treatments-xml/data/C6/45/87/C6458797FFD7FFAB5F97FB2456E9FD8F.xml

<document id="F734A17D1CC91690A5FBC92D11836D6F" ID="10.11646/zootaxa.4175.6.6" ID-DOI="10.11646/zootaxa.4175.6.6" ID-GBIF-Dataset="e36255f4-f061-44d4-99ac-4f14b00e06ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="256165" ID-ZooBank="D5E99C62-D61E-4303-9D1A-BC68EEAD4E5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476867069165" checkinUser="plazi" docAuthor="White, William T., Last, Peter R. &amp; Naylor, Gavin J. P." docDate="2016" docId="C6458797FFD7FFAB5F97FB2456E9FD8F" docLanguage="en" docName="zootaxa.4175.6.6.pdf" docOrigin="Zootaxa 4175 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhinobatos manai White, Last &amp; Naylor, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="597" masterDocId="3A7CFFEFFFD6FFA25F00FF8F5245FFEF" masterDocTitle="Rhinobatos manai sp. nov., a new species of guitarfish (Rhinopristiformes: Rhinobatidae) from New Ireland, Papua New Guinea" masterLastPageNumber="600" masterPageNumber="588" pageNumber="589" updateTime="1698390159425" updateUser="plazi">
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<mods:title id="35B0C37E1229D5C08254EF13EED25DAF">Rhinobatos manai sp. nov., a new species of guitarfish (Rhinopristiformes: Rhinobatidae) from New Ireland, Papua New Guinea</mods:title>
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<mods:namePart id="F8592D4038F03D78859C84C491DE16BF">White, William T.</mods:namePart>
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<mods:namePart id="69DAFBBBAE0939C17F2310E272383666">Last, Peter R.</mods:namePart>
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<mods:namePart id="B8FA7C39D41234A447BC27B2A1D1C0D5">Naylor, Gavin J. P.</mods:namePart>
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<mods:date id="6BA702EC1E9B1A8E3073AD94C62F7D53">2016</mods:date>
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<emphasis id="7C98EA93FFD7FFA35F97FB24539CFB29" bold="true" box="[151,473,1195,1222]" pageId="1" pageNumber="589">
<taxonomicName id="89EC4D02FFD7FFA35F97FB245333FB2A" box="[151,374,1195,1221]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="589" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD7FFA35F97FB245333FB2A" bold="true" box="[151,374,1195,1221]" italics="true" pageId="1" pageNumber="589">Rhinobatos manai</emphasis>
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<taxonomicNameLabel id="67AB57E8FFD7FFA35E7CFB23539CFB29" box="[380,473,1196,1222]" pageId="1" pageNumber="589" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="4E533681FFD7FFA35F97FB405327FB08" blockId="1.[151,473,1195,1291]" box="[151,354,1230,1255]" pageId="1" pageNumber="589">Papuan Guitarfish</paragraph>
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<subSubSection id="06F6650AFFD7FFA35F97FB7D5336FAE4" box="[151,371,1266,1291]" pageId="1" pageNumber="589" type="description">
<paragraph id="4E533681FFD7FFA35F97FB7D5336FAE4" blockId="1.[151,473,1195,1291]" box="[151,371,1266,1291]" pageId="1" pageNumber="589">
(
<figureCitation id="D6D72A04FFD7FFA35F9FFB7D534EFAE4" box="[159,267,1266,1291]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="4.[151,250,1444,1466]" captionStartId-1="5.[151,250,1124,1146]" captionStartId-2="5.[151,250,1849,1871]" captionStartId-3="6.[151,250,1903,1925]" captionStartId-4="7.[151,250,838,860]" captionStartId-5="8.[151,250,1370,1392]" captionTargetBox-0="[163,1418,520,1378]" captionTargetBox-1="[169,1432,336,1090]" captionTargetBox-2="[340,1246,1196,1827]" captionTargetBox-3="[340,1246,517,1881]" captionTargetBox-4="[340,1246,193,816]" captionTargetBox-5="[175,1405,257,1316]" captionTargetId-0="figure@4.[151,1436,483,1420]" captionTargetId-1="figure@5.[154,1433,301,1103]" captionTargetId-2="figure@5.[340,1247,1196,1828]" captionTargetId-3="figure@6.[340,1247,517,1882]" captionTargetId-4="figure@7.[340,1247,193,817]" captionTargetId-5="figure@8.[168,1419,231,1348]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Dorsal view of Rhinobatos manai sp. nov., adult male holotype, 731 mm TL (NTUM 11500): A. freshly caught; B) preserved." captionText-1="FIGURE 2. Head of preserved adult male holotype, 731 mm TL (NTUM 11500): A. dorsal view; B. ventral view." captionText-2="FIGURE 3. Orbital region of Rhinobatos manai sp. nov., preserved adult male holotype, 731 mm TL (NTUM 11500)." captionText-3="FIGURE 4. Radiograph of the chondrocranium of Rhinobatos manai sp. nov., preserved adult male holotype, 731 mm TL (NTUM 11500)." captionText-4="FIGURE 5. Oronasal region of Rhinobatos manai sp. nov., preserved adult male holotype, 731 mm TL (NTUM 11500)." captionText-5="FIGURE 6. Lateral view of the unpaired fins of Rhinobatos manai sp. nov., preserved adult male holotype, 731 mm TL (NTUM 11500): A. first dorsal; B. second dorsal; and C. caudal." httpUri-0="https://zenodo.org/record/256166/files/figure.png" httpUri-1="https://zenodo.org/record/256167/files/figure.png" httpUri-2="https://zenodo.org/record/256168/files/figure.png" httpUri-3="https://zenodo.org/record/256169/files/figure.png" httpUri-4="https://zenodo.org/record/256170/files/figure.png" httpUri-5="https://zenodo.org/record/256171/files/figure.png" pageId="1" pageNumber="589">Figs. 1–6</figureCitation>
; 
<tableCitation id="036E033AFFD7FFA35E17FB7D532DFAE4" box="[279,360,1266,1291]" captionStart="TABLE 1" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1429,497,1966]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data for the holotype of Rhinobatos manai sp. nov. (NTUM 11500) Measurements expressed as percentages of total length." httpUri="http://table.plazi.org/id/1A936609FFD4FFA05F97FEC35390FE6C" pageId="1" pageNumber="589" tableUuid="1A936609FFD4FFA05F97FEC35390FE6C">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="06F6650AFFD7FFA35F97FAB45097FA98" pageId="1" pageNumber="589" type="materials_examined">
<paragraph id="4E533681FFD7FFA35F97FAB45097FA98" blockId="1.[151,1437,1338,2011]" pageId="1" pageNumber="589">
<materialsCitation id="FE843CDCFFD7FFA35F97FAB45097FA98" ID-GBIF-Occurrence="1323872873" collectingDate="2014-09-07" collectionCode="NTUM" country="Papua New Guinea" elevation="240" latitude="-2.5" location="Kavieng" longLatPrecision="1307" longitude="150.73334" pageId="1" pageNumber="589" specimenCode="NTUM 11500" specimenCount="1" stateProvince="New Ireland" typeStatus="holotype">
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<typeStatus id="91578823FFD7FFA35F97FAB45343FABC" box="[151,262,1339,1363]" pageId="1" pageNumber="589" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="1E4A9EFAFFD7FFA35E17FAB453FBFABC" box="[279,446,1338,1363]" collectionCode="NTUM" pageId="1" pageNumber="589">
<collectionCode id="28FDAE44FFD7FFA35E17FAB4532CFABD" box="[279,361,1339,1362]" pageId="1" pageNumber="589">NTUM</collectionCode>
11500
</specimenCode>
, adult male 
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TL, northwest of 
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, 
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, 
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, 
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, 
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, 
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<quantity id="89149B64FFD7FFA35E7BFAD053BFFA99" box="[379,506,1375,1399]" metricMagnitude="2" metricUnit="m" metricValue="2.405" metricValueMax="2.9" metricValueMin="1.91" pageId="1" pageNumber="589" unit="m" value="240.5" valueMax="290.0" valueMin="191.0">191–290 m</quantity>
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depth, 
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<date id="3A521041FFD7FFA35D4DFAD0508AFA98" box="[589,719,1375,1399]" pageId="1" pageNumber="589" value="2014-09-07">7 Sep 2014</date>
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.
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</paragraph>
</subSubSection>
<subSubSection id="06F6650AFFD7FFA35FC7FA0D56C8F8C8" pageId="1" pageNumber="589" type="diagnosis">
<paragraph id="4E533681FFD7FFA35FC7FA0D56C8F8C8" blockId="1.[151,1437,1338,2011]" pageId="1" pageNumber="589">
<emphasis id="7C98EA93FFD7FFA35FC7FA0D5304FA74" bold="true" box="[199,321,1410,1435]" pageId="1" pageNumber="589">Diagnosis.</emphasis>
A 
<taxonomicName id="89EC4D02FFD7FFA35E6AFA0C53AFFA74" box="[362,490,1411,1435]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="1" pageNumber="589" phylum="Chordata" rank="genus">
<emphasis id="7C98EA93FFD7FFA35E6AFA0C53AFFA74" box="[362,490,1411,1435]" italics="true" pageId="1" pageNumber="589">Rhinobatos</emphasis>
</taxonomicName>
distinguished by the following combination of characters: disc wedge-shaped, its dorsal surface covered in minute dermal denticles but without thorns; snout long, snout length 3.6 times interspiracular distance; orbit moderately large, diameter 1.7 times spiracle length; nostrils oblique, length 1.6 times internarial distance; anterior nasal flaps inserted into internarial space well away from nostril margin; posterior nasal flaps broad; ridges of rostral cartilage well-separated dorsally and almost parallel; prebranchial sensory-pore patch narrow, extending to first gill slit; distance between first gill slits 1.5 times distance between fifth gill slits; distance between fifth gill slits 3.4 times in ventral head length; postscapular sensory canal long, not grooved, extending more than three-quarters distance to pectoral-fin insertions; dorsal fins moderately tall; pelvicfin inner margin shorter than its base; interdorsal distance more than 3.1 times first dorsal-fin base; outer spiracular fold distinctly larger than inner fold; dorsal margin of caudal fin ~2 times preventral margin; 171 post-synarcual (free) vertebral centra; 188 total vertebral centra; 52 nasal lamellae; and dorsal disc brownish, covered with welldefined rusty brown spots and blotches and poorly defined white spots with greyish edges.
</paragraph>
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<emphasis id="7C98EA93FFD7FFA35FC7F8BD531CF8A4" bold="true" box="[199,345,1842,1867]" pageId="1" pageNumber="589">Description.</emphasis>
Disc wedge shaped, bluntly angular anteriorly, angle anterior to eyes about 56°; outer margins broadly rounded, more narrowly rounded distally; length 1.37 times width. Pelvic fins elongate, short-based, base length about 0.78 of inner margin; length 1.73 times their base length, 2.75 times width; anterior margin convex, apex broadly rounded, posterior margin weakly convex. Tail elongate, slender, tapering gradually; in cross-section nearly flat ventrally, rounded dorsally; length from anterior cloaca to tail tip 1.39 times precloacal length, 1.34 times disc length, 6.32 times body width at pelvic-fin insertions; tail width 1.99 times depth at pelvic-fin insertions, 2.41 times at first dorsal-fin origin, 1.68 times at second dorsal-fin origin. Dermal fold ventrolateral on tail, originating slightly anterior to free rear tip of pelvic fin, reaching just behind ventral caudal fin origin; fold well developed, maximum width in interdorsal space about a quarter width of spiracle.
</paragraph>
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<emphasis id="7C98EA93FFD4FFA05F97FEC3534FFE8C" bold="true" box="[151,266,332,355]" pageId="2" pageNumber="590">TABLE 1.</emphasis>
Morphometric data for the holotype of 
<taxonomicName id="89EC4D02FFD4FFA05DB7FEC4513FFE8C" box="[695,890,331,355]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="590" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD4FFA05DB7FEC4513FFE8C" box="[695,890,331,355]" italics="true" pageId="2" pageNumber="590">Rhinobatos manai</emphasis>
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<emphasis id="7C98EA93FFD4FFA05C82FEC3519DFE8C" bold="true" box="[898,984,332,355]" pageId="2" pageNumber="590">sp. nov.</emphasis>
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(NTUM 11500) Measurements expressed as percentages of total length.
</paragraph>
</caption>
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<emphasis id="7C98EA93FFD4FFA05C54FE2F56CAFE5B" box="[852,1167,414,437]" italics="true" pageId="2" pageNumber="590">
<taxonomicName id="89EC4D02FFD4FFA05C54FE2F51EAFE5A" box="[852,943,416,437]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="590" phylum="Chordata" rank="species" species="manai">R. manai</taxonomicName>
<taxonomicName id="89EC4D02FFD4FFA05B34FE2F56CAFE5B" box="[1076,1167,414,437]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="2" pageNumber="590" phylum="Chordata" rank="species" species="whitei">R. whitei</taxonomicName>
</emphasis>
</paragraph>
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Adult males 
<emphasis id="7C98EA93FFD4FFA15B5FF84E56CAFF30" italics="true" lastPageId="3" lastPageNumber="591" pageId="2" pageNumber="590">
......continued on the next page 
<taxonomicName id="89EC4D02FFD5FFA15C54FF4551EAFF30" box="[852,943,202,223]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="591" phylum="Chordata" rank="species" species="manai">R. manai</taxonomicName>
<taxonomicName id="89EC4D02FFD5FFA15B34FF4556CAFF30" box="[1076,1167,201,223]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="3" pageNumber="591" phylum="Chordata" rank="species" species="whitei">R. whitei</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="4E533681FFD4FFA05C54FE7E5748F841" pageId="2" pageNumber="590">
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<th id="B30D5DBFFFD4005D5C53FE7E51F7FDE8" box="[851,946,497,519]" gridcol="1" gridrow="0" pageId="2" pageNumber="590">Holotype</th>
<th id="B30D5DBFFFD4005D5B34FE7E5623FDE8" box="[1076,1126,497,519]" gridcol="2" gridrow="0" pageId="2" pageNumber="590">Min.</th>
<th id="B30D5DBFFFD4005D5BEDFE7E57D0FDE8" box="[1261,1429,497,519]" gridcol="3" gridrow="0" pageId="2" pageNumber="590">Max.</th>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFD9557D0FDDF" box="[159,1429,538,560]" gridrow="1" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFD95506AFDDF" box="[159,559,538,560]" gridcol="0" gridrow="1" pageId="2" pageNumber="590">Total length</th>
<td id="B30D5DBFFFD4005D5C53FD9551F7FDDF" box="[851,946,538,560]" gridcol="1" gridrow="1" pageId="2" pageNumber="590">731</td>
<td id="B30D5DBFFFD4005D5B34FD955623FDDF" box="[1076,1126,538,560]" gridcol="2" gridrow="1" pageId="2" pageNumber="590">556</td>
<td id="B30D5DBFFFD4005D5BEDFD9557D0FDDF" box="[1261,1429,538,560]" gridcol="3" gridrow="1" pageId="2" pageNumber="590">641</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFDCB57D0FDB5" box="[159,1429,580,602]" gridrow="2" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFDCB506AFDB5" box="[159,559,580,602]" gridcol="0" gridrow="2" pageId="2" pageNumber="590">Disc width—maximum</th>
<td id="B30D5DBFFFD4005D5C53FDCB51F7FDB5" box="[851,946,580,602]" gridcol="1" gridrow="2" pageId="2" pageNumber="590">31.6</td>
<td id="B30D5DBFFFD4005D5B34FDCB5623FDB5" box="[1076,1126,580,602]" gridcol="2" gridrow="2" pageId="2" pageNumber="590">30.9</td>
<td id="B30D5DBFFFD4005D5BEDFDCB57D0FDB5" box="[1261,1429,580,602]" gridcol="3" gridrow="2" pageId="2" pageNumber="590">32.6</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFDE257D0FD6C" box="[159,1429,621,643]" gridrow="3" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFDE2506AFD6C" box="[159,559,621,643]" gridcol="0" gridrow="3" pageId="2" pageNumber="590">Disc length</th>
<td id="B30D5DBFFFD4005D5C53FDE251F7FD6C" box="[851,946,621,643]" gridcol="1" gridrow="3" pageId="2" pageNumber="590">43.4</td>
<td id="B30D5DBFFFD4005D5B34FDE25623FD6C" box="[1076,1126,621,643]" gridcol="2" gridrow="3" pageId="2" pageNumber="590">41.3</td>
<td id="B30D5DBFFFD4005D5BEDFDE257D0FD6C" box="[1261,1429,621,643]" gridcol="3" gridrow="3" pageId="2" pageNumber="590">42.7</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFD1957D0FD43" box="[159,1429,662,684]" gridrow="4" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFD19506AFD43" box="[159,559,662,684]" gridcol="0" gridrow="4" pageId="2" pageNumber="590">Head length—dorsal</th>
<td id="B30D5DBFFFD4005D5C53FD1951F7FD43" box="[851,946,662,684]" gridcol="1" gridrow="4" pageId="2" pageNumber="590">23.3</td>
<td id="B30D5DBFFFD4005D5B34FD195623FD43" box="[1076,1126,662,684]" gridcol="2" gridrow="4" pageId="2" pageNumber="590">21.7</td>
<td id="B30D5DBFFFD4005D5BEDFD1957D0FD43" box="[1261,1429,662,684]" gridcol="3" gridrow="4" pageId="2" pageNumber="590">23.2</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFD4F57D0FD39" box="[159,1429,704,726]" gridrow="5" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFD4F506AFD39" box="[159,559,704,726]" gridcol="0" gridrow="5" pageId="2" pageNumber="590">Head length—ventral</th>
<td id="B30D5DBFFFD4005D5C53FD4F51F7FD39" box="[851,946,704,726]" gridcol="1" gridrow="5" pageId="2" pageNumber="590">28.6</td>
<td id="B30D5DBFFFD4005D5B34FD4F5623FD39" box="[1076,1126,704,726]" gridcol="2" gridrow="5" pageId="2" pageNumber="590">27.7</td>
<td id="B30D5DBFFFD4005D5BEDFD4F57D0FD39" box="[1261,1429,704,726]" gridcol="3" gridrow="5" pageId="2" pageNumber="590">29.2</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFD6657D0FD10" box="[159,1429,745,767]" gridrow="6" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFD66506AFD10" box="[159,559,745,767]" gridcol="0" gridrow="6" pageId="2" pageNumber="590">Snout length (presocket)</th>
<td id="B30D5DBFFFD4005D5C53FD6651F7FD10" box="[851,946,745,767]" gridcol="1" gridrow="6" pageId="2" pageNumber="590">17.6</td>
<td id="B30D5DBFFFD4005D5B34FD665623FD10" box="[1076,1126,745,767]" gridcol="2" gridrow="6" pageId="2" pageNumber="590">15.4</td>
<td id="B30D5DBFFFD4005D5BEDFD6657D0FD10" box="[1261,1429,745,767]" gridcol="3" gridrow="6" pageId="2" pageNumber="590">17.0</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFC9D57D0FCC7" box="[159,1429,786,808]" gridrow="7" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFC9D506AFCC7" box="[159,559,786,808]" gridcol="0" gridrow="7" pageId="2" pageNumber="590">Orbit diameter</th>
<td id="B30D5DBFFFD4005D5C53FC9D51F7FCC7" box="[851,946,786,808]" gridcol="1" gridrow="7" pageId="2" pageNumber="590">3.5</td>
<td id="B30D5DBFFFD4005D5B34FC9D5623FCC7" box="[1076,1126,786,808]" gridcol="2" gridrow="7" pageId="2" pageNumber="590">4.0</td>
<td id="B30D5DBFFFD4005D5BEDFC9D57D0FCC7" box="[1261,1429,786,808]" gridcol="3" gridrow="7" pageId="2" pageNumber="590">4.2</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFCB357D0FCBD" box="[159,1429,828,850]" gridrow="8" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFCB3506AFCBD" box="[159,559,828,850]" gridcol="0" gridrow="8" pageId="2" pageNumber="590">Spiracle length</th>
<td id="B30D5DBFFFD4005D5C53FCB351F7FCBD" box="[851,946,828,850]" gridcol="1" gridrow="8" pageId="2" pageNumber="590">2.1</td>
<td id="B30D5DBFFFD4005D5B34FCB35623FCBD" box="[1076,1126,828,850]" gridcol="2" gridrow="8" pageId="2" pageNumber="590">2.5</td>
<td id="B30D5DBFFFD4005D5BEDFCB357D0FCBD" box="[1261,1429,828,850]" gridcol="3" gridrow="8" pageId="2" pageNumber="590">2.5</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFCEA57D0FC94" box="[159,1429,869,891]" gridrow="9" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFCEA506AFC94" box="[159,559,869,891]" gridcol="0" gridrow="9" pageId="2" pageNumber="590">Orbit and spiracle length</th>
<td id="B30D5DBFFFD4005D5C53FCEA51F7FC94" box="[851,946,869,891]" gridcol="1" gridrow="9" pageId="2" pageNumber="590">4.6</td>
<td id="B30D5DBFFFD4005D5B34FCEA5623FC94" box="[1076,1126,869,891]" gridcol="2" gridrow="9" pageId="2" pageNumber="590">4.8</td>
<td id="B30D5DBFFFD4005D5BEDFCEA57D0FC94" box="[1261,1429,869,891]" gridcol="3" gridrow="9" pageId="2" pageNumber="590">5.1</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFC0157D0FC4B" box="[159,1429,910,932]" gridrow="10" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFC01506AFC4B" box="[159,559,910,932]" gridcol="0" gridrow="10" pageId="2" pageNumber="590">Interorbital width</th>
<td id="B30D5DBFFFD4005D5C53FC0151F7FC4B" box="[851,946,910,932]" gridcol="1" gridrow="10" pageId="2" pageNumber="590">3.5</td>
<td id="B30D5DBFFFD4005D5B34FC015623FC4B" box="[1076,1126,910,932]" gridcol="2" gridrow="10" pageId="2" pageNumber="590">3.5</td>
<td id="B30D5DBFFFD4005D5BEDFC0157D0FC4B" box="[1261,1429,910,932]" gridcol="3" gridrow="10" pageId="2" pageNumber="590">3.7</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFC3757D0FC21" box="[159,1429,952,974]" gridrow="11" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFC37506AFC21" box="[159,559,952,974]" gridcol="0" gridrow="11" pageId="2" pageNumber="590">Interspiracular width</th>
<td id="B30D5DBFFFD4005D5C53FC3751F7FC21" box="[851,946,952,974]" gridcol="1" gridrow="11" pageId="2" pageNumber="590">4.9</td>
<td id="B30D5DBFFFD4005D5B34FC375623FC21" box="[1076,1126,952,974]" gridcol="2" gridrow="11" pageId="2" pageNumber="590">5.1</td>
<td id="B30D5DBFFFD4005D5BEDFC3757D0FC21" box="[1261,1429,952,974]" gridcol="3" gridrow="11" pageId="2" pageNumber="590">5.2</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFC6E57D0FC18" box="[159,1429,993,1015]" gridrow="12" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFC6E506AFC18" box="[159,559,993,1015]" gridcol="0" gridrow="12" pageId="2" pageNumber="590">Preoral length</th>
<td id="B30D5DBFFFD4005D5C53FC6E51F7FC18" box="[851,946,993,1015]" gridcol="1" gridrow="12" pageId="2" pageNumber="590">20.6</td>
<td id="B30D5DBFFFD4005D5B34FC6E5623FC18" box="[1076,1126,993,1015]" gridcol="2" gridrow="12" pageId="2" pageNumber="590">18.2</td>
<td id="B30D5DBFFFD4005D5BEDFC6E57D0FC18" box="[1261,1429,993,1015]" gridcol="3" gridrow="12" pageId="2" pageNumber="590">19.8</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFB8557D0FBCF" box="[159,1429,1034,1056]" gridrow="13" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFB85506AFBCF" box="[159,559,1034,1056]" gridcol="0" gridrow="13" pageId="2" pageNumber="590">Mouth width</th>
<td id="B30D5DBFFFD4005D5C53FB8551F7FBCF" box="[851,946,1034,1056]" gridcol="1" gridrow="13" pageId="2" pageNumber="590">5.8</td>
<td id="B30D5DBFFFD4005D5B34FB855623FBCF" box="[1076,1126,1034,1056]" gridcol="2" gridrow="13" pageId="2" pageNumber="590">5.7</td>
<td id="B30D5DBFFFD4005D5BEDFB8557D0FBCF" box="[1261,1429,1034,1056]" gridcol="3" gridrow="13" pageId="2" pageNumber="590">6.4</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFBBB57D0FBA5" box="[159,1429,1076,1098]" gridrow="14" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFBBB506AFBA5" box="[159,559,1076,1098]" gridcol="0" gridrow="14" pageId="2" pageNumber="590">Prenarial distance</th>
<td id="B30D5DBFFFD4005D5C53FBBB51F7FBA5" box="[851,946,1076,1098]" gridcol="1" gridrow="14" pageId="2" pageNumber="590">16.4</td>
<td id="B30D5DBFFFD4005D5B34FBBB5623FBA5" box="[1076,1126,1076,1098]" gridcol="2" gridrow="14" pageId="2" pageNumber="590">14.4</td>
<td id="B30D5DBFFFD4005D5BEDFBBB57D0FBA5" box="[1261,1429,1076,1098]" gridcol="3" gridrow="14" pageId="2" pageNumber="590">16.1</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFBD257D0FB9C" box="[159,1429,1117,1139]" gridrow="15" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFBD2506AFB9C" box="[159,559,1117,1139]" gridcol="0" gridrow="15" pageId="2" pageNumber="590">Nostril length</th>
<td id="B30D5DBFFFD4005D5C53FBD251F7FB9C" box="[851,946,1117,1139]" gridcol="1" gridrow="15" pageId="2" pageNumber="590">4.5</td>
<td id="B30D5DBFFFD4005D5B34FBD25623FB9C" box="[1076,1126,1117,1139]" gridcol="2" gridrow="15" pageId="2" pageNumber="590">3.8</td>
<td id="B30D5DBFFFD4005D5BEDFBD257D0FB9C" box="[1261,1429,1117,1139]" gridcol="3" gridrow="15" pageId="2" pageNumber="590">4.0</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFB0957D0FB73" box="[159,1429,1158,1180]" gridrow="16" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFB09506AFB73" box="[159,559,1158,1180]" gridcol="0" gridrow="16" pageId="2" pageNumber="590">Anterior aperture—width</th>
<td id="B30D5DBFFFD4005D5C53FB0951F7FB73" box="[851,946,1158,1180]" gridcol="1" gridrow="16" pageId="2" pageNumber="590">1.4</td>
<td id="B30D5DBFFFD4005D5B34FB095623FB73" box="[1076,1126,1158,1180]" gridcol="2" gridrow="16" pageId="2" pageNumber="590">1.2</td>
<td id="B30D5DBFFFD4005D5BEDFB0957D0FB73" box="[1261,1429,1158,1180]" gridcol="3" gridrow="16" pageId="2" pageNumber="590">1.3</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFB3F57D0FB29" box="[159,1429,1200,1222]" gridrow="17" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFB3F506AFB29" box="[159,559,1200,1222]" gridcol="0" gridrow="17" pageId="2" pageNumber="590">Anterior nasal flap—base length</th>
<td id="B30D5DBFFFD4005D5C53FB3F51F7FB29" box="[851,946,1200,1222]" gridcol="1" gridrow="17" pageId="2" pageNumber="590">2.7</td>
<td id="B30D5DBFFFD4005D5B34FB3F5623FB29" box="[1076,1126,1200,1222]" gridcol="2" gridrow="17" pageId="2" pageNumber="590">2.7</td>
<td id="B30D5DBFFFD4005D5BEDFB3F57D0FB29" box="[1261,1429,1200,1222]" gridcol="3" gridrow="17" pageId="2" pageNumber="590">2.9</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFB5657D0FB00" box="[159,1429,1241,1263]" gridrow="18" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFB56506AFB00" box="[159,559,1241,1263]" gridcol="0" gridrow="18" pageId="2" pageNumber="590">Anterior nasal flap—width</th>
<td id="B30D5DBFFFD4005D5C53FB5651F7FB00" box="[851,946,1241,1263]" gridcol="1" gridrow="18" pageId="2" pageNumber="590">1.4</td>
<td id="B30D5DBFFFD4005D5B34FB565623FB00" box="[1076,1126,1241,1263]" gridcol="2" gridrow="18" pageId="2" pageNumber="590">1.3</td>
<td id="B30D5DBFFFD4005D5BEDFB5657D0FB00" box="[1261,1429,1241,1263]" gridcol="3" gridrow="18" pageId="2" pageNumber="590">1.4</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFA8D57D0FAF7" box="[159,1429,1282,1304]" gridrow="19" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFA8D506AFAF7" box="[159,559,1282,1304]" gridcol="0" gridrow="19" pageId="2" pageNumber="590">Posterolateral nasal flap—total length</th>
<td id="B30D5DBFFFD4005D5C53FA8D51F7FAF7" box="[851,946,1282,1304]" gridcol="1" gridrow="19" pageId="2" pageNumber="590">3.6</td>
<td id="B30D5DBFFFD4005D5B34FA8D5623FAF7" box="[1076,1126,1282,1304]" gridcol="2" gridrow="19" pageId="2" pageNumber="590">3.3</td>
<td id="B30D5DBFFFD4005D5BEDFA8D57D0FAF7" box="[1261,1429,1282,1304]" gridcol="3" gridrow="19" pageId="2" pageNumber="590">3.4</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFAA357D0FAAD" box="[159,1429,1324,1346]" gridrow="20" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFAA3506AFAAD" box="[159,559,1324,1346]" gridcol="0" gridrow="20" pageId="2" pageNumber="590">Posterolateral nasal flap—width</th>
<td id="B30D5DBFFFD4005D5C53FAA351F7FAAD" box="[851,946,1324,1346]" gridcol="1" gridrow="20" pageId="2" pageNumber="590">1.1</td>
<td id="B30D5DBFFFD4005D5B34FAA35623FAAD" box="[1076,1126,1324,1346]" gridcol="2" gridrow="20" pageId="2" pageNumber="590">0.5</td>
<td id="B30D5DBFFFD4005D5BEDFAA357D0FAAD" box="[1261,1429,1324,1346]" gridcol="3" gridrow="20" pageId="2" pageNumber="590">0.6</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFADA57D0FA84" box="[159,1429,1365,1387]" gridrow="21" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFADA506AFA84" box="[159,559,1365,1387]" gridcol="0" gridrow="21" pageId="2" pageNumber="590">Posterior nasal flap— base length</th>
<td id="B30D5DBFFFD4005D5C53FADA51F7FA84" box="[851,946,1365,1387]" gridcol="1" gridrow="21" pageId="2" pageNumber="590">3.0</td>
<td id="B30D5DBFFFD4005D5B34FADA5623FA84" box="[1076,1126,1365,1387]" gridcol="2" gridrow="21" pageId="2" pageNumber="590">2.8</td>
<td id="B30D5DBFFFD4005D5BEDFADA57D0FA84" box="[1261,1429,1365,1387]" gridcol="3" gridrow="21" pageId="2" pageNumber="590">2.9</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFAF157D0FA7B" box="[159,1429,1406,1428]" gridrow="22" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFAF1506AFA7B" box="[159,559,1406,1428]" gridcol="0" gridrow="22" pageId="2" pageNumber="590">Posterior nasal flap—width</th>
<td id="B30D5DBFFFD4005D5C53FAF151F7FA7B" box="[851,946,1406,1428]" gridcol="1" gridrow="22" pageId="2" pageNumber="590">1.2</td>
<td id="B30D5DBFFFD4005D5B34FAF15623FA7B" box="[1076,1126,1406,1428]" gridcol="2" gridrow="22" pageId="2" pageNumber="590">1.0</td>
<td id="B30D5DBFFFD4005D5BEDFAF157D0FA7B" box="[1261,1429,1406,1428]" gridcol="3" gridrow="22" pageId="2" pageNumber="590">1.2</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFA2757D0FA51" box="[159,1429,1448,1470]" gridrow="23" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFA27506AFA51" box="[159,559,1448,1470]" gridcol="0" gridrow="23" pageId="2" pageNumber="590">Distance across anterior nasal apertures</th>
<td id="B30D5DBFFFD4005D5C53FA2751F7FA51" box="[851,946,1448,1470]" gridcol="1" gridrow="23" pageId="2" pageNumber="590">10.6</td>
<td id="B30D5DBFFFD4005D5B34FA275623FA51" box="[1076,1126,1448,1470]" gridcol="2" gridrow="23" pageId="2" pageNumber="590">8.9</td>
<td id="B30D5DBFFFD4005D5BEDFA2757D0FA51" box="[1261,1429,1448,1470]" gridcol="3" gridrow="23" pageId="2" pageNumber="590">9.4</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFA5E57D0FA08" box="[159,1429,1489,1511]" gridrow="24" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFA5E506AFA08" box="[159,559,1489,1511]" gridcol="0" gridrow="24" pageId="2" pageNumber="590">Internarial distance (minimum)</th>
<td id="B30D5DBFFFD4005D5C53FA5E51F7FA08" box="[851,946,1489,1511]" gridcol="1" gridrow="24" pageId="2" pageNumber="590">2.8</td>
<td id="B30D5DBFFFD4005D5B34FA5E5623FA08" box="[1076,1126,1489,1511]" gridcol="2" gridrow="24" pageId="2" pageNumber="590">2.4</td>
<td id="B30D5DBFFFD4005D5BEDFA5E57D0FA08" box="[1261,1429,1489,1511]" gridcol="3" gridrow="24" pageId="2" pageNumber="590">2.7</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FFA7557D0F9FF" box="[159,1429,1530,1552]" gridrow="25" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FFA75506AF9FF" box="[159,559,1530,1552]" gridcol="0" gridrow="25" pageId="2" pageNumber="590">Distance between anterior nasal flaps</th>
<td id="B30D5DBFFFD4005D5C53FA7551F7F9FF" box="[851,946,1530,1552]" gridcol="1" gridrow="25" pageId="2" pageNumber="590">2.9</td>
<td id="B30D5DBFFFD4005D5B34FA755623F9FF" box="[1076,1126,1530,1552]" gridcol="2" gridrow="25" pageId="2" pageNumber="590">2.3</td>
<td id="B30D5DBFFFD4005D5BEDFA7557D0F9FF" box="[1261,1429,1530,1552]" gridcol="3" gridrow="25" pageId="2" pageNumber="590">2.6</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF9AB57D0F9D5" box="[159,1429,1572,1594]" gridrow="26" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF9AB506AF9D5" box="[159,559,1572,1594]" gridcol="0" gridrow="26" pageId="2" pageNumber="590">Distance from nostril to disc margin</th>
<td id="B30D5DBFFFD4005D5C53F9AB51F7F9D5" box="[851,946,1572,1594]" gridcol="1" gridrow="26" pageId="2" pageNumber="590">3.1</td>
<td id="B30D5DBFFFD4005D5B34F9AB5623F9D5" box="[1076,1126,1572,1594]" gridcol="2" gridrow="26" pageId="2" pageNumber="590">3.1</td>
<td id="B30D5DBFFFD4005D5BEDF9AB57D0F9D5" box="[1261,1429,1572,1594]" gridcol="3" gridrow="26" pageId="2" pageNumber="590">3.3</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF9C257D0F98C" box="[159,1429,1613,1635]" gridrow="27" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF9C2506AF98C" box="[159,559,1613,1635]" gridcol="0" gridrow="27" pageId="2" pageNumber="590">Third gill opening—width</th>
<td id="B30D5DBFFFD4005D5C53F9C251F7F98C" box="[851,946,1613,1635]" gridcol="1" gridrow="27" pageId="2" pageNumber="590">1.5</td>
<td id="B30D5DBFFFD4005D5B34F9C25623F98C" box="[1076,1126,1613,1635]" gridcol="2" gridrow="27" pageId="2" pageNumber="590">1.3</td>
<td id="B30D5DBFFFD4005D5BEDF9C257D0F98C" box="[1261,1429,1613,1635]" gridcol="3" gridrow="27" pageId="2" pageNumber="590">1.5</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF9F957D0F963" box="[159,1429,1654,1676]" gridrow="28" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF9F9506AF963" box="[159,559,1654,1676]" gridcol="0" gridrow="28" pageId="2" pageNumber="590">Distance between first gill openings</th>
<td id="B30D5DBFFFD4005D5C53F9F951F7F963" box="[851,946,1654,1676]" gridcol="1" gridrow="28" pageId="2" pageNumber="590">12.3</td>
<td id="B30D5DBFFFD4005D5B34F9F95623F963" box="[1076,1126,1654,1676]" gridcol="2" gridrow="28" pageId="2" pageNumber="590">12.5</td>
<td id="B30D5DBFFFD4005D5BEDF9F957D0F963" box="[1261,1429,1654,1676]" gridcol="3" gridrow="28" pageId="2" pageNumber="590">13.1</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF92F57D0F959" box="[159,1429,1696,1718]" gridrow="29" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF92F506AF959" box="[159,559,1696,1718]" gridcol="0" gridrow="29" pageId="2" pageNumber="590">Distance between fifth gill openings</th>
<td id="B30D5DBFFFD4005D5C53F92F51F7F959" box="[851,946,1696,1718]" gridcol="1" gridrow="29" pageId="2" pageNumber="590">8.4</td>
<td id="B30D5DBFFFD4005D5B34F92F5623F959" box="[1076,1126,1696,1718]" gridcol="2" gridrow="29" pageId="2" pageNumber="590">8.5</td>
<td id="B30D5DBFFFD4005D5BEDF92F57D0F959" box="[1261,1429,1696,1718]" gridcol="3" gridrow="29" pageId="2" pageNumber="590">9.1</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF94657D0F930" box="[159,1429,1737,1759]" gridrow="30" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF946506AF930" box="[159,559,1737,1759]" gridcol="0" gridrow="30" pageId="2" pageNumber="590">Pelvic fin—length</th>
<td id="B30D5DBFFFD4005D5C53F94651F7F930" box="[851,946,1737,1759]" gridcol="1" gridrow="30" pageId="2" pageNumber="590">14.1</td>
<td id="B30D5DBFFFD4005D5B34F9465623F930" box="[1076,1126,1737,1759]" gridcol="2" gridrow="30" pageId="2" pageNumber="590">13.4</td>
<td id="B30D5DBFFFD4005D5BEDF94657D0F930" box="[1261,1429,1737,1759]" gridcol="3" gridrow="30" pageId="2" pageNumber="590">13.8</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF97D57D0F8E7" box="[159,1429,1778,1800]" gridrow="31" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF97D506AF8E7" box="[159,559,1778,1800]" gridcol="0" gridrow="31" pageId="2" pageNumber="590">Pelvic fin—anterior margin length</th>
<td id="B30D5DBFFFD4005D5C53F97D51F7F8E7" box="[851,946,1778,1800]" gridcol="1" gridrow="31" pageId="2" pageNumber="590">8.1</td>
<td id="B30D5DBFFFD4005D5B34F97D5623F8E7" box="[1076,1126,1778,1800]" gridcol="2" gridrow="31" pageId="2" pageNumber="590">7.7</td>
<td id="B30D5DBFFFD4005D5BEDF97D57D0F8E7" box="[1261,1429,1778,1800]" gridcol="3" gridrow="31" pageId="2" pageNumber="590">8.1</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF89357D0F8DD" box="[159,1429,1820,1842]" gridrow="32" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF893506AF8DD" box="[159,559,1820,1842]" gridcol="0" gridrow="32" pageId="2" pageNumber="590">Pelvic fin—width</th>
<td id="B30D5DBFFFD4005D5C53F89351F7F8DD" box="[851,946,1820,1842]" gridcol="1" gridrow="32" pageId="2" pageNumber="590">5.1</td>
<td id="B30D5DBFFFD4005D5B34F8935623F8DD" box="[1076,1126,1820,1842]" gridcol="2" gridrow="32" pageId="2" pageNumber="590">5.6</td>
<td id="B30D5DBFFFD4005D5BEDF89357D0F8DD" box="[1261,1429,1820,1842]" gridcol="3" gridrow="32" pageId="2" pageNumber="590">5.8</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF8CA57D0F8B4" box="[159,1429,1861,1883]" gridrow="33" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF8CA506AF8B4" box="[159,559,1861,1883]" gridcol="0" gridrow="33" pageId="2" pageNumber="590">Pelvic fin—base length</th>
<td id="B30D5DBFFFD4005D5C53F8CA51F7F8B4" box="[851,946,1861,1883]" gridcol="1" gridrow="33" pageId="2" pageNumber="590">8.1</td>
<td id="B30D5DBFFFD4005D5B34F8CA5623F8B4" box="[1076,1126,1861,1883]" gridcol="2" gridrow="33" pageId="2" pageNumber="590">6.6</td>
<td id="B30D5DBFFFD4005D5BEDF8CA57D0F8B4" box="[1261,1429,1861,1883]" gridcol="3" gridrow="33" pageId="2" pageNumber="590">7.0</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF8E157D0F86B" box="[159,1429,1902,1924]" gridrow="34" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF8E1506AF86B" box="[159,559,1902,1924]" gridcol="0" gridrow="34" pageId="2" pageNumber="590">Pelvic fin—inner margin length</th>
<td id="B30D5DBFFFD4005D5C53F8E151F7F86B" box="[851,946,1902,1924]" gridcol="1" gridrow="34" pageId="2" pageNumber="590">6.3</td>
<td id="B30D5DBFFFD4005D5B34F8E15623F86B" box="[1076,1126,1902,1924]" gridcol="2" gridrow="34" pageId="2" pageNumber="590">7.6</td>
<td id="B30D5DBFFFD4005D5BEDF8E157D0F86B" box="[1261,1429,1902,1924]" gridcol="3" gridrow="34" pageId="2" pageNumber="590">7.8</td>
</tr>
<tr id="F0DC34C3FFD4005D5F9FF81757D0F841" box="[159,1429,1944,1966]" gridrow="35" pageId="2" pageNumber="590">
<th id="B30D5DBFFFD4005D5F9FF817506AF841" box="[159,559,1944,1966]" gridcol="0" gridrow="35" pageId="2" pageNumber="590">First dorsal fin—length</th>
<td id="B30D5DBFFFD4005D5C53F81751F7F841" box="[851,946,1944,1966]" gridcol="1" gridrow="35" pageId="2" pageNumber="590">6.5</td>
<td id="B30D5DBFFFD4005D5B34F8175623F841" box="[1076,1126,1944,1966]" gridcol="2" gridrow="35" pageId="2" pageNumber="590">6.7</td>
<td id="B30D5DBFFFD4005D5BEDF81757D0F841" box="[1261,1429,1944,1966]" gridcol="3" gridrow="35" pageId="2" pageNumber="590">7.1</td>
</tr>
</table>
</paragraph>
<caption id="1A936609FFD5FFA15F9FFF2F53C8FF59" ID-Table-UUID="1A936609FFD5FFA15F9FFF2F53C8FF59" box="[159,397,160,182]" httpUri="http://table.plazi.org/id/1A936609FFD5FFA15F9FFF2F53C8FF59" pageId="3" pageNumber="591" targetBox="[159,1305,284,1587]" targetIsTable="true" targetPageId="3">
<paragraph id="4E533681FFD5FFA15F9FFF2F53C8FF59" blockId="3.[159,1199,160,264]" box="[159,397,160,182]" pageId="3" pageNumber="591">
<emphasis id="7C98EA93FFD5FFA15F9FFF2F5349FF59" bold="true" box="[159,268,160,182]" pageId="3" pageNumber="591">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="4E533681FFD5FFA65B34FF7D569EFE42" blockId="3.[159,1199,160,264]" lastBlockId="4.[151,1436,151,429]" lastPageId="4" lastPageNumber="592" pageId="3" pageNumber="591">
Adult males Head elongate, ventral length 28.6% TL; snout moderately long and bluntly pointed; preoral snout length 3.58 times mouth width, 7.32 times internarial distance, 1.44 times dorsal caudal fin margin, 6.69 times distance from nostril to margin of disc; snout length (direct) 3.58 times interspiracular length, 4.99 times orbit diameter, 5.10 times interorbital width; interorbital space weakly concave, relatively broad; eye dorsolateral, not elevated or protruding; orbit relatively small, diameter 1.65 times spiracle length, 1.02 times interorbital width. Spiracle narrowly bean-shaped, moderately large; two weakly compressed spiracular folds on posterior margin, innermost fold half or less length of outer fold, distance between bases of folds subequal to length of inner fold. Nostril moderately large, oblique, nasal flaps well developed; anterior aperture suboval, width slightly exceeding length; nostril length 3.32 times anterior aperture width, 1.69 times anterior nasal-flap base length, 1.45 times distance from nostril to edge of disc, 1.59 times internarial width. Anterior nasal flap relatively well developed with long, slender process anteriorly; flap base 1.84 times its width at process, 1.97 times anterior aperture width; insertion in internarial space well mesial to nostril margin, its distance from nostril about equal to half width of anterior nasal aperture; distance between their insertions 
<quantity id="89149B64FFD2FFA65D80FF335095FF3A" box="[640,720,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="9.3218" pageId="4" pageNumber="592" unit="in" value="3.67">3.67 in</quantity>
distance between lateral margins of anterior apertures, 
<quantity id="89149B64FFD2FFA65A4CFF3357D9FF3A" box="[1356,1436,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.4638" pageId="4" pageNumber="592" unit="in" value="0.97">0.97 in</quantity>
internarial width; process of flap about twice as long as wide at its base, overlapping posteromesial edge of posterolateral nasal flap and determining inner margin of anterior aperture. Posterolateral nasal flap lobe-like, broadest medially, length 3.36 times width; originating at lateral extremity of anterior nasal aperture, extending postero-medially as a free fold almost to medial margin of posterior flap. Posterior nasal flap broadly lobe-like, base length 2.58 times its width, not reaching innermost end of nostril, inserted well forward of posterior tip; width subequal to anterior aperture width, 1.09 times posterolateral nasal-flap width. Nasal lamellae 52.
</paragraph>
<paragraph id="4E533681FFD5FFA15F9FFE9356BFF9DC" pageId="3" pageNumber="591">
<table id="3CECC421FFD5005D5F9FFE93575CF9DC" box="[159,1305,284,1587]" gridcols="4" gridrows="32" pageId="3" pageNumber="591">
<tr id="F0DC34C3FFD5005D5F9FFE93575CFEDD" box="[159,1305,284,306]" gridrow="0" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFE93500EFEDD" box="[159,587,284,306]" gridcol="0" gridrow="0" pageId="3" pageNumber="591">First dorsal fin—anterior margin length</th>
<th id="B30D5DBFFFD5005D5C53FE9351C5FEDD" box="[851,896,284,306]" gridcol="1" gridrow="0" pageId="3" pageNumber="591">9.7</th>
<th id="B30D5DBFFFD5005D5B34FE935624FEDD" box="[1076,1121,284,306]" gridcol="2" gridrow="0" pageId="3" pageNumber="591">10.1</th>
<th id="B30D5DBFFFD5005D5BEDFE93575CFEDD" box="[1261,1305,284,306]" gridcol="3" gridrow="0" pageId="3" pageNumber="591">11.5</th>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFECA575CFEB4" box="[159,1305,325,347]" gridrow="1" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFECA500EFEB4" box="[159,587,325,347]" gridcol="0" gridrow="1" pageId="3" pageNumber="591">First dorsal fin—height</th>
<td id="B30D5DBFFFD5005D5C53FECA51C5FEB4" box="[851,896,325,347]" gridcol="1" gridrow="1" pageId="3" pageNumber="591">7.9</td>
<td id="B30D5DBFFFD5005D5B34FECA5624FEB4" box="[1076,1121,325,347]" gridcol="2" gridrow="1" pageId="3" pageNumber="591">7.5</td>
<td id="B30D5DBFFFD5005D5BEDFECA575CFEB4" box="[1261,1305,325,347]" gridcol="3" gridrow="1" pageId="3" pageNumber="591">7.8</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFEE1575CFE6B" box="[159,1305,366,388]" gridrow="2" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFEE1500EFE6B" box="[159,587,366,388]" gridcol="0" gridrow="2" pageId="3" pageNumber="591">First dorsal fin—base length</th>
<td id="B30D5DBFFFD5005D5C53FEE151C5FE6B" box="[851,896,366,388]" gridcol="1" gridrow="2" pageId="3" pageNumber="591">4.3</td>
<td id="B30D5DBFFFD5005D5B34FEE15624FE6B" box="[1076,1121,366,388]" gridcol="2" gridrow="2" pageId="3" pageNumber="591">4.3</td>
<td id="B30D5DBFFFD5005D5BEDFEE1575CFE6B" box="[1261,1305,366,388]" gridcol="3" gridrow="2" pageId="3" pageNumber="591">4.4</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFE17575CFE41" box="[159,1305,408,430]" gridrow="3" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFE17500EFE41" box="[159,587,408,430]" gridcol="0" gridrow="3" pageId="3" pageNumber="591">First dorsal fin—inner margin length</th>
<td id="B30D5DBFFFD5005D5C53FE1751C5FE41" box="[851,896,408,430]" gridcol="1" gridrow="3" pageId="3" pageNumber="591">2.2</td>
<td id="B30D5DBFFFD5005D5B34FE175624FE41" box="[1076,1121,408,430]" gridcol="2" gridrow="3" pageId="3" pageNumber="591">2.4</td>
<td id="B30D5DBFFFD5005D5BEDFE17575CFE41" box="[1261,1305,408,430]" gridcol="3" gridrow="3" pageId="3" pageNumber="591">2.8</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFE4E575CFE38" box="[159,1305,449,471]" gridrow="4" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFE4E500EFE38" box="[159,587,449,471]" gridcol="0" gridrow="4" pageId="3" pageNumber="591">Second dorsal fin—length</th>
<td id="B30D5DBFFFD5005D5C53FE4E51C5FE38" box="[851,896,449,471]" gridcol="1" gridrow="4" pageId="3" pageNumber="591">6.3</td>
<td id="B30D5DBFFFD5005D5B34FE4E5624FE38" box="[1076,1121,449,471]" gridcol="2" gridrow="4" pageId="3" pageNumber="591">6.8</td>
<td id="B30D5DBFFFD5005D5BEDFE4E575CFE38" box="[1261,1305,449,471]" gridcol="3" gridrow="4" pageId="3" pageNumber="591">7.2</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFE65575CFDEF" box="[159,1305,490,512]" gridrow="5" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFE65500EFDEF" box="[159,587,490,512]" gridcol="0" gridrow="5" pageId="3" pageNumber="591">Second dorsal fin—anterior margin length</th>
<td id="B30D5DBFFFD5005D5C53FE6551C5FDEF" box="[851,896,490,512]" gridcol="1" gridrow="5" pageId="3" pageNumber="591">9.3</td>
<td id="B30D5DBFFFD5005D5B34FE655624FDEF" box="[1076,1121,490,512]" gridcol="2" gridrow="5" pageId="3" pageNumber="591">9.3</td>
<td id="B30D5DBFFFD5005D5BEDFE65575CFDEF" box="[1261,1305,490,512]" gridcol="3" gridrow="5" pageId="3" pageNumber="591">10.5</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFD9B575CFDC5" box="[159,1305,532,554]" gridrow="6" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFD9B500EFDC5" box="[159,587,532,554]" gridcol="0" gridrow="6" pageId="3" pageNumber="591">Second dorsal fin—height</th>
<td id="B30D5DBFFFD5005D5C53FD9B51C5FDC5" box="[851,896,532,554]" gridcol="1" gridrow="6" pageId="3" pageNumber="591">7.5</td>
<td id="B30D5DBFFFD5005D5B34FD9B5624FDC5" box="[1076,1121,532,554]" gridcol="2" gridrow="6" pageId="3" pageNumber="591">6.2</td>
<td id="B30D5DBFFFD5005D5BEDFD9B575CFDC5" box="[1261,1305,532,554]" gridcol="3" gridrow="6" pageId="3" pageNumber="591">7.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFDB2575CFDBC" box="[159,1305,573,595]" gridrow="7" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFDB2500EFDBC" box="[159,587,573,595]" gridcol="0" gridrow="7" pageId="3" pageNumber="591">Second dorsal fin—base length</th>
<td id="B30D5DBFFFD5005D5C53FDB251C5FDBC" box="[851,896,573,595]" gridcol="1" gridrow="7" pageId="3" pageNumber="591">4.6</td>
<td id="B30D5DBFFFD5005D5B34FDB25624FDBC" box="[1076,1121,573,595]" gridcol="2" gridrow="7" pageId="3" pageNumber="591">4.8</td>
<td id="B30D5DBFFFD5005D5BEDFDB2575CFDBC" box="[1261,1305,573,595]" gridcol="3" gridrow="7" pageId="3" pageNumber="591">5.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFDE9575CFD93" box="[159,1305,614,636]" gridrow="8" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFDE9500EFD93" box="[159,587,614,636]" gridcol="0" gridrow="8" pageId="3" pageNumber="591">Second dorsal fin—inner margin length</th>
<td id="B30D5DBFFFD5005D5C53FDE951C5FD93" box="[851,896,614,636]" gridcol="1" gridrow="8" pageId="3" pageNumber="591">1.8</td>
<td id="B30D5DBFFFD5005D5B34FDE95624FD93" box="[1076,1121,614,636]" gridcol="2" gridrow="8" pageId="3" pageNumber="591">2.0</td>
<td id="B30D5DBFFFD5005D5BEDFDE9575CFD93" box="[1261,1305,614,636]" gridcol="3" gridrow="8" pageId="3" pageNumber="591">2.5</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFD1F575CFD49" box="[159,1305,656,678]" gridrow="9" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFD1F500EFD49" box="[159,587,656,678]" gridcol="0" gridrow="9" pageId="3" pageNumber="591">Caudal fin—dorsal margin</th>
<td id="B30D5DBFFFD5005D5C53FD1F51C5FD49" box="[851,896,656,678]" gridcol="1" gridrow="9" pageId="3" pageNumber="591">14.3</td>
<td id="B30D5DBFFFD5005D5B34FD1F5624FD49" box="[1076,1121,656,678]" gridcol="2" gridrow="9" pageId="3" pageNumber="591">14.6</td>
<td id="B30D5DBFFFD5005D5BEDFD1F575CFD49" box="[1261,1305,656,678]" gridcol="3" gridrow="9" pageId="3" pageNumber="591">15.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFD36575CFD20" box="[159,1305,697,719]" gridrow="10" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFD36500EFD20" box="[159,587,697,719]" gridcol="0" gridrow="10" pageId="3" pageNumber="591">Caudal fin—preventral margin</th>
<td id="B30D5DBFFFD5005D5C53FD3651C5FD20" box="[851,896,697,719]" gridcol="1" gridrow="10" pageId="3" pageNumber="591">7.2</td>
<td id="B30D5DBFFFD5005D5B34FD365624FD20" box="[1076,1121,697,719]" gridcol="2" gridrow="10" pageId="3" pageNumber="591">6.5</td>
<td id="B30D5DBFFFD5005D5BEDFD36575CFD20" box="[1261,1305,697,719]" gridcol="3" gridrow="10" pageId="3" pageNumber="591">6.9</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFD6D575CFD17" box="[159,1305,738,760]" gridrow="11" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFD6D500EFD17" box="[159,587,738,760]" gridcol="0" gridrow="11" pageId="3" pageNumber="591">Caudal fin—lower postventral margin</th>
<td id="B30D5DBFFFD5005D5C53FD6D51C5FD17" box="[851,896,738,760]" gridcol="1" gridrow="11" pageId="3" pageNumber="591">8.2</td>
<td id="B30D5DBFFFD5005D5B34FD6D5624FD17" box="[1076,1121,738,760]" gridcol="2" gridrow="11" pageId="3" pageNumber="591">0.0</td>
<td id="B30D5DBFFFD5005D5BEDFD6D575CFD17" box="[1261,1305,738,760]" gridcol="3" gridrow="11" pageId="3" pageNumber="591">0.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFC83575CFCCD" box="[159,1305,780,802]" gridrow="12" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFC83500EFCCD" box="[159,587,780,802]" gridcol="0" gridrow="12" pageId="3" pageNumber="591">Snout to first dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FC8351C5FCCD" box="[851,896,780,802]" gridcol="1" gridrow="12" pageId="3" pageNumber="591">56.2</td>
<td id="B30D5DBFFFD5005D5B34FC835624FCCD" box="[1076,1121,780,802]" gridcol="2" gridrow="12" pageId="3" pageNumber="591">56.1</td>
<td id="B30D5DBFFFD5005D5BEDFC83575CFCCD" box="[1261,1305,780,802]" gridcol="3" gridrow="12" pageId="3" pageNumber="591">56.7</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFCBA575CFCA4" box="[159,1305,821,843]" gridrow="13" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFCBA500EFCA4" box="[159,587,821,843]" gridcol="0" gridrow="13" pageId="3" pageNumber="591">Snout to second dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FCBA51C5FCA4" box="[851,896,821,843]" gridcol="1" gridrow="13" pageId="3" pageNumber="591">73.9</td>
<td id="B30D5DBFFFD5005D5B34FCBA5624FCA4" box="[1076,1121,821,843]" gridcol="2" gridrow="13" pageId="3" pageNumber="591">72.9</td>
<td id="B30D5DBFFFD5005D5BEDFCBA575CFCA4" box="[1261,1305,821,843]" gridcol="3" gridrow="13" pageId="3" pageNumber="591">73.7</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFCD1575CFC9B" box="[159,1305,862,884]" gridrow="14" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFCD1500EFC9B" box="[159,587,862,884]" gridcol="0" gridrow="14" pageId="3" pageNumber="591">Snout to upper caudal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FCD151C5FC9B" box="[851,896,862,884]" gridcol="1" gridrow="14" pageId="3" pageNumber="591">85.5</td>
<td id="B30D5DBFFFD5005D5B34FCD15624FC9B" box="[1076,1121,862,884]" gridcol="2" gridrow="14" pageId="3" pageNumber="591">85.0</td>
<td id="B30D5DBFFFD5005D5BEDFCD1575CFC9B" box="[1261,1305,862,884]" gridcol="3" gridrow="14" pageId="3" pageNumber="591">85.4</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFC07575CFC71" box="[159,1305,904,926]" gridrow="15" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFC07500EFC71" box="[159,587,904,926]" gridcol="0" gridrow="15" pageId="3" pageNumber="591">Snout to lower caudal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FC0751C5FC71" box="[851,896,904,926]" gridcol="1" gridrow="15" pageId="3" pageNumber="591">86.3</td>
<td id="B30D5DBFFFD5005D5B34FC075624FC71" box="[1076,1121,904,926]" gridcol="2" gridrow="15" pageId="3" pageNumber="591">86.5</td>
<td id="B30D5DBFFFD5005D5BEDFC07575CFC71" box="[1261,1305,904,926]" gridcol="3" gridrow="15" pageId="3" pageNumber="591">87.2</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFC3E575CFC28" box="[159,1305,945,967]" gridrow="16" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFC3E500EFC28" box="[159,587,945,967]" gridcol="0" gridrow="16" pageId="3" pageNumber="591">Snout to pelvic-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FC3E51C5FC28" box="[851,896,945,967]" gridcol="1" gridrow="16" pageId="3" pageNumber="591">39.2</td>
<td id="B30D5DBFFFD5005D5B34FC3E5624FC28" box="[1076,1121,945,967]" gridcol="2" gridrow="16" pageId="3" pageNumber="591">38.1</td>
<td id="B30D5DBFFFD5005D5BEDFC3E575CFC28" box="[1261,1305,945,967]" gridcol="3" gridrow="16" pageId="3" pageNumber="591">39.5</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFC55575CFC1F" box="[159,1305,986,1008]" gridrow="17" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFC55500EFC1F" box="[159,587,986,1008]" gridcol="0" gridrow="17" pageId="3" pageNumber="591">Snout to anterior vent</th>
<td id="B30D5DBFFFD5005D5C53FC5551C5FC1F" box="[851,896,986,1008]" gridcol="1" gridrow="17" pageId="3" pageNumber="591">41.9</td>
<td id="B30D5DBFFFD5005D5B34FC555624FC1F" box="[1076,1121,986,1008]" gridcol="2" gridrow="17" pageId="3" pageNumber="591">40.6</td>
<td id="B30D5DBFFFD5005D5BEDFC55575CFC1F" box="[1261,1305,986,1008]" gridcol="3" gridrow="17" pageId="3" pageNumber="591">42.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFB8B575CFBF5" box="[159,1305,1028,1050]" gridrow="18" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFB8B500EFBF5" box="[159,587,1028,1050]" gridcol="0" gridrow="18" pageId="3" pageNumber="591">Pelvic-fin insertion to dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FB8B51C5FBF5" box="[851,896,1028,1050]" gridcol="1" gridrow="18" pageId="3" pageNumber="591">9.8</td>
<td id="B30D5DBFFFD5005D5B34FB8B5624FBF5" box="[1076,1121,1028,1050]" gridcol="2" gridrow="18" pageId="3" pageNumber="591">10.3</td>
<td id="B30D5DBFFFD5005D5BEDFB8B575CFBF5" box="[1261,1305,1028,1050]" gridcol="3" gridrow="18" pageId="3" pageNumber="591">11.2</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFBA2575CFBAC" box="[159,1305,1069,1091]" gridrow="19" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFBA2500EFBAC" box="[159,587,1069,1091]" gridcol="0" gridrow="19" pageId="3" pageNumber="591">Interdorsal distance</th>
<td id="B30D5DBFFFD5005D5C53FBA251C5FBAC" box="[851,896,1069,1091]" gridcol="1" gridrow="19" pageId="3" pageNumber="591">13.3</td>
<td id="B30D5DBFFFD5005D5B34FBA25624FBAC" box="[1076,1121,1069,1091]" gridcol="2" gridrow="19" pageId="3" pageNumber="591">12.2</td>
<td id="B30D5DBFFFD5005D5BEDFBA2575CFBAC" box="[1261,1305,1069,1091]" gridcol="3" gridrow="19" pageId="3" pageNumber="591">12.5</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFBD9575CFB83" box="[159,1305,1110,1132]" gridrow="20" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFBD9500EFB83" box="[159,587,1110,1132]" gridcol="0" gridrow="20" pageId="3" pageNumber="591">Caudal peduncle length (dorsal)</th>
<td id="B30D5DBFFFD5005D5C53FBD951C5FB83" box="[851,896,1110,1132]" gridcol="1" gridrow="20" pageId="3" pageNumber="591">6.8</td>
<td id="B30D5DBFFFD5005D5B34FBD95624FB83" box="[1076,1121,1110,1132]" gridcol="2" gridrow="20" pageId="3" pageNumber="591">7.0</td>
<td id="B30D5DBFFFD5005D5BEDFBD9575CFB83" box="[1261,1305,1110,1132]" gridcol="3" gridrow="20" pageId="3" pageNumber="591">7.2</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFB0F575CFB79" box="[159,1305,1152,1174]" gridrow="21" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFB0F500EFB79" box="[159,587,1152,1174]" gridcol="0" gridrow="21" pageId="3" pageNumber="591">Body width—pelvic insertion (tail)</th>
<td id="B30D5DBFFFD5005D5C53FB0F51C5FB79" box="[851,896,1152,1174]" gridcol="1" gridrow="21" pageId="3" pageNumber="591">9.2</td>
<td id="B30D5DBFFFD5005D5B34FB0F5624FB79" box="[1076,1121,1152,1174]" gridcol="2" gridrow="21" pageId="3" pageNumber="591">9.5</td>
<td id="B30D5DBFFFD5005D5BEDFB0F575CFB79" box="[1261,1305,1152,1174]" gridcol="3" gridrow="21" pageId="3" pageNumber="591">10.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFB26575CFB50" box="[159,1305,1193,1215]" gridrow="22" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFB26500EFB50" box="[159,587,1193,1215]" gridcol="0" gridrow="22" pageId="3" pageNumber="591">Disc width—anterior orbit</th>
<td id="B30D5DBFFFD5005D5C53FB2651C5FB50" box="[851,896,1193,1215]" gridcol="1" gridrow="22" pageId="3" pageNumber="591">18.3</td>
<td id="B30D5DBFFFD5005D5B34FB265624FB50" box="[1076,1121,1193,1215]" gridcol="2" gridrow="22" pageId="3" pageNumber="591">17.4</td>
<td id="B30D5DBFFFD5005D5BEDFB26575CFB50" box="[1261,1305,1193,1215]" gridcol="3" gridrow="22" pageId="3" pageNumber="591">17.7</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFB5D575CFB07" box="[159,1305,1234,1256]" gridrow="23" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFB5D500EFB07" box="[159,587,1234,1256]" gridcol="0" gridrow="23" pageId="3" pageNumber="591">Body width—first dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FB5D51C5FB07" box="[851,896,1234,1256]" gridcol="1" gridrow="23" pageId="3" pageNumber="591">9.2</td>
<td id="B30D5DBFFFD5005D5B34FB5D5624FB07" box="[1076,1121,1234,1256]" gridcol="2" gridrow="23" pageId="3" pageNumber="591">8.9</td>
<td id="B30D5DBFFFD5005D5BEDFB5D575CFB07" box="[1261,1305,1234,1256]" gridcol="3" gridrow="23" pageId="3" pageNumber="591">9.2</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFB73575CFAFD" box="[159,1305,1276,1298]" gridrow="24" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFB73500EFAFD" box="[159,587,1276,1298]" gridcol="0" gridrow="24" pageId="3" pageNumber="591">Body width—second dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FB7351C5FAFD" box="[851,896,1276,1298]" gridcol="1" gridrow="24" pageId="3" pageNumber="591">4.3</td>
<td id="B30D5DBFFFD5005D5B34FB735624FAFD" box="[1076,1121,1276,1298]" gridcol="2" gridrow="24" pageId="3" pageNumber="591">4.6</td>
<td id="B30D5DBFFFD5005D5BEDFB73575CFAFD" box="[1261,1305,1276,1298]" gridcol="3" gridrow="24" pageId="3" pageNumber="591">5.0</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFAAA575CFAD4" box="[159,1305,1317,1339]" gridrow="25" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFAAA500EFAD4" box="[159,587,1317,1339]" gridcol="0" gridrow="25" pageId="3" pageNumber="591">Body depth—maximum (scap)</th>
<td id="B30D5DBFFFD5005D5C53FAAA51C5FAD4" box="[851,896,1317,1339]" gridcol="1" gridrow="25" pageId="3" pageNumber="591">3.8</td>
<td id="B30D5DBFFFD5005D5B34FAAA5624FAD4" box="[1076,1121,1317,1339]" gridcol="2" gridrow="25" pageId="3" pageNumber="591">5.0</td>
<td id="B30D5DBFFFD5005D5BEDFAAA575CFAD4" box="[1261,1305,1317,1339]" gridcol="3" gridrow="25" pageId="3" pageNumber="591">6.5</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFAC1575CFA8B" box="[159,1305,1358,1380]" gridrow="26" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFAC1500EFA8B" box="[159,587,1358,1380]" gridcol="0" gridrow="26" pageId="3" pageNumber="591">Body depth—pelvic-fin insertion</th>
<td id="B30D5DBFFFD5005D5C53FAC151C5FA8B" box="[851,896,1358,1380]" gridcol="1" gridrow="26" pageId="3" pageNumber="591">4.6</td>
<td id="B30D5DBFFFD5005D5B34FAC15624FA8B" box="[1076,1121,1358,1380]" gridcol="2" gridrow="26" pageId="3" pageNumber="591">4.4</td>
<td id="B30D5DBFFFD5005D5BEDFAC1575CFA8B" box="[1261,1305,1358,1380]" gridcol="3" gridrow="26" pageId="3" pageNumber="591">4.6</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFAF7575CFA61" box="[159,1305,1400,1422]" gridrow="27" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFAF7500EFA61" box="[159,587,1400,1422]" gridcol="0" gridrow="27" pageId="3" pageNumber="591">Body depth—first dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FAF751C5FA61" box="[851,896,1400,1422]" gridcol="1" gridrow="27" pageId="3" pageNumber="591">3.8</td>
<td id="B30D5DBFFFD5005D5B34FAF75624FA61" box="[1076,1121,1400,1422]" gridcol="2" gridrow="27" pageId="3" pageNumber="591">3.7</td>
<td id="B30D5DBFFFD5005D5BEDFAF7575CFA61" box="[1261,1305,1400,1422]" gridcol="3" gridrow="27" pageId="3" pageNumber="591">3.7</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFA2E575CFA58" box="[159,1305,1441,1463]" gridrow="28" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFA2E500EFA58" box="[159,587,1441,1463]" gridcol="0" gridrow="28" pageId="3" pageNumber="591">Body depth—second dorsal-fin origin</th>
<td id="B30D5DBFFFD5005D5C53FA2E51C5FA58" box="[851,896,1441,1463]" gridcol="1" gridrow="28" pageId="3" pageNumber="591">2.5</td>
<td id="B30D5DBFFFD5005D5B34FA2E5624FA58" box="[1076,1121,1441,1463]" gridcol="2" gridrow="28" pageId="3" pageNumber="591">2.3</td>
<td id="B30D5DBFFFD5005D5BEDFA2E575CFA58" box="[1261,1305,1441,1463]" gridcol="3" gridrow="28" pageId="3" pageNumber="591">2.4</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFA45575CFA0F" box="[159,1305,1482,1504]" gridrow="29" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFA45500EFA0F" box="[159,587,1482,1504]" gridcol="0" gridrow="29" pageId="3" pageNumber="591">Clasper—outer length</th>
<td id="B30D5DBFFFD5005D5C53FA4551C5FA0F" box="[851,896,1482,1504]" gridcol="1" gridrow="29" pageId="3" pageNumber="591">8.4</td>
<td id="B30D5DBFFFD5005D5B34FA455624FA0F" box="[1076,1121,1482,1504]" gridcol="2" gridrow="29" pageId="3" pageNumber="591">–</td>
<td id="B30D5DBFFFD5005D5BEDFA45575CFA0F" box="[1261,1305,1482,1504]" gridcol="3" gridrow="29" pageId="3" pageNumber="591">–</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FFA7B575CF9E5" box="[159,1305,1524,1546]" gridrow="30" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FFA7B500EF9E5" box="[159,587,1524,1546]" gridcol="0" gridrow="30" pageId="3" pageNumber="591">Clasper—inner length</th>
<td id="B30D5DBFFFD5005D5C53FA7B51C5F9E5" box="[851,896,1524,1546]" gridcol="1" gridrow="30" pageId="3" pageNumber="591">13.1</td>
<td id="B30D5DBFFFD5005D5B34FA7B5624F9E5" box="[1076,1121,1524,1546]" gridcol="2" gridrow="30" pageId="3" pageNumber="591">–</td>
<td id="B30D5DBFFFD5005D5BEDFA7B575CF9E5" box="[1261,1305,1524,1546]" gridcol="3" gridrow="30" pageId="3" pageNumber="591">–</td>
</tr>
<tr id="F0DC34C3FFD5005D5F9FF992575CF9DC" box="[159,1305,1565,1587]" gridrow="31" pageId="3" pageNumber="591">
<th id="B30D5DBFFFD5005D5F9FF992500EF9DC" box="[159,587,1565,1587]" gridcol="0" gridrow="31" pageId="3" pageNumber="591">Clasper—base length</th>
<td id="B30D5DBFFFD5005D5C53F99251C5F9DC" box="[851,896,1565,1587]" gridcol="1" gridrow="31" pageId="3" pageNumber="591">1.2</td>
<td id="B30D5DBFFFD5005D5B34F9925624F9DC" box="[1076,1121,1565,1587]" gridcol="2" gridrow="31" pageId="3" pageNumber="591">–</td>
<td id="B30D5DBFFFD5005D5BEDF992575CF9DC" box="[1261,1305,1565,1587]" gridcol="3" gridrow="31" pageId="3" pageNumber="591">–</td>
</tr>
</table>
</paragraph>
<caption id="1A936609FFD2FFA65F97FA2B5365FA36" httpUri="https://zenodo.org/record/256166/files/figure.png" pageId="4" pageNumber="592" targetBox="[163,1418,520,1378]" targetPageId="4">
<paragraph id="4E533681FFD2FFA65F97FA2B5365FA36" blockId="4.[151,1436,1444,1497]" pageId="4" pageNumber="592">
<emphasis id="7C98EA93FFD2FFA65F97FA2B5351FA55" bold="true" box="[151,276,1444,1466]" pageId="4" pageNumber="592">FIGURE 1.</emphasis>
Dorsal view of 
<taxonomicName id="89EC4D02FFD2FFA65EBAFA2B5036FA54" box="[442,627,1444,1467]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="4" pageNumber="592" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD2FFA65EBAFA2B5036FA54" box="[442,627,1444,1467]" italics="true" pageId="4" pageNumber="592">Rhinobatos manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFD2FFA65D7BFA2B5089FA55" box="[635,716,1444,1466]" pageId="4" pageNumber="592" rank="species">
<emphasis id="7C98EA93FFD2FFA65D7BFA2B5089FA55" bold="true" box="[635,716,1444,1466]" pageId="4" pageNumber="592">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult male holotype, 731 mm TL (NTUM 11500): A. freshly caught; B) preserved.
