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<document id="372AB67B9F9FA148F496E6BDE5C334A8" ID-DOI="10.5281/zenodo.4645780" ID-ISSN="1942-1354" ID-Zenodo-Dep="4645780" ID-ZooBank="CF745EEC-5265-4AA6-A213-EB5AA1F6A323" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617040667453" checkinUser="felipe" docAuthor="Otto, Robert L." docDate="2015" docId="03FCEC7FFFC5FF982CC7FB748E74A1D3" docLanguage="en" docName="InsectaMundi.0404.1-11.pdf" docOrigin="Insecta Mundi 2015 (404)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Bermilloides lumawigi Otto 2015, new species" docType="treatment" docUuid="18AC36AD-1734-4145-9206-7B8A880AFA00" docUuidSource="ZooBank" docVersion="3" lastPageNumber="7" masterDocId="FFC59407FFC3FF902C07FFB58B3FA529" masterDocTitle="A new genus and four new species of false click beetles (Coleoptera: Eucnemidae) from Southeast Asia" masterLastPageNumber="11" masterPageNumber="1" pageNumber="5" updateTime="1698895445649" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="9B622CBE9351A99F8D9622FC0F3F7278">A new genus and four new species of false click beetles (Coleoptera: Eucnemidae) from Southeast Asia</mods:title>
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<mods:namePart id="A3704D8BCC4A03E8532B65515AAF88D4">Otto, Robert L.</mods:namePart>
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<mods:title id="6EED4A1CC447CE9F0684D57F64BAEC95">Insecta Mundi</mods:title>
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<mods:date id="9481B813C60499D50D59DAEE78BD170B">2015</mods:date>
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<mods:number id="0F4507689A3DD6255D1007F88F6973C9">2015-02-06</mods:number>
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<mods:identifier id="38BF9598F0FFA155EFD7050422917B92" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03FCEC7FFFC5FF982CC7FB748E74A1D3" LSID="urn:lsid:zoobank.org:act:18AC36AD-1734-4145-9206-7B8A880AFA00" httpUri="http://treatment.plazi.org/id/03FCEC7FFFC5FF982CC7FB748E74A1D3" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5">
<subSubSection id="C34F0EE2FFC5FF962CC7FB7489CDA1F0" box="[192,754,1217,1242]" pageId="6" pageNumber="5" type="nomenclature">
<paragraph id="8BEA5D69FFC5FF962CC7FB7489CDA1F0" blockId="6.[192,754,1217,1274]" box="[192,754,1217,1242]" pageId="6" pageNumber="5">
<heading id="D0A2EA05FFC5FF962CC7FB7489CDA1F0" bold="true" box="[192,754,1217,1242]" fontSize="10" level="2" pageId="6" pageNumber="5" reason="3">
<emphasis id="B921817BFFC5FF962CC7FB7489CDA1F0" bold="true" box="[192,754,1217,1242]" pageId="6" pageNumber="5">
<taxonomicName id="4C5526EAFFC5FF962CC7FB74897FA1F0" authority="Otto" authorityName="Otto" authorityYear="2015" box="[192,576,1217,1242]" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="lumawigi" status="sp. nov.">
<emphasis id="B921817BFFC5FF962CC7FB748AC6A1F3" bold="true" box="[192,505,1217,1242]" italics="true" pageId="6" pageNumber="5">Bermilloides lumawigi</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC5FF962E4AFB7789CDA1F0" box="[589,754,1217,1242]" pageId="6" pageNumber="5" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFC5FF982CC7FB578A09A4F3" lastPageId="8" lastPageNumber="7" pageId="6" pageNumber="5" type="description">
<paragraph id="8BEA5D69FFC5FF962CC7FB578A1BA1D0" blockId="6.[192,754,1217,1274]" box="[192,292,1249,1274]" pageId="6" pageNumber="5">
<figureCitation id="136E41ECFFC5FF962CC7FB578A1BA1D0" box="[192,292,1249,1274]" captionStart="Figures 15" captionStartId="4.[192,287,1843,1865]" captionTargetBox="[245,1385,214,1794]" captionTargetId="figure-76@4.[221,1407,200,1800]" captionTargetPageId="4" captionText="Figures 15. Two new species of Eucnemidae. 1) Calyptocerus iridis Otto, n. sp., dorsal of holotype. 2) Calyptocerus iridis Otto, n. sp., venter of holotype. 3) Calyptocerus iridis Otto, n. sp., aedeagus of paratype, dorsal view. 4) Bermilloides lumawigi Otto, n. sp., dorsal of holotype. 5) Bermilloides lumawigi Otto, n. sp., venter of holotype." figureDoi="http://doi.org/10.5281/zenodo.4645784" httpUri="https://zenodo.org/record/4645784/files/figure.png" pageId="6" pageNumber="5">Fig. 45</figureCitation>
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CC7FA948A57A073" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CC7FA948962A013" bold="true" box="[192,605,1313,1338]" pageId="6" pageNumber="5">
Description. Female
<typeStatus id="54EEE3CBFFC5FF962DDAFA978967A013" box="[477,600,1314,1338]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
:
</emphasis>
Length, 14.0 mm. Width,
<quantity id="4CADF08CFFC5FF962F9BFA9488C5A010" box="[924,1018,1313,1337]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="5" unit="mm" value="5.5">5.5 mm</quantity>
. Body color uniformly dark brownblack (
<figureCitation id="136E41ECFFC5FF962D15FAF78A65A073" box="[274,346,1346,1370]" captionStart="Figures 15" captionStartId="4.[192,287,1843,1865]" captionTargetBox="[245,1385,214,1794]" captionTargetId="figure-76@4.[221,1407,200,1800]" captionTargetPageId="4" captionText="Figures 15. Two new species of Eucnemidae. 1) Calyptocerus iridis Otto, n. sp., dorsal of holotype. 2) Calyptocerus iridis Otto, n. sp., venter of holotype. 3) Calyptocerus iridis Otto, n. sp., aedeagus of paratype, dorsal view. 4) Bermilloides lumawigi Otto, n. sp., dorsal of holotype. 5) Bermilloides lumawigi Otto, n. sp., venter of holotype." figureDoi="http://doi.org/10.5281/zenodo.4645784" httpUri="https://zenodo.org/record/4645784/files/figure.png" pageId="6" pageNumber="5">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7FAD78A63A093" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7FAD78A7EA053" bold="true" box="[240,321,1378,1402]" pageId="6" pageNumber="5">Head:</emphasis>
Densely rugose; subspherical; surface dullish; delicate, median carina extremely short, confined to the vertex; epistomal part of epicranium with delicate, short median carina eyes slightly protuberant.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7FA748FEDA0F3" blockId="6.[192,1441,1313,1850]" box="[240,1234,1473,1498]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7FA748A42A0F0" bold="true" box="[240,381,1473,1497]" pageId="6" pageNumber="5">Antennae:</emphasis>
Filiform, reaching just beyond the pronotal hind angles; dark brown.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7FA578E38A333" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7FA578ABCA0D3" bold="true" box="[240,387,1506,1530]" pageId="6" pageNumber="5">Pronotum:</emphasis>
Dark brown-black, dullish with short, yellow erect setae; surfaces densely rugose; wider than long, with large sharp, hind angles; lateral sides arcuate; disc convex; base sinuous.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7F9948AAEA373" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7F9948ABBA310" bold="true" box="[240,388,1569,1593]" pageId="6" pageNumber="5">Scutellum:</emphasis>
Dark brown-black, dull, wide, sparsely setose, with median carina, sub-triangular and distally rounded.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7F9D78FEDA393" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7F9D78A5EA353" bold="true" box="[240,353,1634,1658]" pageId="6" pageNumber="5">Elytron:</emphasis>
Convex, elongate, gradually narrowed from humeri to apices; conjoined tightly at apex; somewhat dullish with short, yellow recumbent setae; dark brown-black; length 10.0 mm; width
<quantity id="4CADF08CFFC5FF96296BF9348BD0A390" metricMagnitude="-3" metricUnit="m" metricValue="2.75" pageId="6" pageNumber="5" unit="mm" value="2.75">2.75 mm</quantity>
at humeri; humeri striate; disc striate; interstices elevated, transversely rugose.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7F9778810A3D0" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7F9778A07A3F3" bold="true" box="[240,312,1730,1754]" pageId="6" pageNumber="5">Legs:</emphasis>
Femur and tibiae dark brown; tarsi dark brown and somewhat shiny; surfaces closely punctate to rugose, with short, yellow recumbent setae.
</paragraph>
<paragraph id="8BEA5D69FFC5FF962CF7F8B78A26A213" blockId="6.[192,1441,1313,1850]" pageId="6" pageNumber="5">
<emphasis id="B921817BFFC5FF962CF7F8B78A71A233" bold="true" box="[240,334,1794,1818]" pageId="6" pageNumber="5">Venter</emphasis>
(
<figureCitation id="136E41ECFFC5FF962D5BF8B78AA0A230" box="[348,415,1793,1818]" captionStart="Figures 15" captionStartId="4.[192,287,1843,1865]" captionTargetBox="[245,1385,214,1794]" captionTargetId="figure-76@4.[221,1407,200,1800]" captionTargetPageId="4" captionText="Figures 15. Two new species of Eucnemidae. 1) Calyptocerus iridis Otto, n. sp., dorsal of holotype. 2) Calyptocerus iridis Otto, n. sp., venter of holotype. 3) Calyptocerus iridis Otto, n. sp., aedeagus of paratype, dorsal view. 4) Bermilloides lumawigi Otto, n. sp., dorsal of holotype. 5) Bermilloides lumawigi Otto, n. sp., venter of holotype." figureDoi="http://doi.org/10.5281/zenodo.4645784" httpUri="https://zenodo.org/record/4645784/files/figure.png" pageId="6" pageNumber="5">Fig. 5</figureCitation>
): Somewhat shiny; dark brown-black; surface with elongate, yellow recumbent setae; rugose.
</paragraph>
<caption id="DF2A0DE1FFC4FF972CC7F8838E9EA2AF" ID-DOI="http://doi.org/10.5281/zenodo.4645786" ID-Zenodo-Dep="4645786" httpUri="https://zenodo.org/record/4645786/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,287,1846,1868]" targetBox="[243,1368,262,1818]" targetPageId="7">
<paragraph id="8BEA5D69FFC4FF972CC7F8838E9EA2AF" blockId="7.[192,1441,1846,1926]" pageId="7" pageNumber="6">
<emphasis id="B921817BFFC4FF972CC7F8838A5EA262" bold="true" box="[192,353,1846,1868]" pageId="7" pageNumber="6">Figures 610.</emphasis>
Two new species of
<taxonomicName id="4C5526EAFFC4FF972E3FF88389FAA265" authorityName="Eschscholtz" authorityYear="1829" box="[568,709,1846,1868]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="family">Eucnemidae</taxonomicName>
.
<emphasis id="B921817BFFC4FF972ECBF88389DCA265" bold="true" box="[716,739,1846,1868]" pageId="7" pageNumber="6">6)</emphasis>
<taxonomicName id="4C5526EAFFC4FF972EEEF8838F2CA262" authority="Otto" authorityName=", Otto" authorityYear="2015" box="[745,1043,1846,1867]" class="Insecta" family="Eucnemidae" genus="Spinifornax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="B921817BFFC4FF972EEEF88388E4A262" box="[745,987,1846,1867]" italics="true" pageId="7" pageNumber="6">Spinifornax elongatus</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC4FF97281BF8828F67A265" box="[1052,1112,1847,1868]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, dorsal of holotype.
<emphasis id="B921817BFFC4FF972929F8838E7AA265" bold="true" box="[1326,1349,1846,1868]" pageId="7" pageNumber="6">7)</emphasis>
<taxonomicName id="4C5526EAFFC4FF97294DF8838A46A241" authority="Otto" authorityName=", Otto" authorityYear="2015" class="Insecta" family="Eucnemidae" genus="Ceratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="antennatus" status="sp. nov.">
<emphasis id="B921817BFFC4FF97294DF8838A03A240" italics="true" pageId="7" pageNumber="6">Ceratus antennatus</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC4FF972D82F8E68AFBA241" box="[389,452,1875,1896]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, dorsal of holotype.
<emphasis id="B921817BFFC4FF972EAFF8E689FFA240" bold="true" box="[680,704,1875,1897]" pageId="7" pageNumber="6">8)</emphasis>
<taxonomicName id="4C5526EAFFC4FF972EC0F8E688E0A241" authority="Otto" authorityName=", Otto" authorityYear="2015" box="[711,991,1875,1897]" class="Insecta" family="Eucnemidae" genus="Ceratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="antennatus" status="sp. nov.">
<emphasis id="B921817BFFC4FF972EC0F8E6889DA240" box="[711,930,1875,1897]" italics="true" pageId="7" pageNumber="6">Ceratus antennatus</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC4FF972FECF8E68F15A241" box="[1003,1066,1875,1896]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, antenna of paratype.
<emphasis id="B921817BFFC4FF97292DF8E68E7EA240" bold="true" box="[1322,1345,1875,1897]" pageId="7" pageNumber="6">9)</emphasis>
<taxonomicName id="4C5526EAFFC4FF97294DF8E68A49A2AC" authority="Otto" authorityName=", Otto" authorityYear="2015" class="Insecta" family="Eucnemidae" genus="Ceratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="antennatus" status="sp. nov.">
<emphasis id="B921817BFFC4FF97294DF8E68A03A2AC" italics="true" pageId="7" pageNumber="6">Ceratus antennatus</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC4FF972D78F8C58A84A2AC" box="[383,443,1904,1925]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, venter of paratype.
<emphasis id="B921817BFFC4FF972E9CF8DA89FFA2AC" bold="true" box="[667,704,1903,1925]" pageId="7" pageNumber="6">10)</emphasis>
<taxonomicName id="4C5526EAFFC4FF972EC1F8C588E8A2AC" authority="Otto" authorityName=", Otto" authorityYear="2015" box="[710,983,1904,1925]" class="Insecta" family="Eucnemidae" genus="Ceratus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="antennatus" status="sp. nov.">
<emphasis id="B921817BFFC4FF972EC1F8C588A2A2AC" box="[710,925,1904,1925]" italics="true" pageId="7" pageNumber="6">Ceratus antennatus</emphasis>
Otto
</taxonomicName>
,
<taxonomicNameLabel id="A2123C00FFC4FF972FE6F8C58F22A2AC" box="[993,1053,1904,1925]" pageId="7" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
, aedeagus of paratype, dorsal view.
</paragraph>
</caption>
<paragraph id="8BEA5D69FFCBFF982CC7FF748A09A4F3" blockId="8.[192,1440,193,474]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FF748A73A5F0" bold="true" box="[192,332,193,217]" pageId="8" pageNumber="7">Variation.</emphasis>
Two
<typeStatus id="54EEE3CBFFCBFF982D93FF778931A5F3" box="[404,526,194,218]" pageId="8" pageNumber="7" type="paratype">paratypes</typeStatus>
were examined. The single male
<typeStatus id="54EEE3CBFFCBFF982FBCFF778F16A5F3" box="[955,1065,194,218]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
is smaller than the
<typeStatus id="54EEE3CBFFCBFF982937FF778EA3A5F3" box="[1328,1436,194,218]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
, measuring
<quantity id="4CADF08CFFCBFF982D4DFF548A9BA5D0" box="[330,420,225,249]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="7" unit="mm" value="8.5">8.5 mm</quantity>
long. Antennae are longer, extending beyond the pronotal hind angles by three segments. Antennomeres IIIVII are quadrate, while antennomeres VIIIX are longer than wide in the male
<typeStatus id="54EEE3CBFFCBFF982D04FE978A4AA413" box="[259,373,290,314]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
. Median carina is strongly indicated on the pronotum, but shorter on the head, confined to the vertex in the male specimen. The single female
<typeStatus id="54EEE3CBFFCBFF982F66FEF788F0A473" box="[865,975,322,346]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
is slightly smaller than the
<typeStatus id="54EEE3CBFFCBFF982937FEF78EA3A473" box="[1328,1436,322,346]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
, measuring 11.0 mm long. Exoskeletal structures are similar to the
<typeStatus id="54EEE3CBFFCBFF982FF3FED78F5FA453" box="[1012,1120,354,378]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
, except for the presence of a median fovea above the base of the epistomal part of epicranium. A median carina is feebly indicated on the pronotum, but strong on the head, extending from vertex midway down the frons in the female
<typeStatus id="54EEE3CBFFCBFF982CC7FE778A0DA4F3" box="[192,306,450,474]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFCBFF982CC7FDB7890FA633" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="8BEA5D69FFCBFF982CC7FDB78E9FA773" blockId="8.[192,1440,513,602]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FDB78AB5A730" bold="true" box="[192,394,513,538]" pageId="8" pageNumber="7">Type material.</emphasis>
<materialsCitation id="3B3D5734FFCBFF982D96FDB78F91A773" collectingDate="2005-06-14" collectorName="Det. R. L. Otto &amp; L. Otto" country="Malaysia" county="Borneo" determinerName="Det. R. L. Otto" location="Range" municipality="Crocker" pageId="8" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B921817BFFCBFF982D96FDB78930A733" bold="true" box="[401,527,514,538]" pageId="8" pageNumber="7">
<typeStatus id="54EEE3CBFFCBFF982D96FDB78930A733" box="[401,527,514,538]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
, female, with label data: “
<collectingCountry id="F3421DF9FFCBFF982F56FDB788D6A733" box="[849,1001,514,538]" name="Malaysia" pageId="8" pageNumber="7">MALAYSIA</collectingCountry>
:
<collectingRegion id="4991938BFFCBFF982FF4FDB78F64A733" box="[1011,1115,514,538]" country="Malaysia" name="Sabah" pageId="8" pageNumber="7">SABAH</collectingRegion>
:, (
<collectingCounty id="628B25E5FFCBFF982875FDB78FF4A733" box="[1138,1227,514,538]" pageId="8" pageNumber="7">Borneo</collectingCounty>
)
<collectingMunicipality id="6B8EC713FFCBFF9828DCFDB78E00A733" box="[1243,1343,514,538]" pageId="8" pageNumber="7">Crocker</collectingMunicipality>
,
<location id="8E8A0BB2FFCBFF98294CFDB78EA3A733" LSID="urn:lsid:plazi:treatment:03FCEC7FFFC5FF982CC7FB748E74A1D3:8E8A0BB2FFCBFF98294CFDB78EA3A733" box="[1355,1436,514,538]" country="Malaysia" county="Borneo" municipality="Crocker" name="Range" pageId="8" pageNumber="7" stateProvince="Sabah">Range</location>
;
<date id="FFEB7BA9FFCBFF982CC7FD948A77A710" box="[192,328,545,569]" pageId="8" pageNumber="7" value="2005-06-14">
<collectingDate id="EFAF8241FFCBFF982CC7FD948A77A710" box="[192,328,545,569]" pageId="8" pageNumber="7" value="2005-06-14">14-VI-2005</collectingDate>
</date>
” / “
<taxonomicName id="4C5526EAFFCBFF982D75FD978AD4A713" authorityName="Bonvouloir" authorityYear="1871" box="[370,491,546,570]" class="Insecta" family="Eucnemidae" genus="Bermillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982D75FD978AD4A713" box="[370,491,546,570]" italics="true" pageId="8" pageNumber="7">Bermillus</emphasis>
</taxonomicName>
, sp.,
<determinerName id="7E36B1ADFFCBFF982E2EFD9789E7A713" box="[553,728,546,570]" pageId="8" pageNumber="7">
<collectorName id="26A038BFFFCBFF982E2EFD9789E7A713" box="[553,728,546,570]" pageId="8" pageNumber="7">Det. R.L. Otto</collectorName>
</determinerName>
, 2012” / “
<emphasis id="B921817BFFCBFF982F41FD9488CCA710" bold="true" box="[838,1011,545,569]" pageId="8" pageNumber="7">
<typeStatus id="54EEE3CBFFCBFF982F41FD9488D1A710" box="[838,1006,545,569]" pageId="8" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
:
</emphasis>
,
<taxonomicName id="4C5526EAFFCBFF982806FD978FA3A713" authorityName="Otto" authorityYear="2015" box="[1025,1180,546,570]" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982806FD978FA3A713" box="[1025,1180,546,570]" italics="true" pageId="8" pageNumber="7">Bermilloides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5526EAFFCBFF9828ADFD978A43A773" authority=", Otto, det. R." authorityName="Otto, Det. R." class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="lumawigi">
<emphasis id="B921817BFFCBFF9828ADFD978E20A713" box="[1194,1311,546,570]" italics="true" pageId="8" pageNumber="7">lumawigi</emphasis>
, Otto, det. R.
<collectorName id="26A038BFFFCBFF982CDCFDF78A0AA773" box="[219,309,578,602]" pageId="8" pageNumber="7">L. Otto</collectorName>
, 2014
</taxonomicName>
” (
<specimenCount id="9D5396E0FFCBFF982D90FDF78A94A770" box="[407,427,578,601]" pageId="8" pageNumber="7" type="female"></specimenCount>
handwritten behind species name on label) [red printed label]
</materialsCitation>
.
<materialsCitation id="3B3D5734FFCBFF9828BEFDF78EA3A773" box="[1209,1436,578,602]" collectionCode="FSCA" country="Malaysia" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54EEE3CBFFCBFF9828BEFDF78E17A773" box="[1209,1320,578,602]" pageId="8" pageNumber="7" type="holotype">Holotype</typeStatus>
in
<collectionCode id="ED44C5ACFFCBFF982948FDF78EA3A773" box="[1359,1436,578,602]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="8" pageNumber="7" type="Museum">FSCA</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEA5D69FFCBFF982CC7FD37890FA633" blockId="8.[192,1440,642,794]" pageId="8" pageNumber="7">
<materialsCitation id="3B3D5734FFCBFF982CC7FD37899AA793" collectingDate="2014-02" collectionCode="AAC" country="Philippines" location="COMPOSTELA" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Compostela" typeStatus="paratype">
<emphasis id="B921817BFFCBFF982CC7FD378A6BA7B3" bold="true" box="[192,340,642,666]" pageId="8" pageNumber="7">
<typeStatus id="54EEE3CBFFCBFF982CC7FD378A71A7B3" box="[192,334,642,666]" pageId="8" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
2, each with label data: 1, “
<collectingCountry id="F3421DF9FFCBFF982E9BFD378869A7B3" box="[668,854,642,666]" name="Philippines" pageId="8" pageNumber="7">PHILIPPINES</collectingCountry>
, MINDANAO, MASARA,
<collectingRegion id="4991938BFFCBFF982897FD378E69A7B3" box="[1168,1366,642,666]" country="Philippines" name="Compostela" pageId="8" pageNumber="7">COMPOSTELA</collectingRegion>
, VAL- LEY,
<date id="FFEB7BA9FFCBFF982D0EFD178A5DA793" box="[265,354,674,698]" pageId="8" pageNumber="7" value="2014-02">
<collectingDate id="EFAF8241FFCBFF982D0EFD178A5DA793" box="[265,354,674,698]" pageId="8" pageNumber="7" value="2014-02">ii 2014</collectingDate>
</date>
, lg: I. LUMAWIG” (
<collectionCode id="ED44C5ACFFCBFF982E59FD1789A2A793" box="[606,669,674,698]" pageId="8" pageNumber="7">AAC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3D5734FFCBFF982EB5FD178EA3A7D3" collectingDate="2014-05" collectionCode="GERP" country="Philippines" county="Mindanao" location="Ismael Lumawig" municipality="Panamokan" pageId="8" pageNumber="7" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bukidnon" typeStatus="paratype">
1, “
<collectingCountry id="F3421DF9FFCBFF982EDAFD1788A4A793" box="[733,923,674,698]" name="Philippines" pageId="8" pageNumber="7">PHILIPPINES</collectingCountry>
:,
<collectingCounty id="628B25E5FFCBFF982FAAFD178F13A793" box="[941,1068,674,698]" pageId="8" pageNumber="7">Mindanao</collectingCounty>
,
<collectingMunicipality id="6B8EC713FFCBFF98283CFD178FEDA793" box="[1083,1234,674,698]" pageId="8" pageNumber="7">Panamokan</collectingMunicipality>
,
<collectingRegion id="4991938BFFCBFF9828E7FD178E63A793" box="[1248,1372,674,698]" country="Philippines" name="Bukidnon" pageId="8" pageNumber="7">Bukidnon</collectingRegion>
,
<date id="FFEB7BA9FFCBFF98296DFD178BC1A7F3" pageId="8" pageNumber="7" value="2014-05">
<collectingDate id="EFAF8241FFCBFF98296DFD178BC1A7F3" pageId="8" pageNumber="7" value="2014-05">May 2014</collectingDate>
</date>
,
<location id="8E8A0BB2FFCBFF982D0EFD778AE8A7F3" LSID="urn:lsid:plazi:treatment:03FCEC7FFFC5FF982CC7FB748E74A1D3:8E8A0BB2FFCBFF982D0EFD778AE8A7F3" box="[265,471,706,730]" country="Philippines" county="Mindanao" municipality="Panamokan" name="Ismael Lumawig" pageId="8" pageNumber="7" stateProvince="Bukidnon">Ismael Lumawig</location>
” (
<collectionCode id="ED44C5ACFFCBFF982DF7FD77897DA7F3" box="[496,578,706,730]" pageId="8" pageNumber="7">GERP</collectionCode>
). Each
<typeStatus id="54EEE3CBFFCBFF982E9EFD778838A7F3" box="[665,775,706,730]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
labeled: “
<emphasis id="B921817BFFCBFF982F78FD748F18A7F0" bold="true" box="[895,1063,705,729]" pageId="8" pageNumber="7">
<typeStatus id="54EEE3CBFFCBFF982F78FD748F1CA7F0" box="[895,1059,705,729]" pageId="8" pageNumber="7" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
,
<taxonomicName id="4C5526EAFFCBFF982831FD778FEEA7F3" authorityName="Otto" authorityYear="2015" box="[1078,1233,706,730]" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982831FD778FEEA7F3" box="[1078,1233,706,730]" italics="true" pageId="8" pageNumber="7">Bermilloides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5526EAFFCBFF9828D8FD778A88A7D3" authority=", Otto, Det. R. L. Otto, 2014" authorityName=", Otto, Det. R. L. Otto" authorityYear="2014" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="lumawigi">
<emphasis id="B921817BFFCBFF9828D8FD778E6BA7F3" box="[1247,1364,706,730]" italics="true" pageId="8" pageNumber="7">lumawigi</emphasis>
, Otto, Det. R.L. Otto, 2014
</taxonomicName>
” (either
<specimenCount id="9D5396E0FFCBFF982E24FD578908A7D0" box="[547,567,738,761]" pageId="8" pageNumber="7" type="male"></specimenCount>
or
<specimenCount id="9D5396E0FFCBFF982E58FD57894CA7D0" box="[607,627,738,761]" pageId="8" pageNumber="7" type="female"></specimenCount>
handwritten behind species name on label) [yellow printed label]
</materialsCitation>
.
<typeStatus id="54EEE3CBFFCBFF982CC7FCB78A02A633" box="[192,317,770,794]" pageId="8" pageNumber="7" type="paratype">Paratypes</typeStatus>
in
<collectionCode id="ED44C5ACFFCBFF982D63FCB78AA1A633" box="[356,414,770,794]" pageId="8" pageNumber="7">AAC</collectionCode>
and
<collectionCode id="ED44C5ACFFCBFF982DDCFCB78913A633" box="[475,556,770,794]" pageId="8" pageNumber="7">GERP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFCBFF982CC7FCF48A9BA653" pageId="8" pageNumber="7" type="distribution">
<paragraph id="8BEA5D69FFCBFF982CC7FCF48A9BA653" blockId="8.[192,1440,833,890]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FCF48A4BA670" bold="true" box="[192,372,833,857]" pageId="8" pageNumber="7">Distribution.</emphasis>
The species is known from a single locality in
<collectingCountry id="F3421DF9FFCBFF982FADFCF78F24A673" box="[938,1051,834,858]" name="Malaysia" pageId="8" pageNumber="7">Malaysia</collectingCountry>
and two localities on Mindanao in the
<collectingCountry id="F3421DF9FFCBFF982D17FCD78A9FA653" box="[272,416,866,890]" name="Philippines" pageId="8" pageNumber="7">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFCBFF982CC7FC148EADA6F3" pageId="8" pageNumber="7" type="biology_ecology">
<paragraph id="8BEA5D69FFCBFF982CC7FC148EADA6F3" blockId="8.[192,1440,929,986]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FC148A0EA690" bold="true" box="[192,305,929,953]" pageId="8" pageNumber="7">Biology.</emphasis>
The
<typeStatus id="54EEE3CBFFCBFF982D6AFC178A9DA693" box="[365,418,930,954]" pageId="8" pageNumber="7">type</typeStatus>
locality is an upper montane forested environment. Two Philippine specimens were taken in tropical montane forests and mountainous forests. Developmental stages remain unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFCBFF982CC7FBB78E9FA133" box="[192,1440,1026,1050]" pageId="8" pageNumber="7" type="etymology">
<paragraph id="8BEA5D69FFCBFF982CC7FBB78E9FA133" blockId="8.[192,1440,1026,1050]" box="[192,1440,1026,1050]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FBB78A63A133" bold="true" box="[192,348,1026,1050]" pageId="8" pageNumber="7">Etymology.</emphasis>
The specific epithet is named in honor of my friend Ismael Lumawig from the
<collectingCountry id="F3421DF9FFCBFF98290BFBB78EA3A133" box="[1292,1436,1026,1050]" name="Philippines" pageId="8" pageNumber="7">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F0EE2FFCBFF982CC7FBF78E74A1D3" pageId="8" pageNumber="7" type="discussion">
<paragraph id="8BEA5D69FFCBFF982CC7FBF78E74A1D3" blockId="8.[192,1440,1090,1274]" pageId="8" pageNumber="7">
<emphasis id="B921817BFFCBFF982CC7FBF78A7BA173" bold="true" box="[192,324,1090,1114]" pageId="8" pageNumber="7">Remarks.</emphasis>
Adults of
<taxonomicName id="4C5526EAFFCBFF982DCCFBF78959A173" authorityName="Otto" authorityYear="2015" box="[459,614,1090,1114]" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982DCCFBF78959A173" box="[459,614,1090,1114]" italics="true" pageId="8" pageNumber="7">Bermilloides</emphasis>
</taxonomicName>
superficially resemble the Southeast Asian genus
<taxonomicName id="4C5526EAFFCBFF9828ECFBF78E5BA173" authorityName="Bonvouloir" authorityYear="1871" box="[1259,1380,1090,1114]" class="Insecta" family="Eucnemidae" genus="Bermillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF9828ECFBF78E5BA173" box="[1259,1380,1090,1114]" italics="true" pageId="8" pageNumber="7">Bermillus</emphasis>
</taxonomicName>
, but differ in the structure of the antennae, being quadrate to longer than wide and filiform in
<emphasis id="B921817BFFCBFF982902FBD78E9FA153" box="[1285,1440,1122,1146]" italics="true" pageId="8" pageNumber="7">Bermilliodes</emphasis>
and wider than long in
<taxonomicName id="4C5526EAFFCBFF982DE1FB378960A1B3" authorityName="Bonvouloir" authorityYear="1871" box="[486,607,1154,1178]" class="Insecta" family="Eucnemidae" genus="Bermillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982DE1FB378960A1B3" box="[486,607,1154,1178]" italics="true" pageId="8" pageNumber="7">Bermillus</emphasis>
</taxonomicName>
. The antennal grooves are medially undefined in
<taxonomicName id="4C5526EAFFCBFF9828CAFB378E57A1B3" authorityName="Otto" authorityYear="2015" box="[1229,1384,1154,1178]" class="Insecta" family="Eucnemidae" genus="Bermilloides" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF9828CAFB378E57A1B3" box="[1229,1384,1154,1178]" italics="true" pageId="8" pageNumber="7">Bermilloides</emphasis>
</taxonomicName>
and well defined in
<taxonomicName id="4C5526EAFFCBFF982D87FB178AC6A193" authorityName="Bonvouloir" authorityYear="1871" box="[384,505,1186,1210]" class="Insecta" family="Eucnemidae" genus="Bermillus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982D87FB178AC6A193" box="[384,505,1186,1210]" italics="true" pageId="8" pageNumber="7">Bermillus</emphasis>
</taxonomicName>
. Slender mandibles, as well as straight sex combs along with the length of the prothoracic tarsomere I in males, places the new genus in the tribe
<taxonomicName id="4C5526EAFFCBFF982818FB778F85A1F3" authorityName="Muona" authorityYear="1993" box="[1055,1210,1218,1242]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Anelastidini">Anelastidini</taxonomicName>
. Form and basally toothed tarsal claws will distinguish it from
<taxonomicName id="4C5526EAFFCBFF982EE5FB5788E4A1D3" authority="Du Val" authorityName="Du Val" box="[738,987,1250,1274]" class="Insecta" family="Eucnemidae" genus="Anelastidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="B921817BFFCBFF982EE5FB578842A1D3" box="[738,893,1250,1274]" italics="true" pageId="8" pageNumber="7">Anelastidius</emphasis>
Du Val
</taxonomicName>
within the tribe
<taxonomicName id="4C5526EAFFCBFF9828ABFB578E78A1D3" authorityName="Muona" authorityYear="1993" box="[1196,1351,1250,1274]" class="Insecta" family="Eucnemidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Anelastidini">Anelastidini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>