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<document id="678348BAEDCC951881A20AC8A709787C" ID-DOI="10.11646/zootaxa.5351.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391297" ID-ZooBank="E43B08B1-4DC0-4327-B239-4F47366A4D67" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695988998153" checkinUser="plazi" docAuthor="Bravo, Freddy, Lopes, Priscila Silva &amp; Araújo, Maíra Xavier" docDate="2023" docId="2565122B0174FFCF03B8FE59FE91D61B" docLanguage="en" docName="zootaxa.5351.1.6.pdf" docOrigin="Zootaxa 5351 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coronamyia Bravo &amp; Lopes &amp; Araújo 2023, gen. nov." docType="treatment" docVersion="3" lastPageNumber="141" masterDocId="D95C6A530175FFCC032FFF8AFF87D75A" masterDocTitle="Coronamyia, a new genus of Neotropical Maruinini (Diptera, Psychodidae) with the description of four new species" masterLastPageNumber="150" masterPageNumber="139" pageNumber="140" updateTime="1695990269332" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A57136B357861EB7A9CBD4D61CEE7E3F">Coronamyia, a new genus of Neotropical Maruinini (Diptera, Psychodidae) with the description of four new species</mods:title>
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<mods:namePart id="730189184555BB3E7D88AEEC4CFBE6FC">Bravo, Freddy</mods:namePart>
<mods:affiliation id="C6B01E95C655BAED9E575F9ED1023FAD">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Feira de Santana, Bahia, Brazil</mods:affiliation>
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<mods:namePart id="C7D876CC9655D5F74B3D583AF0D71257">Lopes, Priscila Silva</mods:namePart>
<mods:affiliation id="9A8FD119309B23358B753529E30B7744">Universidade Federal da Bahia, Instituto de Ciências Biológicas, Salvador, Bahia, Brazil</mods:affiliation>
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<mods:namePart id="75B644D26A0CC41028333F7D1303E523">Araújo, Maíra Xavier</mods:namePart>
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<mods:affiliation id="14F7E5E93506F1985F987CC86561D9EE">Universidade Estadual de Feira de Santana, Departamento de Ciências Biológicas, Feira de Santana, Bahia, Brazil &amp; mah. biology @ gmail. com; https: // orcid. org / 0000 - 0003 - 2447 - 9901</mods:affiliation>
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<paragraph id="AD73A33D0174FFCD03B8FE59FE25D6B7" blockId="1.[151,418,467,493]" box="[151,418,467,493]" pageId="1" pageNumber="140">
<heading id="F63B14510174FFCD03B8FE59FE25D6B7" bold="true" box="[151,418,467,493]" fontSize="11" level="1" pageId="1" pageNumber="140" reason="1">
<emphasis id="9FB87F2F0174FFCD03B8FE59FE25D6B7" bold="true" box="[151,418,467,493]" pageId="1" pageNumber="140">
<taxonomicName id="6ACCD8BE0174FFCD03B8FE59FEAAD6B7" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[151,301,467,493]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="9FB87F2F0174FFCD03B8FE59FEAAD6B7" bold="true" box="[151,301,467,493]" italics="true" pageId="1" pageNumber="140">Coronamyia</emphasis>
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<taxonomicNameLabel id="848BC2540174FFCD021BFE59FE25D6B7" box="[308,418,467,493]" pageId="1" pageNumber="140" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="AD73A33D0174FFCD03B8FD9DFCD8D575" blockId="1.[151,863,535,559]" box="[151,863,535,559]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03B8FD9DFEA1D575" bold="true" box="[151,294,535,559]" pageId="1" pageNumber="140">
<typeStatus id="72771D9F0174FFCD03B8FD9DFF4BD575" box="[151,204,535,559]" pageId="1" pageNumber="140">Type</typeStatus>
species:
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<taxonomicName id="6ACCD8BE0174FFCD0203FD92FD94D575" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[300,531,535,559]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="paschoali" status="sp. nov.">
<emphasis id="9FB87F2F0174FFCD0203FD92FD94D575" box="[300,531,535,559]" italics="true" pageId="1" pageNumber="140">Coronamyia paschoali</emphasis>
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<taxonomicNameLabel id="848BC2540174FFCD0136FD92FDEDD575" box="[537,618,536,559]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
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<subSubSection id="E5D6F0B60174FFCD03B8FDD7FE7FD5C1" pageId="1" pageNumber="140" type="etymology">
<paragraph id="AD73A33D0174FFCD03B8FDD7FE7FD5C1" blockId="1.[151,1437,605,2036]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03B8FDD7FE99D52D" bold="true" box="[151,286,605,631]" pageId="1" pageNumber="140">Etymology.</emphasis>
From Latin, Corona, meaning crown, and Greek, Myia, meaning fly, referring to the setous ring around the aedeagus. Feminine gender.
</paragraph>
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<subSubSection id="E5D6F0B60174FFCD03E8FD2FFDEAD4CD" pageId="1" pageNumber="140" type="diagnosis">
<paragraph id="AD73A33D0174FFCD03E8FD2FFDEAD4CD" blockId="1.[151,1437,605,2036]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03E8FD2FFEC5D5E5" bold="true" box="[199,322,677,703]" pageId="1" pageNumber="140">Diagnosis.</emphasis>
Wing with R 
<subScript id="3148A1780174FFCD02F7FD32FE67D59C" attach="left" box="[472,480,696,710]" fontSize="6" pageId="1" pageNumber="140">5</subScript>
ending in a pointed wing apex. Thoracic sensory area with microtrichia in the dorsal area of anepimeron; small circular thoracic gland near the thoracic sensory area with microtrichia. Aedeagus with a lightly sclerotized plate with long, thin setae surrounding it. Gonostyli glabrous, sclerotized, bifurcated, shorter than the gonocoxites, internal arm curved, external arm with basal lobes of different shapes. The hypopod is elongated and lacks an apical tenaculum. It has two groups of accessory tenacula dorsally: a group of short accessories tenacula at the base of the hypopod and one or two strips of long accessory geniculate tenacula in the remaining part of the hypopod, each inserted in a larger alveoli.
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<subSubSection id="E5D6F0B60174FFCD03E8FC2BFE19D039" pageId="1" pageNumber="140" type="description">
<paragraph id="AD73A33D0174FFCD03E8FC2BFB5CD01A" blockId="1.[151,1437,605,2036]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03E8FC2BFEDED4E1" bold="true" box="[199,345,929,955]" pageId="1" pageNumber="140">Description.</emphasis>
male. Head: semicircular eye bridge with three or four rows of facets; eyes separate by 0.2 to 3.0 facet diameter; interocular suture as an inverted V-shaped; frontal patch of alveoli with anterior margin straight and posterior margin bilobed (
<figureCitation id="35F7BFB80174FFCD02C1FC63FDD4D359" box="[494,595,1001,1027]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1A</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD014FFC63FD04D359" box="[608,643,1001,1027]" captionStart="FIGURE 3" captionStartId="4.[152,255,1937,1961]" captionTargetBox="[170,1417,805,1914]" captionTargetId="figure-260@4.[170,1417,805,1914]" captionTargetPageId="4" captionText="FIGURE 3. Coronamyia santosi sp. nov.: (A): Head; (B): Labrum, anterior view, photograph (C): Labrum, anterior view, drawing, (D): Labium, posterior view, drawing; Labium, posterior view, photograph; (F) Antenna and palpus; (G) Palpus. Abbreviations: lbm: labrum, sfs: small spiniform seta, sps: spiniform seta." figureDoi="http://doi.org/10.5281/zenodo.8391307" httpUri="https://zenodo.org/record/8391307/files/figure.png" pageId="1" pageNumber="140">3B</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD01BFFC63FD32D359" box="[656,693,1001,1027]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6A</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD01EEFC60FD61D35E" box="[705,742,1002,1028]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7A</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD01DDFC63FC9FD359" box="[754,792,1001,1027]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8A</figureCitation>
); labrum subtriangular, dorsally with pointed apex (
<figureCitation id="35F7BFB80174FFCD064BFC63FF5BD37D" captionStart="FIGURE 3" captionStartId="4.[152,255,1937,1961]" captionTargetBox="[170,1417,805,1914]" captionTargetId="figure-260@4.[170,1417,805,1914]" captionTargetPageId="4" captionText="FIGURE 3. Coronamyia santosi sp. nov.: (A): Head; (B): Labrum, anterior view, photograph (C): Labrum, anterior view, drawing, (D): Labium, posterior view, drawing; Labium, posterior view, photograph; (F) Antenna and palpus; (G) Palpus. Abbreviations: lbm: labrum, sfs: small spiniform seta, sps: spiniform seta." figureDoi="http://doi.org/10.5281/zenodo.8391307" httpUri="https://zenodo.org/record/8391307/files/figure.png" pageId="1" pageNumber="140">Figs. 3B, C</figureCitation>
); labella bulbous with many setae; epipharynx with two spiniform setae dorsal to labial furca and at least 10 small spiniform setae in the middle of its surface (
<figureCitation id="35F7BFB80174FFCD01E8FBBBFCCDD311" box="[711,842,1073,1099]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1B, C</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0079FBBBFC1ED316" box="[854,921,1073,1100]" captionStart="FIGURE 3" captionStartId="4.[152,255,1937,1961]" captionTargetBox="[170,1417,805,1914]" captionTargetId="figure-260@4.[170,1417,805,1914]" captionTargetPageId="4" captionText="FIGURE 3. Coronamyia santosi sp. nov.: (A): Head; (B): Labrum, anterior view, photograph (C): Labrum, anterior view, drawing, (D): Labium, posterior view, drawing; Labium, posterior view, photograph; (F) Antenna and palpus; (G) Palpus. Abbreviations: lbm: labrum, sfs: small spiniform seta, sps: spiniform seta." figureDoi="http://doi.org/10.5281/zenodo.8391307" httpUri="https://zenodo.org/record/8391307/files/figure.png" pageId="1" pageNumber="140">3D, E</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD008AFBBBFC4ED311" box="[933,969,1073,1099]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7B</figureCitation>
). Antenna with scape longer than pedicel, subcylindrical; pedicel almost spherical; flagellum with 14 fusiform flagellomeres decreasing in size from base to apex; apiculus cylindrical, approximately 0.5 the length of the last flagellomere; ascoids not visible (
<figureCitation id="35F7BFB80174FFCD0624FBF3FAEBD3C9" box="[1291,1388,1145,1171]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1E</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0655FBF3FA10D3C9" box="[1402,1431,1145,1171]" captionStart="FIGURE 3" captionStartId="4.[152,255,1937,1961]" captionTargetBox="[170,1417,805,1914]" captionTargetId="figure-260@4.[170,1417,805,1914]" captionTargetPageId="4" captionText="FIGURE 3. Coronamyia santosi sp. nov.: (A): Head; (B): Labrum, anterior view, photograph (C): Labrum, anterior view, drawing, (D): Labium, posterior view, drawing; Labium, posterior view, photograph; (F) Antenna and palpus; (G) Palpus. Abbreviations: lbm: labrum, sfs: small spiniform seta, sps: spiniform seta." figureDoi="http://doi.org/10.5281/zenodo.8391307" httpUri="https://zenodo.org/record/8391307/files/figure.png" pageId="1" pageNumber="140">3F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03B8FB14FF58D3ED" box="[151,223,1181,1208]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7A, C</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03C3FB17FE95D3ED" box="[236,274,1181,1207]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8A</figureCitation>
). Palpus approximately 1/3 the length of flagellum with four segments, the last segment not annulate (
<figureCitation id="35F7BFB80174FFCD03B0FB4BFF7CD381" box="[159,251,1217,1243]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD022AFB4BFEADD381" box="[261,298,1217,1243]" captionStart="FIGURE 3" captionStartId="4.[152,255,1937,1961]" captionTargetBox="[170,1417,805,1914]" captionTargetId="figure-260@4.[170,1417,805,1914]" captionTargetPageId="4" captionText="FIGURE 3. Coronamyia santosi sp. nov.: (A): Head; (B): Labrum, anterior view, photograph (C): Labrum, anterior view, drawing, (D): Labium, posterior view, drawing; Labium, posterior view, photograph; (F) Antenna and palpus; (G) Palpus. Abbreviations: lbm: labrum, sfs: small spiniform seta, sps: spiniform seta." figureDoi="http://doi.org/10.5281/zenodo.8391307" httpUri="https://zenodo.org/record/8391307/files/figure.png" pageId="1" pageNumber="140">3G</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD021BFB4BFEDED381" box="[308,345,1217,1243]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6A</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD024CFB4BFE0ED381" box="[355,393,1217,1243]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7A</figureCitation>
). Thorax: anepisternum and laterotergite with patch of alveoli, other pleuron sclerites glabrous (
<figureCitation id="35F7BFB80174FFCD03B0FB6FFF7ED3A5" box="[159,249,1253,1279]" captionStart="FIGURE 4" captionStartId="5.[152,255,1441,1465]" captionTargetBox="[170,1416,489,1418]" captionTargetId="figure-159@5.[170,1416,489,1418]" captionTargetPageId="5" captionText="FIGURE 4. Coronamyia santosi sp. nov.: (A) Thorax, lateral view; (B) Thorax with the thoracic gland and sensory area with microtrichia; (C) Wing; (D) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391309" httpUri="https://zenodo.org/record/8391309/files/figure.png" pageId="1" pageNumber="140">Fig. 4A</figureCitation>
); coxa II with mesal anterior protuberance covered with pores. Wing (
<figureCitation id="35F7BFB80174FFCD072DFB6FFBE3D3A5" box="[1026,1124,1253,1279]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1G</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD075FFB6FFB14D3A5" box="[1136,1171,1253,1279]" captionStart="FIGURE 4" captionStartId="5.[152,255,1441,1465]" captionTargetBox="[170,1416,489,1418]" captionTargetId="figure-159@5.[170,1416,489,1418]" captionTargetPageId="5" captionText="FIGURE 4. Coronamyia santosi sp. nov.: (A) Thorax, lateral view; (B) Thorax with the thoracic gland and sensory area with microtrichia; (C) Wing; (D) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391309" httpUri="https://zenodo.org/record/8391309/files/figure.png" pageId="1" pageNumber="140">4C</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD07B1FB6FFB46D3A5" box="[1182,1217,1253,1279]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6B</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD07E3FB6CFB76D25A" box="[1228,1265,1254,1280]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD07D3FB6FFAA7D3A5" box="[1276,1312,1253,1279]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8B</figureCitation>
): base of C vein, after last break, wider than remaining vein; base of costal area with microsetae near C (
<figureCitation id="35F7BFB80174FFCD07B1FA83FA87D279" box="[1182,1280,1289,1315]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1H</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0623FA83FAB5D279" box="[1292,1330,1289,1315]" captionStart="FIGURE 4" captionStartId="5.[152,255,1441,1465]" captionTargetBox="[170,1416,489,1418]" captionTargetId="figure-159@5.[170,1416,489,1418]" captionTargetPageId="5" captionText="FIGURE 4. Coronamyia santosi sp. nov.: (A) Thorax, lateral view; (B) Thorax with the thoracic gland and sensory area with microtrichia; (C) Wing; (D) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391309" httpUri="https://zenodo.org/record/8391309/files/figure.png" pageId="1" pageNumber="140">4D</figureCitation>
); R 
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not reaching R 
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; R 
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ending at pointed apex of wing, not reaching R 
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; R 
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fork Approximately at the same level of M 
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fork, or M 
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basal to R 
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; small circular thoracic gland near the thoracic sensory area with microtrichia (tgl, 
<figureCitation id="35F7BFB80174FFCD064AFADBFF3CD2D5" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03E8FAFFFF69D2D5" box="[199,238,1397,1423]" captionStart="FIGURE 4" captionStartId="5.[152,255,1441,1465]" captionTargetBox="[170,1416,489,1418]" captionTargetId="figure-159@5.[170,1416,489,1418]" captionTargetPageId="5" captionText="FIGURE 4. Coronamyia santosi sp. nov.: (A) Thorax, lateral view; (B) Thorax with the thoracic gland and sensory area with microtrichia; (C) Wing; (D) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391309" httpUri="https://zenodo.org/record/8391309/files/figure.png" pageId="1" pageNumber="140">4A</figureCitation>
); thoracic sensory area with microtrichia in the dorsal area of anepimeron (sam, 
<figureCitation id="35F7BFB80174FFCD075BFAFFFB51D2D5" box="[1140,1238,1397,1423]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD07CDFAFFFA8ED2D5" box="[1250,1289,1397,1423]" captionStart="FIGURE 4" captionStartId="5.[152,255,1441,1465]" captionTargetBox="[170,1416,489,1418]" captionTargetId="figure-159@5.[170,1416,489,1418]" captionTargetPageId="5" captionText="FIGURE 4. Coronamyia santosi sp. nov.: (A) Thorax, lateral view; (B) Thorax with the thoracic gland and sensory area with microtrichia; (C) Wing; (D) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391309" httpUri="https://zenodo.org/record/8391309/files/figure.png" pageId="1" pageNumber="140">4A</figureCitation>
). Terminalia: gonocoxites horn-like, wider distally than basally, with subapical group of setae in external margin (
<figureCitation id="35F7BFB80174FFCD07D6FA13FA1FD2EE" box="[1273,1432,1433,1460]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1B, C, F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03B8FA37FF5AD282" box="[151,221,1469,1496]" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">2B, D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03C7FA37FE8AD28D" box="[232,269,1469,1495]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0236FA34FEB0D282" box="[281,311,1470,1496]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD026DFA37FEE2D28D" box="[322,357,1469,1495]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8E</figureCitation>
); anterior condyles of gonocoxites plate-like, fused at midline forming a gonocoxal bridge, with medium elevation directed to the ejaculatory apodeme (
<figureCitation id="35F7BFB80174FFCD0021FA6BFCF6D2A1" box="[782,881,1505,1531]" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">Figs. 2C</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0051FA6BFC27D2A6" box="[894,928,1505,1532]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5E</figureCitation>
). Gonostyli glabrous, sclerotized, bifurcated, shorter than gonocoxites, internal arm turned outward, external arm with basal or medial short to long lobe (
<figureCitation id="35F7BFB80174FFCD064BF98FFE9CD11E" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">Figs. 2A, B, D, E</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0209F9A3FE6FD119" box="[294,488,1577,1604]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5A, B, C, D, F, G</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD02DCF9A3FDB9D11E" box="[499,574,1577,1604]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6 C, D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0167F9A0FD03D11E" box="[584,644,1578,1604]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7E, F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD01A0F9A3FD55D119" box="[655,722,1577,1603]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8C, E</figureCitation>
). Aedeagus symmetrical. Plate with tuft of long setae around the aedeagus (
<figureCitation id="35F7BFB80174FFCD0222F9C7FE15D132" box="[269,402,1613,1640]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1A, B</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD02B1F9C7FD82D132" box="[414,517,1613,1640]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5B, C, D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD013EF9C7FDF6D132" box="[529,625,1613,1640]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6D, E, F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0152F9C4FD1CD132" box="[637,667,1614,1640]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0189F9C7FD6AD13D" box="[678,749,1613,1639]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8C, D</figureCitation>
). Aedeagal sheath lightly sclerotized with apical microtrichia (
<figureCitation id="35F7BFB80174FFCD03B0F9FBFEA1D1D6" box="[159,294,1649,1676]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1B, D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD021DF9FBFED0D1D1" box="[306,343,1649,1675]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD024BF9FBFE03D1D1" box="[356,388,1649,1675]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7F</figureCitation>
). Ejaculatory apodeme wider than aedeagus, usually much more so (
<figureCitation id="35F7BFB80174FFCD07A0F9FBFA92D1D6" box="[1167,1301,1649,1676]" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">Figs. 2A, B</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD060DF9FBFAEFD1D6" box="[1314,1384,1649,1676]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5A, B</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD065BF9FBFF29D1EA" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6C, D</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0395F91CFF7CD1F5" box="[186,251,1685,1712]" captionStart="FIGURE 7" captionStartId="8.[152,255,1932,1956]" captionTargetBox="[170,1416,624,1908]" captionTargetId="figure-168@8.[170,1416,624,1908]" captionTargetPageId="8" captionText="FIGURE 7. Coronamyia trilobata sp.nov. (A) Head, antenna and palpi; (B) Labium, posterior view, arrows show the spiniform setae; (C)Antenna, last flagellomeres and apiculus; (D) Wing; (E) Male terminalia, dorsal view, photograph; (F) Male terminalia, dorsal view, drawing; (G) Setous ring, dorsal view; (H) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391315" httpUri="https://zenodo.org/record/8391315/files/figure.png" pageId="1" pageNumber="140">7E, F</figureCitation>
). Epandrium bare, subrectangular, wider than long, with one basal foramen and anterior sclerotized band (
<figureCitation id="35F7BFB80174FFCD03B0F933FF7CD189" box="[159,251,1721,1747]" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">Figs. 2F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0229F933FEABD18E" box="[262,300,1721,1748]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5H</figureCitation>
). Hypopod elongated, base wider than apex, without apical tenaculum, dorsally with one or sometimes two rows of big alveoli bearing accessory geniculate tenacula and a group of short accessories tenacula at base (
<figureCitation id="35F7BFB80174FFCD064BF957FF66D046" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1G, H</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD03C0F88BFE93D046" box="[239,276,1793,1820]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5H</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD020DF88BFEC1D041" box="[290,326,1793,1819]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6G</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD027AF88BFEFED041" box="[341,377,1793,1819]" captionStart="FIGURE 8" captionStartId="9.[152,255,1939,1963]" captionTargetBox="[173,1414,849,1913]" captionTargetId="figure-251@9.[173,1414,849,1916]" captionTargetPageId="9" captionText="FIGURE 8. Coronamyia biapicalis (Bravo, Lago and Castro, 2004). (A) Antenna, base; (B) Wing; (C) Male terminalia; (D) Setous ring, dorsal view; (E) Gonostylus, dorsal view; (F) Gonostylus, base, ventral view; (G) Hypopod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.8391319" httpUri="https://zenodo.org/record/8391319/files/figure.png" pageId="1" pageNumber="140">8G</figureCitation>
), ventrally covered by small alveoli bearing setae (
<figureCitation id="35F7BFB80174FFCD00E5F88BFBA2D041" box="[970,1061,1793,1819]" captionStart="FIGURE 1" captionStartId="2.[152,255,1776,1800]" captionTargetBox="[164,1423,597,1751]" captionTargetId="figure-222@2.[164,1423,597,1753]" captionTargetPageId="2" captionText="FIGURE 1. Coronamyia paschoali sp. nov.: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia.Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland." figureDoi="http://doi.org/10.5281/zenodo.8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="1" pageNumber="140">Figs. 1I</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD071BF88BFBC9D046" box="[1076,1102,1793,1820]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5I</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD0773F88BFB05D041" box="[1116,1154,1793,1819]" captionStart="FIGURE 6" captionStartId="7.[152,255,1581,1605]" captionTargetBox="[214,1374,181,1558]" captionTargetId="figure-19@7.[214,1374,181,1558]" captionTargetPageId="7" captionText="FIGURE 6. Coronamyia triangularis sp. nov. (A) Head; (B) Wing; (C) Male terminalia, dorsal view, photograph; (D) Male terminalia, dorsal view, drawing; (E) Ejaculatory apodeme, aedeagus and aedeagal sheath, lateral view, photograph; (F) Ejaculatory apodeme, aedeagus and aedeagal sheat, lateral view, drawing; (G) Hypopods and subepandrial sclerite, dorsal view; (H) Hypopods, hypandrium and subepandrial sclerite, ventral view. Abbreviations: aed: aedeagus, aes: aes: aedeagal sheat, eja: ejaculatory apodeme." figureDoi="http://doi.org/10.5281/zenodo.8391313" httpUri="https://zenodo.org/record/8391313/files/figure.png" pageId="1" pageNumber="140">6H</figureCitation>
). Epiproct not observed, probably reduced. Hypoproct subtriangular in dorsal view, with apical microtrichia (
<figureCitation id="35F7BFB80174FFCD0768F8AFFB23D065" box="[1095,1188,1829,1855]" captionStart="FIGURE 2" captionStartId="3.[152,255,1816,1840]" captionTargetBox="[164,1423,625,1782]" captionTargetId="figure-133@3.[164,1423,625,1793]" captionTargetPageId="3" captionText="FIGURE 2. Coronamyia paschoali sp. nov.: (A) Male terminalia, dorsal view, photograph; (B) Male terminalia, dorsal view, drawing; (C) Gonocoxal apodeme; (D) Gonostylus, dorsal view; (E) Gonostylus, ventral view; (F) Epandrium and hypoproct, ventral view; (G) Hypopod, dorsal view, arrow shows sensory seta; (H) Hypopod, apex, dorsal view, arrow shows sensory seta; (I) Hypopod, ventral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat; eja: ajaculatory apodeme, for: foramen, gca: gonocoxal apodeme, gcx: gonocoxite, ses: subepandrial sclerite." figureDoi="http://doi.org/10.5281/zenodo.8391301" httpUri="https://zenodo.org/record/8391301/files/figure.png" pageId="1" pageNumber="140">Figs. 2F</figureCitation>
, 
<figureCitation id="35F7BFB80174FFCD079FF8AFFB54D01A" box="[1200,1235,1829,1856]" captionStart="FIGURE 5" captionStartId="6.[152,255,1506,1530]" captionTargetBox="[170,1416,345,1483]" captionTargetId="figure-101@6.[170,1416,345,1483]" captionTargetPageId="6" captionText="FIGURE 5. Coronamyia santosi sp. nov.: (A) Male terminalia, dorsal view, photograph; (b) Male terminalia, dorsal view, drawing; (C) Male terminalia, lateral view, photograph; (D) Male terminalia, lateral view, drawing; (E) Gonocoxal apodeme; (F) Gonostylus, dorsal view; (G) Gonostylus, ventral view; (H) Epandrium, hypopods and hypoproct, dorsal view; (I) Aedeagus, apex, and hypopods, lateral view. Abbreviations: aed: aedeagus, aes: aedeagal sheat, eja: ejaculatory apodeme, gca: gonocoxal apodeme, gcx: gonocoxite, gst: gonostylus, hyp: hypandrium, str: setose ring." figureDoi="http://doi.org/10.5281/zenodo.8391311" httpUri="https://zenodo.org/record/8391311/files/figure.png" pageId="1" pageNumber="140">5H</figureCitation>
)
</paragraph>
<paragraph id="AD73A33D0174FFCD03E8F8C3FE19D039" blockId="1.[151,1437,605,2036]" box="[199,414,1865,1891]" pageId="1" pageNumber="140">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="E5D6F0B60174FFCD03E8F8E7FACBD08A" pageId="1" pageNumber="140" type="distribution">
<paragraph id="AD73A33D0174FFCD03E8F8E7FBE4D0DD" blockId="1.[151,1437,605,2036]" box="[199,1123,1901,1928]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03E8F8E7FEE6D0DD" bold="true" box="[199,353,1901,1927]" pageId="1" pageNumber="140">Distribution.</emphasis>
Brazilian Atlantic Rain Forest: 
<collectingRegion id="6F086DDF0174FFCD01EAF8E7FC8ED0DD" box="[709,777,1901,1927]" country="Brazil" name="Bahia" pageId="1" pageNumber="140">Bahia</collectingRegion>
, 
<collectingRegion id="6F086DDF0174FFCD003AF8E7FC33D0DD" box="[789,948,1901,1927]" country="Brazil" name="Espirito Santo" pageId="1" pageNumber="140">Espirito Santo</collectingRegion>
and S„o Paulo.
</paragraph>
<paragraph id="AD73A33D0174FFCD03E8F81BFACBD08A" blockId="1.[151,1437,605,2036]" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03E8F81BFE15D0F1" bold="true" box="[199,402,1937,1963]" pageId="1" pageNumber="140">Included species.</emphasis>
<taxonomicName id="6ACCD8BE0174FFCD02B5F818FB8DD0F1" authority="(Bravo, Lago and Castro, 2004)" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" baseAuthorityName="Bravo, Lago and Castro" baseAuthorityYear="2004" box="[410,1034,1937,1964]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="biapicalis" status="comb. nov.">
<emphasis id="9FB87F2F0174FFCD02B5F818FD1CD0F1" box="[410,667,1937,1964]" italics="true" pageId="1" pageNumber="140">Coronamyia biapicalis</emphasis>
(
<bibRefCitation id="C95DDECC0174FFCD0184F81BFB84D0F1" author="Bravo, F. &amp; Lago, A. &amp; Castro, I." box="[683,1027,1937,1964]" pageId="1" pageNumber="140" pagination="589 - 599" refId="ref5672" refString="Bravo, F., Lago, A. &amp; Castro, I. (2004) Dez especies novas de Alepia Enderlein (Diptera: Psychodidae) do Brasil. Neotropical Entomology, 33, 589 - 599. https: // doi. org / 10.1590 / S 1519 - 566 X 2004000500008" type="journal article" year="2004">Bravo, Lago and Castro, 2004</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="848BC2540174FFCD073DF81BFB16D0F6" box="[1042,1169,1937,1964]" pageId="1" pageNumber="140" rank="species">comb. nov.</taxonomicNameLabel>
, 
<taxonomicName id="6ACCD8BE0174FFCD07B2F818FA1BD0F1" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[1181,1436,1937,1964]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="paschoali" status="sp. nov.">
<emphasis id="9FB87F2F0174FFCD07B2F818FA1BD0F1" box="[1181,1436,1937,1964]" italics="true" pageId="1" pageNumber="140">Coronamyia paschoali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540174FFCD03B8F83CFF77D08A" box="[151,240,1974,2000]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="6ACCD8BE0174FFCD03D4F83CFE5BD095" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[251,476,1974,2000]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="santosi" status="sp. nov.">
<emphasis id="9FB87F2F0174FFCD03D4F83CFE5BD095" box="[251,476,1974,2000]" italics="true" pageId="1" pageNumber="140">Coronamyia santosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540174FFCD02CBF83CFDBAD08A" box="[484,573,1974,2000]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="6ACCD8BE0174FFCD0167F83CFCE7D095" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[584,864,1973,2000]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="9FB87F2F0174FFCD0167F83CFCE7D095" box="[584,864,1973,2000]" italics="true" pageId="1" pageNumber="140">Coronamyia triangularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540174FFCD0049F83CFC38D08A" box="[870,959,1974,2000]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
, and 
<taxonomicName id="6ACCD8BE0174FFCD00D5F83CFB69D095" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[1018,1262,1973,2000]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="trilobata" status="sp. nov.">
<emphasis id="9FB87F2F0174FFCD00D5F83CFB69D095" box="[1018,1262,1973,2000]" italics="true" pageId="1" pageNumber="140">Coronamyia trilobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540174FFCD07DAF83CFACBD08A" box="[1269,1356,1974,2000]" pageId="1" pageNumber="140" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="E5D6F0B60174FFCF03E8F853FE91D61B" lastPageId="3" lastPageNumber="142" pageId="1" pageNumber="140" type="discussion">
<paragraph id="AD73A33D0174FFCE03E8F853FB2BD78F" blockId="1.[151,1437,605,2036]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="141" pageId="1" pageNumber="140">
<emphasis id="9FB87F2F0174FFCD03E8F853FEBCD0A9" bold="true" box="[199,315,2009,2035]" pageId="1" pageNumber="140">Remarks.</emphasis>
The new genus 
<taxonomicName id="6ACCD8BE0174FFCD02D3F850FD00D0AE" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[508,647,2010,2036]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="9FB87F2F0174FFCD02D3F850FD00D0AE" box="[508,647,2010,2036]" italics="true" pageId="1" pageNumber="140">Coronamyia</emphasis>
</taxonomicName>
falls within the tribe 
<taxonomicName id="6ACCD8BE0174FFCD00ABF853FC70D0A9" box="[900,1015,2009,2035]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
, as delimited by 
<bibRefCitation id="C95DDECC0174FFCD0791F853FADBD0A9" author="Kvifte, G. M." box="[1214,1372,2009,2035]" pageId="1" pageNumber="140" pagination="596 - 605" refId="ref6369" refString="Kvifte, G. M. (2018) Molecular phylogeny of moth flies (Diptera, Psychodidae, Psychodinae) revisited, with a revised tribal classification. Systematic Entomology, 43 (3), 596 - 605. https: // doi. org / 10.1111 / syen. 12288" type="journal article" year="2018">Kvifte (2018)</bibRefCitation>
, with fusiform flagellomeres, Rs not pectinate, and radial fork at the same level as the medial fork (in the diagnosis of the tribe “radial fork basal to medial fork”). The other characters are as the diagnosis of the tribe.
</paragraph>
<paragraph id="AD73A33D0177FFCE03E8FF55FE7AD6D3" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="141">
The new genus 
<taxonomicName id="6ACCD8BE0177FFCE0256FF55FD83D7A3" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[377,516,223,249]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="9FB87F2F0177FFCE0256FF55FD83D7A3" box="[377,516,223,249]" italics="true" pageId="2" pageNumber="141">Coronamyia</emphasis>
</taxonomicName>
is morphologically similar to 
<emphasis id="9FB87F2F0177FFCE0075FF55FC2DD7A3" box="[858,938,223,249]" italics="true" pageId="2" pageNumber="141">
<taxonomicName id="6ACCD8BE0177FFCE0075FF55FC20D7A3" authorityName="Enderlein" authorityYear="1937" box="[858,935,223,249]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus">Alepia</taxonomicName>
.
</emphasis>
This is due to the presence of an eye bridge with 3 or 4 facet rows (contiguous or separated); flagellomeres fusiform; pleuron with alveoli only on anepistemum and lalerotergite; the wings generally with infuscated patterns; R 
<subScript id="3148A1780177FFCE0074FEB3FCE4D61D" attach="left" box="[859,867,313,327]" fontSize="6" pageId="2" pageNumber="141">5</subScript>
ending at the wing apex; gonostyli often bifurcated; hypopods without apical tenaculum and with accessory tenacula (characters obtained from 
<bibRefCitation id="C95DDECC0177FFCE0788FEC1FAAAD63F" author="Quate, L. W." box="[1191,1325,331,357]" pageId="2" pageNumber="141" pagination="1 - 81" refId="ref6422" refString="Quate, L. W. (1996) Preliminary taxonomy of Costa Rican Psychodidae (Diptera), exclusive of Phlebotominae. Revista de Biologia Tropical, 44 (Supplement 1), 1 - 81." type="journal article" year="1996">Quate 1996</bibRefCitation>
; 
<bibRefCitation id="C95DDECC0177FFCE0614FEC1FEA1D6D3" author="Quate, L. W. &amp; Brown, B. V." pageId="2" pageNumber="141" pagination="1 - 117" refId="ref6460" refString="Quate, L. W. &amp; Brown, B. V. (2004) Revision of Neotropical Setomimini (Diptera: Psychodidae: Psychodinae). Contribution in Science, Natural History Museum of Los Angeles, 500, 1 - 117. https: // doi. org / 10.5962 / p. 210558" type="journal article" year="2004">Quate &amp; Brown 2004</bibRefCitation>
; 
<bibRefCitation id="C95DDECC0177FFCE021EFEE5FE76D6D3" author="Jezek, J. &amp; Le Pont, F. &amp; Martinez, E. &amp; Molinedo, S." box="[305,497,367,393]" pageId="2" pageNumber="141" pagination="183 - 210" refId="ref6123" refString="Jezek, J., Le Pont, F., Martinez, E. &amp; Molinedo, S. (2011) Three new species of non-biting moth flies (Diptera: Psychodidae: Psychodinae) from Bolivia, with notes on higher taxa of the subfamily. Acta Entomol Musei Natl Pragae, 51, 183 - 210." type="journal article" year="2011">
Ježek 
<emphasis id="9FB87F2F0177FFCE0259FEE5FE28D6D3" box="[374,431,367,393]" italics="true" pageId="2" pageNumber="141">et al.</emphasis>
2011
</bibRefCitation>
).
</paragraph>
<paragraph id="AD73A33D0177FFCE03E8FE19FA88D567" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="141">
However, this new genus can be differentiated from the other genera of 
<taxonomicName id="6ACCD8BE0177FFCE0721FE19FBF9D6F7" box="[1038,1150,403,429]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="tribe" tribe="Maruinini">Maruinini</taxonomicName>
by the above mentioned diagnostic characters, mainly the presence of a circular plate with a tuft of long setae around the aedeagus (observed only in the species included in 
<taxonomicName id="6ACCD8BE0177FFCE02D4FE51FD01D6AF" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[507,646,475,501]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="genus" status="new genus">
<emphasis id="9FB87F2F0177FFCE02D4FE51FD01D6AF" box="[507,646,475,501]" italics="true" pageId="2" pageNumber="141">Coronamyia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540177FFCE01A0FE51FC8CD6AF" box="[655,779,475,501]" pageId="2" pageNumber="141" rank="genus">new genus</taxonomicNameLabel>
), a bifurcated gonostylus with an arched internal arm and lobed external arm; elongated, subovoid hypopods without apical tenacula; and accessory tenacula of two kinds, the basal 
<typeStatus id="72771D9F0177FFCE03F7FDA9FE8FD567" box="[216,264,547,573]" pageId="2" pageNumber="141">type</typeStatus>
with umbellate tips, with the rest of dorsal area having accessory tenacula with clavate tips.
</paragraph>
<caption id="F9B3F3B50177FFCE03B7F97AFE58D02F" ID-DOI="http://doi.org/10.5281/zenodo.8391299" ID-Zenodo-Dep="8391299" httpUri="https://zenodo.org/record/8391299/files/figure.png" pageId="2" pageNumber="141" startId="2.[152,255,1776,1800]" targetBox="[164,1423,597,1751]" targetPageId="2" targetType="figure">
<paragraph id="AD73A33D0177FFCE03B7F97AFE58D02F" blockId="2.[151,1437,1776,1909]" pageId="2" pageNumber="141">
<emphasis id="9FB87F2F0177FFCE03B7F97AFE9BD053" bold="true" box="[152,284,1776,1801]" pageId="2" pageNumber="141">FIGURE 1.</emphasis>
<taxonomicName id="6ACCD8BE0177FFCE0209F97BFD96D053" authority="Bravo &amp; Lopes &amp; Araújo, 2023" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[294,529,1777,1801]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="paschoali" status="sp. nov.">
<emphasis id="9FB87F2F0177FFCE0209F97BFD96D053" box="[294,529,1777,1801]" italics="true" pageId="2" pageNumber="141">Coronamyia paschoali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="848BC2540177FFCE0135F97BFDF7D052" box="[538,624,1777,1800]" pageId="2" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
: (A) Head; (B) Labium, posterior view, arrows show pair of spiniform setae in the epipharynx; (C) Pair of spiniform setae in the epipharynx; (D) Thorax with the thoracic gland and sensory area with microtrichia; (E) Antenna; (F) Palpus; (G) Wing; (H) Wing, costal area with microtrichia. Abbreviations: sam: sensory area with microtrichia, tgl: thoracic gland.
</paragraph>
</caption>
<paragraph id="AD73A33D0177FFCF03E8F835FE91D61B" blockId="2.[199,1436,1983,2009]" lastBlockId="3.[151,1436,151,321]" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
The hypopods of 
<taxonomicName id="6ACCD8BE0177FFCE028FF835FC1DD083" authority="Bravo, Lago &amp; Castro, 2004" authorityName="Bravo, Lago &amp; Castro" authorityYear="2004" box="[416,922,1983,2009]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="clara">
<emphasis id="9FB87F2F0177FFCE028FF835FDB5D083" box="[416,562,1983,2009]" italics="true" pageId="2" pageNumber="141">Alepia clara</emphasis>
<bibRefCitation id="C95DDECC0177FFCE016FF835FC1DD083" author="Bravo, F. &amp; Lago, A. &amp; Castro, I." box="[576,922,1983,2009]" pageId="2" pageNumber="141" pagination="589 - 599" refId="ref5672" refString="Bravo, F., Lago, A. &amp; Castro, I. (2004) Dez especies novas de Alepia Enderlein (Diptera: Psychodidae) do Brasil. Neotropical Entomology, 33, 589 - 599. https: // doi. org / 10.1590 / S 1519 - 566 X 2004000500008" type="journal article" year="2004">Bravo, Lago &amp; Castro, 2004</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="6ACCD8BE0177FFCE00F2F835FAEBD083" authority="Wagner, 1993" authorityName="Wagner" authorityYear="1993" box="[989,1388,1983,2009]" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="martinicana">
<emphasis id="9FB87F2F0177FFCE00F2F835FB3AD083" box="[989,1213,1983,2009]" italics="true" pageId="2" pageNumber="141">Alepia martinicana</emphasis>
<bibRefCitation id="C95DDECC0177FFCE07E5F835FAEBD083" author="Wagner, R." box="[1226,1388,1983,2009]" pageId="2" pageNumber="141" pagination="109 - 127" refId="ref6517" refString="Wagner, R. (1993) On a collection of Psychodidae (Diptera) by Dr. L. Botosaneanu from some Caribbean islands. Aquatic Insects, 15 (2), 109 - 127. https: // doi. org / 10.1080 / 01650429309361506" type="journal article" year="1993">Wagner, 1993</bibRefCitation>
</taxonomicName>
are morphologically similar to those of the described species of 
<taxonomicName id="6ACCD8BE0176FFCF0061FF1DFC5ED7EB" authorityName="Bravo &amp; Lopes &amp; Araújo" authorityYear="2023" box="[846,985,151,177]" class="Insecta" family="Psychodidae" genus="Coronamyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="9FB87F2F0176FFCF0061FF1DFC5ED7EB" box="[846,985,151,177]" italics="true" pageId="3" pageNumber="142">Coronamyia</emphasis>
</taxonomicName>
, although the gonostyli of the first two species lack internal arched and external lobated arms. The gonostylus of 
<taxonomicName id="6ACCD8BE0176FFCF00C8FF31FD85D7A3" authority="Jaume-Schinkel, Kvifte, van der Weele &amp; Mengual, 2022" authorityName="Jaume-Schinkel, Kvifte, van der Weele &amp; Mengual" authorityYear="2022" class="Insecta" family="Psychodidae" genus="Alepia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="142" phylum="Arthropoda" rank="species" species="viatrix">
<emphasis id="9FB87F2F0176FFCF00C8FF31FB05D78E" box="[999,1154,187,213]" italics="true" pageId="3" pageNumber="142">Alepia viatrix</emphasis>
<bibRefCitation id="C95DDECC0176FFCF07A4FF31FD85D7A3" author="Jaume-Schinkel, S. &amp; Kvifte, G. M. &amp; van der Weele, R. &amp; Mengual, X." pageId="3" pageNumber="142" pagination="384 - 396" refId="ref6046" refString="Jaume-Schinkel, S., Kvifte, G. M., van der Weele, R. &amp; Mengual, X. (2022) Alepia viatrix sp. nov. (Diptera: Psychodidae), a new species of a Neotropical genus found on the Azores Archipelago (Portugal). Zootaxa, 5128 (3), 384 - 396. https: // doi. org / 10.11646 / zootaxa. 5128.3.4" type="journal article" year="2022">Jaume-Schinkel, Kvifte, van der Weele &amp; Mengual, 2022</bibRefCitation>
</taxonomicName>
is similar to the species of the new genus, although the shape of the hypopods, the presence of apical tenacula, and the absence of two kinds of tenacula differentiate it from the other species of the new genus.
</paragraph>
</subSubSection>
</treatment>
</document>