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<document ID-DOI="http://doi.org/10.5281/zenodo.3748525" ID-GBIF-Dataset="6384b520-7e9f-4874-a414-76c2e9b01d74" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="3748525" approvalRequired="1" checkinTime="1586358796837" checkinUser="plazi" docAuthor="Burgin, Connor" docDate="2019" docId="885887A2FFD18A37FF65F12CF865DF82" docLanguage="en" docName="hbmw_9_Rhinolophidae.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docStyle="DocumentStyle{}" docTitle="Rhinolophus nereis K. Andersen 1905" docType="treatment" docVersion="22" lastPageNumber="309" masterDocId="7461FFDAFFCF8A29FFCCFFA1FF85D963" masterDocTitle="Rhinolophidae" masterLastPageNumber="332" masterPageNumber="280" pageNumber="309" updateTime="1658949228820" updateUser="felipe">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhinolophidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Burgin, Connor</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:originInfo>
<mods:dateIssued>2019</mods:dateIssued>
<mods:dateOther type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher>Lynx Edicions</mods:publisher>
<mods:place>
<mods:placeTerm>Barcelona</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Wilson, Don E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Editor</mods:roleTerm>
</mods:role>
<mods:namePart>Mittermeier, Russell A.</mods:namePart>
</mods:name>
<mods:titleInfo>
<mods:title>Handbook of the Mammals of the World Volume 9 Bats</mods:title>
</mods:titleInfo>
<mods:part>
<mods:extent unit="page">
<mods:start>280</mods:start>
<mods:end>332</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>book chapter</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3748525</mods:identifier>
<mods:identifier type="GBIF-Dataset">6384b520-7e9f-4874-a414-76c2e9b01d74</mods:identifier>
<mods:identifier type="ISBN">978-84-16728-19-0</mods:identifier>
<mods:identifier type="Zenodo-Dep">3748525</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3808898" ID-GBIF-Taxon="163549958" ID-Zenodo-Dep="3808898" LSID="urn:lsid:plazi:treatment:885887A2FFD18A37FF65F12CF865DF82" httpUri="http://treatment.plazi.org/id/885887A2FFD18A37FF65F12CF865DF82" lastPageNumber="309" pageId="30" pageNumber="309">
<subSubSection box="[169,982,3725,3786]" pageId="30" pageNumber="309" type="type_taxon">
<paragraph blockId="30.[169,1523,3725,3786]" box="[169,982,3725,3786]" pageId="30" pageNumber="309">
<figureCitation box="[169,240,3725,3786]" captionStart="On" captionStartId="26.[175,218,4332,4359]" captionTargetBox="[136,3490,134,4724]" captionTargetPageId="25" captionText="On following pages: 52. Eastern Horseshoe Bat (Rhinolophus megaphyllus}; 53. Insular Horseshoe Bat (Rhinolophus keyensis}; 54. Yellow-faced Horseshoe Bat (Rhinolophus virgo}; 55. Madura Horseshoe Bat (Rhinolophus madurensis}; 56. Sulawesi Horseshoe Bat (Rhinolophus celebensis}; 57. Robinson's Horseshoe Bat (Rhinolophus robinsoni); 58. Indochinese Horseshoe Bat (Rhinolophus chasem); 59. Anamban Horseshoe Bat (Rhinolophus nereis}; 60. Bornean Horseshoe Bat (Rhinolophus borneensis); 61. Malayan Horseshoe Bat (Rhinolophus malayanus}; 62. Acuminate Horseshoe Bat (Rhinolophus acuminatus}; 63. Acuminate Horseshoe Bat (Rhinolophus acuminatus}; 64. Little Japanese Horseshoe Bat (Rhinolophus cornutus}; 65. Yaeyama Horseshoe Bat (Rhinolophus perditus}; 66. Mountain Horseshoe Bat (Rhinolophus monticolus}; 67. Shortridge's Horseshoe Bat (Rhinolophus shortridgei}; 68. Formosan Horseshoe Bat (Rhinolophus monoceros}; 69. Least Horseshoe Bat (Rhinolophus pusillus}; 70. Little Nepalese Horseshoe Bat (Rhinolophus subbadius}; 71. Convex Horseshoe Bat (Rhinolophus convexus}; 72. Andaman Horseshoe Bat (Rhinolophus cognatus}; 73. Glossy Horseshoe Bat (Rhinolophus refulgens}; 74. Blyth's Horseshoe Bat (Rhinolophus lepidus};" figureDoi="http://doi.org/10.5281/zenodo.3758605" httpUri="https://zenodo.org/record/3758605/files/figure.png" pageId="30" pageNumber="309">59</figureCitation>
. Anamban Horseshoe
<emphasis bold="true" box="[887,982,3725,3786]" pageId="30" pageNumber="309">Bat</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[1047,1523,3725,3786]" pageId="30" pageNumber="309" type="nomenclature">
<paragraph blockId="30.[169,1523,3725,3786]" box="[1047,1523,3725,3786]" pageId="30" pageNumber="309">
<taxonomicName authorityName="K. Andersen" authorityYear="1905" box="[1047,1523,3725,3786]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="30" pageNumber="309" phylum="Chordata" rank="species" species="nereis">
<emphasis box="[1047,1523,3725,3786]" italics="true" pageId="30" pageNumber="309">Rhinolophus nereis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="vernacular_names">
<paragraph blockId="30.[168,1664,3799,3880]" box="[169,1664,3799,3826]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[169,270,3799,3826]" pageId="30" pageNumber="309">French:</emphasis>
<emphasis box="[283,433,3799,3826]" italics="true" pageId="30" pageNumber="309">Rhinolophe</emphasis>
des Anambas /
<emphasis bold="true" box="[652,775,3799,3826]" pageId="30" pageNumber="309">German:</emphasis>
<emphasis box="[783,1213,3799,3826]" italics="true" pageId="30" pageNumber="309">Anambas-lnseln-Hufeisennase I</emphasis>
<emphasis bold="true" box="[1221,1344,3799,3826]" pageId="30" pageNumber="309">Spanish:</emphasis>
Herradura de Anambas
</paragraph>
<paragraph blockId="30.[168,1664,3799,3880]" box="[168,807,3853,3880]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[168,501,3853,3880]" pageId="30" pageNumber="309">
Other
<emphasis bold="true" box="[260,387,3853,3880]" italics="true" pageId="30" pageNumber="309">common</emphasis>
names:
</emphasis>
<emphasis box="[512,755,3853,3880]" italics="true" pageId="30" pageNumber="309">Neriad Horseshoe</emphasis>
Bat
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="reference_group">
<paragraph blockId="30.[989,1784,3950,4518]" pageId="30" pageNumber="309">
Taxonomy.
<taxonomicName authority="K. Andersen, 1905" authorityName="K. Andersen" authorityYear="1905" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="30" pageNumber="309" phylum="Chordata" rank="species" species="nereis">
<emphasis box="[1211,1532,3950,3989]" italics="true" pageId="30" pageNumber="309">Rhinolophus nereis</emphasis>
K. Andersen, 1905
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="materials_examined">
<paragraph blockId="30.[989,1784,3950,4518]" pageId="30" pageNumber="309">
<materialsCitation ID-GBIF-Occurrence="2597534007" country="Indonesia" location="Pulo [= Island] Siantan, Anamba Group [= Islands]" pageId="30" pageNumber="309" specimenCount="1" typeStatus="type">
<location LSID="urn:lsid:plazi:treatment:885887A2FFD18A37FF65F12CF865DF82:052E606FFFD18A37FBA5F003FAB6D69D" country="Indonesia" name="Pulo [= Island] Siantan, Anamba Group [= Islands]" pageId="30" pageNumber="309">
Pulo [= Island] Siantan, Anamba Group [=
<emphasis box="[1179,1320,4055,4094]" italics="true" pageId="30" pageNumber="309">Islands</emphasis>
]
</location>
,”
<collectingCountry box="[1369,1558,4055,4094]" name="Indonesia" pageId="30" pageNumber="309">Indonesia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="discussion">
<paragraph blockId="30.[989,1784,3950,4518]" pageId="30" pageNumber="309">
<emphasis box="[989,1231,4109,4148]" italics="true" pageId="30" pageNumber="309">Traditionally</emphasis>
included in the
<emphasis box="[1577,1784,4109,4148]" italics="true" pageId="30" pageNumber="309">megaphyUus</emphasis>
species group, but this has
<emphasis box="[1489,1546,4161,4200]" italics="true" pageId="30" pageNumber="309">yet</emphasis>
to be tested. The species has been included in
<emphasis box="[1642,1783,4214,4253]" italics="true" pageId="30" pageNumber="309">
A
<emphasis box="[992,1088,4267,4306]" italics="true" pageId="30" pageNumber="309">bomeensis</emphasis>
</emphasis>
but is considered distinct, based on its larger size. Monotypic.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="distribution">
<caption ID-DOI="http://doi.org/10.5281/zenodo.3750024" ID-Zenodo-Dep="3750024" httpUri="https://zenodo.org/record/3750024/files/figure.png" inLine="true" pageId="30" pageNumber="309" targetBox="[167,959,3953,4512]" targetPageId="30">
<paragraph blockId="30.[989,1784,3950,4518]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[990,1227,4373,4412]" pageId="30" pageNumber="309">Distribution.</emphasis>
Known only from Siantan I (Anambas Is) and North Natuna (Natuna Is),
<collectingCountry box="[1063,1254,4479,4518]" name="Indonesia" pageId="30" pageNumber="309">Indonesia</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="description">
<paragraph blockId="30.[167,1781,4530,4624]" lastBlockId="30.[1871,3496,345,1763]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[167,508,4530,4569]" pageId="30" pageNumber="309">Descriptive notes.</emphasis>
Tail
<quantity box="[616,751,4530,4569]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="30" pageNumber="309" unit="mm" value="17.0">17 mm</quantity>
, hindfoot 9-
<quantity box="[995,1110,4530,4569]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="309" unit="mm" value="3.0">3 mm</quantity>
, forearm 43-
<quantity box="[1363,1480,4530,4569]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="309" unit="mm" value="5.0">5 mm</quantity>
. The Anamban Horseshoe
<emphasis bold="true" box="[386,448,4585,4624]" pageId="30" pageNumber="309">Bat</emphasis>
is similar to the Bornean Horseshoe Bat (A
<emphasis box="[1317,1515,4585,4624]" italics="true" pageId="30" pageNumber="309">
<taxonomicName box="[1317,1494,4585,4624]" class="Mammalia" family="Rhinolophidae" genus="Rhinolophus" kingdom="Animalia" order="Chiroptera" pageId="30" pageNumber="309" phylum="Chordata" rank="species" species="bomeensis">bomeensis</taxonomicName>
),
</emphasis>
although it is larger overall, and has different wing shape, where third finger has lengthened
<emphasis box="[3426,3494,345,384]" italics="true" pageId="30" pageNumber="309">second</emphasis>
phalanx (more than 150% of first phalanx, as opposed to less than 150% in the Bornean Horseshoe Bat). Pelage of
<typeStatus box="[2586,2751,451,490]" pageId="30" pageNumber="309">holotype</typeStatus>
specimen is apparently “mars-brown” dorsally (base of hairs “ecru-drab”), while ventral aspect is strangely drab yellow (
<emphasis box="[3387,3493,505,544]" italics="true" pageId="30" pageNumber="309">possibly</emphasis>
due to staining from formalin preservation); no orange morph has been recorded. Ears are medium in size. Noseleaf lancet is
<emphasis box="[2683,2852,612,651]" italics="true" pageId="30" pageNumber="309">relatively</emphasis>
short and straight-sided;
<emphasis box="[3330,3493,612,651]" italics="true" pageId="30" pageNumber="309">connecting</emphasis>
process seems to be high, hairy and rounded, connecting approximately halfway up lancet; sella is separated from intemarial cup by sudden deep constriction, and its sides are
<emphasis box="[2047,2181,771,810]" italics="true" pageId="30" pageNumber="309">slightly</emphasis>
converged above middle, being rounded at top; horseshoe is wide at
<quantity box="[1873,1984,824,863]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="309" unit="mm" value="9.0">9 mm</quantity>
, has shallow median emargination, and lateral leaflets visible from above. Lower lip has three well defined mental grooves. Skull is of medium build (zygomatic width is longer than mastoid width); anterior and lateral nasal swellings are moderately developed while posterior compartments are less inflated; sagittal crest is moderately defined; frontal depression is well developed; supraorbital crests are sharp and well defined. P2 is situated nearly in axis of tooth row and has nearly imperceptible cusp; C1 and P4 come into contact; the only examinable specimen is missing the mandible.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="biology_ecology">
<paragraph blockId="30.[1871,3496,345,1763]" box="[1873,2590,1198,1237]" pageId="30" pageNumber="309">Habitat. Presumably forested habitats.</paragraph>
<paragraph blockId="30.[1871,3496,345,1763]" box="[1873,2545,1251,1290]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[1873,2226,1251,1290]" pageId="30" pageNumber="309">Food and Feeding.</emphasis>
No information.
</paragraph>
<paragraph blockId="30.[1871,3496,345,1763]" box="[1873,2374,1304,1343]" pageId="30" pageNumber="309">Breeding. No information.</paragraph>
<paragraph blockId="30.[1871,3496,345,1763]" box="[1871,3127,1356,1395]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[1871,2190,1356,1395]" pageId="30" pageNumber="309">Activity patterns.</emphasis>
The Anamban Horseshoe Bat may roost in caves.
</paragraph>
<paragraph blockId="30.[1871,3496,345,1763]" box="[1873,3137,1409,1448]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[1873,2817,1409,1448]" pageId="30" pageNumber="309">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="conservation">
<paragraph blockId="30.[1871,3496,345,1763]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[1874,2069,1461,1500]" pageId="30" pageNumber="309">Status and</emphasis>
Conservation. Classified as Data Deficient on
<emphasis box="[2938,3270,1461,1500]" italics="true" pageId="30" pageNumber="309">The JCN ed List.</emphasis>
The Anamban Horseshoe Bat is known from only two localities on different islands and virtually nothing is known about the species. Further research into the species ecology, taxonomy, and potential threats is needed.
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="309" type="reference_group">
<paragraph blockId="30.[1871,3496,345,1763]" pageId="30" pageNumber="309">
<emphasis bold="true" box="[1875,2083,1679,1706]" pageId="30" pageNumber="309">Bibliography.</emphasis>
Andersen (1905), Bates, Bumrungsri, Walston &amp; Csorba (2016), Corbet &amp; Hill (1992), Csorba
<emphasis box="[3466,3496,1679,1706]" italics="true" pageId="30" pageNumber="309">et</emphasis>
al. (2003).
</paragraph>
</subSubSection>
</treatment>
</document>