" pageId="5" pageNumber="496">2+3</subScript>
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and R
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<subScript attach="left" box="[1248,1278,706,723]" fontSize="7" pageId="5" pageNumber="496">4+5</subScript>
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at wing margin. Hindwings (
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<figureCitation box="[947,1031,725,752]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2C</figureCitation>
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) hyaline.
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</paragraph>
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<paragraph blockId="5.[810,1458,789,1868]" pageId="5" pageNumber="496">
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<emphasis bold="true" box="[813,959,789,815]" pageId="5" pageNumber="496">Morphology.</emphasis>
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Outer cusp of lacinial tips broad, with seven short denticles (
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<figureCitation box="[975,1056,821,847]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2A</figureCitation>
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). Right forewing (
|
||
<figureCitation box="[1242,1319,821,847]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2B</figureCitation>
|
||
) pterostigma narrow proximally, wider in the middle; areola postica wide, tall, apically rounded; Rs shorter than its branches, slightly concave, R
|
||
<subScript attach="left" box="[932,963,930,947]" fontSize="7" pageId="5" pageNumber="496">2+3</subScript>
|
||
almost straight, R
|
||
<subScript attach="left" box="[1165,1196,929,946]" fontSize="7" pageId="5" pageNumber="496">4+5</subScript>
|
||
with a median concave curve; M stem concave proximally, then almost straight, with six primary branches. Right hindwing (
|
||
<figureCitation box="[1244,1329,980,1007]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2C</figureCitation>
|
||
) Rs almost straight, R
|
||
<subScript attach="left" box="[930,960,1025,1042]" fontSize="7" pageId="5" pageNumber="496">2+3</subScript>
|
||
straight, R
|
||
<subScript attach="left" box="[1086,1116,1025,1042]" fontSize="7" pageId="5" pageNumber="496">4+5</subScript>
|
||
with a median concave curve; M with four primary branches, M
|
||
<subScript attach="left" box="[1200,1210,1057,1074]" fontSize="7" pageId="5" pageNumber="496">2</subScript>
|
||
forked. Left forewing (
|
||
<figureCitation box="[822,913,1076,1102]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2D</figureCitation>
|
||
) differing from the right forewing in having M stem with five primary branches, M
|
||
<subScript attach="left" box="[1220,1230,1121,1138]" fontSize="7" pageId="5" pageNumber="496">5</subScript>
|
||
forked, resulting in M
|
||
<subScript attach="left" box="[839,857,1153,1170]" fontSize="7" pageId="5" pageNumber="496">5a</subScript>
|
||
and M
|
||
<subScript attach="left" box="[940,960,1153,1170]" fontSize="7" pageId="5" pageNumber="496">5b</subScript>
|
||
. Left hindwing (
|
||
<figureCitation box="[1152,1236,1140,1166]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2E</figureCitation>
|
||
) differing from the right hindwing in having the M stem with four primary branches. Hypandrium (
|
||
<figureCitation box="[1086,1173,1204,1230]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2G</figureCitation>
|
||
) of three sclerites, central sclerite with with fields of long macrosetae on each side of the longitudinal midline, that do not reach the apices of the postero-lateral processes. Phallosome (
|
||
<figureCitation box="[1240,1324,1299,1326]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2F</figureCitation>
|
||
) side struts slender, V-shaped, anteriorly independent, long, narrow, distal ends almost triangular as an outward directed hook, not fused posteriorly to external parameres, these slender, proximally dilated, elongated, narrowing distally, with a field of pores and an apical end inwardly directed. Four pairs of endophallic sclerites, an anterior small pair with two acuminate projections, the longest directed inward almost touching in the middle and the short, directed outward; a lateral bow-shaped pair curved inward, proximally wide, triangular, distally narrow, with a radular end area; a posterior pair, long, slightly wide proximally, almost straight, slightly curved outward distally and a mesal triangular sclerite, with two wide lateral process distally rounded. Epiproct (
|
||
<figureCitation box="[1360,1445,1714,1740]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2H</figureCitation>
|
||
) rhomboid, with three short central setae, two long lateral and a set of posterior ones. Paraprocts straight based, broad, sensory fields with 33 trichobothria on basal rosettes, setae as illustrated (
|
||
<figureCitation box="[965,1055,1841,1868]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="5" pageNumber="496">Fig. 2H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[813,1457,1905,2028]" pageId="5" pageNumber="496">
|
||
<emphasis bold="true" box="[813,1145,1905,1931]" pageId="5" pageNumber="496">Measurements (in microns).</emphasis>
|
||
FW: 5503, HW: 3618, F: 1310, T: 2249, t1: 879, t2: 67, t3: 165, f1: 927, f2: 904, f3: 567, Mx4: 270, IO: 545, D: 489, d: 317, IO/d:1.71, PO: 0.58.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7304132" ID-Zenodo-Dep="7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="6" pageNumber="497" startId="6.[132,143,1962,1980]" targetBox="[166,1420,216,1901]" targetPageId="6">
|
||
<paragraph blockId="6.[132,1455,1962,2025]" pageId="6" pageNumber="497">
|
||
FIG. 2. —
|
||
<taxonomicName authority="Cutrim & M & Silva-Neto & Rafael & García & Aldrete, 2022" authorityName="Cutrim & M & Silva-Neto & Rafael & García & Aldrete" authorityYear="2022" box="[212,378,1962,1980]" class="Insecta" family="Ptiloneuridae" genus="Ptiloneura" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="497" phylum="Arthropoda" rank="species" species="leonardoi" status="n. sp.">
|
||
<emphasis box="[212,378,1962,1980]" italics="true" pageId="6" pageNumber="497">Ptiloneura leonardoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[383,431,1962,1980]" pageId="6" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
|
||
holotype:
|
||
<emphasis bold="true" box="[517,530,1962,1980]" pageId="6" pageNumber="497">A</emphasis>
|
||
, front view of head;
|
||
<emphasis bold="true" box="[699,712,1962,1980]" pageId="6" pageNumber="497">B</emphasis>
|
||
, right forewing;
|
||
<emphasis bold="true" box="[847,861,1962,1980]" pageId="6" pageNumber="497">C</emphasis>
|
||
, right hindwing;
|
||
<emphasis bold="true" box="[1000,1014,1962,1980]" pageId="6" pageNumber="497">D</emphasis>
|
||
, left forewing;
|
||
<emphasis bold="true" box="[1136,1148,1962,1980]" pageId="6" pageNumber="497">E</emphasis>
|
||
, left hindwing;
|
||
<emphasis bold="true" box="[1275,1286,1962,1980]" pageId="6" pageNumber="497">F</emphasis>
|
||
, phallosome;
|
||
<emphasis bold="true" box="[1403,1417,1962,1980]" pageId="6" pageNumber="497">G</emphasis>
|
||
, hypandrium;
|
||
<emphasis bold="true" box="[222,236,1985,2003]" pageId="6" pageNumber="497">H</emphasis>
|
||
, left paraproct and epiproct. Abbreviations:
|
||
<emphasis bold="true" box="[602,633,1985,2003]" pageId="6" pageNumber="497">aes</emphasis>
|
||
, anterior endophallic sclerites;
|
||
<emphasis bold="true" box="[893,915,1985,2003]" pageId="6" pageNumber="497">ep</emphasis>
|
||
, external parameres;
|
||
<emphasis bold="true" box="[1096,1122,1985,2003]" pageId="6" pageNumber="497">les</emphasis>
|
||
, lateral endophallic sclerites;
|
||
<emphasis bold="true" box="[1368,1395,1985,2003]" pageId="6" pageNumber="497">ms</emphasis>
|
||
, mesal sclerite;
|
||
<emphasis bold="true" box="[202,240,2007,2025]" pageId="6" pageNumber="497">msp</emphasis>
|
||
, mesal sclerite processes;
|
||
<emphasis bold="true" box="[467,499,2007,2025]" pageId="6" pageNumber="497">pes</emphasis>
|
||
, posterior endophallic sclerites;
|
||
<emphasis bold="true" box="[770,787,2007,2025]" pageId="6" pageNumber="497">st</emphasis>
|
||
, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[131,776,215,398]" pageId="7" pageNumber="498">
|
||
VARIATION. — There are variations in the fore and hindwing venation in this specimen as seen in the right forewing (
|
||
<figureCitation box="[642,716,241,266]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="7" pageNumber="498">Fig. 2B</figureCitation>
|
||
) with six primary branches differing from the left forewing (
|
||
<figureCitation box="[645,720,268,292]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="7" pageNumber="498">Fig. 2D</figureCitation>
|
||
) with five primary branches with M5 forked, resulting in M5
|
||
<subScript attach="left" box="[668,675,305,321]" fontSize="7" pageId="7" pageNumber="498">a</subScript>
|
||
and M5
|
||
<subScript attach="none" box="[760,769,305,322]" fontSize="7" pageId="7" pageNumber="498">b</subScript>
|
||
, and in the right hindwing (
|
||
<figureCitation box="[408,485,321,345]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="7" pageNumber="498">Fig. 2C</figureCitation>
|
||
) with four primary branches with M2 forked, resulting in M2
|
||
<subScript attach="left" box="[454,461,358,374]" fontSize="7" pageId="7" pageNumber="498">a</subScript>
|
||
and M2
|
||
<subScript attach="left" box="[547,556,358,375]" fontSize="7" pageId="7" pageNumber="498">b</subScript>
|
||
differing from the left hindwing (
|
||
<figureCitation box="[239,312,374,398]" captionStart="FIG" captionStartId="6.[132,143,1962,1980]" captionTargetBox="[166,1420,216,1901]" captionTargetId="figure-52@6.[685,853,639,1224]" captionTargetPageId="6" captionText="FIG. 2. — Ptiloneura leonardoi n. sp. holotype: A, front view of head; B, right forewing; C, right hindwing; D, left forewing; E, left hindwing; F, phallosome; G, hypandrium; H, left paraproct and epiproct. Abbreviations: aes, anterior endophallic sclerites; ep, external parameres; les, lateral endophallic sclerites; ms, mesal sclerite; msp, mesal sclerite processes; pes, posterior endophallic sclerites; st, side struts. Scale bars: A, F-H, 0.1 mm; B-E, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7304132" httpUri="https://zenodo.org/record/7304132/files/figure.png" pageId="7" pageNumber="498">Fig. 2E</figureCitation>
|
||
) with four simple primary branches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |