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<document id="6A8EE13EDCE5C24FC76EB44AA258EF86" ID-CLB-Dataset="22343" ID-DOI="10.5852/ejt.2015.167" ID-GBIF-Dataset="72b00f76-b85d-47c0-877f-511110a6a0f4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3805815" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588788296119" checkinUser="carolina" docAuthor="Hughes, Mark, Girmansyah, Deden &amp; Ardi, Wisnu Handoyo" docDate="2015" docId="03FCF055FFF8FA4AD8FE3C35FE626E84" docLanguage="en" docName="ejt-2015-167.pdf.imf" docOrigin="European Journal of Taxonomy 167" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia halabanensis M. Hughes 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="FFC5882DFFF3FA46D942382BFFB66D01" masterDocTitle="Further discoveries in the ever-expanding genus Begonia (Begoniaceae): fifteen new species from Sumatra" masterLastPageNumber="40" masterPageNumber="1" pageNumber="12" updateTime="1698838759931" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="27E7213FB2A5AF04C1EC1046667168FD">Further discoveries in the ever-expanding genus Begonia (Begoniaceae): fifteen new species from Sumatra</mods:title>
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<mods:namePart id="AC79DD4C69468B02528A66F6633AAC8B">Hughes, Mark</mods:namePart>
<mods:affiliation id="E7CBAC105FFBB93E644488B2F95BF6DD">Royal Botanic Garden Edinburgh, 20 a Inverleith Row, EH 3 5 LR, UK. &amp; Corresponding author: m. hughes @ rbge. ac. uk</mods:affiliation>
<mods:nameIdentifier id="B1FEFA727B97C4CBF808D588B22BCA78" type="email">m.hughes@rbge.ac.uk</mods:nameIdentifier>
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<mods:namePart id="E995C3383CDC022A6E3B0BAA9A918786">Girmansyah, Deden</mods:namePart>
<mods:affiliation id="06CB27AC53E46174DF275A7BADE4CDE0">Herbarium Bogoriense, Botany Division, Research Center for Biology, Indonesian Institute of Sciences, Cibinong Science Center (CSC), Jl. Raya Jakarta-Bogor Km 46, Cibinong, Bogor 16911, Indonesia. &amp; Email: deden _ bo @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="B53280BFC1F159115111A9BE12B219F2" type="email">deden_bo@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="4AF8F5A132F0533DC86B40A1A7FAF53B">Ardi, Wisnu Handoyo</mods:namePart>
<mods:affiliation id="0E0058E539C4BBEFC3818A3CD1057AF5">Center For Plant Conservation-Bogor Botanic Garden, Jl. Ir. H. Juanda No. 13, P. O. Box 309, Bogor 16003, Indonesia. &amp; Email: wisnu. handoyo. ardi @ lipi. go. id</mods:affiliation>
<mods:nameIdentifier id="73CBBA4F33C5DE685460468AC09497AB" type="email">handoyo.ardi@lipi.go.id</mods:nameIdentifier>
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<mods:title id="331B1867FEFE8461A00560F4B7569655">European Journal of Taxonomy</mods:title>
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<mods:date id="CA1892BB096E44EA32AB49C1102327D1">2015</mods:date>
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<mods:number id="0D368BBC687D6D3EB1DFE10497370D91">2015-12-21</mods:number>
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<treatment id="03FCF055FFF8FA4AD8FE3C35FE626E84" ID-DOI="http://doi.org/10.5281/zenodo.3815853" ID-GBIF-Taxon="163970620" ID-Zenodo-Dep="3815853" LSID="urn:lsid:plazi:treatment:03FCF055FFF8FA4AD8FE3C35FE626E84" httpUri="http://treatment.plazi.org/id/03FCF055FFF8FA4AD8FE3C35FE626E84" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="C34F12C8FFF8FA4DD8FE3C35FBC16939" box="[444,1143,1054,1081]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BEA4143FFF8FA4DD8FE3C35FBC16939" blockId="11.[444,1143,1054,1152]" box="[444,1143,1054,1081]" pageId="11" pageNumber="12">
<heading id="D0A2F62FFFF8FA4DD8FE3C35FBC16939" box="[444,1143,1054,1081]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<taxonomicName id="4C553AC0FFF8FA4DD8FE3C35FCFD6939" ID-CoL="7SFT5" authority="M.Hughes" authorityName="M. Hughes" authorityYear="2015" box="[444,843,1054,1081]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="halabanensis" status="sp. nov.">
<emphasis id="B9219D51FFF8FA4DD8FE3C35FD7E6938" bold="true" box="[444,712,1054,1081]" italics="true" pageId="11" pageNumber="12">Begonia halabanensis</emphasis>
M.Hughes
</taxonomicName>
<taxonomicNameLabel id="A212202AFFF8FA4DDA103C34FC1A6938" box="[850,940,1055,1081]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
§
<taxonomicName id="4C553AC0FFF8FA4DDA8B3C35FBC16939" box="[969,1143,1054,1080]" class="Magnoliopsida" family="Begoniaceae" genus="Reichenheimia" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="genus">
<emphasis id="B9219D51FFF8FA4DDA8B3C35FBC16939" box="[969,1143,1054,1080]" italics="true" pageId="11" pageNumber="12">Reichenheimia</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFF8FA4DDB053C69FCFB6981" pageId="11" pageNumber="12" type="description">
<paragraph id="8BEA4143FFF8FA4DDB053C69FC5B695C" blockId="11.[444,1143,1054,1152]" box="[583,1005,1090,1117]" pageId="11" pageNumber="12">urn:lsid:ipni.org:names:77151640-1</paragraph>
<paragraph id="8BEA4143FFF8FA4DDBA53C4EFCFB6981" blockId="11.[444,1143,1054,1152]" box="[743,845,1125,1152]" pageId="11" pageNumber="12">
<figureCitation id="136E5DC6FFF8FA4DDBA53C4EFC84697E" box="[743,818,1125,1151]" captionStart="Fig" captionStartId="8.[189,232,1965,1991]" captionTargetBox="[189,1398,586,1923]" captionTargetId="figure@8.[189,1398,585,1923]" captionTargetPageId="8" captionText="Fig. 2. Map of Sumatra showing the full known species distributions based on collection localities. Each point may represent more than one collection." figureDoi="http://doi.org/10.5281/zenodo.3805823" httpUri="https://zenodo.org/record/3805823/files/figure.png" pageId="11" pageNumber="12">Figs 2</figureCitation>
,
<figureCitation id="136E5DC6FFF8FA4DDA7C3C4EFCFB6981" box="[830,845,1125,1152]" captionStart="Fig" captionStartId="12.[189,232,1930,1956]" captionTargetBox="[206,1388,950,1888]" captionTargetId="figure@12.[189,1398,943,1892]" captionTargetPageId="12" captionText="Fig. 5. Begonia halabanensis M.Hughes sp. nov. A. Holotype [Meijer 7550 (L)]. B. Portion of leaf lamina underside showing margin hairs and recurved teeth. C. Anthers. D. Fruit. E. Male flower. Drawings by M. Hughes." figureDoi="http://doi.org/10.5281/zenodo.3805833" httpUri="https://zenodo.org/record/3805833/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFF8FA4DD9FF3C8CFC0D685B" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BEA4143FFF8FA4DD9FF3C8CFE8069C0" blockId="11.[189,310,1191,1217]" box="[189,310,1191,1217]" pageId="11" pageNumber="12">
<heading id="D0A2F62FFFF8FA4DD9FF3C8CFE8069C0" bold="true" box="[189,310,1191,1217]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9219D51FFF8FA4DD9FF3C8CFE8069C0" bold="true" box="[189,310,1191,1217]" pageId="11" pageNumber="12">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFF8FA4DD9FF3CFDFC0D685B" blockId="11.[189,1399,1238,1371]" pageId="11" pageNumber="12">
The peltate and succulent leaves are also found in
<taxonomicName id="4C553AC0FFF8FA4DDA4C3CFCFC1569F1" box="[782,931,1238,1264]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="kudoensis">
<emphasis id="B9219D51FFF8FA4DDA4C3CFCFC1569F1" box="[782,931,1238,1264]" italics="true" pageId="11" pageNumber="12">B. kudoensis</emphasis>
</taxonomicName>
, however
<taxonomicName id="4C553AC0FFF8FA4DDD5C3CFCFB6C69F1" authorityName="M. Hughes" authorityYear="2015" box="[1054,1242,1238,1264]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="halabanensis">
<emphasis id="B9219D51FFF8FA4DDD5C3CFCFB6C69F1" box="[1054,1242,1238,1264]" italics="true" pageId="11" pageNumber="12">B. halabanensis</emphasis>
</taxonomicName>
has a thickly hairy rhizome and petioles (not glabrous), male flowers with 2 tepals (not 4), larger inflorescences (with
<emphasis id="B9219D51FFF8FA4DD9FF3D35FF676836" box="[189,209,1310,1335]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
80 flowers, not
<emphasis id="B9219D51FFF8FA4DD8D53D35FE1D6836" box="[407,427,1310,1335]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
40) and fruit with a truncate (not obtuse) apex. The key from couplet 6 onwards deals with all 5 currently described peltate species from
<collectingRegion id="49918FA1FFF8FA4DDA103D6BFC0E685B" box="[850,952,1344,1370]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="12">Sumatra</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFF8FA4DD9FF3DA9FA8A68CA" box="[189,1340,1410,1483]" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BEA4143FFF8FA4DD9FF3DA9FEF2689D" blockId="11.[189,324,1410,1436]" box="[189,324,1410,1436]" pageId="11" pageNumber="12">
<heading id="D0A2F62FFFF8FA4DD9FF3DA9FEF2689D" bold="true" box="[189,324,1410,1436]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9219D51FFF8FA4DD9FF3DA9FEF2689D" bold="true" box="[189,324,1410,1436]" pageId="11" pageNumber="12">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFF8FA4DD9FF3D9AFA8A68CA" blockId="11.[189,1340,1457,1483]" box="[189,1340,1457,1483]" pageId="11" pageNumber="12">
The epithet refers to the distribution of the species in Halaban in
<collectingRegion id="49918FA1FFF8FA4DDAF53D9AFBEC68CA" box="[951,1114,1457,1483]" country="Indonesia" name="Sumatera Barat" pageId="11" pageNumber="12">West Sumatra</collectingRegion>
near Payakumbuh.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFF8FA4DD9FF3DD8FA9C6B3D" box="[189,1322,1523,1596]" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BEA4143FFF8FA4DD9FF3DD8FF4C6B0C" blockId="11.[189,250,1523,1549]" box="[189,250,1523,1549]" pageId="11" pageNumber="12">
<heading id="D0A2F62FFFF8FA4DD9FF3DD8FF4C6B0C" bold="true" box="[189,250,1523,1549]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9219D51FFF8FA4DD9FF3DD8FF4C6B0C" bold="true" box="[189,250,1523,1549]" pageId="11" pageNumber="12">
<typeStatus id="54EEFFE1FFF8FA4DD9FF3DD8FF4C6B0C" box="[189,250,1523,1549]" pageId="11" pageNumber="12">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFF8FA4DD9FF3E0AFA9C6B3D" blockId="11.[189,1322,1569,1596]" box="[189,1322,1569,1596]" pageId="11" pageNumber="12">
<collectingRegion id="49918FA1FFF8FA4DD9FF3E0AFEF86B3A" box="[189,334,1569,1595]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="12">SUMATRA</collectingRegion>
:
<collectingRegion id="49918FA1FFF8FA4DD81A3E09FE486B3A" box="[344,510,1569,1596]" country="Indonesia" name="Sumatera Barat" pageId="11" pageNumber="12">West Sumatra</collectingRegion>
, Pajakumbuh, Halaban,
<quantity id="4CADECA6FFF8FA4DDA5E3E0AFCDE6B3D" box="[796,872,1569,1596]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="12" unit="m" value="800.0">800 m</quantity>
,
<date id="FFEB6783FFF8FA4DDA363E0AFBBA6B3D" box="[884,1036,1569,1596]" pageId="11" pageNumber="12" value="1950-01-29">29 Jan. 1950</date>
,
<emphasis id="B9219D51FFF8FA4DDD5A3E09FB1F6B3A" box="[1048,1193,1569,1595]" italics="true" pageId="11" pageNumber="12">Meijer 7550</emphasis>
(holo-:
<collectionCode id="ED44D986FFF8FA4DDC453E09FAAB6B3D" box="[1287,1309,1570,1596]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="11" pageNumber="12" type="Herbarium">L</collectionCode>
).
</paragraph>
</subSubSection>
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<heading id="D0A2F62FFFF8FA4DD9FF3E48FEF86B7C" bold="true" box="[189,334,1635,1661]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B9219D51FFF8FA4DD9FF3E48FEF86B7C" bold="true" box="[189,334,1635,1661]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFF8FA4AD9FF3EB9FBD36CE1" blockId="11.[189,1399,1681,2027]" lastBlockId="12.[189,1399,267,481]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Erect acaulescent herb
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tall; stem rhizomatous, stout, internodes
<emphasis id="B9219D51FFF8FA4DDD5F3EB9FB876BAA" box="[1053,1073,1682,1707]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
<quantity id="4CADECA6FFF8FA4DDD7A3EB9FBC76BAD" box="[1080,1137,1682,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="cm" value="1.0">1 cm</quantity>
long, densely covered in a matted indumentum of reddish brown multi-branched thick hairs
<emphasis id="B9219D51FFF8FA4DDAB33E9DFBB36BCE" box="[1009,1029,1718,1743]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
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long. Stipules lanceolate, stout, 23 ×
<quantity id="4CADECA6FFF8FA4DD8093EF3FE2A6BF2" box="[331,412,1752,1779]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="cm" value="1.5">1.5 cm</quantity>
, fibrous, persistent, with thick, long hairs. Leaves: petiole
<emphasis id="B9219D51FFF8FA4DDD783EF2FBF86BF3" box="[1082,1102,1753,1778]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
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long, cross section unknown, with quite dense
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long hairs; lamina peltate, point of petiole attachment placed to within
<quantity id="4CADECA6FFF8FA4DD9FF3F0BFF406A3B" box="[189,246,1824,1850]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="cm" value="3.0">3 cm</quantity>
of the margin, suborbicular, 1015 ×
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, thick and succulent, base rounded, margin entire in outline, with evenly spaced
<quantity id="4CADECA6FFF8FA4DDB423F68FDF56A5C" box="[512,579,1859,1885]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="mm" value="2.0">2 mm</quantity>
long stiff rounded teeth that are recurved underneath, margin also with a fringe of
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long hairs, apex acute, underside with many pale dots of stomatal clusters, venation palmate, main veins 8, quite widely spaced over most of the lamina. Inflorescences axillary,
<emphasis id="B9219D51FFF8FA4DDC4D3FA0FA956AA5" box="[1295,1315,1931,1956]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
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long and exceeding the leaves, cymose, many-flowered, branching
<emphasis id="B9219D51FFF8FA4DDAA53F85FC4D6AC6" box="[999,1019,1966,1991]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
8 times, bisexual, protandrous; bracts elliptic, minute, glabrous, entire,
<emphasis id="B9219D51FFF8FA4DDBC83FF9FD286AEA" box="[650,670,2002,2027]" italics="true" pageId="11" pageNumber="12">c.</emphasis>
2 ×
<quantity id="4CADECA6FFF8FA4DDB923FFBFC9F6AEA" box="[720,809,2000,2027]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="mm" value="1.5">1.5 mm</quantity>
towards the terminal branches, basal pair not seen. Male flowers: pedicel
<quantity id="4CADECA6FFFFFA4AD8843920FDBD6C27" box="[454,523,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="mm" value="5.0">5 mm</quantity>
long, slender, glabrous; tepals 2, orbicular, entire, glabrous,
<emphasis id="B9219D51FFFFFA4ADD9E3927FB466C24" box="[1244,1264,268,293]" italics="true" pageId="12" pageNumber="13">c.</emphasis>
8 ×
<quantity id="4CADECA6FFFFFA4ADC6B3920FAC46C27" box="[1321,1394,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="mm" value="8.0">8 mm</quantity>
; androecium globose, symmetric, with
<emphasis id="B9219D51FFFFFA4ADB3E3919FD266C4A" box="[636,656,306,331]" italics="true" pageId="12" pageNumber="13">c.</emphasis>
40 stamens, filaments unequal, fused at base, shorter to slightly longer than the anther, anther
<quantity id="4CADECA6FFFFFA4ADB57397DFDCA6C70" box="[533,636,342,369]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="12" pageNumber="13" unit="mm" value="0.75">0.75 mm</quantity>
long, ellipsoid-obtriangular, dehiscing through short slits near the apex, apex retuse, not hooded. Female flowers unknown. Fruits pendent on a
<quantity id="4CADECA6FFFFFA4ADD133950FB666C97" box="[1105,1232,379,406]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="12" pageNumber="13" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">1520 mm</quantity>
long hair-like pedicel, borne in a cyme of
<emphasis id="B9219D51FFFFFA4AD8BF3989FDA66CBA" box="[509,528,418,443]" italics="true" pageId="12" pageNumber="13">c.</emphasis>
50, total size 9 ×
<quantity id="4CADECA6FFFFFA4ADB9E398AFC866CBD" box="[732,816,417,444]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="13" unit="mm" value="21.0">21 mm</quantity>
; capsule globose,
<quantity id="4CADECA6FFFFFA4ADD40398AFBF36CBD" box="[1026,1093,417,444]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="mm" value="6.0">6 mm</quantity>
in diameter, three-locular, placentae entire; wings 3, subequal, 810 ×
<quantity id="4CADECA6FFFFFA4ADB8339ECFC936CE0" box="[705,805,455,481]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="12" pageNumber="13" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
, tips rounded; apex obtuse.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFFFFA4AD9FF3A21FE4A6FA1" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BEA4143FFFFFA4AD9FF3A21FE586F25" blockId="12.[189,494,522,548]" box="[189,494,522,548]" pageId="12" pageNumber="13">
<heading id="D0A2F62FFFFFFA4AD9FF3A21FE586F25" bold="true" box="[189,494,522,548]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B9219D51FFFFFA4AD9FF3A21FE586F25" bold="true" box="[189,494,522,548]" pageId="12" pageNumber="13">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFFFFA4AD9FF3A10FE4A6FA1" blockId="12.[189,1399,571,672]" pageId="12" pageNumber="13">
Found on limestone, endemic to the
<typeStatus id="54EEFFE1FFFFFA4ADB303A17FD136F57" box="[626,677,572,598]" pageId="12" pageNumber="13">type</typeStatus>
locality in Halaban,
<collectingRegion id="49918FA1FFFFFA4ADAE23A17FBF26F54" box="[928,1092,571,598]" country="Indonesia" name="Sumatera Barat" pageId="12" pageNumber="13">West Sumatra</collectingRegion>
(
<figureCitation id="136E5DC6FFFFFA4ADD143A10FB296F54" box="[1110,1183,571,597]" captionStart="Fig" captionStartId="8.[189,232,1965,1991]" captionTargetBox="[189,1398,586,1923]" captionTargetId="figure@8.[189,1398,585,1923]" captionTargetPageId="8" captionText="Fig. 2. Map of Sumatra showing the full known species distributions based on collection localities. Each point may represent more than one collection." figureDoi="http://doi.org/10.5281/zenodo.3805823" httpUri="https://zenodo.org/record/3805823/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
). The thick fleshy leaves and densely hairy rhizome of this very distinctive species imply adaptation to dryer habitats. Known only from the
<typeStatus id="54EEFFE1FFFFFA4AD8803AADFE4E6FA1" box="[450,504,646,672]" pageId="12" pageNumber="13">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFFFFA4AD9FF3AE2FE626E84" pageId="12" pageNumber="13" type="conservation">
<paragraph id="8BEA4143FFFFFA4AD9FF3AE2FE036FE5" blockId="12.[189,437,713,740]" box="[189,437,713,740]" pageId="12" pageNumber="13">
<heading id="D0A2F62FFFFFFA4AD9FF3AE2FE036FE5" bold="true" box="[189,437,713,740]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B9219D51FFFFFA4AD9FF3AE2FE036FE5" bold="true" box="[189,437,713,740]" pageId="12" pageNumber="13">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFFFFA4AD9FF3AD1FE626E84" blockId="12.[189,1399,762,901]" pageId="12" pageNumber="13">
The distribution of limestone habitat in the Halaban area is not very well known, and the exact location of the
<typeStatus id="54EEFFE1FFFFFA4AD8443B0BFE8F6E3B" box="[262,313,800,826]" pageId="12" pageNumber="13">type</typeStatus>
collection is not clear. Whether the species is already extinct, or in fact thriving in a naturally small population needs further fieldwork in the Halaban region; we assess
<taxonomicName id="4C553AC0FFFFFA4ADD743B6DFB456E5E" authorityName="M. Hughes" authorityYear="2015" box="[1078,1267,837,863]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="halabanensis">
<emphasis id="B9219D51FFFFFA4ADD743B6DFB456E5E" box="[1078,1267,837,863]" italics="true" pageId="12" pageNumber="13">B. halabanensis</emphasis>
</taxonomicName>
to be Data Deficient (
<bibRefCitation id="EFC43CB2FFFFFA4AD87A3B40FE736E84" author="IUCN" box="[312,453,875,901]" pageId="12" pageNumber="13" refId="ref18379" refString="IUCN 2012. IUCN Red List Categories and Criteria: Version 3.1. Second Edition. IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>