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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.83571" ID-Pensoft-Pub="2747-8106-96-103" ID-Pensoft-UUID="4E0188A4D3D7563C980CB7C5B9424F3C" ID-ZooBank="5B10FFF604074E09B986F931635E9BCC" ModsDocID="2747-8106-96-103" checkinTime="1654860115112" checkinUser="pensoft" docAuthor="Schwarzhans, Werner, Ohe, Fumio, Tsuchiya, Yuki &amp; Ujihara, Atsushi" docDate="2022" docId="8D0F54062A2B5D7890185E7725703F1D" docLanguage="en" docName="Zitteliana 96: 103-134" docOrigin="Zitteliana 96" docPubDate="2022-06-10" docSource="http://dx.doi.org/10.3897/zitteliana.96.83571" docTitle="Diaphus angulatus Ohe &amp; Araki 1973" docType="treatment" docVersion="1" id="4E0188A4D3D7563C980CB7C5B9424F3C" lastPageNumber="103" masterDocId="4E0188A4D3D7563C980CB7C5B9424F3C" masterDocTitle="Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan" masterLastPageNumber="134" masterPageNumber="103" pageNumber="103" updateTime="1654860115112" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Lanternfish otoliths (Myctophidae, Teleostei) from the Miocene of Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schwarzhans, Werner</mods:namePart>
<mods:affiliation>Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark &amp; Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">wwschwarz@aol.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ohe, Fumio</mods:namePart>
<mods:affiliation>Nara National Research Institute for Cultural Properties, Nara 630 - 8577, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tsuchiya, Yuki</mods:namePart>
<mods:affiliation>Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ujihara, Atsushi</mods:namePart>
<mods:affiliation>Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zitteliana</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>96</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>103</mods:start>
<mods:end>134</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zitteliana.96.83571</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.83571</mods:identifier>
<mods:identifier type="Pensoft-Pub">2747-8106-96-103</mods:identifier>
<mods:identifier type="ZooBank">5B10FFF604074E09B986F931635E9BCC</mods:identifier>
<mods:identifier type="Pensoft-UUID">4E0188A4D3D7563C980CB7C5B9424F3C</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:8D0F54062A2B5D7890185E7725703F1D" httpUri="http://treatment.plazi.org/id/8D0F54062A2B5D7890185E7725703F1D" lastPageNumber="103" pageId="0" pageNumber="103">
<subSubSection pageId="0" pageNumber="103" type="nomenclature">
<paragraph pageId="0" pageNumber="103">
<taxonomicName LSID="8D0F5406-2A2B-5D78-9018-5E7725703F1D" authority="Ohe &amp; Araki, 1973" authorityName="Ohe &amp; Araki" authorityYear="1973" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus angulatus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="angulatus">Diaphus angulatus Ohe &amp; Araki, 1973</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">Fig. 8A-F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="reference_group">
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="Ohe &amp; Araki" authorityYear="1973" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus angulatus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="angulatus">Diaphus angulatus</taxonomicName>
1973
<taxonomicName authorityName="Ohe &amp; Araki" authorityYear="1973" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus angulatus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="angulatus">Diaphus angulatus</taxonomicName>
- Ohe &amp; Araki: pl. 49, figs 1-2, 20-21.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="materials_examined">
<paragraph pageId="0" pageNumber="103">Material.</paragraph>
<paragraph pageId="0" pageNumber="103">
<specimenCount type="generic">2 specimens</specimenCount>
;
<specimenCount type="generic">1 specimen</specimenCount>
(SMF PO 101.130)
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Katada FM, Higashibessho FM" location="Nagano River" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:8D0F54062A2B5D7890185E7725703F1D:94221341F7683775DFB9C4BC1E6ED9D9" name="Nagano River">Nagano River</location>
at Inabacho, Tsu City, Mie Prefecture,
<collectorName>Katada FM</collectorName>
, level I 2, late Burdigalian, lower part of planktonic foraminifera biozone N8; 1 small, tentatively assigned specimen (SMF PO 101.131), Osedani, Toyama City, Toyama Prefecture,
<collectorName>Higashibessho FM</collectorName>
, level H 20, late Burdigalian to early Langhian, diatom zone 3A-4A
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="remarks">
<paragraph pageId="0" pageNumber="103">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="Ohe &amp; Araki" authorityYear="1973" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus angulatus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="angulatus">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus angulatus</emphasis>
</taxonomicName>
was described based on a single incomplete specimen from the Yakuoji Formation, a partial lateral equivalent of the Katada Formation (see
<bibRefCitation DOI="https://doi.org/10.5575/geosoc.2018.0032" author="Oshida, H" journalOrPublisher="Journal of the Geological Sciety of Japan" pageId="0" pageNumber="103" pagination="919 - 933" refId="B52" refString="Oshida, H, Hayashi, H, Yanagisawa, Y, Kurihara, Y, Hoshi, H, 2018. Planktonic foraminiferal and diatom biostratigraphy of the upper part of the Miocene Ichishi Group in Mie Prefecture, central Honshu, Japan. Journal of the Geological Sciety of Japan 124: 919 - 933, DOI: https://doi.org/10.5575/geosoc.2018.0032" title="Planktonic foraminiferal and diatom biostratigraphy of the upper part of the Miocene Ichishi Group in Mie Prefecture, central Honshu, Japan." url="https://doi.org/10.5575/geosoc.2018.0032" volume="124" year="2018">Oshida et al. 2018</bibRefCitation>
). Due to the incomplete preservation,
<taxonomicName genus="D." lsidName="D. angulatus" pageId="0" pageNumber="103" rank="species" species="angulatus">
<emphasis italics="true" pageId="0" pageNumber="103">D. angulatus</emphasis>
</taxonomicName>
was considered a doubtful species by
<bibRefCitation author="Nolf, D" journalOrPublisher="Royal Belgian Institute of Natural Sciences, Brussels" pageId="0" pageNumber="103" refId="B43" refString="Nolf, D, 2013. The diversity of fish otoliths, past and present. Royal Belgian Institute of Natural Sciences, Brussels" title="The diversity of fish otoliths, past and present." year="2013">Nolf (2013)</bibRefCitation>
. Now, we have found a second specimen from the Katada FM which perfectly resembles the holotype but again lacks a rostrum (size 4.3+ mm; Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8A-C</figureCitation>
). A complete smaller specimen of 2.6 mm in length from the Higashibessho FM (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8D-F</figureCitation>
) somewhat deviates from the larger specimens in the shape of the posterior rim and is therefore only tentatively assigned to the species. This difference is here considered to reflect an ontogenetic effect. These new finds are here used to revalidate and redefine the species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="diagnosis">
<paragraph pageId="0" pageNumber="103">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">(emended).</emphasis>
OL:OH = 1.2-1.3. Ventral rim regularly curved with 6 denticles of variable strength. Rostrum long, massive; antirostrum and excisura small. Dorsal rim with prominent postdorsal angle; predorsal portion depressed. Inner face flat. Caudal colliculum shorter and much narrower than ostial colliculum (OCL:CCL = 1.5-1.8; OCH:CCH = 1.6-1.8).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">Description</paragraph>
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">(emended).</emphasis>
Large, thin otoliths reaching at least 5.5 mm in length (holotype recorded by
<bibRefCitation author="Ohe, F" journalOrPublisher="Science reports of the Tohoku University, second series (Geology)" pageId="0" pageNumber="103" pagination="1 - 15" refId="B49" refString="Ohe, F, Araki, Y, 1973. Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture. Science reports of the Tohoku University, second series (Geology) 38: 1 - 15" title="Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture." volume="38" year="1973">Ohe and Araki 1973</bibRefCitation>
); OH:OT = 4.0 (4.5 in tentatively assigned specimen). Rostrum long, pointed, 20% of OL in tentatively assigned small specimen with completely preserved rostrum (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8D</figureCitation>
). Antirostrum distinct, but very short; excisura small. Ventral rim relatively shallow and regularly curved, with 6 denticles of varying strength. Dorsal rim smooth, anteriorly depressed, posteriorly with prominent postdorsal angle positioned above termination of cauda. Posterior rim obliquely cut in large specimens (holotype figured by
<bibRefCitation author="Ohe, F" journalOrPublisher="Science reports of the Tohoku University, second series (Geology)" pageId="0" pageNumber="103" pagination="1 - 15" refId="B49" refString="Ohe, F, Araki, Y, 1973. Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture. Science reports of the Tohoku University, second series (Geology) 38: 1 - 15" title="Some Miocene fish otoliths from the Yakuoji Formation, Tsu, Mie Prefecture." volume="38" year="1973">Ohe and Araki 1973</bibRefCitation>
, and Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8A</figureCitation>
), rounded and without postventral angle in tentatively assigned small specimen of Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A-F: Diaphus angulatus Ohe &amp; Araki, 1973; A-C: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305; D-E: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20; G-N: Diaphus epipedus sp. nov.; G-I: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15; K-J: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15; L-N: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21; O-T: Diaphus metopoclampoides Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f; U-AB: Diaphus watatsumi sp. nov., Tsuyama City, Takakura Formation; U-X: holotype, SMF PO 101.136, level MS 52; Y-AB: paratypes, SMF PO 101.137, levels gh, j; AC-AE: Diaphus sp. 1, col. Tsuchiya, Ichishi Group." figureDoi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103">8D</figureCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="103">Inner face flat with distinctly supramedian sulcus. Ostial colliculum about twice as long and wide as caudal colliculum; caudal pseudocolliculum long and distinct. Dorsal margin of ostium straight; cauda slightly bent upwards. Ventral furrow distinct, moderately close to ventral rim of otolith; dorsal depression large, rather deep but with indistinct margins except for crista superior towards sulcus. Outer face slightly convex with shallow postcentral umbo, smooth.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="discussion">
<paragraph pageId="0" pageNumber="103">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="103">
A typical and rather large otolith-based species of the
<taxonomicName authorityName="Eigenmann &amp; Eigenmann" authorityYear="1890" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus theta" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="theta">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus theta</emphasis>
</taxonomicName>
Otolith Group that resembles the coeval
<taxonomicName genus="D." lsidName="D. haereticus" pageId="0" pageNumber="103" rank="species" species="haereticus">
<emphasis italics="true" pageId="0" pageNumber="103">D. haereticus</emphasis>
</taxonomicName>
(Brzobohatý &amp; Schultz, 1978) but differs in the far more backward positioned postdorsal angle and the much narrower caudal colliculum. In the latter aspect, it resembles
<taxonomicName genus="D." lsidName="D. audax" pageId="0" pageNumber="103" rank="species" species="audax">
<emphasis italics="true" pageId="0" pageNumber="103">D. audax</emphasis>
</taxonomicName>
Schwarzhans, 2019 from the early Miocene of New Zealand and Chile but again differs in the distinctly further backward positioned postdorsal angle and the complete lack of a postdorsal concavity.
</paragraph>
<caption doi="10.3897/zitteliana.96.83571.figure8" httpUri="https://binary.pensoft.net/fig/694121" pageId="0" pageNumber="103" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 8.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="103">A-F</emphasis>
:
<taxonomicName authorityName="Ohe &amp; Araki" authorityYear="1973" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus angulatus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="angulatus">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus angulatus</emphasis>
</taxonomicName>
Ohe &amp; Araki, 1973;
<emphasis bold="true" pageId="0" pageNumber="103">A-C</emphasis>
: SMF PO 101.130, Inabacho, Tsu City, Katada Formation, level ML 2305;
<emphasis bold="true" pageId="0" pageNumber="103">D-E</emphasis>
: SMF PO 101.131, Fuchu machi, Toyama City, Higashibessho Formation, level H 20;
<emphasis bold="true" pageId="0" pageNumber="103">G-N</emphasis>
:
<taxonomicName authorityName="Schwarzhans &amp; Ohe &amp; Tsuchiya &amp; Ujihara" authorityYear="2022" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus epipedus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="epipedus">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus epipedus</emphasis>
</taxonomicName>
sp. nov.;
<emphasis bold="true" pageId="0" pageNumber="103">G-I</emphasis>
: holotype, SMF PO 101.132, Tsuyama City, Takakura Formation, level MS 15;
<emphasis bold="true" pageId="0" pageNumber="103">K-J</emphasis>
: paratype, SMF PO 101.133, Tsuyama City, Takakura Formation, level MS 15;
<emphasis bold="true" pageId="0" pageNumber="103">L-N</emphasis>
: paratype, SMF PO 101.134, Yatsuo, Toyama City, Higashibessho Formation, level H 21;
<emphasis bold="true" pageId="0" pageNumber="103">O-T</emphasis>
:
<taxonomicName authorityName="Steurbaut" authorityYear="1983" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus metopoclampoides" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="metopoclampoides">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus metopoclampoides</emphasis>
</taxonomicName>
Steurbaut, 1983, SMF PO 101.135, Tsuyama City, Takakura Formation level MS 06, f;
<emphasis bold="true" pageId="0" pageNumber="103">U-AB</emphasis>
:
<taxonomicName authorityName="Schwarzhans &amp; Ohe &amp; Tsuchiya &amp; Ujihara" authorityYear="2022" class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus watatsumi" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="species" species="watatsumi">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus watatsumi</emphasis>
</taxonomicName>
sp. nov., Tsuyama City, Takakura Formation;
<emphasis bold="true" pageId="0" pageNumber="103">U-X</emphasis>
: holotype, SMF PO 101.136, level MS 52;
<emphasis bold="true" pageId="0" pageNumber="103">Y-AB</emphasis>
: paratypes, SMF PO 101.137, levels gh, j;
<emphasis bold="true" pageId="0" pageNumber="103">AC-AE</emphasis>
:
<taxonomicName class="Actinopterygii" family="Myctophidae" genus="Diaphus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diaphus" order="Myctophiformes" pageId="0" pageNumber="103" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Diaphus</emphasis>
</taxonomicName>
sp.1, col. Tsuchiya, Ichishi Group.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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