</paragraph>
</caption>
<paragraph id="4E533681FFD2FFA65FC7F98F5183F8F9" blockId="4.[151,1437,1536,2030]" pageId="4" pageNumber="592">
Mouth moderately wide, width 1.29 times nostril length, 
<quantity id="89149B64FFD2FFA65C48F98F51D3F9F6" box="[840,918,1536,1561]" metricMagnitude="-1" metricUnit="m" metricValue="1.84404" pageId="4" pageNumber="592" unit="in" value="7.26">7.26 in</quantity>
precloacal length; positioned about level with hind margin of orbit; jaws not greatly thickened. Upper jaw weakly convex, upper lip arched slightly, no preoral groove; lower lip pronounced, not separated from post-oral groove by ridges of strongly corrugated skin; short lateral grooves around corners of mouth. Teeth small, blunt, crowns rhomboidal; teeth quincuncial, ~92 rows in upper jaw and ~86 rows in lower jaw; upper and lower jaw teeth similar in shape and size. Gill openings weakly sshaped, fifth less so; length of third gill slit 
<quantity id="89149B64FFD2FFA65D86F93A5091F921" box="[646,724,1717,1742]" metricMagnitude="-2" metricUnit="m" metricValue="7.4676" pageId="4" pageNumber="592" unit="in" value="2.94">2.94 in</quantity>
nostril length, 
<quantity id="89149B64FFD2FFA65C80F93A518AF921" box="[896,975,1717,1742]" metricMagnitude="-1" metricUnit="m" metricValue="1.40462" pageId="4" pageNumber="592" unit="in" value="5.53">5.53 in</quantity>
distance between fifth gill slits; distance between first gill slits 1.47 times distance between fifth gill slits; distance between fifth gill slits 2.98 times internarial distance, 1.46 times mouth width, 
<quantity id="89149B64FFD2FFA65D93F97250A4F8F9" box="[659,737,1789,1814]" metricMagnitude="-2" metricUnit="m" metricValue="8.636" pageId="4" pageNumber="592" unit="in" value="3.4">3.40 in</quantity>
ventral head length.
</paragraph>
<paragraph id="4E533681FFD2FFA75FC7F8AF5327FF17" blockId="4.[151,1437,1536,2030]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="593" pageId="4" pageNumber="592">Dorsal fins of moderate size, upright, relatively narrow, not falcate, apices narrowly rounded to almost angular; anterior margins convex distally, posterior margins nearly straight; free rear tips forming right angle, not produced; first dorsal-fin slightly taller than second, length of first dorsal fin 0.82 times its height, base length 1.99 times inner margin length; second dorsal-fin length 0.85 times its height, base length 2.58 times inner margin length. First dorsal-fin origin well behind pelvic-fin rear tip, interspace 0.73 times interdorsal distance; interdorsal space relatively short, 1.78 times second dorsal-fin height, 3.08 times base length of first dorsal-fin, 1.95 times interspace between second dorsal-fin insertion and upper origin of caudal fin. Caudal fin small, dorsal caudal margin 1.98 times preventral margin length. Mature clasper slender, relatively short, inner length 13.1% TL; tip acute, glans weakly expanded.</paragraph>
<caption id="1A936609FFD3FFA75F97FBEB575BFB95" box="[151,1310,1124,1146]" httpUri="https://zenodo.org/record/256167/files/figure.png" pageId="5" pageNumber="593" targetBox="[169,1432,336,1090]" targetPageId="5">
<paragraph id="4E533681FFD3FFA75F97FBEB575BFB95" blockId="5.[151,1310,1124,1146]" box="[151,1310,1124,1146]" pageId="5" pageNumber="593">
<emphasis id="7C98EA93FFD3FFA75F97FBEB5357FB95" bold="true" box="[151,274,1124,1146]" pageId="5" pageNumber="593">FIGURE 2.</emphasis>
Head of preserved adult male holotype, 731 mm TL (NTUM 11500): A. dorsal view; B. ventral view.
</paragraph>
</caption>
<caption id="1A936609FFD3FFA75F97F8B6570EF8A0" box="[151,1355,1849,1871]" httpUri="https://zenodo.org/record/256168/files/figure.png" pageId="5" pageNumber="593" targetBox="[340,1246,1196,1827]" targetPageId="5">
<paragraph id="4E533681FFD3FFA75F97F8B6570EF8A0" blockId="5.[151,1355,1849,1871]" box="[151,1355,1849,1871]" pageId="5" pageNumber="593">
<emphasis id="7C98EA93FFD3FFA75F97F8B65357F8A0" bold="true" box="[151,274,1849,1871]" pageId="5" pageNumber="593">FIGURE 3.</emphasis>
Orbital region of 
<taxonomicName id="89EC4D02FFD3FFA75EC9F8B650C7F8A0" box="[457,642,1849,1871]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="5" pageNumber="593" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD3FFA75EC9F8B650C7F8A0" box="[457,642,1849,1871]" italics="true" pageId="5" pageNumber="593">Rhinobatos manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFD3FFA75D89F8B6509CF8A0" box="[649,729,1849,1871]" pageId="5" pageNumber="593" rank="species">
<emphasis id="7C98EA93FFD3FFA75D89F8B6509CF8A0" bold="true" box="[649,729,1849,1871]" pageId="5" pageNumber="593">sp. nov.</emphasis>
</taxonomicNameLabel>
, preserved adult male holotype, 731 mm TL (NTUM 11500).
</paragraph>
</caption>
<paragraph id="4E533681FFD3FFA45FC7F8F350E9FE8A" blockId="5.[151,1436,1916,2013]" lastBlockId="6.[151,1437,151,464]" lastPageId="6" lastPageNumber="594" pageId="5" pageNumber="593">Dermal denticles minute, close-set, covering entire body and fins; thorns and tubercles absent; dorsal surface with narrow series of slightly enlarged, seed-like denticles around orbit, along midline, and on scapular region; around orbit, enlarged denticles most pronounced anteriorly at preorbit and posteriorly, with posterior patch extending over upper spiracle margin; along midline, enlarged denticles most pronounced above abdomen; weakly represented between dorsal fins and barely evident on caudal peduncle; enlarged denticles irregular in size and shape, largest with crenulate anterior margins; dorsal surface of claspers naked at tip and near pelvic-fin insertion. Ventral surface uniformly covered with minute denticles, including upper lip edges, near insertion of anterior nasal flap, below posterolateral and posterior nasal flaps, and on tail beneath pelvic fins, and most of claspers; a dense covering of small denticles over nasal lamellae.</paragraph>
<paragraph id="4E533681FFD0FFA45FC7FEE050AEFE3F" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="594">Prebranchial sensory pore patch relatively narrow, extending posteriorly to level of first gill slit. Postscapular sensory canal long, notched near fifth gill slits, terminating about an orbit diameter from pectoral-fin insertions; canal deeply embedded, not forming shallow groove.</paragraph>
<caption id="1A936609FFD0FFA45F97F8E0537BF84B" httpUri="https://zenodo.org/record/256169/files/figure.png" pageId="6" pageNumber="594" targetBox="[340,1246,517,1881]" targetPageId="6">
<paragraph id="4E533681FFD0FFA45F97F8E0537BF84B" blockId="6.[151,1436,1903,1956]" pageId="6" pageNumber="594">
<emphasis id="7C98EA93FFD0FFA45F97F8E05350F86A" bold="true" box="[151,277,1903,1925]" pageId="6" pageNumber="594">FIGURE 4.</emphasis>
Radiograph of the chondrocranium of 
<taxonomicName id="89EC4D02FFD0FFA45DAFF8E0512FF869" box="[687,874,1903,1926]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="6" pageNumber="594" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD0FFA45DAFF8E0512FF869" box="[687,874,1903,1926]" italics="true" pageId="6" pageNumber="594">Rhinobatos manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFD0FFA45C73F8E05183F86A" box="[883,966,1903,1925]" pageId="6" pageNumber="594" rank="species">
<emphasis id="7C98EA93FFD0FFA45C73F8E05183F86A" bold="true" box="[883,966,1903,1925]" pageId="6" pageNumber="594">sp. nov.</emphasis>
</taxonomicNameLabel>
, preserved adult male holotype, 731 mm TL (NTUM 11500).
</paragraph>
</caption>
<caption id="1A936609FFD1FFA55F97FCC9571BFCB3" box="[151,1374,838,861]" httpUri="https://zenodo.org/record/256170/files/figure.png" pageId="7" pageNumber="595" targetBox="[340,1246,193,816]" targetPageId="7">
<paragraph id="4E533681FFD1FFA55F97FCC9571BFCB3" blockId="7.[151,1374,838,861]" box="[151,1374,838,861]" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55F97FCC95357FCB3" bold="true" box="[151,274,838,860]" pageId="7" pageNumber="595">FIGURE 5.</emphasis>
Oronasal region of 
<taxonomicName id="89EC4D02FFD1FFA55EDDFCC950D0FCB2" box="[477,661,838,861]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="595" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFD1FFA55EDDFCC950D0FCB2" box="[477,661,838,861]" italics="true" pageId="7" pageNumber="595">Rhinobatos manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFD1FFA55D9CFCC950A8FCB3" box="[668,749,838,860]" pageId="7" pageNumber="595" rank="species">
<emphasis id="7C98EA93FFD1FFA55D9CFCC950A8FCB3" bold="true" box="[668,749,838,860]" pageId="7" pageNumber="595">sp. nov.</emphasis>
</taxonomicNameLabel>
, preserved adult male holotype, 731 mm TL (NTUM 11500).
</paragraph>
</caption>
<paragraph id="4E533681FFD1FFA55FC7FC0B5607FB77" blockId="7.[151,1436,900,2005]" pageId="7" pageNumber="595">
Rostral cartilage in 
<typeStatus id="91578823FFD1FFA55EA8FC0B504FFC72" box="[424,522,900,925]" pageId="7" pageNumber="595" type="holotype">holotype</typeStatus>
long and broad, its shaft not increasing in width in a posterior direction; rostral node broadly expanded and elongate, rounded apically, not angular, axis at widest part of node 9.4% of length of rostral cartilage from snout tip; precerebral cavity broad and uniformly convex posteriorly, narrowly rounded anteriorly at rostral node; rostral cartilage length ~67% of length of neurocranium, ventral edges of rostral cartilage united; nasal capsules large, their transverse axes anterolaterally directed; maximum width across capsules 1.52 times nasobasal length of cranium (base of rostrum to occipital condyles); nasal capsules slightly wider than long; basal plate narrow, its minimum width 5.04 times in nasobasal length; cranial roof with small, narrowly oval fenestra, located well behind precerebral cavity (separated by ~1.4 times its length).
</paragraph>
<paragraph id="4E533681FFD1FFA55FC7FB2B5376FA80" blockId="7.[151,1436,900,2005]" pageId="7" pageNumber="595">Pectoral skeleton with 30 propterygial, 7 mesopterygial, 2 neopterygial, 27 metapterygial, 66 total radials; anterior radials of propterygium extending forward of nasal capsules by about 10.6% of rostral length. Total pelvic radials 1+24+1; first greatly enlarged, on puboischiadic bar; 24 basipterygial radials; clasper calcified. Vertebral column with 188 total centra (synarcual and free), 171 post-synarcual centra; 17 synarcual centra; 25 monospondylous centra, all centra with ribs; 108 diplospondylous precaudal centra, about 38 diplospondylous caudal centra.</paragraph>
<paragraph id="4E533681FFD1FFA55FC7FAF45352F983" blockId="7.[151,1436,900,2005]" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55FC7FAF45365FA7B" bold="true" box="[199,288,1403,1428]" pageId="7" pageNumber="595">Colour.</emphasis>
<emphasis id="7C98EA93FFD1FFA55E28FAF25394FA7B" box="[296,465,1405,1428]" italics="true" pageId="7" pageNumber="595">In preservative</emphasis>
: Body pale yellowish brown dorsally covered with a complex pattern of small, rusty brown spots and blotches, and small whitish spots with greyish edges; rusty brown spots diffuse-edged and somewhat irregular in size, mostly circular, larger near disc edges and on lateral trunk; whitish spots sparser and more diffuse, poorly defined anteriorly and posteriorly on body; paler near hind margin of pectoral and pelvic fins; paler yellowish brown on translucent areas of snout, lateral cutaneous fold of tail, and between ridges of rostral cartilage; dorsal and caudal fins also with several brownish spots. Ventral surface uniformly white; no dark tip on snout apex.
</paragraph>
<paragraph id="4E533681FFD1FFA55FC7F9F851CEF97F" blockId="7.[151,1436,900,2005]" box="[199,907,1655,1680]" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55FC7F9F852BBF97F" bold="true" box="[199,254,1655,1680]" pageId="7" pageNumber="595">Size.</emphasis>
Only known from the 
<typeStatus id="91578823FFD1FFA55EFEF9F85026F97F" box="[510,611,1655,1680]" pageId="7" pageNumber="595" type="holotype">holotype</typeStatus>
, a 
<quantity id="89149B64FFD1FFA55D81F9F7509BF960" box="[641,734,1656,1680]" metricMagnitude="-1" metricUnit="m" metricValue="7.31" pageId="7" pageNumber="595" unit="mm" value="731.0">731 mm</quantity>
TL adult male.
</paragraph>
</subSubSection>
<subSubSection id="06F6650AFFD1FFA55FC7F9145173F937" pageId="7" pageNumber="595" type="distribution">
<paragraph id="4E533681FFD1FFA55FC7F9145173F937" blockId="7.[151,1436,900,2005]" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55FC7F9145325F95B" bold="true" box="[199,352,1691,1716]" pageId="7" pageNumber="595">Distribution.</emphasis>
<typeStatus id="91578823FFD1FFA55E6AF9135397F95A" box="[362,466,1692,1717]" pageId="7" pageNumber="595" type="holotype">Holotype</typeStatus>
collected northwest of Kavieng (
<geoCoordinate id="2BD85046FFD1FFA55C54F91351F6F95B" box="[852,947,1692,1716]" direction="south" orientation="latitude" pageId="7" pageNumber="595" precision="925" value="-2.5">02°30’S</geoCoordinate>
, 
<geoCoordinate id="2BD85046FFD1FFA55CC2F9135675F95A" box="[962,1072,1692,1717]" direction="east" orientation="longitude" pageId="7" pageNumber="595" precision="925" value="150.73334">150°44’E</geoCoordinate>
), 
<collectingRegion id="8C28F863FFD1FFA55B47F912569DF95A" box="[1095,1240,1692,1717]" country="Papua New Guinea" name="New Ireland" pageId="7" pageNumber="595">
New 
<collectingCountry id="36FB7611FFD1FFA55B86F913569DF95A" box="[1158,1240,1692,1717]" name="Ireland" pageId="7" pageNumber="595">Ireland</collectingCountry>
</collectingRegion>
, in the 
<collectingCountry id="36FB7611FFD1FFA55A31F9135367F937" name="Papua New Guinea" pageId="7" pageNumber="595">Bismarck Archipelago</collectingCountry>
of 
<collectingCountry id="36FB7611FFD1FFA55E46F94E505AF937" box="[326,543,1727,1752]" name="Papua New Guinea" pageId="7" pageNumber="595">
<collectingRegion id="8C28F863FFD1FFA55E46F94E53CEF937" box="[326,395,1729,1752]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="595">Papua</collectingRegion>
New Guinea
</collectingCountry>
at depths of 
<quantity id="89149B64FFD1FFA55DB0F94F5177F937" box="[688,818,1728,1752]" metricMagnitude="2" metricUnit="m" metricValue="2.405" metricValueMax="2.9" metricValueMin="1.91" pageId="7" pageNumber="595" unit="m" value="240.5" valueMax="290.0" valueMin="191.0">
191– 
<specimenCount id="58EAFD08FFD1FFA55DE6F94F5177F937" box="[742,818,1728,1752]" pageId="7" pageNumber="595" type="generic">290 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="06F6650AFFD1FFAA5FC7F96B533CFF3A" lastPageId="8" lastPageNumber="596" pageId="7" pageNumber="595" type="etymology">
<paragraph id="4E533681FFD1FFA55FC7F96B50ABF8AA" blockId="7.[151,1436,900,2005]" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55FC7F96B530BF913" bold="true" box="[199,334,1764,1788]" pageId="7" pageNumber="595">Etymology.</emphasis>
Epithet in recognition of Dr Ralph Mana of the University of 
<collectingCountry id="36FB7611FFD1FFA55B5FF96A5703F912" box="[1119,1350,1764,1789]" name="Papua New Guinea" pageId="7" pageNumber="595">
<collectingRegion id="8C28F863FFD1FFA55B5FF96A56E1F913" box="[1119,1188,1765,1788]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="595">Papua</collectingRegion>
New Guinea
</collectingCountry>
whose invaluable work on the BioPapua projects throughout 
<collectingCountry id="36FB7611FFD1FFA55DF8F8865197F8CF" box="[760,978,1799,1824]" name="Papua New Guinea" pageId="7" pageNumber="595">
<collectingRegion id="8C28F863FFD1FFA55DF8F8865179F8CF" box="[760,828,1801,1824]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="595">Papua</collectingRegion>
New Guinea
</collectingCountry>
has led to a considerable increase in our knowledge of the deepwater fish fauna of this region.
</paragraph>
<paragraph id="4E533681FFD1FFAA5FC7F8DF533CFF3A" blockId="7.[151,1436,900,2005]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="596" pageId="7" pageNumber="595">
<emphasis id="7C98EA93FFD1FFA55FC7F8DF53EBF887" bold="true" box="[199,430,1871,1896]" pageId="7" pageNumber="595">Molecular analysis.</emphasis>
The analysis of the NADH2 data suggests that 
<taxonomicName id="89EC4D02FFD1FFA55CBDF8DF56CFF888" box="[957,1162,1872,1896]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="595" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFD1FFA55CBDF8DF56CFF888" box="[957,1162,1872,1896]" italics="true" pageId="7" pageNumber="595">Rhinobatos manai</emphasis>
</taxonomicName>
represents a lineage that is distinct from, but closely related to, 
<taxonomicName id="89EC4D02FFD1FFA55D66F8FA50BBF863" box="[614,766,1908,1932]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="595" phylum="Chordata" rank="species" species="sainsburyi">
<emphasis id="7C98EA93FFD1FFA55D66F8FA50BBF863" box="[614,766,1908,1932]" italics="true" pageId="7" pageNumber="595">R. sainsburyi</emphasis>
</taxonomicName>
and that these two species, in turn, are sister to a clade containing two recently described species from Borneo and the 
<collectingCountry id="36FB7611FFD1FFA55C6CF81751A8F85E" box="[876,1005,1944,1969]" name="Philippines" pageId="7" pageNumber="595">Philippines</collectingCountry>
, 
<taxonomicName id="89EC4D02FFD1FFA55CFAF8165751F85F" authority="Last, Seret" authorityName="Last, Seret" box="[1018,1300,1944,1968]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="595" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="7C98EA93FFD1FFA55CFAF81656D6F85F" box="[1018,1171,1944,1968]" italics="true" pageId="7" pageNumber="595">R. borneensis</emphasis>
Last, Séret
</taxonomicName>
&amp; Manjaji- Matsumoto, 2016 and 
<taxonomicName id="89EC4D02FFD1FFA55EA7F83251CAF83B" authority="Last, Corrigan &amp; Naylor, 2014" authorityName="Last, Corrigan &amp; Naylor" authorityYear="2014" box="[423,911,1980,2005]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="7" pageNumber="595" phylum="Chordata" rank="species" species="whitei">
<emphasis id="7C98EA93FFD1FFA55EA7F832504EF83B" box="[423,523,1980,2004]" italics="true" pageId="7" pageNumber="595">R. whitei</emphasis>
<bibRefCitation id="2A7D4B70FFD1FFA55D18F83251CAF83B" author="Last" box="[536,911,1980,2005]" pageId="7" pageNumber="595" refString="Last, P. R., Corrigan, S. &amp; Naylor, G. J. P. (2014) Rhinobatos whitei, a new shovelnose ray (Batoidea: Rhinobatidae) from the Philippine Archipelago. Zootaxa, 3872 (1), 31 - 47. http: // dx. doi. org / 10.11646 / zootaxa. 3872.1.3" type="journal article" year="2014">Last, Corrigan &amp; Naylor, 2014</bibRefCitation>
</taxonomicName>
, respectively. We caution however that this inference is based on a single mitochondrial marker. Inclusion of multiple nuclear markers could affect the presented inference.
</paragraph>
</subSubSection>
<caption id="1A936609FFDEFFAA5F97FAD55160FA61" httpUri="https://zenodo.org/record/256171/files/figure.png" pageId="8" pageNumber="596" targetBox="[175,1405,257,1316]" targetPageId="8">
<paragraph id="4E533681FFDEFFAA5F97FAD55160FA61" blockId="8.[151,1436,1370,1422]" pageId="8" pageNumber="596">
<emphasis id="7C98EA93FFDEFFAA5F97FAD55353FA9F" bold="true" box="[151,278,1370,1392]" pageId="8" pageNumber="596">FIGURE 6.</emphasis>
Lateral view of the unpaired fins of 
<taxonomicName id="89EC4D02FFDEFFAA5DA5FAD55124FA9F" box="[677,865,1370,1392]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFDEFFAA5DA5FAD55124FA9F" box="[677,865,1370,1392]" italics="true" pageId="8" pageNumber="596">Rhinobatos manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFDEFFAA5C6BFAD551FAFA9F" box="[875,959,1370,1392]" pageId="8" pageNumber="596" rank="species">
<emphasis id="7C98EA93FFDEFFAA5C6BFAD551FAFA9F" bold="true" box="[875,959,1370,1392]" pageId="8" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
, preserved adult male holotype, 731 mm TL (NTUM 11500): A. first dorsal; B. second dorsal; and C. caudal.
</paragraph>
</caption>
<subSubSection id="06F6650AFFDEFFAB5FC7FA3556E9FD8F" lastPageId="9" lastPageNumber="597" pageId="8" pageNumber="596" type="description">
<paragraph id="4E533681FFDEFFAA5FC7FA3550B2F847" blockId="8.[151,1437,1466,2032]" pageId="8" pageNumber="596">
<emphasis id="7C98EA93FFDEFFAA5FC7FA355328FA3C" bold="true" box="[199,365,1466,1491]" pageId="8" pageNumber="596">Comparisons.</emphasis>
The distinctive dorsal colour pattern of small, rusty brown spots and whitish spots with greyish edges distinguishes this species from all other species of 
<taxonomicName id="89EC4D02FFDEFFAA5C6FFA5051ABFA18" box="[879,1006,1503,1527]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="genus">
<emphasis id="7C98EA93FFDEFFAA5C6FFA5051ABFA18" box="[879,1006,1503,1527]" italics="true" pageId="8" pageNumber="596">Rhinobatos</emphasis>
</taxonomicName>
. In comparison, 
<taxonomicName id="89EC4D02FFDEFFAA5BC9FA6F5729FA18" box="[1225,1388,1503,1527]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="7C98EA93FFDEFFAA5BC9FA6F5729FA18" box="[1225,1388,1503,1527]" italics="true" pageId="8" pageNumber="596">R. borneensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5A84FA6F53B1F9F3" authority="Norman, 1922" authorityName="Norman" authorityYear="1922" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="holcorhynchus">
<emphasis id="7C98EA93FFDEFFAA5A84FA6F5379F9F4" italics="true" pageId="8" pageNumber="596">R. holcorhynchus</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5E49F98B53B1F9F3" author="Norman" box="[329,500,1540,1564]" pageId="8" pageNumber="596" refString="Norman, J. R. (1922) Three new fishes from Zululand and Natal, collected by Mr. H. W. Bell Marley; with additions to the fish fauna of Natal. Annals and Magazine of Natural History, Series 9, 9 (52), 318 - 322." type="journal article" year="1922">Norman, 1922</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5D05F98B566DF9F3" authority="Last, White &amp; Fahmi, 2006" authorityName="Last, White &amp; Fahmi" authorityYear="2006" box="[517,1064,1539,1564]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="jimbaranensis">
<emphasis id="7C98EA93FFDEFFAA5D05F98B508FF9F4" box="[517,714,1539,1563]" italics="true" pageId="8" pageNumber="596">R. jimbaranensis</emphasis>
Last, 
<bibRefCitation id="2A7D4B70FFDEFFAA5C1CF98C566DF9F3" author="White" box="[796,1064,1539,1564]" pageId="8" pageNumber="596" refString="Last, P. R., White, W. T. &amp; Fahmi (2006) Rhinobatos jimbaranensis and R. penggali, two new shovelnose rays (Batoidea: Rhinobatidae) from eastern Indonesia. Cybium, 30 (3), 261 - 271." type="journal article" year="2006">White &amp; Fahmi, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5B3AF98B5737F9F3" authority="Norman, 1926" authorityName="Norman" authorityYear="1926" box="[1082,1394,1539,1564]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7C98EA93FFDEFFAA5B3AF98B56FCF9F4" box="[1082,1209,1539,1563]" italics="true" pageId="8" pageNumber="596">R. lionotus</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5BC7F98B5737F9F3" author="Norman" box="[1223,1394,1539,1564]" pageId="8" pageNumber="596" refString="Norman, J. R. (1926) A synopsis of the rays of the family Rhinobatidae, with a revision of the genus Rhinobatus. Proceedings of the Zoological Society of London, 1926 (4), 941 - 982. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1926. tb 02228. x" type="journal article" year="1926">Norman, 1926</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5A84F98B50E7F9AF" authority="Last, Compagno &amp; Nakaya, 2004" authorityName="Last, Compagno &amp; Nakaya" authorityYear="2004" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="nudidorsalis">
<emphasis id="7C98EA93FFDEFFAA5A84F98B5366F9D0" italics="true" pageId="8" pageNumber="596">R. nudidorsalis</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5E29F9A750E7F9AF" author="Last" box="[297,674,1575,1600]" pageId="8" pageNumber="596" refString="Last, P. R., Compagno, L. J. V. &amp; Nakaya, K. (2004) Rhinobatos nudidorsalis, a new species of shovelnose ray (Batoidea: Rhinobatidae) from the Mascarene Ridge, central Indian Ocean. Ichthyological Research, 51 (2), 153 - 158. http: // dx. doi. org / 10.1007 / s 10228 - 004 - 0211 - 0" type="journal article" year="2004">Last, Compagno &amp; Nakaya, 2004</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5DACF9A75104F9D0" box="[684,833,1575,1599]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="sainsburyi">
<emphasis id="7C98EA93FFDEFFAA5DACF9A75104F9D0" box="[684,833,1575,1599]" italics="true" pageId="8" pageNumber="596">R. sainsburyi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="89EC4D02FFDEFFAA5C7CF9A75743F9AF" authority="Muller &amp; Henle, 1841" authorityName="Muller &amp; Henle" authorityYear="1841" box="[892,1286,1575,1600]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="schlegelii">
<emphasis id="7C98EA93FFDEFFAA5C7CF9A75643F9D0" box="[892,1030,1575,1599]" italics="true" pageId="8" pageNumber="596">R. schlegelii</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5B0DF9A85743F9AF" author="Muller" box="[1037,1286,1575,1600]" pageId="8" pageNumber="596" refString="Muller, J. &amp; Henle, F. G. J. (1841) Systematische Beschreibung der Plagiostomen. Veit und Comp., Berlin, pp. 103 - 200. http: // dx. doi. org / 10.5962 / bhl. title. 6906" type="book chapter" year="1841">Müller &amp; Henle, 1841</bibRefCitation>
</taxonomicName>
, all lack such prominent spotting (dorsal coloration uniform or with larger irregular blotches). 
<taxonomicName id="89EC4D02FFDEFFAA5B69F9C45379F967" authority="Norman, 1930" authorityName="Norman" authorityYear="1930" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="albomaculatus">
<emphasis id="7C98EA93FFDEFFAA5B69F9C457D9F98C" box="[1129,1436,1611,1635]" italics="true" pageId="8" pageNumber="596">Rhinobatos albomaculatus</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5F97F9FF5379F967" author="Norman" box="[151,316,1648,1672]" pageId="8" pageNumber="596" refString="Norman, J. R. (1930) A new ray of the genus Rhinobatus from the Gold Coast. Annals and Magazine of Natural History, Series 10, 6 (32), 226 - 228." type="journal article" year="1930">Norman, 1930</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5E47F9FF50D5F967" authority="Norman, 1926" authorityName="Norman" authorityYear="1926" box="[327,656,1647,1672]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="annandalei">
<emphasis id="7C98EA93FFDEFFAA5E47F9FF53A1F968" box="[327,484,1647,1671]" italics="true" pageId="8" pageNumber="596">R. annandalei</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5EEBF9FF50D5F967" author="Norman" box="[491,656,1647,1672]" pageId="8" pageNumber="596" refString="Norman, J. R. (1926) A synopsis of the rays of the family Rhinobatidae, with a revision of the genus Rhinobatus. Proceedings of the Zoological Society of London, 1926 (4), 941 - 982. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1926. tb 02228. x" type="journal article" year="1926">Norman, 1926</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5D9BF9FF561EF967" authority="Last, White &amp; Fahmi, 2006" authorityName="Last, White &amp; Fahmi" authorityYear="2006" box="[667,1115,1647,1672]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="penggali">
<emphasis id="7C98EA93FFDEFFAA5D9BF9FF5159F968" box="[667,796,1647,1671]" italics="true" pageId="8" pageNumber="596">R. penggali</emphasis>
Last, 
<bibRefCitation id="2A7D4B70FFDEFFAA5C62F9E0561EF967" author="White" box="[866,1115,1647,1672]" pageId="8" pageNumber="596" refString="Last, P. R., White, W. T. &amp; Fahmi (2006) Rhinobatos jimbaranensis and R. penggali, two new shovelnose rays (Batoidea: Rhinobatidae) from eastern Indonesia. Cybium, 30 (3), 261 - 271." type="journal article" year="2006">White &amp; Fahmi, 2006</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="89EC4D02FFDEFFAA5B96F9FF5315F943" authority="Compagno &amp; Randall, 1987" authorityName="Compagno &amp; Randall" authorityYear="1987" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="punctifer">
<emphasis id="7C98EA93FFDEFFAA5B96F9FF575FF967" box="[1174,1306,1648,1672]" italics="true" pageId="8" pageNumber="596">R. punctifer</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5A21F9FF5315F943" author="Compagno" pageId="8" pageNumber="596" refString="Compagno, L. J. V. &amp; Randall, J. E. (1987) Rhinobatos punctifer, a new species of guitarfish (Rhinobatiformes: Rhinobatidae) from the Red Sea, with notes on the Red Sea batoid fauna. Proceedings of the California Academy of Sciences, Series 4, 44 (14), 335 - 342." type="journal article" year="1987">Compagno &amp; Randall, 1987</bibRefCitation>
</taxonomicName>
have a colour pattern consisting primarily of white spots, but none of these have a combination of whitish and brownish spots. 
<taxonomicName id="89EC4D02FFDEFFAA5ED6F938518EF93F" authority="Richardson, 1846" authorityName="Richardson" authorityYear="1846" box="[470,971,1719,1744]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="hynnicephalus">
<emphasis id="7C98EA93FFDEFFAA5ED6F93850BAF920" box="[470,767,1719,1743]" italics="true" pageId="8" pageNumber="596">Rhinobatos hynnicephalus</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5C05F938518EF93F" author="Richardson" box="[773,971,1719,1744]" pageId="8" pageNumber="596" refString="Richardson, J. (1846) Report on the ichthyology of the seas of China and Japan. Report of the British Association for the Advancement of Science 15 th meeting, [1845], 187 - 320. http: // dx. doi. org / 10.5962 / bhl. title. 59530" type="journal article" year="1846">Richardson, 1846</bibRefCitation>
</taxonomicName>
has a variable pattern of small dark spots on the dorsal surface, either aggregated in small to large clusters or free. The dorsal surface of 
<taxonomicName id="89EC4D02FFDEFFAA5BC9F953528AF8F7" authority="Norman, 1931" authorityName="Norman" authorityYear="1931" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="irvinei">
<emphasis id="7C98EA93FFDEFFAA5BC9F9535774F91C" box="[1225,1329,1756,1779]" italics="true" pageId="8" pageNumber="596">R. irvinei</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5A39F953528AF8F7" author="Norman" pageId="8" pageNumber="596" refString="Norman, J. R. (1931) Four new fishes from the Gold Coast. Annals and Magazine of Natural History, Series 10, 7 (40), 352 - 359. http: // dx. doi. org / 10.1080 / 00222933108673322" type="journal article" year="1931">Norman, 1931</bibRefCitation>
</taxonomicName>
has a distinctive colour pattern of pale orange blotches with dark margins and black spots. 
<taxonomicName id="89EC4D02FFDEFFAA5A1CF970538EF8D3" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="rhinobatos">
<emphasis id="7C98EA93FFDEFFAA5A1CF9705355F8D4" italics="true" pageId="8" pageNumber="596">Rhinobatos rhinobatos</emphasis>
<bibRefCitation id="2A7D4B70FFDEFFAA5E1AF8AC538EF8D3" author="Linnaeus" box="[282,459,1827,1852]" pageId="8" pageNumber="596" refString="Linnaeus, C. (1758) Systema Naturae, Ed. X. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Vol. 1. Impensis Direct. Laurentii Salvii, Holmiae, 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
lacks spots and often has faint bluish grey stripes and blotches on the dorsal body. 
<taxonomicName id="89EC4D02FFDEFFAA5F97F8C85326F8B0" box="[151,355,1863,1887]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="whitei">
<emphasis id="7C98EA93FFDEFFAA5F97F8C85326F8B0" box="[151,355,1863,1887]" italics="true" pageId="8" pageNumber="596">Rhinobatos whitei</emphasis>
</taxonomicName>
has the most similar dorsal coloration to 
<taxonomicName id="89EC4D02FFDEFFAA5C3DF8C751E1F8B0" box="[829,932,1864,1887]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="manai" status="sp. nov.">
<emphasis id="7C98EA93FFDEFFAA5C3DF8C751E1F8B0" box="[829,932,1864,1887]" italics="true" pageId="8" pageNumber="596">R. manai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67AB57E8FFDEFFAA5CACF8C75642F88F" box="[940,1031,1864,1888]" pageId="8" pageNumber="596" rank="species">
<emphasis id="7C98EA93FFDEFFAA5CACF8C75642F88F" bold="true" box="[940,1031,1864,1888]" pageId="8" pageNumber="596">sp. nov.</emphasis>
</taxonomicNameLabel>
, consisting of poorly defined white spots and large diffuse dusky and orange blotches. However, 
<taxonomicName id="89EC4D02FFDEFFAA5C4BF8E351F4F86C" box="[843,945,1900,1923]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFDEFFAA5C4BF8E351F4F86C" box="[843,945,1900,1923]" italics="true" pageId="8" pageNumber="596">R. manai</emphasis>
</taxonomicName>
differs in having much better defined rusty brown spots and faint white spots with a greyish edge.
</paragraph>
<paragraph id="4E533681FFDEFFAA5FC7F83C57D9F81F" blockId="8.[151,1437,1466,2032]" pageId="8" pageNumber="596">
Based on its NADH2 sequence, 
<taxonomicName id="89EC4D02FFDEFFAA5D49F83C515FF824" box="[585,794,1971,1995]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFDEFFAA5D49F83C515FF824" box="[585,794,1971,1995]" italics="true" pageId="8" pageNumber="596">Rhinobatos manai</emphasis>
</taxonomicName>
belongs to a subgroup of 
<collectingRegion id="8C28F863FFDEFFAA5B5AF83B56EBF824" box="[1114,1198,1972,1995]" country="Ghana" name="Eastern" pageId="8" pageNumber="596">Eastern</collectingRegion>
Indian and 
<collectingRegion id="8C28F863FFDEFFAA5A41F83B57D9F824" box="[1345,1436,1972,1995]" country="Papua New Guinea" name="Western" pageId="8" pageNumber="596">Western</collectingRegion>
<collectingRegion id="8C28F863FFDEFFAA5F97F85852AFF81F" box="[151,234,2007,2032]" country="Papua New Guinea" name="Central" pageId="8" pageNumber="596">Central</collectingRegion>
Pacific species, i.e. 
<taxonomicName id="89EC4D02FFDEFFAA5ED5F8575035F800" box="[469,624,2007,2031]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="borneensis">
<emphasis id="7C98EA93FFDEFFAA5ED5F8575035F800" box="[469,624,2007,2031]" italics="true" pageId="8" pageNumber="596">R. borneensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5D80F8575105F800" box="[640,832,2007,2031]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="jimbaranensis">
<emphasis id="7C98EA93FFDEFFAA5D80F8575105F800" box="[640,832,2007,2031]" italics="true" pageId="8" pageNumber="596">R. jimbaranensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5C50F85751B4F800" box="[848,1009,2007,2032]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="lionotus">
<emphasis id="7C98EA93FFDEFFAA5C50F857512CF800" box="[848,873,2008,2031]" italics="true" pageId="8" pageNumber="596">R.</emphasis>
cf. 
<emphasis id="7C98EA93FFDEFFAA5C97F85851B4F800" box="[919,1009,2007,2031]" italics="true" pageId="8" pageNumber="596">lionotus</emphasis>
</taxonomicName>
, 
<taxonomicName id="89EC4D02FFDEFFAA5B02F85756D0F800" box="[1026,1173,2007,2031]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="sainsburyi">
<emphasis id="7C98EA93FFDEFFAA5B02F85756D0F800" box="[1026,1173,2007,2031]" italics="true" pageId="8" pageNumber="596">R. sainsburyi</emphasis>
</taxonomicName>
and 
<taxonomicName id="89EC4D02FFDEFFAA5BD0F8575770F800" box="[1232,1333,2007,2031]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="8" pageNumber="596" phylum="Chordata" rank="species" species="whitei">
<emphasis id="7C98EA93FFDEFFAA5BD0F8575770F800" box="[1232,1333,2007,2031]" italics="true" pageId="8" pageNumber="596">R. whitei</emphasis>
</taxonomicName>
(
<figureCitation id="D6D72A04FFDEFFAA5A47F85857C8F81F" box="[1351,1421,2007,2032]" captionStart="FIGURE 7" captionStartId="9.[151,250,1682,1704]" captionTargetBox="[226,1360,666,1655]" captionTargetId="figure@9.[226,1360,663,1659]" captionTargetPageId="9" captionText="FIGURE 7. Neighbour-joining tree based on p-distances derived from an alignment of mitochondrial NADH 2 sequences (1044 sites)." httpUri="https://zenodo.org/record/256172/files/figure.png" pageId="8" pageNumber="596">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="4E533681FFDFFFAB5F97FF1852BCFE43" blockId="9.[151,1436,151,608]" pageId="9" pageNumber="597">
This subgroup of species typically have weak colour patterns with variably developed rusty brown spots or blotches (sometimes plain), with only 
<taxonomicName id="89EC4D02FFDFFFAB5D45FF3250EEFF3B" box="[581,683,189,212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFDFFFAB5D45FF3250EEFF3B" box="[581,683,189,212]" italics="true" pageId="9" pageNumber="597">R. manai</emphasis>
</taxonomicName>
and 
<taxonomicName id="89EC4D02FFDFFFAB5DE2FF325100FF3B" box="[738,837,188,212]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="whitei">
<emphasis id="7C98EA93FFDFFFAB5DE2FF325100FF3B" box="[738,837,188,212]" italics="true" pageId="9" pageNumber="597">R. whitei</emphasis>
</taxonomicName>
also possessing whitish spots. Although closest to 
<taxonomicName id="89EC4D02FFDFFFAB5A84FF325349FF17" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="sainsburyi">
<emphasis id="7C98EA93FFDFFFAB5A84FF325349FF17" italics="true" pageId="9" pageNumber="597">R. sainsburyi</emphasis>
</taxonomicName>
based on the NADH2 sequence data, 
<taxonomicName id="89EC4D02FFDFFFAB5DC4FF6E516EFF18" box="[708,811,224,248]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFDFFFAB5DC4FF6E516EFF18" box="[708,811,224,248]" italics="true" pageId="9" pageNumber="597">R. manai</emphasis>
</taxonomicName>
is clearly distinct from this species in having: a much longer snout (preorbital length 17.6 
<emphasis id="7C98EA93FFDFFFAB5D2DFE8A5001FEF3" box="[557,580,261,284]" italics="true" pageId="9" pageNumber="597">vs</emphasis>
. 13.1–14.7% TL; preoral length 20.6 
<emphasis id="7C98EA93FFDFFFAB5CEFFE8A5643FEF3" box="[1007,1030,261,284]" italics="true" pageId="9" pageNumber="597">vs</emphasis>
. 15.5–17.9% TL; prenarial distance 16.4 
<emphasis id="7C98EA93FFDFFFAB5FD0FEA652AAFEAF" box="[208,239,297,320]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
12.0–13.6% TL), larger distance between anterior nasal flap insertions (2.9 
<emphasis id="7C98EA93FFDFFFAB5B53FEA65634FEAF" box="[1107,1137,297,320]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
2.0–2.3% TL), more total free vertebrae (188 
<emphasis id="7C98EA93FFDFFFAB5E72FEC253D5FE8B" box="[370,400,333,356]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
175–185), slightly more pectoral radials (66 
<emphasis id="7C98EA93FFDFFFAB5C89FEC251E2FE8B" box="[905,935,333,356]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
59–65), and in coloration (large rusty brown blotches and no white spots 
<emphasis id="7C98EA93FFDFFFAB5EEAFEFE504DFE67" box="[490,520,369,392]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
smaller, more regular and better defined rusty brown spots and whitish spots present).
</paragraph>
<paragraph id="4E533681FFDFFFAB5FC7FE3856E9FD8F" blockId="9.[151,1436,151,608]" pageId="9" pageNumber="597">
Morphologically, 
<taxonomicName id="89EC4D02FFDFFFAB5E93FE3653BEFE20" box="[403,507,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="manai">
<emphasis id="7C98EA93FFDFFFAB5E93FE3653BEFE20" box="[403,507,440,464]" italics="true" pageId="9" pageNumber="597">R. manai</emphasis>
</taxonomicName>
is closest to the recently described 
<taxonomicName id="89EC4D02FFDFFFAB5CA0FE365641FE3F" box="[928,1028,440,464]" class="Elasmobranchii" family="Rhinobatidae" genus="Rhinobatos" kingdom="Animalia" order="Rajiformes" pageId="9" pageNumber="597" phylum="Chordata" rank="species" species="whitei">
<emphasis id="7C98EA93FFDFFFAB5CA0FE365641FE3F" box="[928,1028,440,464]" italics="true" pageId="9" pageNumber="597">R. whitei</emphasis>
</taxonomicName>
from the 
<collectingCountry id="36FB7611FFDFFFAB5B7CFE3856BFFE3F" box="[1148,1274,439,464]" name="Philippines" pageId="9" pageNumber="597">Philippines</collectingCountry>
and seems to differ only in the following subtle morphological characteristics: distance across anterior nasal apertures (10.6 
<emphasis id="7C98EA93FFDFFFAB5A7DFE5257D9FE1B" box="[1405,1436,477,500]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
8.5–9.7% TL); slightly larger nostrils (nostril length 4.5 
<emphasis id="7C98EA93FFDFFFAB5C2DFD8E510EFDF7" box="[813,843,513,536]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
3.4–4.1% TL); dorsal fins slightly more separated (interdorsal distance 13.2 
<emphasis id="7C98EA93FFDFFFAB5EBBFDAA539FFDD3" box="[443,474,549,572]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
11.3–12.9% TL); and a slightly taller second dorsal fin (7.2 
<emphasis id="7C98EA93FFDFFFAB5B86FDAA56E0FDD3" box="[1158,1189,549,572]" italics="true" pageId="9" pageNumber="597">vs.</emphasis>
5.5–7.0% TL). These two species are most readily distinguished based on their coloration as mentioned previously.
</paragraph>
</subSubSection>
</treatment>
</document>