treatments-xml/data/A2/65/47/A2654718677FFFC9723BFCFD8EC8FE77.xml
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<document id="BEA8F45BC64AD0C4744F1305970E0CE6" ID-CLB-Dataset="27046" ID-DOI="10.5281/zenodo.2645967" ID-GBIF-Dataset="df49d6da-ad36-4b37-a437-c69a6edf61d7" ID-ISSN="1175-5326" ID-Zenodo-Dep="2645967" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555587711137" checkinUser="plazi" docAuthor="Anker, Arthur, Poddoubtchenko, Denis &amp; Wehrtmann, Ingo S." docDate="2006" docId="A2654718677FFFC9723BFCFD8EC8FE77" docLanguage="en" docName="zt01183p041.pdf" docOrigin="Zootaxa 1183" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Leslibetaeus Anker &amp; Poddoubtchenko &amp; Wehrtmann 2006, n. gen." docType="treatment" docVersion="11" lastPageNumber="41" masterDocId="5E5C3F60677DFFC77333FFE38D5CFC44" masterDocTitle="Leslibetaeus coibita, n. gen., n. sp., a new alpheid shrimp from the Pacific coast of Panama (Crustacea: Decapoda)" masterLastPageNumber="41" masterPageNumber="27" pageNumber="29" updateTime="1698722741082" updateUser="ExternalLinkService">
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<mods:title id="AD8E40B7FECBDF39B78A8A8CAFACFA18">Leslibetaeus coibita, n. gen., n. sp., a new alpheid shrimp from the Pacific coast of Panama (Crustacea: Decapoda)</mods:title>
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<mods:namePart id="CF056FD8EA17722A5DE83B0240DEAEE5">Anker, Arthur</mods:namePart>
</mods:name>
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<mods:namePart id="708610AF260F39F0FA5465D43AD50E8D">Poddoubtchenko, Denis</mods:namePart>
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<mods:namePart id="993696060A6AF6C0C62EE58CC2DA090B">Wehrtmann, Ingo S.</mods:namePart>
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<mods:part id="7769B9CE590EAEE966A9086B031A2B9E">
<mods:date id="D02D8C37CA08112CA348F5527B6DE629">2006</mods:date>
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<mods:number id="62B2012483A78F035191A0ACE717A23B">2006-04-24</mods:number>
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<treatment id="A2654718677FFFC9723BFCFD8EC8FE77" ID-GBIF-Taxon="156789108" LSID="urn:lsid:plazi:treatment:A2654718677FFFC9723BFCFD8EC8FE77" httpUri="http://treatment.plazi.org/id/A2654718677FFFC9723BFCFD8EC8FE77" lastPageId="14" lastPageNumber="41" pageId="2" pageNumber="29">
<subSubSection id="62D6A585677FFFC5723BFCFD885FFA8F" pageId="2" pageNumber="29" type="nomenclature">
<paragraph id="2A73F60E677FFFC5723BFCFD8CA2FF7C" blockId="2.[264,510,798,824]" box="[264,510,798,824]" pageId="2" pageNumber="29">
<heading id="713B4162677FFFC5723BFCFD8CA2FF7C" bold="true" box="[264,510,798,824]" fontSize="11" level="1" pageId="2" pageNumber="29" reason="1">
<emphasis id="18B82A1C677FFFC5723BFCFD8CA2FF7C" bold="true" box="[264,510,798,824]" pageId="2" pageNumber="29">
<taxonomicName id="EDCC8D8D677FFFC5723BFCFD8CC1FF7C" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[264,413,798,824]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C677FFFC5723BFCFD8CC1FF7C" bold="true" box="[264,413,798,824]" italics="true" pageId="2" pageNumber="29">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B9767677FFFC5729FFCFD8CA2FF7C" box="[428,510,798,824]" pageId="2" pageNumber="29" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2A73F60E677FFFC5723BFC8A885FFA8F" blockId="2.[264,1323,873,1899]" pageId="2" pageNumber="29">Body moderately stout. Frontal margin rounded, without rostrum and orbital teeth. Pterygostomial angle rounded, not protruding. Eyes partly concealed in dorsal and lateral view, visible in frontal view; cornea well developed. Antennular peduncle moderately slender; first segment without ventromesial tooth; lateral antennular flagellum with poorly developed secondary ramus. Antenna with basicerite robust; carpocerite short, reaching far beyond scaphocerite. Mouthparts typical for family; mandible with bisegmented palp and well developed molar and incisor processes. Third maxilliped pediform, lateral plate feebly developed, rounded; tip of ultimate segment with stout spines. Chelipeds enlarged, equal in size, symmetrical in shape, robust, carried extended; ischium with spines on dorsodistal margin; merus ventrally flattened; carpus very stout, cup­shaped, with strong distal lobes, with two comb­like rows of setae mesially; chela stout, inflated; palm smooth, linea impressa and adhesive discs absent; cutting edges of fingers with teeth, without snapping mechanism. Second pereiopod with five­segmented carpus, second segment longest. Third pereiopod with ischium lacking spine; merus unarmed; carpus with distoventral spine; propodus with slender spines on ventral margin; dactylus simple. Sixth abdominal somite without articulated plate at posterolateral angle. Second pleopod with appendix interna and appendix masculina. Uropod with sympodite bearing subacute tooth; diaeresis with broad subacute tooth in mid­portion. Telson with two pairs of dorsal spines and two pairs of posterolateral spines, lateral much shorter and directed upwards; posterior margin reduced; anal tubercles absent. Gill/exopod formula: 5 pleurobranchs (P15); 1 arthrobranch (Mxp3); 0 podobranch; 2 lobe­shaped epipods (Mxp12); 5 strap­like epipods or mastigobranchs (Mxp3, P14); 5 setobranchs (P15); 3 exopods (Mxp13).</paragraph>
</subSubSection>
<subSubSection id="62D6A585677FFFC5720BF93A8851FAB7" box="[312,1293,1753,1779]" pageId="2" pageNumber="29" type="materials_examined">
<paragraph id="2A73F60E677FFFC5720BF93A8851FAB7" blockId="2.[264,1323,873,1899]" box="[312,1293,1753,1779]" pageId="2" pageNumber="29">
<emphasis id="18B82A1C677FFFC5720BF93A8C8CFAB7" bold="true" box="[312,464,1753,1779]" pageId="2" pageNumber="29">
<typeStatus id="F57748AC677FFFC5720BF93A8C29FAB7" box="[312,373,1753,1779]" pageId="2" pageNumber="29">Type</typeStatus>
species
</emphasis>
. —
<taxonomicName id="EDCC8D8D677FFFC57135F93A8FADFAB6" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[518,753,1753,1778]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="29" phylum="Arthropoda" rank="species" species="coibita" status="sp. nov.">
<emphasis id="18B82A1C677FFFC57135F93A8FADFAB6" box="[518,753,1753,1778]" italics="true" pageId="2" pageNumber="29">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B9767677FFFC57033F93A8E1CFAB7" box="[768,832,1753,1779]" pageId="2" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
, by monotypy and present designation.
</paragraph>
</subSubSection>
<subSubSection id="62D6A585677FFFC4720BF8E28870FD72" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29" type="etymology">
<paragraph id="2A73F60E677FFFC4720BF8E28870FD72" blockId="2.[264,1323,873,1899]" lastBlockId="3.[264,1324,284,350]" lastPageId="3" lastPageNumber="30" pageId="2" pageNumber="29">
<emphasis id="18B82A1C677FFFC5720BF8E28CE1FB5F" bold="true" box="[312,445,1793,1819]" pageId="2" pageNumber="29">Etymology</emphasis>
. — We are more than pleased to dedicate this genus to our dear colleague and one of the collectors of this shrimp, Leslie Harris. The generic name is a combination of the collectors first name, Leslie, and
<taxonomicName id="EDCC8D8D677FFFC571DEF8B28E1BFB2E" authorityName="Dana" authorityYear="1852" box="[749,839,1873,1898]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C677FFFC571DEF8B28E1BFB2E" box="[749,839,1873,1898]" italics="true" pageId="2" pageNumber="29">betaeus</emphasis>
</taxonomicName>
, referring to the superficial resemblance of the new genus to the alpheid genus
<taxonomicName id="EDCC8D8D677EFFC471F4FEFF8EEFFD72" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[711,947,284,310]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C677EFFC471F4FEFF8E78FD71" box="[711,804,284,309]" italics="true" pageId="3" pageNumber="30">Betaeus</emphasis>
Dana, 1852
</taxonomicName>
(see below). Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="62D6A585677EFFC4720BFEA08F1BFD91" pageId="3" pageNumber="30" type="discussion">
<paragraph id="2A73F60E677EFFC4720BFEA08F00FD1A" blockId="3.[264,1324,284,350]" box="[312,604,323,350]" pageId="3" pageNumber="30">
<emphasis id="18B82A1C677EFFC4720BFEA08CF6FD19" bold="true" box="[312,426,323,349]" pageId="3" pageNumber="30">Remarks</emphasis>
. — See below.
</paragraph>
<paragraph id="2A73F60E677EFFC4723BFE5F8F1BFD91" blockId="3.[264,583,443,470]" box="[264,583,443,470]" pageId="3" pageNumber="30">
<emphasis id="18B82A1C677EFFC4723BFE5F8F1BFD91" bold="true" box="[264,583,443,470]" pageId="3" pageNumber="30">
<taxonomicName id="EDCC8D8D677EFFC4723BFE5F8CA9FD92" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[264,501,444,470]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="30" phylum="Arthropoda" rank="species" species="coibita" status="sp. nov.">
<emphasis id="18B82A1C677EFFC4723BFE5F8CA9FD92" bold="true" box="[264,501,444,470]" italics="true" pageId="3" pageNumber="30">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B9767677EFFC47137FE588F1BFD91" box="[516,583,443,469]" pageId="3" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="62D6A585677EFFC4723BFDE58E1BFF55" pageId="3" pageNumber="30" type="materials_examined">
<paragraph id="2A73F60E677EFFC4723BFDE58E1BFF55" blockId="3.[264,1325,518,1905]" pageId="3" pageNumber="30">
<emphasis id="18B82A1C677EFFC4723BFDE58CA4FE64" bold="true" box="[264,504,518,544]" pageId="3" pageNumber="30">Material examined</emphasis>
. —
<materialsCitation id="9AA4FC53677EFFC47101FDE4897BFE85" ID-GBIF-Occurrence="2417032650" collectingDate="2005-03-12" collectionCode="LACM, STRI" collectorName="Pacific &amp; L. Harris &amp; I. S. Wehrtmann" country="Iceland" county="Coiba National Park" latitude="81.7" location="Coibita" municipality="Smithsonian Tropical Research Instititute" pageId="3" pageNumber="30" specimenCode="CR 2005, 013.2" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="F57748AC677EFFC47101FDE48FFFFE65" box="[562,675,519,545]" pageId="3" pageNumber="30" type="holotype">Holotype</typeStatus>
:
<collectionCode id="4CDD6ECB677EFFC47182FDE48E5AFE65" box="[689,774,519,545]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">LACM</collectionCode>
<specimenCode id="7A6A5E75677EFFC47022FDE48E22FE65" box="[785,894,519,545]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">CR 2005</specimenCode>
­
<specimenCode id="7A6A5E75677EFFC470B5FDE48E97FE65" box="[902,971,519,545]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">013.2</specimenCode>
,
<specimenCount id="3CCA3D87677EFFC470E9FDE4891FFE65" box="[986,1091,519,545]" pageId="3" pageNumber="30" type="female">1 female</specimenCount>
(CL 3.4 mm), Isla
<location id="2F13A0D5677EFFC4723BFDCC8C3DFE0D" LSID="urn:lsid:plazi:treatment:A2654718677FFFC9723BFCFD8EC8FE77:2F13A0D5677EFFC4723BFDCC8C3DFE0D" box="[264,353,559,585]" country="Iceland" county="Coiba National Park" latitude="81.7" municipality="Smithsonian Tropical Research Instititute" name="Coibita" pageId="3" pageNumber="30">Coibita</location>
, on side facing
<collectingCountry id="52DBB69E677EFFC4712FFDCC8F1BFE0D" box="[540,583,559,585]" name="Iceland" pageId="3" pageNumber="30">Isla</collectingCountry>
<location id="2F13A0D5677EFFC4717DFDCC8FCBFE0D" LSID="urn:lsid:plazi:treatment:A2654718677FFFC9723BFCFD8EC8FE77:2F13A0D5677EFFC4717DFDCC8FCBFE0D" box="[590,663,559,585]" country="Iceland" county="Coiba National Park" latitude="81.7" municipality="Smithsonian Tropical Research Instititute" name="Coiba" pageId="3" pageNumber="30">Coiba</location>
, north side of
<collectingMunicipality id="CA176C74677EFFC47071FDCC8876FE0D" box="[834,1322,559,585]" pageId="3" pageNumber="30">Smithsonian Tropical Research Instititute</collectingMunicipality>
(
<collectionCode id="4CDD6ECB677EFFC47223FDB48C0FFE35" box="[272,339,599,625]" country="Panama" lsid="urn:lsid:biocol.org:col:14898" name="Smithsonian Tropical Research Institute" pageId="3" pageNumber="30" type="Herbarium">STRI</collectionCode>
) beach house, first small island connected by causeway,
<geoCoordinate id="4FF890C9677EFFC470D8FDB4887BFE35" box="[1003,1319,599,625]" pageId="3" pageNumber="30">7°3880” N 81°42159” W</geoCoordinate>
, Isla
<collectingCounty id="C3128E82677EFFC47209FD9C8F70FEDD" box="[314,556,639,665]" pageId="3" pageNumber="30">Coiba National Park</collectingCounty>
,
<collectorName id="873993D8677EFFC4710BFD9C8FD5FEDD" box="[568,649,639,665]" pageId="3" pageNumber="30">Pacific</collectorName>
coast of
<collectingCountry id="52DBB69E677EFFC471C7FD9C8E08FEDD" box="[756,852,639,665]" name="Panama" pageId="3" pageNumber="30">Panama</collectingCountry>
, extracted by hand/forceps from crevice of a shale rock, coll.
<collectorName id="873993D8677EFFC47134FD448F2FFE85" box="[519,627,679,705]" pageId="3" pageNumber="30">L. Harris</collectorName>
&amp;
<collectorName id="873993D8677EFFC471AEFD448E06FE85" box="[669,858,679,705]" pageId="3" pageNumber="30">I.S. Wehrtmann</collectorName>
,
<date id="5E72D0CE677EFFC47054FD44897EFE85" box="[871,1058,679,705]" pageId="3" pageNumber="30" value="2005-03-12">
<collectingDate id="4E362926677EFFC47054FD44897EFE85" box="[871,1058,679,705]" pageId="3" pageNumber="30" value="2005-03-12">12 March 2005</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="9AA4FC53677EFFC47703FD448E1BFF55" ID-GBIF-Occurrence="2417032651" collectingDate="2005-03-12" collectionCode="LACM" collectorName="Pacific &amp; L. Harris &amp; I. S. Wehrtmann" country="Iceland" county="Coiba National Park" latitude="81.7" location="Coibita" municipality="Smithsonian Tropical Research Instititute" pageId="3" pageNumber="30" specimenCode="CR 2005, 013.1" specimenCount="1" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="F57748AC677EFFC47703FD4489C7FE85" box="[1072,1179,679,705]" pageId="3" pageNumber="30" type="paratype">Paratype</typeStatus>
:
<collectionCode id="4CDD6ECB677EFFC47794FD4489A0FE85" box="[1191,1276,679,705]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">LACM</collectionCode>
<specimenCode id="7A6A5E75677EFFC47635FD448C1AFEAD" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">CR 2005</specimenCode>
­
<specimenCode id="7A6A5E75677EFFC4727DFD2C8CCFFEAD" box="[334,403,719,745]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="3" pageNumber="30" type="Museum">013.1</specimenCode>
.
<specimenCount id="3CCA3D87677EFFC472ADFD2C8CB2FEAD" box="[414,494,719,745]" pageId="3" pageNumber="30" type="male">1 male</specimenCount>
(CL 3.2 mm), same data as for holotype [specimen poorly preserved, broken in two parts, most appendages dissected].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="62D6A585677EFFCD720BFCFD8FCDFDBA" lastPageId="10" lastPageNumber="37" pageId="3" pageNumber="30" type="description">
<paragraph id="2A73F60E677EFFC4720BFCFD8F71F83D" blockId="3.[264,1325,518,1905]" pageId="3" pageNumber="30">
<emphasis id="18B82A1C677EFFC4720BFCFD8C95FF7C" bold="true" box="[312,457,798,824]" pageId="3" pageNumber="30">Description</emphasis>
. — Body moderately elongated (Fig, 1), not particularly compressed. Carapace smooth, with very shallow, poorly defined grooves on flanks, without sutures, glabrous except for minute, scarce, erect setae; anterior portion somewhat inflated in lateral view (
<figureCitation id="B2F7EA8B677EFFC472F2FC748F49FFF5" box="[449,533,919,945]" captionStart="FIGURE 1" captionStartId="4.[264,367,1318,1342]" captionTargetBox="[304,1279,684,1289]" captionTargetId="figure@4.[264,1323,667,1322]" captionTargetPageId="4" captionText="FIGURE 1. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2): habitus. Scale 1 mm." httpUri="https://zenodo.org/record/2645969/files/figure.png" pageId="3" pageNumber="30">Fig. 1</figureCitation>
); pterygostomial angle rounded, not protruding (
<figureCitation id="B2F7EA8B677EFFC47787FC748840FFF5" box="[1204,1308,919,945]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2b</figureCitation>
); branchiostegial margin with a few setae; cardiac notch well developed (
<figureCitation id="B2F7EA8B677EFFC47758FC5C899FFF9D" box="[1131,1219,959,985]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2c</figureCitation>
); frontal margin of carapace broadly rounded (
<figureCitation id="B2F7EA8B677EFFC4718EFC048E4FF845" box="[701,787,999,1025]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2a</figureCitation>
), rostrum and orbital teeth absent. Eyes partly concealed, anterior portion visible in dorsal and lateral view (
<figureCitation id="B2F7EA8B677EFFC470EBFBEC8916F86D" box="[984,1098,1039,1065]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2a, b</figureCitation>
), without tubercles or spines, cornea moderately developed. Ocellar beak not conspicuous. Epistomial sclerite without acute projection.
</paragraph>
<paragraph id="2A73F60E677EFFC4720BFB648E7BFA1D" blockId="3.[264,1325,518,1905]" pageId="3" pageNumber="30">
Antennular peduncle not particularly stout, moderately elongated, slender (
<figureCitation id="B2F7EA8B677EFFC477F0FB648841F8E5" box="[1219,1309,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2a</figureCitation>
); first segment with stylocerite somewhat apressed, reaching distal margin of first segment, distally subacute (
<figureCitation id="B2F7EA8B677EFFC472D9FB348F36F8B5" box="[490,618,1239,1265]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2a, d</figureCitation>
), ventromesial carina unarmed (
<figureCitation id="B2F7EA8B677EFFC47733FB3489DCF8B5" box="[1024,1152,1239,1265]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2d, e</figureCitation>
), dorsomesial surface proximal to eye with onetwo small spines (
<figureCitation id="B2F7EA8B677EFFC47047FB1C8EBBF95D" box="[884,999,1279,1305]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2a, e</figureCitation>
); second segment subequal to visible portion of first segment, much longer than third segment; lateral flagellum with secondary ramus reduced to a small knob with one (male) or two (female) tufts of aesthetascs, latter longer and more numerous developed in female (
<figureCitation id="B2F7EA8B677EFFC4772FFA9489F7F9D5" box="[1052,1195,1399,1425]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Fig. 2b, d, e</figureCitation>
), proximal fused portion composed of five­six segments. Antenna with basicerite relatively stout, armed with strong ventrolateral tooth (
<figureCitation id="B2F7EA8B677EFFC471EDFA248E1FF9A5" box="[734,835,1479,1505]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Figs. 2b</figureCitation>
,
<figureCitation id="B2F7EA8B677EFFC47060FA248E28F9A5" box="[851,884,1479,1505]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">3b</figureCitation>
); scaphocerite subrectangular, short, reaching only slightly beyond half length of carpocerite, anterior margin of blade convex, not extending beyond distolateral tooth (
<figureCitation id="B2F7EA8B677EFFC471D8F9F48E11FA75" box="[747,845,1559,1585]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="3" pageNumber="30">Figs. 2a</figureCitation>
,
<figureCitation id="B2F7EA8B677EFFC47068F9F48E27FA75" box="[859,891,1559,1585]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">3b</figureCitation>
); carpocerite robust, elongated (
<figureCitation id="B2F7EA8B677EFFC477C8F9F48C7BFA1D" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">Fig. 3a</figureCitation>
); tubercle of antennal gland not protruding.
</paragraph>
<paragraph id="2A73F60E677EFFC3720BF9848F3CFE32" blockId="3.[264,1325,518,1905]" lastBlockId="4.[264,1324,284,630]" lastPageId="4" lastPageNumber="31" pageId="3" pageNumber="30">
Mouthparts typical for family. Mandible with bisegmented palp; molar process slightly broadened distally (
<figureCitation id="B2F7EA8B677EFFC4716AF96C8FEEFAED" box="[601,690,1679,1705]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">Fig. 3c</figureCitation>
); incisor process distally with five­six subtriangular teeth (
<figureCitation id="B2F7EA8B677EFFC47262F9548C98FA95" box="[337,452,1719,1745]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">Fig. 3d, e</figureCitation>
). Maxillule with palp bilobed, dorsal lobe with one seta, ventral lobe with several setae (
<figureCitation id="B2F7EA8B677EFFC4729CF93C8F58FABD" box="[431,516,1759,1785]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">Fig. 3f</figureCitation>
). Maxilla with scaphognathite narrow (
<figureCitation id="B2F7EA8B677EFFC470EBF93C896CFABD" box="[984,1072,1759,1785]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">
Fig.
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); endopod small, not segmented; dorsal endite not deeply notched (
<figureCitation id="B2F7EA8B677EFFC47013F8E48E2BFB65" box="[800,887,1799,1825]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">
Fig.
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</figureCitation>
); ventral endite with elongated setae. First maxilliped with caridean lobe on exopod narrow, with a few setae (
<figureCitation id="B2F7EA8B677EFFC4776BF8CC89ECFB0D" box="[1112,1200,1839,1865]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="3" pageNumber="30">Fig. 3h</figureCitation>
); endopod not segmented, with elongated setae distally; epipod relatively small, oval to subtriangular. Second maxilliped with small oval­rounded epipod (
<figureCitation id="B2F7EA8B6779FFC370B9FEFF8E82FD72" box="[906,990,284,310]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">
Fig.
<date id="5E72D0CE6779FFC370F6FEFF8E82FD72" box="[965,990,284,310]" pageId="4" pageNumber="31">3i</date>
</figureCitation>
); endopod without specific features; exopod somewhat broadened proximally. Third maxilliped pediform, elongated but stout (
<figureCitation id="B2F7EA8B6779FFC372B4FE8F8C87FDC2" box="[391,475,364,390]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">Fig. 3j</figureCitation>
); coxa with lateral plate feebly developed, ear­shaped, without acute projections (
<figureCitation id="B2F7EA8B6779FFC372AEFE778F46FDEA" box="[413,538,404,430]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">Fig. 3l, m</figureCitation>
), mesial surface with row of spinules (
<figureCitation id="B2F7EA8B6779FFC370C7FE77890BFDEA" box="[1012,1111,404,430]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">
Fig.
<quantity id="ED345BEB6779FFC3771DFE77890BFDEA" box="[1070,1111,404,430]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="31" unit="m" value="3.0">3m</quantity>
</figureCitation>
); antepenultimate segment somewhat broadened, triangular in cross section; penultimate segment about twice as long as wide (
<figureCitation id="B2F7EA8B6779FFC37123FE078F03FDBA" box="[528,607,484,510]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">Fig. 3j</figureCitation>
); ultimate segment tapering distally, dorsal margin with short spines or spiniform setae, tip with two stout subterminal spines and one stout terminal spine (
<figureCitation id="B2F7EA8B6779FFC37253FDD78C9DFE0A" box="[352,449,564,590]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">Fig. 3k</figureCitation>
); exopod reaching 3/4 length of antepenultimate segment (
<figureCitation id="B2F7EA8B6779FFC377F2FDD78847FE0A" box="[1217,1307,564,590]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="4" pageNumber="31">Fig. 3j</figureCitation>
); arthrobranch well developed.
</paragraph>
<caption id="7EB3A6866779FFC3723BFAC58CB4F924" httpUri="https://zenodo.org/record/2645969/files/figure.png" pageId="4" pageNumber="31" startId="4.[264,367,1318,1342]" targetBox="[304,1279,684,1289]" targetPageId="4">
<paragraph id="2A73F60E6779FFC3723BFAC58CB4F924" blockId="4.[264,1324,1318,1376]" pageId="4" pageNumber="31">
<emphasis id="18B82A1C6779FFC3723BFAC58CD6F97A" bold="true" box="[264,394,1318,1342]" pageId="4" pageNumber="31">FIGURE 1</emphasis>
.
<taxonomicName id="EDCC8D8D6779FFC372ADFAC48F27F97A" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[414,635,1319,1342]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="31" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6779FFC372ADFAC48F27F97A" box="[414,635,1319,1342]" italics="true" pageId="4" pageNumber="31">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676779FFC371BCFAC48E6EF97A" box="[655,818,1319,1342]" pageId="4" pageNumber="31" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Holotype, female (LACM CR 2005­013.2): habitus. Scale
<quantity id="ED345BEB6779FFC37290FAAA8CB8F924" box="[419,484,1353,1376]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="31" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="2A73F60E6779FFC2720BFA5F8CCDFDC2" blockId="4.[264,1323,1468,1894]" lastBlockId="5.[264,1323,284,390]" lastPageId="5" lastPageNumber="32" pageId="4" pageNumber="31">
Chelipeds equal in size, symmetrical in shape, carried extended, robust (
<figureCitation id="B2F7EA8B6779FFC37797FA5F8841F992" box="[1188,1309,1468,1494]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4a, b</figureCitation>
); coxa robust, with row of spinules ventromesially (
<figureCitation id="B2F7EA8B6779FFC37057FA078EE2F9BA" box="[868,958,1508,1534]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4h</figureCitation>
); basis small, with tuft of stiff setae mesially; ischium very stout, dorsal margin distally produced and furnished with two­three stout spines (
<figureCitation id="B2F7EA8B6779FFC37113F9D78FECFA0A" box="[544,688,1588,1614]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4c, h, i</figureCitation>
); merus stout, flattened ventrally, distolateral margin with rounded lobe; carpus very stout, cup­shaped, distal margin with several rounded lobes ­ one dorsolateral, one dorsomesial and one, particularly developed, ventral lobe, latter partially embracing palm (
<figureCitation id="B2F7EA8B6779FFC3710EF94F8E58FA82" box="[573,772,1708,1734]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4a, b, d, e, h</figureCitation>
); mesial face with two comb­like rows of setae (
<figureCitation id="B2F7EA8B6779FFC37222F9378CD1FAAA" box="[273,397,1748,1774]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4d, g</figureCitation>
); chela relatively short, inflated; palm smooth, linea impressa and adhesive discs absent; fingers stout, slightly more than half length of palm (
<figureCitation id="B2F7EA8B6779FFC3771EF91F89D8FB52" box="[1069,1156,1788,1814]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4e</figureCitation>
, gj), slightly gaping when closed; cutting edge of dactylus unarmed, with hiatus anterior to mid­portion (
<figureCitation id="B2F7EA8B6779FFC37222F8AF8C3BFB22" box="[273,359,1868,1894]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="4" pageNumber="31">Fig. 4e</figureCitation>
, hj); cutting edge of pollex armed with blunt tooth opposed to hiatus, and with one minute subdistal tooth (
<figureCitation id="B2F7EA8B6778FFC2716EFEFF8F8DFD72" box="[605,721,284,310]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="5" pageNumber="32">Fig. 4h, i</figureCitation>
); snapping mechanism absent; finger tips simple, crossing when chela closed (
<figureCitation id="B2F7EA8B6778FFC27150FEA78F87FD1A" box="[611,731,324,350]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="5" pageNumber="32">Fig. 4e, h</figureCitation>
); finger armature similar on right and left chelae (
<figureCitation id="B2F7EA8B6778FFC27222FE8F8CD8FDC2" box="[273,388,364,390]" captionStart="FIGURE 4" captionStartId="7.[264,367,1426,1450]" captionTargetBox="[94,1315,286,1400]" captionTargetId="figure@7.[264,1323,314,1426]" captionTargetPageId="7" captionText="FIGURE 4. Leslibetaeus coibita, n. gen., n. sp. Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj)." httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="5" pageNumber="32">Fig. 4e, h</figureCitation>
).
</paragraph>
<caption id="7EB3A6866778FFC2723BF972898DFB77" httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="5" pageNumber="32" startId="5.[264,367,1681,1705]" targetBox="[272,1295,460,1649]" targetPageId="5">
<paragraph id="2A73F60E6778FFC2723BF972898DFB77" blockId="5.[264,1323,1681,1843]" pageId="5" pageNumber="32">
<emphasis id="18B82A1C6778FFC2723BF9728CD9FAED" bold="true" box="[264,389,1681,1705]" pageId="5" pageNumber="32">FIGURE 2</emphasis>
.
<taxonomicName id="EDCC8D8D6778FFC272A6F9718F30FAED" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[405,620,1682,1705]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="32" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6778FFC272A6F9718F30FAED" box="[405,620,1682,1705]" italics="true" pageId="5" pageNumber="32">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676778FFC2714FF9728E53FAEC" box="[636,783,1681,1704]" pageId="5" pageNumber="32" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di).
</paragraph>
</caption>
<caption id="7EB3A686677BFFC1723BF9E08CB3FB17" httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="6" pageNumber="33" startId="6.[264,367,1539,1563]" targetBox="[283,1493,285,1518]" targetPageId="6">
<paragraph id="2A73F60E677BFFC1723BF9E08CB3FB17" blockId="6.[264,1323,1539,1875]" pageId="6" pageNumber="33">
<emphasis id="18B82A1C677BFFC1723BF9E08CDFFA5F" bold="true" box="[264,387,1539,1563]" pageId="6" pageNumber="33">FIGURE 3</emphasis>
.
<taxonomicName id="EDCC8D8D677BFFC172A1F9E78F34FA5F" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[402,616,1540,1563]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C677BFFC172A1F9E78F34FA5F" box="[402,616,1540,1563]" italics="true" pageId="6" pageNumber="33">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B9767677BFFC17145F9E78E59FA5F" box="[630,773,1540,1563]" pageId="6" pageNumber="33" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale).
</paragraph>
</caption>
<caption id="7EB3A686677AFFC0723BFA718925FA77" httpUri="https://zenodo.org/record/2645975/files/figure.svg" pageId="7" pageNumber="34" startId="7.[264,367,1426,1450]" targetBox="[94,1315,286,1400]" targetPageId="7">
<paragraph id="2A73F60E677AFFC0723BFA718925FA77" blockId="7.[264,1323,1426,1587]" pageId="7" pageNumber="34">
<emphasis id="18B82A1C677AFFC0723BFA718CDFF9EE" bold="true" box="[264,387,1426,1450]" pageId="7" pageNumber="34">FIGURE 4</emphasis>
.
<taxonomicName id="EDCC8D8D677AFFC072A2FA718F3BF9ED" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[401,615,1426,1449]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="34" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C677AFFC072A2FA718F3BF9ED" box="[401,615,1426,1449]" italics="true" pageId="7" pageNumber="34">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B9767677AFFC07146FA718E5FF9ED" box="[629,771,1426,1449]" pageId="7" pageNumber="34" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Paratype male (LACM CR 2005­013.1): chelipeds, a — left cheliped, lateral view; b — same, mesial view; c — same, ischium and basis, lateral view; d — same, carpus, mesial view; e — same, chela and carpus, lateral view; f — same, chela fingers open, lateral view; g — same, mesial view; h — right cheliped, coxa to proximal merus, mesial view; i — lateral view; j — same, chela, lateral view. Scale 1 mm (A: a, b, B: cj).
</paragraph>
</caption>
<paragraph id="2A73F60E677AFFCF720BF96C8F42FDBA" blockId="7.[264,1323,1679,1905]" lastBlockId="8.[264,1323,284,510]" lastPageId="8" lastPageNumber="35" pageId="7" pageNumber="34">
Second pereiopod slender; ischium subequal to merus; carpus with five segments, second longest, segment length ratio approximately equal to 1: 1.5: 1.2: 0.8: 1 (
<figureCitation id="B2F7EA8B677AFFC077F6F9548840FA95" box="[1221,1308,1719,1745]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="7" pageNumber="34">Fig. 3n</figureCitation>
); chela simple, fingers slightly shorter than palm, with tufts of setae. Third pereiopod with all endopodal articles strongly compressed; ischium without ventrolateral spine, with row of stiff setae ventrally (
<figureCitation id="B2F7EA8B677AFFC0711AF8CC8FD9FB0D" box="[553,645,1839,1865]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="7" pageNumber="34">Fig. 3o</figureCitation>
); merus broadened, dorsal and ventral margins slightly convex, unarmed; carpus distinctly shorter and narrower than merus, with stiff seta ventrodistally (
<figureCitation id="B2F7EA8B6775FFCF7288FEFF8F4FFD72" box="[443,531,284,310]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="8" pageNumber="35">Fig. 3o</figureCitation>
); propodus subequal in length and width to carpus, with about six slender spines on ventral margin and pair of stouter distoventral spines adjacent to dactylus (
<figureCitation id="B2F7EA8B6775FFCF7248FE8F8C88FDC2" box="[379,468,364,390]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="8" pageNumber="35">Fig. 3o</figureCitation>
); dactylus simple, moderately slender, slightly curved, about 1/3 length of propodus, furnished with setae (
<figureCitation id="B2F7EA8B6775FFCF71FFFE778E77FDEA" box="[716,811,404,430]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="8" pageNumber="35">Fig. 3o</figureCitation>
). Fourth pereiopod similar to third, but distinctly more slender (
<figureCitation id="B2F7EA8B6775FFCF710AFE5F8FD9FD92" box="[569,645,444,470]" captionStart="FIGURE 1" captionStartId="4.[264,367,1318,1342]" captionTargetBox="[304,1279,684,1289]" captionTargetId="figure@4.[264,1323,667,1322]" captionTargetPageId="4" captionText="FIGURE 1. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2): habitus. Scale 1 mm." httpUri="https://zenodo.org/record/2645969/files/figure.png" pageId="8" pageNumber="35">Fig. 1</figureCitation>
). Fifth pereiopod unknown (missing on both sides in
<typeStatus id="F57748AC6775FFCF723BFE078C32FDBA" box="[264,366,484,510]" pageId="8" pageNumber="35" type="holotype">holotype</typeStatus>
and
<typeStatus id="F57748AC6775FFCF7294FE078F4EFDBA" box="[423,530,484,510]" pageId="8" pageNumber="35" type="paratype">paratype</typeStatus>
).
</paragraph>
<caption id="7EB3A6866775FFCF723BF9E68E8EFA04" httpUri="https://zenodo.org/record/2645977/files/figure.png" pageId="8" pageNumber="35" startId="8.[264,367,1541,1565]" targetBox="[264,1322,566,1517]" targetPageId="8">
<paragraph id="2A73F60E6775FFCF723BF9E68E8EFA04" blockId="8.[264,1323,1541,1600]" pageId="8" pageNumber="35">
<emphasis id="18B82A1C6775FFCF723BF9E68CD8FA59" bold="true" box="[264,388,1541,1565]" pageId="8" pageNumber="35">FIGURE 5</emphasis>
.
<taxonomicName id="EDCC8D8D6775FFCF72A1F9E58F34FA59" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[402,616,1542,1565]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="35" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6775FFCF72A1F9E58F34FA59" box="[402,616,1542,1565]" italics="true" pageId="8" pageNumber="35">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676775FFCF7145F9E58E5AFA59" box="[630,774,1542,1565]" pageId="8" pageNumber="35" rank="species">n. gen., n. sp.</taxonomicNameLabel>
Holotype, female (LACM CR 2005­013.2): colour pattern, a — right side; b — left side. Photograph by Leslie Harris.
</paragraph>
</caption>
<paragraph id="2A73F60E6775FFCE720BF9788C21FF22" blockId="8.[264,1323,1691,1877]" lastBlockId="9.[264,1324,284,870]" lastPageId="9" lastPageNumber="36" pageId="8" pageNumber="35">
Abdominal somites IV with posterolateral angles of pleura rounded (
<figureCitation id="B2F7EA8B6775FFCF77BFF9788989FAF1" box="[1164,1237,1691,1717]" captionStart="FIGURE 1" captionStartId="4.[264,367,1318,1342]" captionTargetBox="[304,1279,684,1289]" captionTargetId="figure@4.[264,1323,667,1322]" captionTargetPageId="4" captionText="FIGURE 1. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2): habitus. Scale 1 mm." httpUri="https://zenodo.org/record/2645969/files/figure.png" pageId="8" pageNumber="35">Fig. 1</figureCitation>
); sixth somite rounded posteriorly and without distinct articulated plate at ventroposterior angle (
<figureCitation id="B2F7EA8B6775FFCF7222F9088C30FB41" box="[273,364,1771,1797]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="8" pageNumber="35">Fig. 2f</figureCitation>
); preanal plate slightly projecting posteriorly, subrectangular (
<figureCitation id="B2F7EA8B6775FFCF774DF908898BFB41" box="[1150,1239,1771,1797]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="8" pageNumber="35">
Fig.
<date id="5E72D0CE6775FFCF778EF908898BFB41" box="[1213,1239,1771,1797]" pageId="8" pageNumber="35">2i</date>
</figureCitation>
). First pleopod with endopod less than half length of exopod, oval, fringed with setae (
<figureCitation id="B2F7EA8B6775FFCF77F7F8F08841FB69" box="[1220,1309,1811,1837]" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="8" pageNumber="35">Fig. 3p</figureCitation>
). Male second pleopod with appendix masculina short, slightly longer than appendix interna, with three­four slender setae on apex and three setae along mesial margin (
<figureCitation id="B2F7EA8B6774FFCE77FCFEFF8C49FD1A" captionStart="FIGURE 3" captionStartId="6.[264,367,1539,1563]" captionTargetBox="[283,1493,285,1518]" captionTargetId="figure@6.[264,1323,316,1538]" captionTargetPageId="6" captionText="FIGURE 3. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (p) and paratype male (LACM CR 2005­013.1) (ao, q, r), a — antenna, ventral view; b — same, dorsal view; c — right mandible, mesial view; d — same, lateral view (palp detached); e — left mandible, lateral view (palp detached); f — maxillule, lateral view; g — maxilla, lateral view; h — first maxilliped, lateral view; i — second maxilliped, lateral view; j — third maxilliped, lateral view (arthrobranch detached, position indicated); k — same, tip of ultimate segment, lateral view; l — same, proximal region, lateral view; m — same, ventrolateral view; n — second pereiopod, lateral view; o — third pereiopod, lateral view; p — first pleopod, mesial view; q — second pleopod, mesial view; r — same, detail of appendices externa and interna. Scale 1 mm (A: j, B: ah, k; r drawn without scale)." httpUri="https://zenodo.org/record/2645973/files/figure.svg" pageId="9" pageNumber="36">Fig. 3q, r</figureCitation>
); female second pleopod with appendix interna only. Uropods slightly exceeding telson (
<figureCitation id="B2F7EA8B6774FFCE7222FE8F8C39FDC2" box="[273,357,364,390]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">Fig. 2f</figureCitation>
); lateral lobe of sympodite distally acute (
<figureCitation id="B2F7EA8B6774FFCE7064FE8F8EF3FDC2" box="[855,943,364,390]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">
Fig.
<quantity id="ED345BEB6774FFCE70BCFE8F8EF3FDC2" box="[911,943,364,390]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="9" pageNumber="36" unit="g" value="2.0">2g</quantity>
</figureCitation>
); endopod and exopod subequal in length; exopod with diaeresis bearing three teeth ­ one small triangular tooth near distolateral spine, one broad subacute tooth in mid­portion and one broadly rounded tooth closer to mesial margin (
<figureCitation id="B2F7EA8B6774FFCE7100FE078FD0FDBA" box="[563,652,484,510]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">
Fig.
<quantity id="ED345BEB6774FFCE715FFE078FD0FDBA" box="[620,652,484,510]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="9" pageNumber="36" unit="g" value="2.0">2g</quantity>
</figureCitation>
); distolateral spine relatively small but robust (
<figureCitation id="B2F7EA8B6774FFCE77F0FE078841FDBA" box="[1219,1309,484,510]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">
Fig.
<quantity id="ED345BEB6774FFCE77CFFE078841FDBA" box="[1276,1309,484,510]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="9" pageNumber="36" unit="g" value="2.0">2g</quantity>
</figureCitation>
). Telson broad, tapering distally, subtriangular, about 1.5 times as long as wide proximally, with lateral margins convex (
<figureCitation id="B2F7EA8B6774FFCE7151FDD78FE7FE0A" box="[610,699,564,590]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">Fig. 2h</figureCitation>
); proximal width about five times width of posterior margin; dorsal surface with two pairs of small spines situated at some distance from lateral margin, one pair at mid­length, another pair at distal 1/3 of telson length (
<figureCitation id="B2F7EA8B6774FFCE778EFD678840FEDA" box="[1213,1308,644,670]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">Fig. 2h</figureCitation>
); posterior margin reduced, straight, with two pairs of posterolateral spines, lateral much shorter than mesial and directed upwards (
<figureCitation id="B2F7EA8B6774FFCE702BFD378ECDFEAA" box="[792,913,724,750]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">Fig. 2f, h</figureCitation>
), median portion between mesial spines with two long, non­plumose setae and two shorter setae departing obliquely (missing in one specimen); anal tubercles absent (
<figureCitation id="B2F7EA8B6774FFCE706CFCC78EEEFF7A" box="[863,946,804,830]" captionStart="FIGURE 2" captionStartId="5.[264,367,1681,1705]" captionTargetBox="[272,1295,460,1649]" captionTargetId="figure@5.[264,1323,446,1657]" captionTargetPageId="5" captionText="FIGURE 2. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2) (a, b) and paratype male (LACM CR 2005­013.1) (ci), a — frontal region, dorsal view; b — same, lateral view; c — carapace, lateral view; d — antennule, lateral view; e — same, mesial view; f — posterior region of abdomen and telson, lateral view; g — uropod, dorsal view; h — telson, dorsal view; i — telson and sixth abdominal somite, ventral view. Scale 1 mm (A: c, B: a, b, di)." httpUri="https://zenodo.org/record/2645971/files/figure.png" pageId="9" pageNumber="36">
Fig.
<date id="5E72D0CE6774FFCE70ABFCC78EEEFF7A" box="[920,946,804,830]" pageId="9" pageNumber="36">2i</date>
</figureCitation>
). Gill/exopod formula as given for genus.
</paragraph>
<caption id="7EB3A6866774FFCE723BF9338E41FB0B" httpUri="https://zenodo.org/record/2645979/files/figure.png" pageId="9" pageNumber="36" startId="9.[264,367,1744,1768]" targetBox="[264,1323,1089,1720]" targetPageId="9">
<paragraph id="2A73F60E6774FFCE723BF9338E41FB0B" blockId="9.[264,1323,1744,1871]" pageId="9" pageNumber="36">
<emphasis id="18B82A1C6774FFCE723BF9338CD8FAAC" bold="true" box="[264,388,1744,1768]" pageId="9" pageNumber="36">FIGURE 6</emphasis>
. Locality of collection of
<taxonomicName id="EDCC8D8D6774FFCE71ACF9338E2AFAA3" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[671,886,1744,1767]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6774FFCE71ACF9338E2AFAA3" box="[671,886,1744,1767]" italics="true" pageId="9" pageNumber="36">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676774FFCE70B7F933894AFAA3" box="[900,1046,1744,1767]" pageId="9" pageNumber="36" rank="species">
<taxonomicNameLabel id="038B97676774FFCE70B7F9338E93FAA3" box="[900,975,1744,1767]" pageId="9" pageNumber="36" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</taxonomicNameLabel>
, intertidal rocky shore on Isla Coibita, Isla Coiba National Park, Panama, with Leslie Harris extracting animals from rock crevices. The red arrow indicates approximate position of the shale rock where the two specimens of
<taxonomicName id="EDCC8D8D6774FFCE7216F8DB8CA7FB0B" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[293,507,1848,1871]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="36" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6774FFCE7216F8DB8CA7FB0B" box="[293,507,1848,1871]" italics="true" pageId="9" pageNumber="36">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676774FFCE713BF8DB8FCBFB0B" box="[520,663,1848,1871]" pageId="9" pageNumber="36" rank="species">
<taxonomicNameLabel id="038B97676774FFCE713BF8DB8F0FFB0B" box="[520,595,1848,1871]" pageId="9" pageNumber="36" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</taxonomicNameLabel>
were found.
</paragraph>
</caption>
<paragraph id="2A73F60E6777FFCD720BFEF88FE0FDEA" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFEF88CD2FD71" bold="true" box="[312,398,283,309]" pageId="10" pageNumber="37">Colour</emphasis>
. — Semitranslucent with grey tinge; anterolateral and posterolateral portions of carapace, lateral portions of the first five abdominal somites, antennular peduncle and basicerite of antenna with bright red chromatophores; chelae hyaline whitish; cornea brown; ovaries bright yellow (
<figureCitation id="B2F7EA8B6777FFCD7158FE778FF0FDEA" box="[619,684,404,430]" captionStart="FIGURE 5" captionStartId="8.[264,367,1541,1565]" captionTargetBox="[264,1322,566,1517]" captionTargetId="figure@8.[264,1322,566,1517]" captionTargetPageId="8" captionText="FIGURE 5. Leslibetaeus coibita, n. gen., n. sp. Holotype, female (LACM CR 2005­013.2): colour pattern, a — right side; b — left side. Photograph by Leslie Harris." httpUri="https://zenodo.org/record/2645977/files/figure.png" pageId="10" pageNumber="37">Fig 5</figureCitation>
).
</paragraph>
<paragraph id="2A73F60E6777FFCD720BFE588FCDFDBA" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFE588C36FD91" bold="true" box="[312,362,443,469]" pageId="10" pageNumber="37">Size</emphasis>
. — Small sized alpheid, with CL ranging from 3.2 mm in male to 3.4 mm in female, latter with TL of 9.1 mm.
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856777FFCD720BFDE88CD6FE32" pageId="10" pageNumber="37" type="etymology">
<paragraph id="2A73F60E6777FFCD720BFDE88CD6FE32" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFDE88CE1FE61" bold="true" box="[312,445,523,549]" pageId="10" pageNumber="37">Etymology</emphasis>
. — The name of the new species refers to the
<typeStatus id="F57748AC6777FFCD70E5FDEF8955FE62" box="[982,1033,524,550]" pageId="10" pageNumber="37">type</typeStatus>
locality, Coibita, a small island off
<collectingCountry id="52DBB69E6777FFCD724CFDD78CF5FE0A" box="[383,425,564,590]" name="Iceland" pageId="10" pageNumber="37">Isla</collectingCountry>
Ranchería in the
<collectingCountry id="52DBB69E6777FFCD7149FDD78FF9FE0A" box="[634,677,564,590]" name="Iceland" pageId="10" pageNumber="37">Isla</collectingCountry>
Coiba National Park. Coibita is here used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856777FFCD720BFD678F5EFF52" pageId="10" pageNumber="37" type="biology_ecology">
<paragraph id="2A73F60E6777FFCD720BFD678F5EFF52" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFD678CC5FEDA" bold="true" box="[312,409,644,670]" pageId="10" pageNumber="37">Ecology</emphasis>
. — The
<typeStatus id="F57748AC6777FFCD7121FD678F24FEDA" box="[530,632,644,670]" pageId="10" pageNumber="37" type="holotype">holotype</typeStatus>
specimens of
<taxonomicName id="EDCC8D8D6777FFCD7001FD67897EFED9" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[818,1058,644,669]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="species" species="coibita" status="n. gen., n. sp.">
<emphasis id="18B82A1C6777FFCD7001FD67897EFED9" box="[818,1058,644,669]" italics="true" pageId="10" pageNumber="37">Leslibetaeus coibita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676777FFCD7704FD6789B8FEDA" box="[1079,1252,644,670]" pageId="10" pageNumber="37" rank="species">n. gen., n. sp.</taxonomicNameLabel>
were collected at low tide on the rocky mid­intertidal (
<figureCitation id="B2F7EA8B6777FFCD7051FD4F8EFAFE82" box="[866,934,684,710]" captionStart="FIGURE 6" captionStartId="9.[264,367,1744,1768]" captionTargetBox="[264,1323,1089,1720]" captionTargetId="figure@9.[264,1323,926,1720]" captionTargetPageId="9" captionText="FIGURE 6. Locality of collection of Leslibetaeus coibita, n. gen., n. sp., intertidal rocky shore on Isla Coibita, Isla Coiba National Park, Panama, with Leslie Harris extracting animals from rock crevices. The red arrow indicates approximate position of the shale rock where the two specimens of Leslibetaeus coibita, n. gen., n. sp. were found." httpUri="https://zenodo.org/record/2645979/files/figure.png" pageId="10" pageNumber="37">Fig 6</figureCitation>
), from crevices of a large shale rock. It is noteworthy that the bedrock and most loose rocks in the area were basaltic (L. Harris, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856777FFCD720BFCC78EE6FF22" pageId="10" pageNumber="37" type="distribution">
<paragraph id="2A73F60E6777FFCD720BFCC78EE6FF22" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFCC78C8DFF7A" bold="true" box="[312,465,804,830]" pageId="10" pageNumber="37">Distribution</emphasis>
. — Presently known only from the
<typeStatus id="F57748AC6777FFCD704AFCC78EF0FF7A" box="[889,940,804,830]" pageId="10" pageNumber="37">type</typeStatus>
locality:
<collectingCountry id="52DBB69E6777FFCD772FFCC7891AFF7A" box="[1052,1094,804,830]" name="Iceland" pageId="10" pageNumber="37">Isla</collectingCountry>
Coibita,
<collectingCountry id="52DBB69E6777FFCD7780FCC78981FF7A" box="[1203,1245,804,830]" name="Iceland" pageId="10" pageNumber="37">Isla</collectingCountry>
Coiba National Park, Golfo de
<collectingRegion id="E80838EC6777FFCD711BFCAF8FD1FF22" box="[552,653,844,870]" country="Panama" name="Chiriqui" pageId="10" pageNumber="37">Chiriqui</collectingRegion>
, Pacific coast of
<collectingCountry id="52DBB69E6777FFCD7066FCAF8EEAFF22" box="[853,950,844,870]" name="Panama" pageId="10" pageNumber="37">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856777FFCC720BFC978E79FD1A" lastPageId="11" lastPageNumber="38" pageId="10" pageNumber="37" type="discussion">
<paragraph id="2A73F60E6777FFCD720BFC978FA9F902" blockId="10.[264,1325,283,1870]" pageId="10" pageNumber="37">
<emphasis id="18B82A1C6777FFCD720BFC978CF6FFCA" bold="true" box="[312,426,884,910]" pageId="10" pageNumber="37">Remarks</emphasis>
. —
<taxonomicName id="EDCC8D8D6777FFCD72D3FC978F2FFFC9" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[480,627,884,909]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6777FFCD72D3FC978F2FFFC9" box="[480,627,884,909]" italics="true" pageId="10" pageNumber="37">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676777FFCD71B0FC978F88FFCA" box="[643,724,884,910]" pageId="10" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
is defined by the combination of several features: the frontal margin of the carapace without orbital teeth and distinct rostrum; the incomplete orbital hoods, only partially concealing the eyes; the sixth abdominal somite without articulated plate; the symmetrical stout chelipeds, with ischium bearing spines dorsally, carpus distally strongly lobed and bearing two comb­like rows of setae mesially, and short, smooth chela, without snapping mechanism on the finger cutting edges; the carpus of the second pereiopod five­segmented, with second segment distinctly longer than first; the ischium and merus of the third pereiopod without spines, ischium with a row of setae ventrally; the first segment of the antennular peduncle without tooth on the ventromesial carina; the scaphocerite being only half as long as carpocerite; the tip of the third maxilliped with stout spines; and the strap­like epipods present on the third maxilliped and the first to fifth pereiopod.
</paragraph>
<paragraph id="2A73F60E6777FFCC720BFAB78E79FD1A" blockId="10.[264,1325,283,1870]" lastBlockId="11.[264,1325,284,1030]" lastPageId="11" lastPageNumber="38" pageId="10" pageNumber="37">
The characters of the frontal region, chelipeds and walking legs suggest that
<taxonomicName id="EDCC8D8D6777FFCD723BFA9F8CC7F9D1" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[264,411,1404,1429]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6777FFCD723BFA9F8CC7F9D1" box="[264,411,1404,1429]" italics="true" pageId="10" pageNumber="37">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676777FFCD7286FA9F8F4CF9D2" box="[437,528,1404,1430]" pageId="10" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
is not closely related to any described alpheid genus. The symmetrical chelipeds, without snapping mechanism on the chela fingers, and with rows of setae on the mesial side of the carpus, as well as the imperfect orbital hoods, suggest that
<taxonomicName id="EDCC8D8D6777FFCD7208FA178C91FA49" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[315,461,1524,1549]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6777FFCD7208FA178C91FA49" box="[315,461,1524,1549]" italics="true" pageId="10" pageNumber="37">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676777FFCD72E8FA178F76FA4A" box="[475,554,1524,1550]" pageId="10" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
is probably a relatively basal genus. Five other alpheid genera are characterized by the equal and symmetrical chelipeds, with setae arranged in two or more rows on the mesial face of the cheliped carpus:
<taxonomicName id="EDCC8D8D6777FFCD70FBF9A7887AFA1A" authority="Kensley, 1988" authorityName="Kensley" authorityYear="1988" box="[968,1318,1604,1630]" class="Malacostraca" family="Alpheidae" genus="Yagerocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD70FBF9A7890DFA19" box="[968,1105,1604,1629]" italics="true" pageId="10" pageNumber="37">Yagerocaris</emphasis>
<bibRefCitation id="4E5D8BFF6777FFCD7758F9A7887AFA1A" author="Kensley, B. F." box="[1131,1318,1604,1630]" pageId="10" pageNumber="37" refId="ref4823" refString="Kensley, B. F. (1988) New species and records of cave shrimps from the Yucatan Peninsula (Deca-" type="book" year="1988">Kensley, 1988</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="EDCC8D8D6777FFCD723BF98F8F2BFAC2" authority="Powell, 1979" authorityName="Powell" authorityYear="1979" box="[264,631,1644,1670]" class="Malacostraca" family="Alpheidae" genus="Potamalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD723BF98F8CE2FAC1" box="[264,446,1644,1669]" italics="true" pageId="10" pageNumber="37">Potamalpheops</emphasis>
<bibRefCitation id="4E5D8BFF6777FFCD72FCF98F8F2BFAC2" author="Powell, C. B." box="[463,631,1644,1670]" pageId="10" pageNumber="37" pagination="116 - 150" refId="ref5129" refString="Powell, C. B. (1979) Three alpheid shrimps of a new genus from West African fresh and brackish waters: taxonomy and ecological zonations (Crustacea Decapoda Natantia). Revue de Zoologie Africaine, 93 (1), 116 - 150." type="journal article" year="1979">Powell, 1979</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="EDCC8D8D6777FFCD71BFF98F8E97FAC2" authority="Miya, 1997" authorityName="Miya" authorityYear="1997" box="[652,971,1644,1670]" class="Malacostraca" family="Alpheidae" genus="Stenalpheops" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD71BFF98F8E7BFAC1" box="[652,807,1644,1669]" italics="true" pageId="10" pageNumber="37">Stenalpheops</emphasis>
<bibRefCitation id="4E5D8BFF6777FFCD700BF98F8E97FAC2" author="Miya, Y." box="[824,971,1644,1670]" pageId="10" pageNumber="37" pagination="145 - 161" refId="ref5074" refString="Miya, Y. (1997) Stenalpheops anacanthus, new genus, new species (Crustacea, Decapoda, Alpheidae) from the Seto Inland Sea and the Sea of Ariake, South Japan. Bulletin of Faculty of Liberal Arts, Nagasaki University, 38 (1), 145 - 161." type="journal article" year="1997">Miya, 1997</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="EDCC8D8D6777FFCD70D2F98F8CC5FAEA" authority="Anker and Felder 2005" authorityName="Anker and Felder" authorityYear="2005" class="Malacostraca" family="Alpheidae" genus="Coutieralpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD70D2F98F89CAFAC1" box="[993,1174,1644,1669]" italics="true" pageId="10" pageNumber="37">Coutieralpheus</emphasis>
<bibRefCitation id="4E5D8BFF6777FFCD7795F98F8CC5FAEA" author="Anker, A. &amp; Felder, D. L." pageId="10" pageNumber="37" pagination="40 - 52" refId="ref4061" refString="Anker, A. &amp; Felder, D. L. (2005) Description of Coutieralpheus setirostris, new genus, new species, an infaunal alpheid shrimp (Crustacea, Decapoda) from Florida, U. S. A. Crustacean Research, 34, 40 - 52." type="journal article" year="2005">Anker and Felder 2005</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="EDCC8D8D6777FFCD72E9F9778E23FAEA" authority="Anker &amp; Dworschak" authorityName="Anker &amp; Dworschak" authorityYear="2007" box="[474,895,1684,1710]" class="Malacostraca" family="Alpheidae" genus="Jengalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD72E9F9778F27FAE9" box="[474,635,1684,1709]" italics="true" pageId="10" pageNumber="37">Jengalpheops</emphasis>
Anker &amp; Dworschak
</taxonomicName>
, in press. However, all these genera differ from
<taxonomicName id="EDCC8D8D6777FFCD72A1F95F8F79FA91" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[402,549,1724,1749]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6777FFCD72A1F95F8F79FA91" box="[402,549,1724,1749]" italics="true" pageId="10" pageNumber="37">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676777FFCD7107F95F8FD8FA92" box="[564,644,1724,1750]" pageId="10" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
in many other aspects, e.g., the presence of spines on the ischium/merus of the third and fourth pereiopod; the presence of a well developed rostrum (
<taxonomicName id="EDCC8D8D6777FFCD7221F8EF8CC7FB61" authorityName="Kensley" authorityYear="1988" box="[274,411,1804,1829]" class="Malacostraca" family="Alpheidae" genus="Yagerocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD7221F8EF8CC7FB61" box="[274,411,1804,1829]" italics="true" pageId="10" pageNumber="37">Yagerocaris</emphasis>
</taxonomicName>
,
<taxonomicName id="EDCC8D8D6777FFCD729CF8EF8F39FB61" authorityName="Powell" authorityYear="1979" box="[431,613,1804,1829]" class="Malacostraca" family="Alpheidae" genus="Potamalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD729CF8EF8F39FB61" box="[431,613,1804,1829]" italics="true" pageId="10" pageNumber="37">Potamalpheops</emphasis>
</taxonomicName>
,
<taxonomicName id="EDCC8D8D6777FFCD714BF8EF8E71FB61" authorityName="Anker and Felder" authorityYear="2005" box="[632,813,1804,1829]" class="Malacostraca" family="Alpheidae" genus="Coutieralpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD714BF8EF8E71FB61" box="[632,813,1804,1829]" italics="true" pageId="10" pageNumber="37">Coutieralpheus</emphasis>
</taxonomicName>
); and the sixth abdominal somite with an articulated plate (all but
<taxonomicName id="EDCC8D8D6777FFCD710FF8D78F9BFB09" authorityName="Kensley" authorityYear="1988" box="[572,711,1844,1869]" class="Malacostraca" family="Alpheidae" genus="Yagerocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6777FFCD710FF8D78F9BFB09" box="[572,711,1844,1869]" italics="true" pageId="10" pageNumber="37">Yagerocaris</emphasis>
</taxonomicName>
) (cf.
<bibRefCitation id="4E5D8BFF6777FFCD7022F8D78E9EFB0A" author="Kensley, B. F." box="[785,962,1844,1870]" pageId="10" pageNumber="37" refId="ref4823" refString="Kensley, B. F. (1988) New species and records of cave shrimps from the Yucatan Peninsula (Deca-" type="book" year="1988">Kensley, 1988</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6777FFCD70E0F8D7892AFB0A" author="Powell, C. B." box="[979,1142,1844,1870]" pageId="10" pageNumber="37" pagination="116 - 150" refId="ref5129" refString="Powell, C. B. (1979) Three alpheid shrimps of a new genus from West African fresh and brackish waters: taxonomy and ecological zonations (Crustacea Decapoda Natantia). Revue de Zoologie Africaine, 93 (1), 116 - 150." type="journal article" year="1979">Powell, 1979</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6777FFCD77BBF8D7887BFB0A" author="Hobbs, H. H. Jr." box="[1160,1319,1844,1870]" pageId="10" pageNumber="37" pagination="221 - 224" refId="ref4688" refString="Hobbs, H. H. Jr. (1983) The African shrimp genus Potamalpheops in Mexico (Decapoda: Alpheidae). Crustaceana, 44 (2), 221 - 224." type="journal article" year="1983">Hobbs, 1983</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6776FFCC723BFEFF8CCEFD72" author="Miya, Y." box="[264,402,284,310]" pageId="11" pageNumber="38" pagination="145 - 161" refId="ref5074" refString="Miya, Y. (1997) Stenalpheops anacanthus, new genus, new species (Crustacea, Decapoda, Alpheidae) from the Seto Inland Sea and the Sea of Ariake, South Japan. Bulletin of Faculty of Liberal Arts, Nagasaki University, 38 (1), 145 - 161." type="journal article" year="1997">Miya, 1997</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6776FFCC72ADFEFF8FFFFD72" author="Anker, A. &amp; Felder, D. L." box="[414,675,284,310]" pageId="11" pageNumber="38" pagination="40 - 52" refId="ref4061" refString="Anker, A. &amp; Felder, D. L. (2005) Description of Coutieralpheus setirostris, new genus, new species, an infaunal alpheid shrimp (Crustacea, Decapoda) from Florida, U. S. A. Crustacean Research, 34, 40 - 52." type="journal article" year="2005">Anker &amp; Felder, 2005</bibRefCitation>
; Anker &amp; Dworschak, in press). Therefore, they do not have close affinities with
<taxonomicName id="EDCC8D8D6776FFCC7106FEA78F9BFD19" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[565,711,324,349]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC7106FEA78F9BFD19" box="[565,711,324,349]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC71E6FEA78E79FD1A" box="[725,805,324,350]" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856776FFCC720BFE8F8CB1F83A" pageId="11" pageNumber="38" type="reference_group">
<paragraph id="2A73F60E6776FFCC720BFE8F8E47F842" blockId="11.[264,1325,284,1030]" pageId="11" pageNumber="38">
<taxonomicName id="EDCC8D8D6776FFCC720BFE8F8C96FDC1" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[312,458,364,389]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC720BFE8F8C96FDC1" box="[312,458,364,389]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC72EFFE8F8F73FDC2" box="[476,559,364,390]" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
shares with
<taxonomicName id="EDCC8D8D6776FFCC71FFFE8F8EE3FDC2" ID-CoL="84LT5" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[716,959,364,390]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC71FFFE8F8E75FDC1" box="[716,809,364,389]" italics="true" pageId="11" pageNumber="38">Betaeus</emphasis>
Dana, 1852
</taxonomicName>
and
<taxonomicName id="EDCC8D8D6776FFCC70CDFE8F8C8DFDEA" ID-CoL="85H3R" authority="De Grave and Anker, 2000" authorityName="De Grave and Anker" authorityYear="2000" class="Malacostraca" family="Alpheidae" genus="Orygmalpheus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC70CDFE8F89F7FDC1" box="[1022,1195,364,389]" italics="true" pageId="11" pageNumber="38">Orygmalpheus</emphasis>
<bibRefCitation id="4E5D8BFF6776FFCC7785FE8F8C8DFDEA" author="De Grave, S. &amp; Anker, A." pageId="11" pageNumber="38" pagination="249 - 256" refId="ref4496" refString="De Grave, S. &amp; Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. Raffles Bulletin of Zoology, 48 (2), 249 - 256." type="journal article" year="2000">De Grave and Anker, 2000</bibRefCitation>
</taxonomicName>
the symmetrical (or subsymmetrical) chelipeds, and the frontal margin devoid of rostrum and orbital hoods, but differs from those genera in the position of the dactylus on the chela, and in the features on the third maxilliped, third and second pereiopods, sixth abdominal somite etc. (cf.
<bibRefCitation id="4E5D8BFF6776FFCC7008FDEF8E9EFE62" author="Hart, J. F. L." box="[827,962,524,550]" pageId="11" pageNumber="38" pagination="431 - 466" refId="ref4593" refString="Hart, J. F. L. (1964) Shrimps of the genus Betaeus on the Pacific coast of North America with descriptions of three new species. Proceedings of the United States National Museum, 115, 431 - 466." type="journal article" year="1964">Hart, 1964</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6776FFCC70E5FDEF8843FE62" author="De Grave, S. &amp; Anker, A." box="[982,1311,524,550]" pageId="11" pageNumber="38" pagination="249 - 256" refId="ref4496" refString="De Grave, S. &amp; Anker, A. (2000) A new genus and new species of alpheid shrimp (Crustacea: Decapoda) from Papua New Guinea. Raffles Bulletin of Zoology, 48 (2), 249 - 256." type="journal article" year="2000">De Grave &amp; Anker, 2000</bibRefCitation>
). Despite a very superficial resemblance (symmetrical chelipeds carried extended and frontal margin without rostrum and orbital teeth) between
<taxonomicName id="EDCC8D8D6776FFCC70E1FDBF8973FE31" ID-CoL="84LT5" authorityName="Dana" authorityYear="1852" box="[978,1071,604,629]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC70E1FDBF8973FE31" box="[978,1071,604,629]" italics="true" pageId="11" pageNumber="38">Betaeus</emphasis>
</taxonomicName>
and
<taxonomicName id="EDCC8D8D6776FFCC775CFDBF885EFE31" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[1135,1282,604,629]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC775CFDBF885EFE31" box="[1135,1282,604,629]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC7626FDBF8C60FEDA" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
, these two genera are not closely related.
<taxonomicName id="EDCC8D8D6776FFCC707DFD678EBDFED9" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[846,993,644,669]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC707DFD678EBDFED9" box="[846,993,644,669]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC70C6FD678915FEDA" box="[1013,1097,644,670]" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
also has common features with
<taxonomicName id="EDCC8D8D6776FFCC728EFD4F8E69FE82" ID-CoL="84Q9B" authority="Borradaile, 1898" authorityName="Borradaile" authorityYear="1898" box="[445,821,684,710]" class="Malacostraca" family="Alpheidae" genus="Metabetaeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC728EFD4F8F0FFE81" box="[445,595,684,709]" italics="true" pageId="11" pageNumber="38">Metabetaeus</emphasis>
Borradaile, 1898
</taxonomicName>
and
<taxonomicName id="EDCC8D8D6776FFCC70B1FD4F89F0FE82" ID-CoL="85JKM" authority="Xuan, 2001" authorityName="Xuan" authorityYear="2001" box="[898,1196,684,710]" class="Malacostraca" family="Alpheidae" genus="Thuylamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC70B1FD4F8958FE81" box="[898,1028,684,709]" italics="true" pageId="11" pageNumber="38">Thuylamea</emphasis>
<bibRefCitation id="4E5D8BFF6776FFCC7725FD4F89F0FE82" author="Xuan, N. V." box="[1046,1196,684,710]" pageId="11" pageNumber="38" pagination="217 - 228" refId="ref5895" refString="Xuan, N. V. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75 (1 - 15), 217 - 228." type="journal article" year="2001">Xuan, 2001</bibRefCitation>
</taxonomicName>
, e.g., the symmetrical chelipeds and the absence of an articulated plate on the sixth abdominal somite, but differs greatly from those genera in the cheliped shape and the configuration of the frontal margin (cf.
<bibRefCitation id="4E5D8BFF6776FFCC7121FCC78E70FF7A" author="Banner, A. H. &amp; Banner, D. M." box="[530,812,804,830]" pageId="11" pageNumber="38" pagination="299 - 303" refId="ref4113" refString="Banner, A. H. &amp; Banner, D. M. (1960) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean, part VII: On Metabetaeus Borradaile, with a new species from Hawaii. Pacific Science, 15 (3), 299 - 303." type="journal article" year="1960">Banner &amp; Banner, 1960</bibRefCitation>
;
<bibRefCitation id="4E5D8BFF6776FFCC700BFCC78E99FF7A" author="Xuan, N. V." box="[824,965,804,830]" pageId="11" pageNumber="38" pagination="217 - 228" refId="ref5895" refString="Xuan, N. V. (2001) A new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from South Vietnam. Zoologische Mededelingen, Leiden, 75 (1 - 15), 217 - 228." type="journal article" year="2001">Xuan, 2001</bibRefCitation>
). A close relationship between
<taxonomicName id="EDCC8D8D6776FFCC723BFCAF8CC7FF21" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[264,411,844,869]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC723BFCAF8CC7FF21" box="[264,411,844,869]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC729AFCAF8CA5FF22" box="[425,505,844,870]" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
and
<taxonomicName id="EDCC8D8D6776FFCC7102FCAF8E0FFF22" ID-CoL="84LPX" authority="De Man, 1888" authorityName="De Man" authorityYear="1888" box="[561,851,844,870]" class="Malacostraca" family="Alpheidae" genus="Automate" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC7102FCAF8FFDFF21" box="[561,673,844,869]" italics="true" pageId="11" pageNumber="38">Automate</emphasis>
De Man, 1888
</taxonomicName>
,
<taxonomicName id="EDCC8D8D6776FFCC7053FCAF89C9FF22" ID-CoL="84LB4" authority="Coutiere, 1896" authorityName="Coutiere" authorityYear="1897" box="[864,1173,844,870]" class="Malacostraca" family="Alpheidae" genus="Alpheopsis" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="18B82A1C6776FFCC7053FCAF8EBDFF21" box="[864,993,844,869]" italics="true" pageId="11" pageNumber="38">Alpheopsis</emphasis>
Coutière, 1896
</taxonomicName>
or any other genus present in the eastern Pacific is also unlikely. Thus the phylogenetic affinities of
<taxonomicName id="EDCC8D8D6776FFCC723BFC7F8CC7FFF1" ID-CoL="85FVN" authority="Anker &amp; Poddoubtchenko &amp; Wehrtmann, 2006" authorityName="Anker &amp; Poddoubtchenko &amp; Wehrtmann" authorityYear="2006" box="[264,411,924,949]" class="Malacostraca" family="Alpheidae" genus="Leslibetaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="18B82A1C6776FFCC723BFC7F8CC7FFF1" box="[264,411,924,949]" italics="true" pageId="11" pageNumber="38">Leslibetaeus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="038B97676776FFCC729DFC7F8F5DFFF2" box="[430,513,924,950]" pageId="11" pageNumber="38" rank="genus">n. gen.</taxonomicNameLabel>
are rather enigmatic. The finding of a completely new taxon in a relatively accessible habitat indicates a potentially high biodiversity of the
<collectingCountry id="52DBB69E6776FFCC779EFC278984FF9A" box="[1197,1240,964,990]" name="Iceland" pageId="11" pageNumber="38">Isla</collectingCountry>
Coiba National Park, which remains to be explored.
</paragraph>
<paragraph id="2A73F60E6776FFCC723BFB878CB1F83A" blockId="11.[264,493,1124,1150]" box="[264,493,1124,1150]" pageId="11" pageNumber="38">
<emphasis id="18B82A1C6776FFCC723BFB878CB1F83A" bold="true" box="[264,493,1124,1150]" pageId="11" pageNumber="38">Acknowledgments</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="62D6A5856776FFC9723BFB4C8EC8FE77" lastPageId="14" lastPageNumber="41" pageId="11" pageNumber="38" type="description">
<paragraph id="2A73F60E6776FFCC723BFB4C8CCEFA75" blockId="11.[264,1325,1199,1585]" pageId="11" pageNumber="38">
We would like to thank Leslie Harris (Natural History Museum of Los Angeles County, Los Angeles,
<collectingCountry id="52DBB69E6776FFCC7287FB348CAFF8B5" box="[436,499,1239,1265]" name="United States of America" pageId="11" pageNumber="38">USA</collectingCountry>
), who contributed in the collection of the present material. ISW is grateful to Dr. Peter W. Glynn (University of Miami, The Rosenstiel School,
<collectingCountry id="52DBB69E6776FFCC779FFB1C89B4F95D" box="[1196,1256,1279,1305]" name="United States of America" pageId="11" pageNumber="38">USA</collectingCountry>
) who invited him to participate in his project activities (
<date id="5E72D0CE6776FFCC7054FAC48EA0F905" box="[871,1020,1319,1345]" pageId="11" pageNumber="38" value="2005-03">March 2005</date>
) in
<collectingCountry id="52DBB69E6776FFCC7703FAC489CDF905" box="[1072,1169,1319,1345]" name="Panama" pageId="11" pageNumber="38">Panama</collectingCountry>
. The present study was supported by Dr. Nancy Knowlton (Scripps Institution of Oceanography, La Jolla,
<collectingCountry id="52DBB69E6776FFCC7260FA948CCCF9D5" box="[339,400,1399,1425]" name="United States of America" pageId="11" pageNumber="38">USA</collectingCountry>
) in form of a Smithsonian Institutions postdoctoral fellowship to the first author (AA), and was accomplished in the Smithsonian Tropical Research Institution (STRI) facilities at Naos laboratory in
<collectingRegion id="E80838EC6776FFCC71EBFA248E2EF9A5" box="[728,882,1479,1505]" country="Panama" name="Panama" pageId="11" pageNumber="38">Panama City</collectingRegion>
,
<collectingCountry id="52DBB69E6776FFCC704CFA248929F9A5" box="[895,1141,1479,1505]" name="Panama" pageId="11" pageNumber="38">Republic of Panama</collectingCountry>
. We also thank two anonymous reviewers whose suggestions and remarks improved the quality of the manuscript.
</paragraph>
<paragraph id="2A73F60E6776FFCC723BF96D8C91FAEC" blockId="11.[264,461,1678,1704]" box="[264,461,1678,1704]" pageId="11" pageNumber="38">
<emphasis id="18B82A1C6776FFCC723BF96D8C91FAEC" bold="true" box="[264,461,1678,1704]" pageId="11" pageNumber="38">Literature cited</emphasis>
</paragraph>
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The species of
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and
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On the status of
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