185 lines
22 KiB
XML
185 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. & Titarenko, Andrey Y." docDate="2014" docId="BB7D0359B82E5AED8A71A86FF311C80F" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. duripshensis Kurnakov 1972" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="29" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="29" updateTime="1668159733340" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
|
||
<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
|
||
<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
|
||
<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
|
||
<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
|
||
<mods:affiliation>Publicly Traded Company " Morpho Absoloni ", Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2014</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2014-12-12</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>463</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>21</mods:start>
|
||
<mods:end>56</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.463.8499</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-463-21</mods:identifier>
|
||
<mods:identifier type="ZooBank">71C1AE92B0CE4D26B1E5076E480180AD</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">3E5CFFC6FFC91628FFF6E355951DFFF8</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">578654</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="182234816" LSID="urn:lsid:plazi:treatment:BB7D0359B82E5AED8A71A86FF311C80F" httpUri="http://treatment.plazi.org/id/BB7D0359B82E5AED8A71A86FF311C80F" lastPageNumber="29" pageId="8" pageNumber="29">
|
||
<subSubSection pageId="8" pageNumber="29" type="nomenclature">
|
||
<paragraph pageId="8" pageNumber="29">
|
||
<taxonomicName LSID="BB7D0359-B82E-5AED-8A71-A86FF311C80F" authority="Kurnakov, 1972" authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis Kurnakov, 1972</taxonomicName>
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">Figs 1</figureCitation>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">, 2</figureCitation>
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="8" pageNumber="29">, 7</figureCitation>
|
||
<figureCitation captionStart="Figures 22–27" captionStartId="F10" captionText="Figures 22 - 27. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus duripshensis and Carabus (Archiplectes) besleticus napraensis. 22 - 24 Carabus (Archiplectes) besleticus duripshensis 25 - 27 Carabus (Archiplectes) besleticus napraensis." pageId="8" pageNumber="29">, 22-24</figureCitation>
|
||
<figureCitation captionStart="Figures 58–61" captionStartId="F15" captionText="Figures 58 - 61. Dorsal habitus, Carabus (Archiplectes) besleticus duripshensis. 58 - 59 Abkhazia, S slope of Bsybian Mt. Range, valley of River Khipsta 60 - 61 Abkhazia, Bsybian Mt. Range, Duriph env. (after Retezar 2008)." pageId="8" pageNumber="29">, 58-61</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="reference_group">
|
||
<paragraph pageId="8" pageNumber="29">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Neoplectes) satyrus subsp. duripshensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Neoplectes" subSpecies="duripshensis">Carabus (Neoplectes) satyrus duripshensis</taxonomicName>
|
||
Kurnakov, 1972: 114 (
|
||
<normalizedToken originalValue="“Chaîne">"Chaine</normalizedToken>
|
||
Bzybienne
|
||
<normalizedToken originalValue="près">pres</normalizedToken>
|
||
du village de Douripch")
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. duripshensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) satyrus duripshensis</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Gottwald, J" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="23" pageNumber="44" pagination="278 - 314" refId="B5" refString="Gottwald, J, 1985. Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae). Acta Entomologica Bohemoslovaca 824: 278 - 314" title="Revision der Untergattung Archiplectes Gottwald (Plectes auct.) der Gattung Carabus (Coleoptera, Carabidae)." volume="824" year="1985">Gottwald 1985</bibRefCitation>
|
||
: 310;
|
||
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Catalogue of Palaearctic Coleoptera" pageId="23" pageNumber="44" pagination="118 - 201" refId="B2" refString="Bousquet, Y, Brezina, B, Davies, A, Farkac, J, Smetana, A, 2003. Tribe Carabini Latreille, 1802. Catalogue of Palaearctic Coleoptera Vol. 1: 118 - 201" title="Tribe Carabini Latreille, 1802." volume="Vol. 1" year="2003">Bousquet et al. 2003</bibRefCitation>
|
||
: 132;
|
||
<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
|
||
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
|
||
2008
|
||
</bibRefCitation>
|
||
: 40.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) satyrus subsp. duripshensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Tribax" subSpecies="duripshensis">Carabus (Tribax) satyrus duripshensis</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Deuve, T" journalOrPublisher="Pensoft Publishers, Sofia-Moscow" pageId="23" pageNumber="44" refId="B3" refString="Deuve, T, 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers, Sofia-Moscow" title="Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae)." year="2004">Deuve 2004</bibRefCitation>
|
||
: 275.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="comparative material examined">
|
||
<paragraph pageId="8" pageNumber="29">Comparative material examined.</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
69 specimens were examined (4 specimens measured, 2 male genitalia preparations studied): 15 male, 18 female, Abkhazia, Gudauta Distr. near Village Khuap, V-VII.1993, leg. A. Zamotajlov, A. Miroshnikov (cZAM, cFDD); 1 female, Abkhazia, Bzybian Mt. Range, vicinities of Village Duripsh, 04.VI.1988, leg. V. Karmanian (cSIA); 1 male, Abkhazia, Gudauta Distr. near Village Khuap, 25.V-05.VIII.1986 (cSIA); 1 male, Abkhazia, Gudauta Distr., Khipsta Gorge, 580 m (cPNYu); 1 male, Abkhazia, Gudauta Distr., valley of River Khipsta, vicinities of Village Tvanaarkhu, karst crater with cave, beech, blackberry site, 425 m,
|
||
<geoCoordinate degrees="43" direction="north" minutes="14" orientation="latitude" precision="925" value="43.233334">N43°14'</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="040" direction="east" minutes="39" orientation="longitude" precision="15" seconds="9" value="40.6525">E040°39'9"</geoCoordinate>
|
||
, 08.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova, S. Pavlyuchuk (cSIA); 1 male, 1 female, Abkhazia, Gudauta Distr., valley of River Khipsta, vicinities of Village Tvanaarkhu, valley of small brook, 380-425 m, hornbeam, beech, box forest site,
|
||
<geoCoordinate degrees="43" direction="north" minutes="13" orientation="latitude" precision="925" value="43.216667">N43°13'</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="040" direction="east" minutes="39" orientation="longitude" precision="925" value="40.65">E040°39'</geoCoordinate>
|
||
, 08.V-05.VII.2013, leg. I. Solodovnikov, S. Solodovnikova, S. Pavlyuchuk (cSIA, cPSM); 13 males, 17 females, Abkhazia, Gudauta Distr. near Village Khuap, 700 m, 11.IV-6.VI.2014, leg. D. Fominykh and A. Bondarenko (cFDD, cBAS).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="description">
|
||
<paragraph pageId="8" pageNumber="29">Description.</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
The medium-sized form, males 32.5-32.5 (32.5) mm, females 32.5-32.6 (32.6) mm long (according to
|
||
<bibRefCitation author="Kurnakov, VN" journalOrPublisher="Nouvelle Revue d'entomologie II" pageId="23" pageNumber="44" pagination="111 - 115" refId="B7" refString="Kurnakov, VN, 1972. Contribution a la faune des Carabiques du Caucase. VI. Nouveaux Carabes du Caucase. Nouvelle Revue d'entomologie II 2: 111 - 115" title="Contribution a la faune des Carabiques du Caucase. VI. Nouveaux Carabes du Caucase." volume="2" year="1972">Kurnakov 1972</bibRefCitation>
|
||
- 28-33 mm long). Underside black, dorsum normally with bright metallic lustre, violet, black-violet, black, rather seldom green, bronze, crimson, females matte. Body slender.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
Head not inflated. Pronotum variable in shape, mainly transverse. PW/PL = 1.30-1.40 (1.35) in males and 1.27-1.28 (1.28) in females, with approximately same PW/PB as in the nominotypical subspecies: 1.27-1.32 (1.30) in males and 1.16-1.26 (1.21) in females, hind angles strongly protruding backwards and somewhat sidewards, pointed apically. Median groove distinct. Disk transversally rugose, rugosity gradually strengthening towards median groove. Elytra oblong-ovate, more elongate and somewhat convex in males, with quite inconspicuous depression in the middle in females. EL/EW = 1.77-1.80 (1.79) in males and 1.81-1.86 (1.83) in females. Elytral sculpture nearly identical in males and females, forming precise series of short links. The main morphometric measurements are presented in Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="8" pageNumber="29" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
The endophallus differs from that of
|
||
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
|
||
in the shape of its dorsal appendix, which is more elongate and possesses a more extended form, characteristic for populations dwelling in an interfluve of r. Kealasur and Bzyb.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="differential diagnosis">
|
||
<paragraph pageId="8" pageNumber="29">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
Habitually resembles
|
||
<taxonomicName authorityName="Belousov & Zamotajlov" authorityYear="1993" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. napraensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="napraensis">Carabus (Archiplectes) besleticus napraensis</taxonomicName>
|
||
from which it differs in the more elongate and less ovate elytra, cordate pronotum with more elevated and sharper lateral borders, and stronger protruding hind angles. Elytral sculpture is more coarse, often with faintly developed tertiary interspaces, especially in females, body size on average larger. It also differs in habitat:
|
||
<taxonomicName authorityName="Kurnakov" authorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. duripshensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="duripshensis">Carabus (Archiplectes) besleticus duripshensis</taxonomicName>
|
||
populates forest belts, rising up to the subalpine belt, while
|
||
<taxonomicName authorityName="Belousov & Zamotajlov" authorityYear="1993" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. napraensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="napraensis">Carabus (Archiplectes) besleticus napraensis</taxonomicName>
|
||
dwells exclusively at alpestrine and alpine zones. Morphometric characters are illustrated in Figures
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">1</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="8" pageNumber="29">2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="distribution">
|
||
<paragraph pageId="8" pageNumber="29">Distribution.</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">Populates southern slopes of Bzybian karstic figau at altitudes ranging from 380 to 1800 m, to the west from the River Khipsta.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="8" pageNumber="29" type="habitat">
|
||
<paragraph pageId="8" pageNumber="29">Habitat.</paragraph>
|
||
<paragraph pageId="8" pageNumber="29">
|
||
Prefers beech and beech-chestnut forest sites, rich in ground litter on karstic landforms. Activity of imago proceeds from April until July.
|
||
<taxonomicName authority="polychrous" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) polychrous subsp. polychrous" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="polychrous" subGenus="Archiplectes" subSpecies="polychrous">Carabus (Archiplectes) polychrous polychrous</taxonomicName>
|
||
Rost, 1892,
|
||
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) apollo subsp. tenebricosus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="apollo" subGenus="Archiplectes" subSpecies="tenebricosus">Carabus (Archiplectes) apollo tenebricosus</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Mandl" authorityYear="1955" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Microplectes) argonautarum subsp. reischitzi" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="argonautarum" subGenus="Microplectes" subSpecies="reischitzi">Carabus (Microplectes) argonautarum reischitzi</taxonomicName>
|
||
,
|
||
<taxonomicName authority="apschuanus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) apschuanus subsp. apschuanus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="apschuanus" subGenus="Tribax" subSpecies="apschuanus">Carabus (Tribax) apschuanus apschuanus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
|
||
,
|
||
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
|
||
Ganglbauer, 1886 (natio
|
||
<taxonomicName class="Insecta" family="Carabidae" form="abasinus" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="form">
|
||
<emphasis italics="true" pageId="8" pageNumber="29">abasinus</emphasis>
|
||
</taxonomicName>
|
||
Rost, 1893),
|
||
<taxonomicName authorityName="Motschulsky" authorityYear="1840" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Megodontus) septemcarinatus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="septemcarinatus" subGenus="Megodontus">Carabus (Megodontus) septemcarinatus</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Sphodristocarabus) armeniacus subsp. dvorschaki" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="armeniacus" subGenus="Sphodristocarabus" subSpecies="dvorschaki">Carabus (Sphodristocarabus) armeniacus dvorschaki</taxonomicName>
|
||
Mandl, 1975, and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procerus) caucasicus subsp. colchicus" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="caucasicus" subGenus="Procerus" subSpecies="colchicus">Carabus (Procerus) caucasicus colchicus</taxonomicName>
|
||
occur together with this subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |