299 lines
33 KiB
XML
299 lines
33 KiB
XML
<document id="F455348DCF957E62BB05B03AED6D6A0C" ID-DOI="10.1649/1082.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5370086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268477416" checkinUser="valdenar" docAuthor="Carlton, Christopher E." docDate="2008" docId="8D2B87E4FF89DC58FE24FCEDB53BFA55" docLanguage="en" docName="1082.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1082.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Arianops hiltenae Carlton 2008, new species" docType="treatment" docVersion="3" lastPageNumber="323" masterDocId="7112FF9CFF9DDC4EFFBCFFE8B638FFD8" masterDocTitle="Eight New Species ofArianopsBrendel from the Southeastern United States with an Updated Key and Notes on Additional Species (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="323" masterPageNumber="297" pageNumber="297" updateTime="1699205358136" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1A45E27C7722C01FB2406DAD8775158F">Eight New Species ofArianopsBrendel from the Southeastern United States with an Updated Key and Notes on Additional Species (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:name id="145A33C86A93074146A5048B32727945" type="personal">
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<mods:namePart id="D3AB6B46C40117CEB8DCA8EA27FF8A6B">Carlton, Christopher E.</mods:namePart>
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<mods:typeOfResource id="F645C1107F40F24B5DC8A984EB527929">text</mods:typeOfResource>
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<mods:title id="FC4D3C7C840707ED80A9CD12D05BF510">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="9B24FDEBF6298919A0691159C32B52E5">
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<mods:date id="1DD6C7D2EDACF846DFB3EB9BA07A4DCB">2008</mods:date>
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<mods:number id="3DD5551D6F6CA23E3F1243688F432F46">2008-06-30</mods:number>
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<mods:number id="514BD30CBE515051D38B2091018E8AEA">62</mods:number>
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<mods:url id="C132F8C77FFCD703E9C165E6706E42AE">http://dx.doi.org/10.1649/1082.1</mods:url>
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<mods:identifier id="30928A38C6D078BBC9552AB43A9B82FD" type="DOI">10.1649/1082.1</mods:identifier>
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<mods:identifier id="EF65DDA8E18222484995477CD49CED6F" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="4D986579FF89DC5AFE24FCEDB4F1FCC3" box="[408,713,773,795]" pageId="20" type="nomenclature">
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<paragraph id="053D36F2FF89DC5AFE24FCEDB4F1FCC3" blockId="20.[408,713,773,822]" box="[408,713,773,795]" pageId="20">
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<heading id="5E75819EFF89DC5AFE24FCEDB4F1FCC3" box="[408,713,773,795]" centered="true" fontSize="9" level="2" pageId="20" reason="2">
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<emphasis id="37F6EAE0FF89DC5AFE24FCEDB4F1FCC3" bold="true" box="[408,713,773,795]" pageId="20">
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<taxonomicName id="C2824D71FF89DC5AFE24FCEDB470FCC3" ID-CoL="5VR8Z" authority="Carlton, 2008" authorityName="Carlton" authorityYear="2008" box="[408,584,773,795]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="species" species="hiltenae" status="sp. nov.">
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<emphasis id="37F6EAE0FF89DC5AFE24FCEDB470FCC3" bold="true" box="[408,584,773,795]" italics="true" pageId="20">Arianops hiltenae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="2CC5579BFF89DC5AFDEAFCEDB4F1FCC3" box="[598,713,773,795]" pageId="20" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4D986579FF89DC5AFE09FCC8B494FCEE" box="[437,684,800,822]" pageId="20" type="description">
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<paragraph id="053D36F2FF89DC5AFE09FCC8B494FCEE" blockId="20.[408,713,773,822]" box="[437,684,800,822]" pageId="20">
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(
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<figureCitation id="9DB92A77FF89DC5AFE01FCC8B40DFCEE" box="[445,565,800,822]" captionStart="Figs" captionStartId="18.[167,206,1368,1387]" captionTargetBox="[177,945,194,1346]" captionTargetId="figure-66@18.[177,945,194,1346]" captionTargetPageId="18" captionText="Figs. 13–16. Dorsal and lateral views, Arianops new species holotypes. 13–14) A. gorei;" figureDoi="http://doi.org/10.5281/zenodo.5370094" httpUri="https://zenodo.org/record/5370094/files/figure.png" pageId="20">Figs. 15–16</figureCitation>
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,
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<figureCitation id="9DB92A77FF89DC5AFDFEFCC8B4B8FCEE" box="[578,640,800,822]" captionStart="Figs" captionStartId="20.[167,206,697,716]" captionTargetBox="[211,909,194,673]" captionTargetId="figure-290@20.[211,909,194,674]" captionTargetPageId="20" captionText="Figs. 28–33. Arianops new species. 28) A. folkertsi, head; 29) A. folkertsi, ventrites 3–4; 30) A. gorei, head; 31) A. gorei, ventrites 3–4; 32) A. hiltenae, head; 33) A. hiltenae, ventrites 3–4." figureDoi="http://doi.org/10.5281/zenodo.5370098" httpUri="https://zenodo.org/record/5370098/files/figure.png" pageId="20">32–33</figureCitation>
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,
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<figureCitation id="9DB92A77FF89DC5AFD30FCC8B49DFCEE" box="[652,677,800,822]" captionStart="Figs" captionStartId="21.[196,235,1117,1136]" captionTargetBox="[192,987,194,1094]" captionTargetId="figure-132@21.[192,987,194,1094]" captionTargetPageId="21" captionText="Figs. 34–41. Arianops new species aedeagi, dorsal views. 34) A. carteri; 35) A. clintoni;" figureDoi="http://doi.org/10.5281/zenodo.5370100" httpUri="https://zenodo.org/record/5370100/files/figure.png" pageId="20">40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="4D986579FF89DC5AFF16FCBEB4BDFC64" pageId="20" type="materials_examined">
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<paragraph id="053D36F2FF89DC5AFF16FCBEB4BDFC64" blockId="20.[146,976,854,1570]" pageId="20">
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<materialsCitation id="B5EA3CAFFF89DC5AFF16FCBEB465FC79" collectingDate="2006-06-16" collectorName="Sokolov, I. M." county="Blount Co." elevation="620" latitude="83.73583" location="White Oak Sink" pageId="20" specimenCount="1" typeStatus="holotype">
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<emphasis id="37F6EAE0FF89DC5AFF16FCBEB779FCB4" bold="true" box="[170,321,854,876]" pageId="20">
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<typeStatus id="DA398850FF89DC5AFF16FCBEB73EFCB4" box="[170,262,854,876]" pageId="20" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(
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<figureCitation id="9DB92A77FF89DC5AFEE9FCBEB7F4FCB4" box="[341,460,854,876]" captionStart="Figs" captionStartId="18.[167,206,1368,1387]" captionTargetBox="[177,945,194,1346]" captionTargetId="figure-66@18.[177,945,194,1346]" captionTargetPageId="18" captionText="Figs. 13–16. Dorsal and lateral views, Arianops new species holotypes. 13–14) A. gorei;" figureDoi="http://doi.org/10.5281/zenodo.5370094" httpUri="https://zenodo.org/record/5370094/files/figure.png" pageId="20">Figs. 15–16</figureCitation>
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). /TN:
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<collectingCounty id="EC5C4E7EFF89DC5AFD9FFCBEB4A4FCB4" box="[547,668,854,876]" pageId="20">Blount Co.</collectingCounty>
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, GSMNP,
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<location id="005D6029FF89DC5AFCA5FCBEB5F3FCB4" LSID="urn:lsid:plazi:treatment:8D2B87E4FF89DC58FE24FCEDB53BFA55:005D6029FF89DC5AFCA5FCBEB5F3FCB4" box="[793,971,854,876]" county="Blount Co." latitude="83.73583" name="White Oak Sink" pageId="20">White Oak Sink</location>
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, 35
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<emphasis id="37F6EAE0FF89DC5AFF16FC99B68BFC5E" bold="true" box="[170,179,881,902]" italics="true" pageId="20">°</emphasis>
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38
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<emphasis id="37F6EAE0FF89DC5AFF77FC98B6EBFC5E" bold="true" box="[203,211,880,902]" pageId="20">9</emphasis>
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07
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<emphasis id="37F6EAE0FF89DC5AFF57FC98B6CBFC5E" bold="true" box="[235,243,880,902]" pageId="20">0</emphasis>
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<geoCoordinate id="60B65035FF89DC5AFF4FFC99B76BFC5E" box="[243,339,880,903]" degrees="83" direction="north" minutes="44" orientation="latitude" pageId="20" precision="15" seconds="9" value="83.73583">
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N 83
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<emphasis id="37F6EAE0FF89DC5AFE96FC99B70BFC5E" bold="true" box="[298,307,881,902]" italics="true" pageId="20">°</emphasis>
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44
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<emphasis id="37F6EAE0FF89DC5AFEF7FC98B76BFC5E" bold="true" box="[331,339,880,902]" pageId="20">9</emphasis>
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</geoCoordinate>
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49
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<quantity id="C27A9B17FF89DC5AFED7FC98B7FFFC5F" box="[363,455,880,903]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="20" unit="m" value="620.0">
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||
<elevation id="8EAFD1C1FF89DC5AFED7FC98B7FFFC5F" box="[363,455,880,903]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="20" unit="m" value="620.0">
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<emphasis id="37F6EAE0FF89DC5AFED7FC98B74BFC5E" bold="true" box="[363,371,880,902]" pageId="20">0</emphasis>
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, 620 m
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</elevation>
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</quantity>
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, mixed forest, litter hand sifting,
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<collectorName id="A8775324FF89DC5AFC86FC99B5F7FC5F" box="[826,975,881,903]" pageId="20">Sokolov, I.M.</collectorName>
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||
<date id="713C1032FF89DC5AFF2EFC63B739FC79" box="[146,257,907,929]" pageId="20" value="2006-06-16">
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<collectingDate id="6178E9DAFF89DC5AFF2EFC63B739FC79" box="[146,257,907,929]" pageId="20" value="2006-06-16">16.06.2006</collectingDate>
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</date>
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||
/
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||
<taxonomicName id="C2824D71FF89DC5AFEB5FC63B465FC79" authority="Carlton, 2007" authorityName="Carlton" authorityYear="2007" box="[265,605,907,929]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="20" phylum="Arthropoda" rank="species" species="hiltenae">Arianops hiltenae Carlton, 2007</taxonomicName>
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</materialsCitation>
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<materialsCitation id="B5EA3CAFFF89DC5AFDD5FC63B4B9FC64" collectorName="Chicago, IL" location="Type" municipality="Field Museum of Natural History" pageId="20" specimenCount="1" typeStatus="holotype">
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||
<typeStatus id="DA398850FF89DC5AFDD5FC63B4F1FC79" box="[617,713,907,929]" pageId="20" type="holotype">Holotype</typeStatus>
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male/.
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||
<location id="005D6029FF89DC5AFCA2FC63B56AFC79" LSID="urn:lsid:plazi:treatment:8D2B87E4FF89DC58FE24FCEDB53BFA55:005D6029FF89DC5AFCA2FC63B56AFC79" box="[798,850,907,929]" municipality="Field Museum of Natural History" name="Type" pageId="20">Type</location>
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deposition,
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<collectingMunicipality id="E559AC88FF89DC5AFF2EFC4EB7CDFC64" box="[146,501,934,956]" pageId="20">Field Museum of Natural History</collectingMunicipality>
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,
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<collectorName id="A8775324FF89DC5AFDBEFC4EB4B9FC64" box="[514,641,934,956]" pageId="20">Chicago, IL</collectorName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D986579FF89DC5AFF16FC29B57AFBD4" pageId="20" type="etymology">
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<paragraph id="053D36F2FF89DC5AFF16FC29B57AFBD4" blockId="20.[146,976,854,1570]" pageId="20">
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<emphasis id="37F6EAE0FF89DC5AFF16FC29B725FC0F" bold="true" box="[170,285,961,983]" pageId="20">Etymology.</emphasis>
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This species is named after Jeanie Hilten, Director Emeritus of Discover Life in America, Inc., supporter, and promoter of the All Taxa Biodiversity Inventory in Great Smoky Mountains National Park.
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</paragraph>
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</subSubSection>
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<subSubSection id="4D986579FF89DC5BFF16FBF9B7B7FA43" lastPageId="21" pageId="20" type="description">
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<paragraph id="053D36F2FF89DC5AFF16FBF9B505FB49" blockId="20.[146,976,854,1570]" pageId="20">
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<emphasis id="37F6EAE0FF89DC5AFF16FBF9B71BFBFF" bold="true" box="[170,291,1041,1063]" pageId="20">Description.</emphasis>
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<typeStatus id="DA398850FF89DC5AFE95FBF9B7B1FBFF" box="[297,393,1041,1063]" pageId="20" type="holotype">Holotype</typeStatus>
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male. Measurements (in mm): head 0.50 long, 0.50 wide; pronotum 0.45 long, 0.55 wide; elytra 0.60 long, 0.70 wide; median lengths of visible abdominal tergites 1–5, respectively 0.35, 0.10, 0.08, 0.30, 0.08. Antennomeres 1–11, respectively 0.11, 0.10, 0.10, 0.10, 0.08, 0.08, 0.08, 0.06, 0.07, 0.06, 0.22. Aedeagus 0.30 long, 0.25 wide. Total length 2.30.
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</paragraph>
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<paragraph id="053D36F2FF89DC5AFF16FB7FB7ECFA59" blockId="20.[146,976,854,1570]" pageId="20">
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Head (
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<figureCitation id="9DB92A77FF89DC5AFF4AFB7FB77BFB75" box="[246,323,1175,1197]" captionStart="Figs" captionStartId="20.[167,206,697,716]" captionTargetBox="[211,909,194,673]" captionTargetId="figure-290@20.[211,909,194,674]" captionTargetPageId="20" captionText="Figs. 28–33. Arianops new species. 28) A. folkertsi, head; 29) A. folkertsi, ventrites 3–4; 30) A. gorei, head; 31) A. gorei, ventrites 3–4; 32) A. hiltenae, head; 33) A. hiltenae, ventrites 3–4." figureDoi="http://doi.org/10.5281/zenodo.5370098" httpUri="https://zenodo.org/record/5370098/files/figure.png" pageId="20">Fig. 32</figureCitation>
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). Lateral vertexal carinae weak, discernable above antennal tubercles, becoming obsolete at a point well before vertexal foveae. Traces of median occipital carina visible, discontinuous between base of occiput and vertex. Vertexal foveae small, shallow, not connected via circumambient sulcus, vertex evenly convex, almost flat between antennal tubercles. Carinae to antennal tubercles not extending onto clypeus; clypeus not carinate medially. Labral margin bearing four minute denticles. Lateral carinae from clypeus to ocular spines well-developed. Ocular spine moderate, triangularly acute, oriented laterally. Mentum not toothed.
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</paragraph>
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<paragraph id="053D36F2FF89DC5AFF16FA6EB5E8FA09" blockId="20.[146,976,854,1570]" pageId="20">Pronotum. Basomedian fovea absent, submedian spines and bumps absent. Basolateral foveoid depression barely discernable at certain angles of lighting. Pronotum weakly transversely rugose along basolateral margin, smooth medially.</paragraph>
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<paragraph id="053D36F2FF89DC5AFF16FA3FB7E9F9DF" blockId="20.[146,976,854,1570]" pageId="20">Elytra. Basal margin of each elytron straight to humeral declivity. Basal depressions and foveae absent.</paragraph>
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<paragraph id="053D36F2FF89DC5AFF16F9E4B766F9FA" blockId="20.[146,976,854,1570]" box="[170,350,1548,1570]" pageId="20">Legs unmodified.</paragraph>
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<caption id="51FD667AFF88DC5BFF78FBB5B5D4FBA8" ID-DOI="http://doi.org/10.5281/zenodo.5370100" ID-Zenodo-Dep="5370100" box="[196,1004,1117,1136]" httpUri="https://zenodo.org/record/5370100/files/figure.png" pageId="21" startId="21.[196,235,1117,1136]" targetBox="[192,987,194,1094]" targetPageId="21">
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||
<paragraph id="053D36F2FF88DC5BFF78FBB5B5D4FBA8" blockId="21.[175,1004,1117,1160]" box="[196,1004,1117,1136]" pageId="21">
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<emphasis id="37F6EAE0FF88DC5BFF78FBB5B70DFBA8" bold="true" box="[196,309,1117,1136]" pageId="21">Figs. 34–41.</emphasis>
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<taxonomicName id="C2824D71FF88DC5BFEF6FBB5B7A2FBA8" authority="Carlton, 2008" authorityName="Carlton" authorityYear="2008" box="[330,410,1117,1136]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="genus" status="sp. nov.">
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||
<emphasis id="37F6EAE0FF88DC5BFEF6FBB5B7A2FBA8" box="[330,410,1117,1136]" italics="true" pageId="21">Arianops</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="2CC5579BFF88DC5BFE1DFBB5B432FBA8" box="[417,522,1117,1136]" pageId="21" rank="species">new species</taxonomicNameLabel>
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aedeagi, dorsal views.
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<emphasis id="37F6EAE0FF88DC5BFD63FBB5B4C3FBA8" bold="true" box="[735,763,1117,1136]" pageId="21">34)</emphasis>
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<taxonomicName id="C2824D71FF88DC5BFCBEFBB5B560FBA8" authorityName="Carlton" authorityYear="2007" box="[770,856,1117,1136]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="carteri">
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<emphasis id="37F6EAE0FF88DC5BFCBEFBB5B560FBA8" box="[770,856,1117,1136]" italics="true" pageId="21">A. carteri</emphasis>
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</taxonomicName>
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;
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<emphasis id="37F6EAE0FF88DC5BFCDAFBB5B5BAFBA8" bold="true" box="[870,898,1117,1136]" pageId="21">35)</emphasis>
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<taxonomicName id="C2824D71FF88DC5BFC35FBB5B5DEFBA8" authorityName="Carlton" authorityYear="2007" box="[905,998,1117,1136]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="clintoni">
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<emphasis id="37F6EAE0FF88DC5BFC35FBB5B5DEFBA8" box="[905,998,1117,1136]" italics="true" pageId="21">A. clintoni</emphasis>
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</taxonomicName>
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;
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</paragraph>
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</caption>
|
||
<paragraph id="053D36F2FF88DC5BFF13FB9DB500FB50" blockId="21.[175,1004,1117,1160]" box="[175,824,1141,1160]" pageId="21">
|
||
<emphasis id="37F6EAE0FF88DC5BFF13FB9DB6F4FB50" bold="true" box="[175,204,1141,1160]" pageId="21">36)</emphasis>
|
||
<taxonomicName id="C2824D71FF88DC5BFF6FFB9DB723FB50" authorityName="Carlton" authorityYear="2007" box="[211,283,1141,1160]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="ashei">
|
||
<emphasis id="37F6EAE0FF88DC5BFF6FFB9DB723FB50" box="[211,283,1141,1160]" italics="true" pageId="21">A. ashei</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="37F6EAE0FF88DC5BFE94FB9DB77CFB50" bold="true" box="[296,324,1141,1160]" pageId="21">37)</emphasis>
|
||
<taxonomicName id="C2824D71FF88DC5BFEF7FB9DB7AAFB50" authorityName="Carlton" authorityYear="2007" box="[331,402,1141,1160]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="barri">
|
||
<emphasis id="37F6EAE0FF88DC5BFEF7FB9DB7AAFB50" box="[331,402,1141,1160]" italics="true" pageId="21">A. barri</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="37F6EAE0FF88DC5BFE23FB9DB784FB50" bold="true" box="[415,444,1141,1160]" pageId="21">38)</emphasis>
|
||
<taxonomicName id="C2824D71FF88DC5BFE7FFB9DB410FB50" authorityName="Carlton" authorityYear="2007" box="[451,552,1141,1160]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="folkertsi">
|
||
<emphasis id="37F6EAE0FF88DC5BFE7FFB9DB410FB50" box="[451,552,1141,1160]" italics="true" pageId="21">A. folkertsi</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="37F6EAE0FF88DC5BFD89FB9DB469FB50" bold="true" box="[565,593,1141,1160]" pageId="21">39)</emphasis>
|
||
<taxonomicName id="C2824D71FF88DC5BFDE4FB9DB498FB50" authorityName="Carlton" authorityYear="2007" box="[600,672,1141,1160]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="gorei">
|
||
<emphasis id="37F6EAE0FF88DC5BFDE4FB9DB498FB50" box="[600,672,1141,1160]" italics="true" pageId="21">A. gorei</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="37F6EAE0FF88DC5BFD11FB9DB4F2FB50" bold="true" box="[685,714,1141,1160]" pageId="21">40)</emphasis>
|
||
<taxonomicName id="C2824D71FF88DC5BFD6DFB9DB50AFB50" authorityName="Carlton" authorityYear="2007" box="[721,818,1141,1160]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="hiltenae">
|
||
<emphasis id="37F6EAE0FF88DC5BFD6DFB9DB50AFB50" box="[721,818,1141,1160]" italics="true" pageId="21">A. hiltenae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="053D36F2FF88DC5BFF7BFB46B758FB21" blockId="21.[175,1005,1198,1570]" pageId="21">
|
||
Abdomen. Dorsally weakly rounded, evenly, gently declivous at tergite 4. Visible ventrite 3 with a short, broad, median, marginal shelf (
|
||
<figureCitation id="9DB92A77FF88DC5BFC95FB21B54DFB07" box="[809,885,1225,1247]" captionStart="Figs" captionStartId="20.[167,206,697,716]" captionTargetBox="[211,909,194,673]" captionTargetId="figure-290@20.[211,909,194,674]" captionTargetPageId="20" captionText="Figs. 28–33. Arianops new species. 28) A. folkertsi, head; 29) A. folkertsi, ventrites 3–4; 30) A. gorei, head; 31) A. gorei, ventrites 3–4; 32) A. hiltenae, head; 33) A. hiltenae, ventrites 3–4." figureDoi="http://doi.org/10.5281/zenodo.5370098" httpUri="https://zenodo.org/record/5370098/files/figure.png" pageId="21">Fig. 33</figureCitation>
|
||
). Ventrite 6 broadly concave.
|
||
</paragraph>
|
||
<paragraph id="053D36F2FF88DC5BFF7BFB17B7B9FAA7" blockId="21.[175,1005,1198,1570]" pageId="21">
|
||
Aedeagus (
|
||
<figureCitation id="9DB92A77FF88DC5BFE84FB17B7BEFACD" box="[312,390,1279,1301]" captionStart="Figs" captionStartId="21.[196,235,1117,1136]" captionTargetBox="[192,987,194,1094]" captionTargetId="figure-132@21.[192,987,194,1094]" captionTargetPageId="21" captionText="Figs. 34–41. Arianops new species aedeagi, dorsal views. 34) A. carteri; 35) A. clintoni;" figureDoi="http://doi.org/10.5281/zenodo.5370100" httpUri="https://zenodo.org/record/5370100/files/figure.png" pageId="21">Fig. 40</figureCitation>
|
||
). Parameres absent. Ventral shelf acutely pointed left due to prolongation of right margin and emargination of left, giving aedeagus the overall shape of a comma. Right base of shelf bearing a loose group of five setae, apex with two setae. Short triangular process from left interior of phallobase bearing three terminal setae.
|
||
</paragraph>
|
||
<paragraph id="053D36F2FF88DC5BFF7BFA6DB7B7FA43" blockId="21.[175,1005,1198,1570]" box="[199,399,1413,1435]" pageId="21">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D986579FF88DC5BFF7BFA48B485FA6E" box="[199,701,1440,1462]" pageId="21" type="materials_examined">
|
||
<paragraph id="053D36F2FF88DC5BFF7BFA48B485FA6E" blockId="21.[175,1005,1198,1570]" box="[199,701,1440,1462]" pageId="21">
|
||
<emphasis id="37F6EAE0FF88DC5BFF7BFA48B7B5FA6E" bold="true" box="[199,397,1440,1462]" pageId="21">Material examined.</emphasis>
|
||
Only the
|
||
<typeStatus id="DA398850FF88DC5BFE4AFA48B468FA6E" box="[502,592,1440,1462]" pageId="21" type="holotype">holotype</typeStatus>
|
||
is known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D986579FF88DC5BFF7BFA54B741FA34" pageId="21" type="distribution">
|
||
<paragraph id="053D36F2FF88DC5BFF7BFA54B741FA34" blockId="21.[175,1005,1198,1570]" pageId="21">
|
||
<emphasis id="37F6EAE0FF88DC5BFF7BFA54B77EFA0A" bold="true" box="[199,326,1468,1490]" pageId="21">Distribution.</emphasis>
|
||
The species is only known from the
|
||
<typeStatus id="DA398850FF88DC5BFD4FFA54B527FA0A" box="[755,799,1468,1490]" pageId="21">type</typeStatus>
|
||
locality in Blount County,
|
||
<collectingRegion id="C746F810FF88DC5BFEB6FA3EB74CFA34" box="[266,372,1494,1516]" country="United States of America" name="Tennessee" pageId="21">Tennessee</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D986579FF88DC58FF7BFA1AB53BFA55" lastPageId="22" pageId="21" type="discussion">
|
||
<paragraph id="053D36F2FF88DC58FF7BFA1AB5B5FBCD" blockId="21.[175,1005,1198,1570]" lastBlockId="22.[146,976,811,1365]" lastPageId="22" pageId="21">
|
||
<emphasis id="37F6EAE0FF88DC5BFF7BFA1AB70FF9D0" bold="true" box="[199,311,1522,1544]" pageId="21">Comments.</emphasis>
|
||
External characters of
|
||
<taxonomicName id="C2824D71FF88DC5BFD90FA1AB4AFF9DF" authorityName="Carlton" authorityYear="2007" box="[556,663,1522,1543]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="hiltenae">
|
||
<emphasis id="37F6EAE0FF88DC5BFD90FA1AB4AFF9DF" box="[556,663,1522,1543]" italics="true" pageId="21">A. hiltenae</emphasis>
|
||
</taxonomicName>
|
||
are consistent with
|
||
<bibRefCitation id="61134B03FF88DC5BFCD6FA1AB5D5F9D0" author="Barr" box="[874,1005,1522,1544]" pageId="21" refId="ref16021" refString="Barr, T. C., Jr. 1974. The eyeless beetles of the genus Arianops Brendel (Coleoptera, Pselaphidae). Bulletin of the American Museum of Natural History 154, Article 1, 51 pp. (accessible online: http: // digitallibrary. amnh. org / dspace / handle / 2246 / 1142, accessed 27 November 2007)." type="book" year="1974">Barr’s (1974)</bibRefCitation>
|
||
cavernensis species group. The species is similar to
|
||
<taxonomicName id="C2824D71FF88DC5BFD1FF9E5B59DF9FA" authority="Park (1951)" authorityName="Park" authorityYear="1951" box="[675,933,1548,1570]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="21" phylum="Arthropoda" rank="species" species="cavernensis">
|
||
<emphasis id="37F6EAE0FF88DC5BFD1FF9E5B514F9FA" box="[675,812,1549,1570]" italics="true" pageId="21">A. cavernensis</emphasis>
|
||
<bibRefCitation id="61134B03FF88DC5BFC8EF9E4B59DF9FA" author="Park" box="[818,933,1548,1570]" pageId="21" refId="ref16546" refString="Park, O. 1951. Cavernicolous pselaphid beetles of Alabama and Tennessee, with observations on the taxonomy of the family. Geological Survey of Alabama, Museum Paper 31, 107 pp." type="book" year="1951">Park (1951)</bibRefCitation>
|
||
</taxonomicName>
|
||
in that 318 the median vertexal carina is almost completely absent, and the pronotal foveae, tubercles and spines are absent. These are the only two species having this degree of reduction of these features. The lateral vertexal carinae of
|
||
<taxonomicName id="C2824D71FF8BDC58FD51FC88B560FCAD" authorityName="Carlton" authorityYear="2007" box="[749,856,864,885]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="hiltenae">
|
||
<emphasis id="37F6EAE0FF8BDC58FD51FC88B560FCAD" box="[749,856,864,885]" italics="true" pageId="22">A. hiltenae</emphasis>
|
||
</taxonomicName>
|
||
are weakly developed whereas they are absent on
|
||
<taxonomicName id="C2824D71FF8BDC58FDBCFC93B4BDFC48" box="[512,645,891,912]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="cavernensis">
|
||
<emphasis id="37F6EAE0FF8BDC58FDBCFC93B4BDFC48" box="[512,645,891,912]" italics="true" pageId="22">A. cavernensis</emphasis>
|
||
</taxonomicName>
|
||
. The vertexal foveae of
|
||
<taxonomicName id="C2824D71FF8BDC58FCD5FC93B5F7FC48" authorityName="Carlton" authorityYear="2007" box="[873,975,891,912]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="hiltenae">
|
||
<emphasis id="37F6EAE0FF8BDC58FCD5FC93B5F7FC48" box="[873,975,891,912]" italics="true" pageId="22">A. hiltenae</emphasis>
|
||
</taxonomicName>
|
||
are small, but obvious, whereas those of
|
||
<taxonomicName id="C2824D71FF8BDC58FD97FC7DB48DFC72" box="[555,693,917,938]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="cavernensis">
|
||
<emphasis id="37F6EAE0FF8BDC58FD97FC7DB48DFC72" box="[555,693,917,938]" italics="true" pageId="22">A. cavernensis</emphasis>
|
||
</taxonomicName>
|
||
are vestigial and reduced to small pits. Unfortunately,
|
||
<taxonomicName id="C2824D71FF8BDC58FE29FC58B426FC1D" box="[405,542,944,965]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="cavernensis">
|
||
<emphasis id="37F6EAE0FF8BDC58FE29FC58B426FC1D" box="[405,542,944,965]" italics="true" pageId="22">A. cavernensis</emphasis>
|
||
</taxonomicName>
|
||
is based only on the female
|
||
<typeStatus id="DA398850FF8BDC58FC8BFC58B5B7FC1E" box="[823,911,944,966]" pageId="22" type="holotype">holotype</typeStatus>
|
||
, so an aedeagal comparison is not possible, and these comparisons are based on
|
||
<bibRefCitation id="61134B03FF8BDC58FCEEFC22B5F7FC38" author="Barr" box="[850,975,970,992]" pageId="22" refId="ref16021" refString="Barr, T. C., Jr. 1974. The eyeless beetles of the genus Arianops Brendel (Coleoptera, Pselaphidae). Bulletin of the American Museum of Natural History 154, Article 1, 51 pp. (accessible online: http: // digitallibrary. amnh. org / dspace / handle / 2246 / 1142, accessed 27 November 2007)." type="book" year="1974">Barr’s (1974)</bibRefCitation>
|
||
description. The differences in the lateral vertexal carinae and vertexal foveae, and the widely separated localities (,
|
||
<quantity id="C27A9B17FF8BDC58FE0EFC17B7C5FBCD" box="[434,509,1023,1045]" metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="22" unit="km" value="250.0">250 km</quantity>
|
||
) support placement as a distinct species.
|
||
</paragraph>
|
||
<caption id="51FD667AFF8BDC58FF1BFD15B5E8FCC8" ID-DOI="http://doi.org/10.5281/zenodo.5370102" ID-Zenodo-Dep="5370102" box="[167,976,765,784]" httpUri="https://zenodo.org/record/5370102/files/figure.png" pageId="22" startId="22.[167,198,765,784]" targetBox="[149,972,194,742]" targetPageId="22">
|
||
<paragraph id="053D36F2FF8BDC58FF1BFD15B5E8FCC8" blockId="22.[167,976,765,784]" box="[167,976,765,784]" pageId="22">
|
||
<emphasis id="37F6EAE0FF8BDC58FF1BFD15B6D7FCC8" bold="true" box="[167,239,765,784]" pageId="22">Fig. 41.</emphasis>
|
||
<taxonomicName id="C2824D71FF8BDC58FEB9FD15B7F7FCC8" authority="Barr" authorityName="Barr" authorityYear="1974" box="[261,463,765,784]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="37F6EAE0FF8BDC58FEB9FD15B7A5FCC8" box="[261,413,765,784]" italics="true" pageId="22">Arianops alticola</emphasis>
|
||
Barr
|
||
</taxonomicName>
|
||
, female abdominal apex and putative male internal sac.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="053D36F2FF8BDC58FF16FBF2B56CFA8D" blockId="22.[146,976,811,1365]" pageId="22">
|
||
White Oak Sink, near the northwest boundary of Great Smoky Mountains National Park, is part of a small area within the park having a karst substrate with a network of interconnected limestone caves. While a number of
|
||
<taxonomicName id="C2824D71FF8BDC58FD47FBB8B5A4FBBD" box="[763,924,1103,1125]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="37F6EAE0FF8BDC58FD47FBB8B56BFBBD" box="[763,851,1104,1125]" italics="true" pageId="22">Arianops</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have been collected in caves and may be obligate troglobites, especially within the cavernensis species group, the
|
||
<typeStatus id="DA398850FF8BDC58FE06FB6DB429FB43" box="[442,529,1157,1179]" pageId="22" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="C2824D71FF8BDC58FD93FB6DB4AEFB42" authorityName="Carlton" authorityYear="2007" box="[559,662,1157,1178]" class="Insecta" family="Staphylinidae" genus="Arianops" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="hiltenae">
|
||
<emphasis id="37F6EAE0FF8BDC58FD93FB6DB4AEFB42" box="[559,662,1157,1178]" italics="true" pageId="22">A. hiltenae</emphasis>
|
||
</taxonomicName>
|
||
was collected sifting forest litter,
|
||
<quantity id="C27A9B17FF8BDC58FF1AFB77B6E1FB6D" box="[166,217,1183,1205]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="22" unit="m" value="10.0">10 m</quantity>
|
||
from the nearest cave opening (I. Sokolov, pers. comm.). Litter collections from small caves and sinkholes in the immediate vicinity have thus far failed to produce additional specimens.
|
||
<bibRefCitation id="61134B03FF8BDC58FE7AFB3CB401FB32" author="Barr" box="[454,569,1236,1258]" pageId="22" refId="ref16021" refString="Barr, T. C., Jr. 1974. The eyeless beetles of the genus Arianops Brendel (Coleoptera, Pselaphidae). Bulletin of the American Museum of Natural History 154, Article 1, 51 pp. (accessible online: http: // digitallibrary. amnh. org / dspace / handle / 2246 / 1142, accessed 27 November 2007)." type="book" year="1974">Barr (1974)</bibRefCitation>
|
||
discussed habitats where other members of the species group have been taken. Most seem to be edaphobites of deep litter in karst landscapes, and their association with caves may be incidental. The troglophilic carabid
|
||
<taxonomicName id="C2824D71FF8BDC58FF58FACCB7D2FAE2" authority="Barr" authorityName="Barr" authorityYear="1962" box="[228,490,1316,1338]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="22" phylum="Arthropoda" rank="species" species="tuckaleechee">
|
||
<emphasis id="37F6EAE0FF8BDC58FF58FACCB78AFAE1" box="[228,434,1316,1337]" italics="true" pageId="22">Trechus tuckaleechee</emphasis>
|
||
Barr
|
||
</taxonomicName>
|
||
, originally described from Tuckaleechee Cave, a short distance outside the park, has also been collected at this locality.
|
||
</paragraph>
|
||
<paragraph id="053D36F2FF8BDC58FEE3FA91B53BFA55" blockId="22.[351,771,1401,1423]" box="[351,771,1401,1423]" pageId="22">
|
||
<emphasis id="37F6EAE0FF8BDC58FEE3FA91B53BFA55" bold="true" box="[351,771,1401,1423]" pageId="22">
|
||
NOTES ON PREVIOUSLY DESCRIBED
|
||
<emphasis id="37F6EAE0FF8BDC58FD0AFA91B53BFA55" bold="true" box="[694,771,1401,1423]" pageId="22">SPECIES</emphasis>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |