293 lines
27 KiB
XML
293 lines
27 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1129.95245" ID-GBIF-Dataset="4d57ec62-e4ca-4afe-9ba5-c9de955f94da" ID-Pensoft-Pub="1313-2970-1129-37" ID-Pensoft-UUID="FBBD05B1376B5FB695A40A1C568236B4" ID-ZooBank="B19E8A19D8A04CA38620D8532B447D17" ModsDocID="1313-2970-1129-37" checkinTime="1668159808446" checkinUser="pensoft" docAuthor="Yin, Zi-Wei & Zhao, Tie-Xiong" docDate="2022" docId="9C307C40F6D7554FBBAFD9AFC8979F96" docLanguage="en" docName="ZooKeys 1129: 37-44" docOrigin="ZooKeys 1129" docPubDate="2022-11-10" docSource="http://dx.doi.org/10.3897/zookeys.1129.95245" docTitle="Pseudophanias tianmuensis Yin & Zhao 2022, sp. nov." docType="treatment" docUuid="4476E74C-644F-4667-8AF5-45C75D21638A" docUuidSource="ZooBank" docVersion="2" id="FBBD05B1376B5FB695A40A1C568236B4" lastPageNumber="37" masterDocId="FBBD05B1376B5FB695A40A1C568236B4" masterDocTitle="An unusual species of Pseudophanias Raffray from West Tianmu Mountain, China (Coleoptera, Staphylinidae, Pselaphinae)" masterLastPageNumber="44" masterPageNumber="37" pageNumber="37" updateTime="1668247212472" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>An unusual species of Pseudophanias Raffray from West Tianmu Mountain, China (Coleoptera, Staphylinidae, Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yin, Zi-Wei</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6659-9448</mods:nameIdentifier>
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<mods:affiliation>Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, Shanghai 200234, China</mods:affiliation>
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<mods:nameIdentifier type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Tie-Xiong</mods:namePart>
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<mods:affiliation>44 East Genta Road, Zhuji, Shaoxing, Zhejiang 311800, China</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-11-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1129</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>37</mods:start>
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<mods:end>44</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1129.95245</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1129.95245</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1129-37</mods:identifier>
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<mods:identifier type="ZooBank">B19E8A19D8A04CA38620D8532B447D17</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FBBD05B1376B5FB695A40A1C568236B4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="204928542" LSID="urn:lsid:zoobank.org:act:4476E74C-644F-4667-8AF5-45C75D21638A" httpUri="http://treatment.plazi.org/id/9C307C40F6D7554FBBAFD9AFC8979F96" lastPageNumber="37" pageId="0" pageNumber="37">
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<subSubSection pageId="0" pageNumber="37" type="nomenclature">
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<paragraph pageId="0" pageNumber="37">
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<taxonomicName LSID="https://zoobank.org/4476E74C-644F-4667-8AF5-45C75D21638A" authority="Yin & Zhao" authorityName="Yin & Zhao" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias tianmuensis" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="tianmuensis" status="sp. nov.">Pseudophanias tianmuensis Yin & Zhao</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="37">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="37" type="description">
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<paragraph pageId="0" pageNumber="37">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudophanias tianmuensis sp. nov. A male B female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1129.95245.figure1" httpUri="https://binary.pensoft.net/fig/768275" pageId="0" pageNumber="37">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology and habitat of Pseudophanias tianmuensis sp. nov. A-C aedeagus A dorsal B lateral C ventral D paramere, lateral E general environment of the collecting site F decomposing tree branches inhabited by the beetle. Scale bars: 0.2 mm (A-C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1129.95245.figure3" httpUri="https://binary.pensoft.net/fig/768277" pageId="0" pageNumber="37">
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, 3 Chinese common name:
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<normalizedToken originalValue="天目隐须蚁甲">天目隐须蚁甲</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="37" type="type material">
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<paragraph pageId="0" pageNumber="37">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="37">
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(3 exx.).
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<emphasis bold="true" pageId="0" pageNumber="37">
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<emphasis italics="true" pageId="0" pageNumber="37">Holotype</emphasis>
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: China
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</emphasis>
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: ♂, 'China: Zhejiang, Linan District, West Tianmu Mountain, 1100 m, 23.viii.2014,
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<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Cryptomeria" kingdom="Plantae" lsidName="Cryptomeria" order="Pinales" pageId="0" pageNumber="37" phylum="Tracheophyta" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="37">Cryptomeria</emphasis>
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</taxonomicName>
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leaf, sift, Tie-Xiong Zhao leg.,
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<normalizedToken originalValue="浙江省临安市西天目山(此路不通)’">浙江省临安市西天目山(此路不通)'</normalizedToken>
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(in SNUC).
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<emphasis bold="true" pageId="0" pageNumber="37">
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<emphasis italics="true" pageId="0" pageNumber="37">Paratypes</emphasis>
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: China
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</emphasis>
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: 1 ♂ [completely disarticulated specimen], China: Zhejiang, Linan District, West Tianmu Mountain, 1150 m, 2.vii.2006, decomposing log, sift, Xiao-Bin Song leg.,
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<normalizedToken originalValue="浙江省临安市西天目山">浙江省临安市西天目山</normalizedToken>
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(停车场); 1 ♀, 'China: Zhejiang, Linan District, West Tianmu Mountain, 1180 m, 18.viii.2019, decomposing log, sift, Zi-Wei Yin leg.,
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<normalizedToken originalValue="浙江省临安市西天目山(消防道)’">浙江省临安市西天目山(消防道)'</normalizedToken>
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(both in SNUC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="37" type="diagnosis">
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<paragraph pageId="0" pageNumber="37">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="37">
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Body length 2.8-2.9 mm; dorsal vestiture coarsely punctate and densely setose. Head subrounded at base, with small vertexal and frontal foveae; antennomeres moniliform. Elytra with discal and lateral longitudinal ridges, areas between ridges impressed. Tergite 1 (IV) longer than 2 (V) at middle but clearly shorter than 2 and 3 (VI) combined.
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<emphasis bold="true" pageId="0" pageNumber="37">Male</emphasis>
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: antennal clubs formed by apical four enlarged, cylindrical antennomeres. Aedeagus symmetric dorso-ventrally; median lobe C-shaped laterally; parameres thin and elongate, each with two long apical and one preapical seta.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="37" type="description">
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<paragraph pageId="0" pageNumber="37">Description.</paragraph>
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<paragraph pageId="0" pageNumber="37">
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<emphasis bold="true" pageId="0" pageNumber="37">Male.</emphasis>
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Body (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudophanias tianmuensis sp. nov. A male B female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1129.95245.figure1" httpUri="https://binary.pensoft.net/fig/768275" pageId="0" pageNumber="37">1A</figureCitation>
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) length 2.91 mm; color red-brown, tarsi and mouthparts lighter. Dorsal vestiture coarsely punctate and covered with short, dense setae.
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</paragraph>
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<caption doi="10.3897/zookeys.1129.95245.figure1" httpUri="https://binary.pensoft.net/fig/768275" pageId="0" pageNumber="37" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="37">
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<emphasis bold="true" pageId="0" pageNumber="37">Figure 1.</emphasis>
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Habitus of
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<taxonomicName authorityName="Yin & Zhao" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias tianmuensis" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="tianmuensis">
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<emphasis italics="true" pageId="0" pageNumber="37">Pseudophanias tianmuensis</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
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male
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<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
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female. Scale bars: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="37">
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Head (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2A</figureCitation>
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) subrounded at base, longer than wide, length 0.63 mm, width 0.49 mm; vertex weakly convex, with two small, asetose vertexal foveae (dorsal tentorial pits); antennal tubercles indistinct; frons with single small fovea, protruding anteriorly to form rostrum; clypeus sharply descending, its surface coarse, anterior margin rounded and slightly carinate; lacking ocular-mandibular carina. Venter with two small, well-separated gular foveae (posterior tentorial pits), lacking median carina/sulcus. Maxillary palpus (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2B</figureCitation>
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) small in size, palpomere 1 minute, 2 pedunculate basally and broadened apically, 3 roundly trapezoidal, 4 broadly at approximately basal 1/3 and gradually narrowing apically. Compound eyes moderately prominent, reniform in lateral view, each composed of approximately 55 ommatidia. Antenna elongate, length 1.75 mm, distinct club (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2C</figureCitation>
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) formed by greatly enlarged antennomeres 8-11 and approximately as long as antennomeres 1-7 combined (0.88 mm
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<emphasis italics="true" pageId="0" pageNumber="37">vs</emphasis>
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0.87 mm); antennomere 1 large, subcylindrical, 2-7 moniliform, of similar shape, 8, 9 and 10 each much larger than 7, subquadrate, 11 longest, slightly narrower than 10 and as long as 9 and 10 combined.
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</paragraph>
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<caption doi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="37">
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<emphasis bold="true" pageId="0" pageNumber="37">Figure 2.</emphasis>
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Morphology of
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<taxonomicName authorityName="Yin & Zhao" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias tianmuensis" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="tianmuensis">
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<emphasis italics="true" pageId="0" pageNumber="37">Pseudophanias tianmuensis</emphasis>
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</taxonomicName>
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sp. nov., male
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<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
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head and pronotum
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<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
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maxillary palpus
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<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
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antennal club
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<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
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elytron
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<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
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meso- and metaventrite
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<emphasis bold="true" pageId="0" pageNumber="37">F</emphasis>
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Mesotarsal claws. Scale bars: 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="37">A, D, E</emphasis>
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); 0.1 mm (
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<emphasis bold="true" pageId="0" pageNumber="37">B, F</emphasis>
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); 0.3 mm (
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<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="37">
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Pronotum (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2A</figureCitation>
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) approximately as long as wide, length 0.66 mm, width 0.63 mm, widest anterior middle; sides from widest point greatly convergent anteriorly and subparallel posteriorly; disc distinctly convex to form hump at middle, lacking carina or sulcus; with small asetose median and lateral antebasal foveae; lacking basolateral foveae. Prosternum with anterior part much shorter than coxal part at middle, with small, asetose lateral procoxal foveae; hypomeron lacking ridge that demarcating it from pronotum; margin of coxal cavity non-carinate.
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</paragraph>
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<paragraph pageId="0" pageNumber="37">
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Elytra much wider than long, length 0.96 mm, width 1.24 mm, truncate at bases; each elytron (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2D</figureCitation>
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) with two moderately large, asetose basal foveae; with one discal and one lateral ridge, discal stria formed by broad impressed area between ridges; humeral region lacking subhumeral fovea nor marginal stria; epipleural area broad; posterolateral margin with broad notch. Metathoracic wings fully developed.
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</paragraph>
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<paragraph pageId="0" pageNumber="37">
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Mesoventrite (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2E</figureCitation>
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) with rough surface, with pair of thick admesal carinae, lateral area fully demarcated from metaventrite, openings of median mesoventral foveae widely separated, foveae greatly extended internally, lateral mesoventral foveae unforked, extended internally and overlapped with median foveae; intercoxal process short. Metaventrite (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2E</figureCitation>
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) coarsely punctate, greatly prominent medially and with small crest; with large, asetose lateral mesocoxal foveae; posterior margin roundly emarginate at middle.
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</paragraph>
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<paragraph pageId="0" pageNumber="37">
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Legs elongate; femora coarsely punctate; each tarsus (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Morphology of Pseudophanias tianmuensis sp. nov., male A head and pronotum B maxillary palpus C antennal club D elytron E meso- and metaventrite F Mesotarsal claws. Scale bars: 0.5 mm (A, D, E); 0.1 mm (B, F); 0.3 mm (C)." figureDoi="10.3897/zookeys.1129.95245.figure2" httpUri="https://binary.pensoft.net/fig/768276" pageId="0" pageNumber="37">2F</figureCitation>
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) with one major and one reduced pretarsal claw.
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</paragraph>
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<paragraph pageId="0" pageNumber="37">Abdomen widest at lateral margins of tergite 1 (IV), length 0.86 mm, width 1.15 mm. Tergite 1 (IV) slightly less than twice as long as 2 (V), with broad, deep basal sulcus and two basolateral foveae at lateral ends of sulcus, 2 and 3 (VI) successively shorter, lacking basal sulcus nor fovea, 4 (VII), much longer than 3, narrowed posteriorly, with one pair of basolateral foveae; tergites 1-3 each with pair of quadrangular and 4 with pair of triangular paratergites, 5 (VIII) broadly rounded, with one pair of small basolateral foveae, posterior margin roundly emarginate at middle. Sternite 2 (IV) longest, with densely setose basal sulcus and pair of basolateral foveae at lateral ends of sulcus, 3 (V) and 4 (VI) each short at middle, combined approximately as long as 5 (VII), 3-5 lacking sulcus nor fovea at base, 6 (VIII) transverse, posterior margin slightly protruding at middle, 7 (IX) composed of pair of membranous lamellae.</paragraph>
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<paragraph pageId="0" pageNumber="37">
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Aedeagus (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology and habitat of Pseudophanias tianmuensis sp. nov. A-C aedeagus A dorsal B lateral C ventral D paramere, lateral E general environment of the collecting site F decomposing tree branches inhabited by the beetle. Scale bars: 0.2 mm (A-C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1129.95245.figure3" httpUri="https://binary.pensoft.net/fig/768277" pageId="0" pageNumber="37">3A-C</figureCitation>
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) 0.51 mm long, dorso-ventrally symmetric; median lobe C-shaped in lateral view and broadened in apical half in dorso-ventral view, with large basal capsule and oval foramen; parameres (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology and habitat of Pseudophanias tianmuensis sp. nov. A-C aedeagus A dorsal B lateral C ventral D paramere, lateral E general environment of the collecting site F decomposing tree branches inhabited by the beetle. Scale bars: 0.2 mm (A-C); 0.1 mm (D)." figureDoi="10.3897/zookeys.1129.95245.figure3" httpUri="https://binary.pensoft.net/fig/768277" pageId="0" pageNumber="37">3D</figureCitation>
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) each thin, elongate and curved ventrally in lateral view, with two apical and one preapical macroseta and four short setae near base.
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</paragraph>
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<caption doi="10.3897/zookeys.1129.95245.figure3" httpUri="https://binary.pensoft.net/fig/768277" pageId="0" pageNumber="37" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="37">
|
||
<emphasis bold="true" pageId="0" pageNumber="37">Figure 3.</emphasis>
|
||
Morphology and habitat of
|
||
<taxonomicName authorityName="Yin & Zhao" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias tianmuensis" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="tianmuensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="37">Pseudophanias tianmuensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="37">A-C</emphasis>
|
||
aedeagus
|
||
<emphasis bold="true" pageId="0" pageNumber="37">A</emphasis>
|
||
dorsal
|
||
<emphasis bold="true" pageId="0" pageNumber="37">B</emphasis>
|
||
lateral
|
||
<emphasis bold="true" pageId="0" pageNumber="37">C</emphasis>
|
||
ventral
|
||
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
|
||
paramere, lateral
|
||
<emphasis bold="true" pageId="0" pageNumber="37">E</emphasis>
|
||
general environment of the collecting site
|
||
<emphasis bold="true" pageId="0" pageNumber="37">F</emphasis>
|
||
decomposing tree branches inhabited by the beetle. Scale bars: 0.2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="37">A-C</emphasis>
|
||
); 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="37">D</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="37">
|
||
<emphasis bold="true" pageId="0" pageNumber="37">Female.</emphasis>
|
||
Similar to male in external morphology (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of Pseudophanias tianmuensis sp. nov. A male B female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1129.95245.figure1" httpUri="https://binary.pensoft.net/fig/768275" pageId="0" pageNumber="37">1B</figureCitation>
|
||
). Antenna slightly shorter, club less enlarged (0.77 mm for antennomeres 1-7 vs. 0.82 mm for antennomeres 8-11); each compound eye composed of approximately 43 ommatidia; elytra slightly shorter; metathoracic wings also well developed; metaventrite lacking crest at middle. Measurements (as for male): body length 2.84 mm; length/width of head 0.62/0.49 mm, pronotum 0.63/0.65 mm, elytra 0.91/1.22 mm; abdomen 0.83/1.13 mm; length of antenna 1.59 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="37" type="comparisons">
|
||
<paragraph pageId="0" pageNumber="37">Comparative notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">This species is unique among all known members of the genus in the coarsely punctate body surface, and a rather different type of the male antennal modifications.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="37" type="bionomics">
|
||
<paragraph pageId="0" pageNumber="37">Bionomics.</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">
|
||
The male holotype was collected by sifting
|
||
<taxonomicName class="Pinopsida" family="Cupressaceae" genus="Cryptomeria" kingdom="Plantae" lsidName="Cryptomeria" order="Pinales" pageId="0" pageNumber="37" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="37">Cryptomeria</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName class="Pinopsida" family="Cupressaceae" kingdom="Plantae" lsidName="" order="Pinales" pageId="0" pageNumber="37" phylum="Tracheophyta" rank="family">Cupressaceae</taxonomicName>
|
||
) leaves, while the other two individuals were found inside decomposing wood with termites. Although indicated by a set of morphological traits such as the compact body segments and moniliform antennomeres, it is still inconclusive as to whether the beetle has an obligate association with social insects, pending further observations in the field or at the laboratory.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="37" type="distribution">
|
||
<paragraph pageId="0" pageNumber="37">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">East China: Zhejiang.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="37" type="etymology">
|
||
<paragraph pageId="0" pageNumber="37">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">The specific epithet refers to West Tianmu Mountain, the type locality of this species.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="37" type="remarks">
|
||
<paragraph pageId="0" pageNumber="37">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">
|
||
To accommodate the new species, the key to East and South Asian species of
|
||
<taxonomicName authorityName="Raffray" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="37">Pseudophanias</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.987.53648" author="Inoue, S" journalOrPublisher="ZooKeys" pageId="0" pageNumber="37" pagination="135 - 156" refId="B5" refString="Inoue, S, Nomura, S, Yin, Z-W, 2020. Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys 987: 135 - 156, DOI: https://doi.org/10.3897/zookeys.987.53648" title="Three new species of Pseudophanias Raffray from Japan and Taiwan Island, and synonymy of Chandleriella Hlavac with Pseudophanias (Coleoptera, Staphylinidae, Pselaphinae)." url="https://doi.org/10.3897/zookeys.987.53648" volume="987" year="2020">Inoue et al. (2020)</bibRefCitation>
|
||
is modified as follow:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="37">
|
||
<table inLine="true" pageId="0" pageNumber="37">
|
||
<tr pageId="0" pageNumber="37">
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">0</td>
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">Whole body surface roughly punctate and with dense, short setae; antennal club formed by enlarged, subcylindrical antennomeres 8-11; elytra roundly quadrate, broadly truncate at bases</td>
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="37">
|
||
<taxonomicName authorityName="Yin & Zhao" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudophanias tianmuensis" order="Coleoptera" pageId="0" pageNumber="37" phylum="Arthropoda" rank="species" species="tianmuensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="37">Pseudophanias tianmuensis</emphasis>
|
||
</taxonomicName>
|
||
Yin & Zhao, sp. nov. (East China: Zhejiang)
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="37">
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">
|
||
Body surface usually finely punctate (head with coarse punctation in
|
||
<taxonomicName lsidName="P. excavatus" pageId="0" pageNumber="37" rank="species" species="excavatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="37">P. excavatus</emphasis>
|
||
</taxonomicName>
|
||
Inoue, Nomura & Yin) and with moderately long pubescence; antennal club formed by antennomere 11 alone, or by variously modified antennomeres 5-11, 8-11, or 9-11; elytra roundly triangular, and more strongly constricted at bases
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="37" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="37">1</emphasis>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |