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<document id="B8EF47C6C69E70192C372DA6BA13666C" ID-DOI="10.11646/zootaxa.5318.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8181850" ID-ZooBank="10C3728D-B718-4074-AC83-142A98A88895" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1690274897597" checkinUser="plazi" docAuthor="Schneider, Clément &amp; Panina, Ksenia" docDate="2023" docId="03FC6644DC452C7AFF79FA446A54FA47" docLanguage="en" docName="zootaxa.5318.4.2.pdf" docOrigin="Zootaxa 5318 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megalothorax boerneri Schneider &amp; Panina 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="484" masterDocId="FFC51E3CDC432C70FFEEFFEA6B34FFF7" masterDocTitle="Revision of Megalothorax incertus Börner, 1903 reveals it to be another widespread Palearctic species of the genus (Collembola, Neelidae)" masterLastPageNumber="488" masterPageNumber="474" pageNumber="480" updateTime="1690275829247" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8E9C387E603DBDB27B5E715FED88988F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BBFBA38FA28B4BBD22B03DA51ACF5306">
<mods:title id="72893EB58624D5D793785617F30C20DF">Revision of Megalothorax incertus Börner, 1903 reveals it to be another widespread Palearctic species of the genus (Collembola, Neelidae)</mods:title>
</mods:titleInfo>
<mods:name id="0E26A94DBA6EEC2C1016B4DFAAA25BDF" type="personal">
<mods:role id="1FF61F5C06F4F32CB17F5AD518657231">
<mods:roleTerm id="1272AE0B71AC0586BD69DC743AFDB634">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="254CA09BAEAC88FD13FF98373DF1C7A4">Schneider, Clément</mods:namePart>
<mods:nameIdentifier id="F2080204B9F30DDBD20CBC3DF84AA077" type="ORCID">0000-0003-3743-9319</mods:nameIdentifier>
<mods:affiliation id="24709CBB203B800622E9624686C87328">Abteilung Bodenzoologie, Senckenberg Gesellschaft für Naturforschung, 02826 Görlitz, Germany clement. schneider @ senckenberg. de; https: // orcid. org / 0000 - 0003 - 3743 - 9319</mods:affiliation>
<mods:nameIdentifier id="9440D766680BC815A3B112681403D349" type="email">clement.schneider@senckenberg.de</mods:nameIdentifier>
</mods:name>
<mods:name id="77AB2A584072B9D3A774EE1919141498" type="personal">
<mods:role id="F89FAF389726951422739E5F618767EF">
<mods:roleTerm id="10EB79297BF36CD400A6E14AE5B100C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33A4F945C3B27FF532083C081EC0792D">Panina, Ksenia</mods:namePart>
<mods:affiliation id="F46EB05BB12EEE23C578E83CD00ABD6C">Moscow State Pedagogical University, Kibalchicha str., 6, korp. 3, Moscow, 129278, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8F89D42786BA3A9D011F8C5A617E03A4">text</mods:typeOfResource>
<mods:relatedItem id="EFCC5B85C75ABE0585ADB03DCF3CB72E" type="host">
<mods:titleInfo id="91E01A5B1ABD7F682FE4283FE2A6CA9A">
<mods:title id="6E6D33263B72EC179A3CE272E1869673">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E715297413118D17D69C9AD42014463B">
<mods:date id="8C5A4DC02C3F28333669FC9DEAECD842">2023</mods:date>
<mods:detail id="74D8105CEECDB1F1BD759FF50E4EE04A" type="pubDate">
<mods:number id="C8760684AE5A15092A8224B9D8309013">2023-07-21</mods:number>
</mods:detail>
<mods:detail id="146611A7702C1B0E566E6420D2FA044C" type="volume">
<mods:number id="F72026907138B169093854619F007789">5318</mods:number>
</mods:detail>
<mods:detail id="B5B8A6FFE17F5CE34A69C2AC410BC820" type="issue">
<mods:number id="562D1DC7FE3DED49A0A47917F475B12D">4</mods:number>
</mods:detail>
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<mods:start id="E35258311ACEBF70F7EC6653D2C7E316">474</mods:start>
<mods:end id="2892CBF109D6ECE715AAB8170577614F">488</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="55EDFD00166450E9428FB67127588E85">
<mods:url id="18DAD41A5B763A5966A40C678EFF1DFD">http://dx.doi.org/10.11646/zootaxa.5318.4.2</mods:url>
</mods:location>
<mods:classification id="5D2DA4190DB213167A50F76B41EFC979">journal article</mods:classification>
<mods:identifier id="FD013FD1EAB55086B62236B57D0D9587" type="DOI">10.11646/zootaxa.5318.4.2</mods:identifier>
<mods:identifier id="03ABCF3A6BB7D58790CAC069DE46BF31" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E39E431AE34498A4C74C9548E0334888" type="Zenodo-Dep">8181850</mods:identifier>
<mods:identifier id="595933B431EF307B69A78D27E0D69CA6" type="ZooBank">10C3728D-B718-4074-AC83-142A98A88895</mods:identifier>
</mods:mods>
<treatment id="03FC6644DC452C7AFF79FA446A54FA47" LSID="urn:lsid:plazi:treatment:03FC6644DC452C7AFF79FA446A54FA47" httpUri="http://treatment.plazi.org/id/03FC6644DC452C7AFF79FA446A54FA47" lastPageId="10" lastPageNumber="484" pageId="6" pageNumber="480">
<subSubSection id="C34F84D9DC452C76FF79FA446921FA3E" box="[151,533,1454,1481]" pageId="6" pageNumber="480" type="nomenclature">
<paragraph id="8BEAD752DC452C76FF79FA446921FA3E" blockId="6.[151,533,1454,1517]" box="[151,533,1454,1481]" pageId="6" pageNumber="480">
<heading id="D0A2603EDC452C76FF79FA446921FA3E" bold="true" box="[151,533,1454,1481]" fontSize="11" level="1" pageId="6" pageNumber="480" reason="1">
<emphasis id="B9210B40DC452C76FF79FA446921FA3E" bold="true" box="[151,533,1454,1481]" pageId="6" pageNumber="480">
<taxonomicName id="4C55ACD1DC452C76FF79FA446A85FA3E" authority="Schneider &amp; Panina, 2023" authorityName="Schneider &amp; Panina" authorityYear="2023" box="[151,433,1454,1481]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="6" pageNumber="480" phylum="Arthropoda" rank="species" species="boerneri" status="sp. nov.">
<emphasis id="B9210B40DC452C76FF79FA446A85FA3E" bold="true" box="[151,433,1454,1481]" italics="true" pageId="6" pageNumber="480">Megalothorax boerneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212B63BDC452C76FE56FA456921FA3E" box="[440,533,1455,1481]" pageId="6" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F84D9DC452C76FF79FA38686AF954" pageId="6" pageNumber="480" type="description">
<paragraph id="8BEAD752DC452C76FF79FA386AE5FA1A" blockId="6.[151,533,1454,1517]" box="[151,465,1490,1517]" pageId="6" pageNumber="480">
<figureCitation id="136ECBD7DC452C76FF79FA386BCFFA1A" box="[151,251,1490,1517]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,250,1437,1461]" captionStartId-1="8.[151,250,1018,1042]" captionStartId-2="9.[151,250,1743,1767]" captionTargetBox-0="[156,1432,186,1408]" captionTargetBox-1="[156,1431,186,989]" captionTargetBox-2="[156,1432,186,1714]" captionTargetId-0="figure-19@7.[151,1436,181,1413]" captionTargetId-1="figure-17@8.[151,1436,181,994]" captionTargetId-2="figure-19@9.[151,1436,181,1719]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." captionText-1="FIGURE 6. Megalothorax boerneri sp. nov., trunk tergites chaetotaxy (Th. I–Abd. V)." captionText-2="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi-0="http://doi.org/10.5281/zenodo.8181862" figureDoi-1="http://doi.org/10.5281/zenodo.8181864" figureDoi-2="http://doi.org/10.5281/zenodo.8181866" httpUri-0="https://zenodo.org/record/8181862/files/figure.png" httpUri-1="https://zenodo.org/record/8181864/files/figure.png" httpUri-2="https://zenodo.org/record/8181866/files/figure.png" pageId="6" pageNumber="480">Figs 5–7</figureCitation>
,
<figureCitation id="136ECBD7DC452C76FEE8FA396A78FA1A" box="[262,332,1491,1517]" captionStart="FIGURE 8" captionStartId="10.[151,250,1251,1275]" captionTargetBox="[156,1431,186,1223]" captionTargetId="figure-17@10.[151,1436,181,1228]" captionTargetPageId="10" captionText="FIGURE 8. Chaetotaxy diagrams: M. incertus sensu nov. (A) head, (B) trunk, M. boerneri sp. nov. (A) head, (B) trunk." figureDoi="http://doi.org/10.5281/zenodo.8181868" httpUri="https://zenodo.org/record/8181868/files/figure.png" pageId="6" pageNumber="480">8C, D</figureCitation>
;
<tableCitation id="C6D7E2E9DC452C76FEB6FA386AE5FA1A" box="[344,465,1490,1517]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="3.[151,239,1164,1188]" captionStartId-1="6.[151,239,548,572]" captionText-0="TABLE 1. Genbank accession numbers." captionText-1="TABLE 2. Differential diagnosis of the closest relatives of M. incertus sensu nov." pageId="6" pageNumber="480">Tables 1–2</tableCitation>
</paragraph>
<paragraph id="8BEAD752DC452C76FF79F9F0686AF954" blockId="6.[151,1437,1562,2021]" pageId="6" pageNumber="480">
<materialsCitation id="3B3DDD0FDC452C76FF79F9F0686EF954" collectingDate="2011-04-10" collectorName="C. Schneider &amp; Fr. &amp; All &amp; Apterygota &amp; Senckenberg Museum &amp; Naturkunde G" country="France" county="Charmes" latitude="47.9262" location="Material" longLatPrecision="6" longitude="5.342" municipality="Haute-Marne" pageId="6" pageNumber="480" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<location id="8E8A8189DC452C76FF79F9F06BC2F9C3" LSID="urn:lsid:plazi:treatment:03FC6644DC452C7AFF79FA446A54FA47:8E8A8189DC452C76FF79F9F06BC2F9C3" box="[151,246,1562,1588]" country="France" county="Charmes" latitude="47.9262" longLatPrecision="6" longitude="5.342" municipality="Haute-Marne" name="Material" pageId="6" pageNumber="480">Material</location>
examined.
<typeStatus id="54EE69F0DC452C76FE92F9F06ADDF9C3" box="[380,489,1562,1588]" pageId="6" pageNumber="480">Holotype</typeStatus>
: female on slide (SMNG-APT-AA00023),
<collectingCountry id="F34297C2DC452C76FC3DF9F16F16F9C2" box="[979,1058,1563,1589]" name="France" pageId="6" pageNumber="480">France</collectingCountry>
,
<collectingMunicipality id="6B8E4D28DC452C76FBC1F9F16FF3F9C2" box="[1071,1223,1563,1589]" pageId="6" pageNumber="480">Haute-Marne</collectingMunicipality>
,
<collectingCounty id="628BAFDEDC452C76FB3AF9F06E0EF9C3" box="[1236,1338,1562,1588]" pageId="6" pageNumber="480">Charmes</collectingCounty>
,
<date id="FFEBF192DC452C76FAA6F9F16BE5F9AE" pageId="6" pageNumber="480" value="2011-04-10">
<collectingDate id="EFAF087ADC452C76FAA6F9F16BE5F9AE" pageId="6" pageNumber="480" value="2011-04-10">10 Apr. 2011</collectingDate>
</date>
,
<geoCoordinate id="EE61B195DC452C76FF34F9D46A63F9AF" box="[218,343,1598,1624]" degrees="47.9262" direction="north" orientation="latitude" pageId="6" pageNumber="480" precision="5" value="47.9262">47.9262°N</geoCoordinate>
,
<geoCoordinate id="EE61B195DC452C76FE8EF9D46AF8F9AE" box="[352,460,1598,1625]" degrees="5.3420" direction="east" orientation="longitude" pageId="6" pageNumber="480" precision="5" value="5.342">5.3420°E</geoCoordinate>
, mosses growing on a rock, leg.
<collectorName id="26A0B284DC452C76FCC4F9D5688EF9AF" box="[810,954,1598,1625]" pageId="6" pageNumber="480">C. Schneider</collectorName>
, original collection code:
<collectorName id="26A0B284DC452C76FB3DF9D56FC0F9AE" box="[1235,1268,1599,1625]" pageId="6" pageNumber="480">Fr.</collectorName>
021.
<typeStatus id="54EE69F0DC452C76FAC9F9D56EACF9AE" box="[1319,1432,1599,1625]" pageId="6" pageNumber="480">Paratypes</typeStatus>
:
<specimenCount id="9D531CDBDC452C76FF79F9886A18F98B" box="[151,300,1634,1660]" pageId="6" pageNumber="480" type="female">three females</specimenCount>
on slides (SMNG-APT-AA0024–26), same data as the holotype.
<collectorName id="26A0B284DC452C76FBEEF9886F10F98B" box="[1024,1060,1634,1660]" pageId="6" pageNumber="480">All</collectorName>
types deposited in the
<collectorName id="26A0B284DC452C76FAF0F9896EA8F98A" box="[1310,1436,1635,1661]" pageId="6" pageNumber="480">Apterygota</collectorName>
collection of the
<collectorName id="26A0B284DC452C76FEBAF96C6979F956" box="[340,589,1670,1697]" pageId="6" pageNumber="480">Senckenberg Museum</collectorName>
f̧r
<collectorName id="26A0B284DC452C76FD92F96C6829F954" box="[636,797,1670,1699]" pageId="6" pageNumber="480">Naturkunde G</collectorName>
̂rlitz
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F84D9DC452C76FF29F94068DDF8A3" pageId="6" pageNumber="480" type="materials_examined">
<paragraph id="8BEAD752DC452C76FF29F9406904F8FA" blockId="6.[151,1437,1562,2021]" pageId="6" pageNumber="480">
<emphasis id="B9210B40DC452C76FF29F9406AB1F933" bold="true" box="[199,389,1706,1732]" pageId="6" pageNumber="480">Other material.</emphasis>
<materialsCitation id="3B3DDD0FDC452C76FE60F9416904F8FA" collectingDate="2011-06-26" collectorName="Mosses &amp; C. Schneider &amp; Fr." country="France" county="Soulan" latitude="42.8268" location="Two" longLatPrecision="6" longitude="0.3114" municipality="Hautes-Pyrenees" pageId="6" pageNumber="480" specimenCount="1">
<location id="8E8A8189DC452C76FE60F9416A8BF932" LSID="urn:lsid:plazi:treatment:03FC6644DC452C7AFF79FA446A54FA47:8E8A8189DC452C76FE60F9416A8BF932" box="[398,447,1707,1733]" country="France" county="Soulan" latitude="42.8268" longLatPrecision="6" longitude="0.3114" municipality="Hautes-Pyrenees" name="Two" pageId="6" pageNumber="480">Two</location>
females on slides (SMNG-APT-AA00027, 28),
<collectingCountry id="F34297C2DC452C76FC1EF9416F0BF932" box="[1008,1087,1707,1733]" name="France" pageId="6" pageNumber="480">France</collectingCountry>
,
<collectingMunicipality id="6B8E4D28DC452C76FBA0F9416E39F932" box="[1102,1293,1707,1733]" pageId="6" pageNumber="480">Hautes-Pyrénées</collectingMunicipality>
, Saint-Lary-
<collectingCounty id="628BAFDEDC452C76FF79F9246BDCF91F" box="[151,232,1742,1768]" pageId="6" pageNumber="480">Soulan</collectingCounty>
,
<date id="FFEBF192DC452C76FF18F9246AB9F91E" box="[246,397,1742,1769]" pageId="6" pageNumber="480" value="2011-06-26">
<collectingDate id="EFAF087ADC452C76FF18F9246AB9F91E" box="[246,397,1742,1769]" pageId="6" pageNumber="480" value="2011-06-26">26 Jun. 2011</collectingDate>
</date>
,
<geoCoordinate id="EE61B195DC452C76FE75F924692CF91F" box="[411,536,1742,1768]" degrees="42.8268" direction="north" orientation="latitude" pageId="6" pageNumber="480" precision="5" value="42.8268">42.8268°N</geoCoordinate>
,
<geoCoordinate id="EE61B195DC452C76FDC8F92569A6F91E" box="[550,658,1743,1769]" degrees="0.3114" direction="east" orientation="longitude" pageId="6" pageNumber="480" precision="5" value="0.3114">0.3114°E</geoCoordinate>
,
<collectorName id="26A0B284DC452C76FD71F92569C7F91E" box="[671,755,1743,1769]" pageId="6" pageNumber="480">Mosses</collectorName>
and litter on the side of a path, leg.
<collectorName id="26A0B284DC452C76FB4CF9256E0CF91F" box="[1186,1336,1742,1769]" pageId="6" pageNumber="480">C. Schneider</collectorName>
, original collection codes:
<collectorName id="26A0B284DC452C76FEB5F9196A4EF8FA" box="[347,378,1779,1805]" pageId="6" pageNumber="480">Fr.</collectorName>
031py,
<collectorName id="26A0B284DC452C76FE22F9196ADEF8FA" box="[460,490,1779,1805]" pageId="6" pageNumber="480">Fr.</collectorName>
033py
</materialsCitation>
</paragraph>
<paragraph id="8BEAD752DC452C76FF29F8FD68DDF8A3" blockId="6.[151,1437,1562,2021]" pageId="6" pageNumber="480">
<emphasis id="B9210B40DC452C76FF29F8FD6ABEF8C6" bold="true" box="[199,394,1815,1841]" pageId="6" pageNumber="480">Molecular data.</emphasis>
One individual from Fr.021 (same data as
<typeStatus id="54EE69F0DC452C76FC7BF8FC68CDF8C7" box="[917,1017,1814,1840]" pageId="6" pageNumber="480">holotype</typeStatus>
) and one individual from Fr.033py (Genbank accession number in
<tableCitation id="C6D7E2E9DC452C76FE16F8D0697DF8A2" box="[504,585,1850,1877]" captionStart="TABLE 1" captionStartId="3.[151,239,1164,1188]" captionText="TABLE 1. Genbank accession numbers." pageId="6" pageNumber="480">Table 1</tableCitation>
). Vouchers have not been recovered.
</paragraph>
</subSubSection>
<subSubSection id="C34F84D9DC452C76FF29F8B46E7EF813" pageId="6" pageNumber="480" type="diagnosis">
<paragraph id="8BEAD752DC452C76FF29F8B46E7EF813" blockId="6.[151,1437,1562,2021]" pageId="6" pageNumber="480">
<emphasis id="B9210B40DC452C76FF29F8B46A76F88F" bold="true" box="[199,322,1886,1912]" pageId="6" pageNumber="480">Diagnosis.</emphasis>
Colour whitish (in 96 % ethanol), connection of head channels with
<emphasis id="B9210B40DC452C76FBD1F8B46FEBF88F" box="[1087,1247,1886,1912]" italics="true" pageId="6" pageNumber="480">linea ventralis</emphasis>
crossed, without channels on trunk, basomedian fields of labium with 3 + 3 chaetae, basolateral fields of labium with 2 + 2 chaetae, abdomen without
<emphasis id="B9210B40DC452C76FE86F84D6AB0F836" box="[360,388,1959,1985]" italics="true" pageId="6" pageNumber="480">β2</emphasis>
; but
<emphasis id="B9210B40DC452C76FE2DF84D6AEBF836" box="[451,479,1959,1985]" italics="true" pageId="6" pageNumber="480">β4</emphasis>
and
<emphasis id="B9210B40DC452C76FDF2F84C6901F837" box="[540,565,1958,1984]" italics="true" pageId="6" pageNumber="480">ζ4</emphasis>
present, abdominal s-chaetae
<emphasis id="B9210B40DC452C76FC7AF84D6899F836" box="[916,941,1959,1985]" italics="true" pageId="6" pageNumber="480">s3</emphasis>
present, only slightly larger than
<emphasis id="B9210B40DC452C76FAD6F84D6E65F836" box="[1336,1361,1959,1985]" italics="true" pageId="6" pageNumber="480">s2</emphasis>
, 3 + 3 chaetae on Abd. IV sternite, retinaculum with 4 + 4 teeth, posterior lamellae of the mucro thin and serrated.
</paragraph>
</subSubSection>
<caption id="DF2A87DADC442C77FF79FA776F05FA2E" ID-DOI="http://doi.org/10.5281/zenodo.8181862" ID-Zenodo-Dep="8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481" startId="7.[151,250,1437,1461]" targetBox="[156,1432,186,1408]" targetPageId="7" targetType="figure">
<paragraph id="8BEAD752DC442C77FF79FA776F05FA2E" blockId="7.[151,1436,1437,1497]" pageId="7" pageNumber="481">
<emphasis id="B9210B40DC442C77FF79FA776A27FA43" bold="true" box="[151,275,1437,1461]" pageId="7" pageNumber="481">FIGURE 5.</emphasis>
<taxonomicName id="4C55ACD1DC442C77FEF4FA776933FA42" authority="Schneider &amp; Panina, 2023" authorityName="Schneider &amp; Panina" authorityYear="2023" box="[282,519,1437,1461]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="7" pageNumber="481" phylum="Arthropoda" rank="species" species="boerneri" status="sp. nov.">
<emphasis id="B9210B40DC442C77FEF4FA776933FA42" box="[282,519,1437,1461]" italics="true" pageId="7" pageNumber="481">Megalothorax boerneri</emphasis>
</taxonomicName>
<emphasis id="B9210B40DC442C77FDE0FA776954FA42" bold="true" box="[526,608,1437,1461]" pageId="7" pageNumber="481">
<taxonomicNameLabel id="A212B63BDC442C77FDE0FA776954FA42" box="[526,608,1437,1461]" pageId="7" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna.
</paragraph>
</caption>
<subSubSection id="C34F84D9DC442C7AFF29F9C16A54FA47" lastPageId="10" lastPageNumber="484" pageId="7" pageNumber="481" type="description">
<paragraph id="8BEAD752DC442C77FF29F9C16948F99E" blockId="7.[151,1437,1579,2038]" pageId="7" pageNumber="481">
<emphasis id="B9210B40DC442C77FF29F9C16A6DF9B2" bold="true" box="[199,345,1579,1605]" pageId="7" pageNumber="481">Description.</emphasis>
General aspect. Habitus and segmentation typical of the genus. Length from labrum to anus: up to 400 µm. Colour whitish (in 96 % ethanol).
</paragraph>
<paragraph id="8BEAD752DC442C77FF29F99E6AD0F90E" blockId="7.[151,1437,1579,2038]" pageId="7" pageNumber="481">
Integument. Secondary granulation present dorsally on the head (
<figureCitation id="136ECBD7DC442C77FC47F9996F37F979" box="[937,1027,1651,1678]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481">Fig. 5A</figureCitation>
), the thoracic region, the abdominal region and Abd. VI sternite (
<figureCitation id="136ECBD7DC442C77FE34F97D6901F946" box="[474,565,1687,1713]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="7" pageNumber="481">Fig. 7A</figureCitation>
). Integumentary channels limited to the lateral and posterior part of the head, cycling around the postero-lateral chaetae (
<figureCitation id="136ECBD7DC442C77FD6EF95169EFF921" box="[640,731,1723,1750]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481">Fig. 5A</figureCitation>
); no anterior channel, connection with
<emphasis id="B9210B40DC442C77FB79F9516E0DF922" box="[1175,1337,1723,1749]" italics="true" pageId="7" pageNumber="481">linea ventralis</emphasis>
crossed. Channels absent on the trunk.
</paragraph>
<paragraph id="8BEAD752DC442C77FF29F8E96BC6F87E" blockId="7.[151,1437,1579,2038]" pageId="7" pageNumber="481">
Sensory fields and wax rods. Ordinary distribution of sensory fields and wax rods secretory crypts: 2 + 2
<emphasis id="B9210B40DC442C77FA9DF8EE6EA8F8EA" box="[1395,1436,1796,1821]" italics="true" pageId="7" pageNumber="481">wrc</emphasis>
on head, 12 + 12
<emphasis id="B9210B40DC442C77FEB3F8C26AB2F8B6" box="[349,390,1832,1857]" italics="true" pageId="7" pageNumber="481">wrc</emphasis>
on body; including the ones associated with the 6 + 6 sensory fields (
<figureCitation id="136ECBD7DC442C77FB4EF8CD6FD1F8B6" box="[1184,1253,1831,1857]" captionStart="FIGURE 6" captionStartId="8.[151,250,1018,1042]" captionTargetBox="[156,1431,186,989]" captionTargetId="figure-17@8.[151,1436,181,994]" captionTargetPageId="8" captionText="FIGURE 6. Megalothorax boerneri sp. nov., trunk tergites chaetotaxy (Th. I–Abd. V)." figureDoi="http://doi.org/10.5281/zenodo.8181864" httpUri="https://zenodo.org/record/8181864/files/figure.png" pageId="7" pageNumber="481">Fig. 6</figureCitation>
). Sensory fields include the swollen inner chaetae, all globular:
<emphasis id="B9210B40DC442C77FD46F8A169D1F892" box="[680,741,1867,1893]" italics="true" pageId="7" pageNumber="481">sf1–6</emphasis>
: 0, 1, 3, 2, 1, 1.
<emphasis id="B9210B40DC442C77FC77F8A668E4F891" box="[921,976,1868,1894]" italics="true" pageId="7" pageNumber="481">wrc5</emphasis>
and
<emphasis id="B9210B40DC442C77FBE8F8A16F20F892" box="[1030,1044,1867,1893]" italics="true" pageId="7" pageNumber="481">6</emphasis>
separated from
<emphasis id="B9210B40DC442C77FB27F8A16FDDF892" box="[1225,1257,1867,1893]" italics="true" pageId="7" pageNumber="481">sf5</emphasis>
by ~ 4 granules (
<figureCitation id="136ECBD7DC442C77FF71F8856BD0F87E" box="[159,228,1903,1929]" captionStart="FIGURE 6" captionStartId="8.[151,250,1018,1042]" captionTargetBox="[156,1431,186,989]" captionTargetId="figure-17@8.[151,1436,181,994]" captionTargetPageId="8" captionText="FIGURE 6. Megalothorax boerneri sp. nov., trunk tergites chaetotaxy (Th. I–Abd. V)." figureDoi="http://doi.org/10.5281/zenodo.8181864" httpUri="https://zenodo.org/record/8181864/files/figure.png" pageId="7" pageNumber="481">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BEAD752DC442C77FF29F8796E10F802" blockId="7.[151,1437,1579,2038]" pageId="7" pageNumber="481">
Head chaetotaxy. Number of chaetae: 12 +
<quantity id="4CAD7AB7DC442C77FD48F87969E9F85A" box="[678,733,1939,1965]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="7" pageNumber="481" unit="in" value="12.0">12 in</quantity>
the postero-dorsal region, 10 + 10 and 2 unpaired in the antero-dorsal region, 1 +
<quantity id="4CAD7AB7DC442C77FE8FF85D6ABFF826" box="[353,395,1975,2001]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="481" unit="in" value="1.0">1 in</quantity>
the antero-lateral region (
<figureCitation id="136ECBD7DC442C77FD46F85D6834F825" box="[680,768,1975,2002]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481">Fig. 5A</figureCitation>
,
<figureCitation id="136ECBD7DC442C77FCE4F85D681AF826" box="[778,814,1975,2001]" captionStart="FIGURE 8" captionStartId="10.[151,250,1251,1275]" captionTargetBox="[156,1431,186,1223]" captionTargetId="figure-17@10.[151,1436,181,1228]" captionTargetPageId="10" captionText="FIGURE 8. Chaetotaxy diagrams: M. incertus sensu nov. (A) head, (B) trunk, M. boerneri sp. nov. (A) head, (B) trunk." figureDoi="http://doi.org/10.5281/zenodo.8181868" httpUri="https://zenodo.org/record/8181868/files/figure.png" pageId="7" pageNumber="481">8C</figureCitation>
). With 7 + 7 postero-dorsal chaetae thickened (
<figureCitation id="136ECBD7DC442C77FAD9F85D6EBAF825" box="[1335,1422,1975,2002]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481">Fig. 5A</figureCitation>
), anterior chaetae short and ordinary (
<figureCitation id="136ECBD7DC442C77FDC3F83169B3F801" box="[557,647,2011,2038]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="7" pageNumber="481">Fig. 5A</figureCitation>
). Ventrally with 3 + 3 post-labial chaetae sub-equally sized.
</paragraph>
<caption id="DF2A87DADC4B2C78FF79FC106F3EFBE5" ID-DOI="http://doi.org/10.5281/zenodo.8181864" ID-Zenodo-Dep="8181864" box="[151,1034,1018,1042]" httpUri="https://zenodo.org/record/8181864/files/figure.png" pageId="8" pageNumber="482" startId="8.[151,250,1018,1042]" targetBox="[156,1431,186,989]" targetPageId="8" targetType="figure">
<paragraph id="8BEAD752DC4B2C78FF79FC106F3EFBE5" blockId="8.[151,1034,1018,1042]" box="[151,1034,1018,1042]" pageId="8" pageNumber="482">
<emphasis id="B9210B40DC4B2C78FF79FC106A27FBE5" bold="true" box="[151,275,1018,1042]" pageId="8" pageNumber="482">FIGURE 6.</emphasis>
<taxonomicName id="4C55ACD1DC4B2C78FEF7FC106932FBE5" authority="Schneider &amp; Panina, 2023" authorityName="Schneider &amp; Panina" authorityYear="2023" box="[281,518,1018,1042]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="8" pageNumber="482" phylum="Arthropoda" rank="species" species="boerneri" status="sp. nov.">
<emphasis id="B9210B40DC4B2C78FEF7FC106932FBE5" box="[281,518,1018,1042]" italics="true" pageId="8" pageNumber="482">Megalothorax boerneri</emphasis>
</taxonomicName>
<emphasis id="B9210B40DC4B2C78FDE3FC106969FBE5" bold="true" box="[525,605,1018,1042]" pageId="8" pageNumber="482">
<taxonomicNameLabel id="A212B63BDC4B2C78FDE3FC106969FBE5" box="[525,605,1018,1042]" pageId="8" pageNumber="482" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, trunk tergites chaetotaxy (Th. I–Abd. V).
</paragraph>
</caption>
<paragraph id="8BEAD752DC4B2C78FF29FB8E696AFB54" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Labium. Basomedian fields of labium with 3 + 3 chaetae, basolateral fields of labium with 2 + 2 chaetae (
<figureCitation id="136ECBD7DC4B2C78FA81FB8E6B8FFB54" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5B</figureCitation>
). Labial palps ordinary, as is
<figureCitation id="136ECBD7DC4B2C78FDEDFB62696EFB54" box="[515,602,1160,1187]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5B</figureCitation>
.
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29FB466F95FB30" blockId="8.[151,1437,1124,1943]" box="[199,1185,1196,1223]" pageId="8" pageNumber="482">
Labrum. Chaetae
<emphasis id="B9210B40DC4B2C78FE61FB466A9FFB31" box="[399,427,1196,1222]" italics="true" pageId="8" pageNumber="482">a1</emphasis>
,
<emphasis id="B9210B40DC4B2C78FE57FB466AE1FB31" box="[441,469,1196,1222]" italics="true" pageId="8" pageNumber="482">a2</emphasis>
with external teeth,
<emphasis id="B9210B40DC4B2C78FD54FB4669E2FB31" box="[698,726,1196,1222]" italics="true" pageId="8" pageNumber="482">a1</emphasis>
with apical outward curvature (
<figureCitation id="136ECBD7DC4B2C78FBD2FB466FA1FB30" box="[1084,1173,1196,1223]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29FB3A6F0FFAF8" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Other mouthparts. Oral fold with 2 + 2 chaetae (
<figureCitation id="136ECBD7DC4B2C78FD15FB3A6863FB1C" box="[763,855,1232,1259]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5D</figureCitation>
). Maxilla outer lobe with two chaetae (apical and basal), sublobal plate with a hair (
<figureCitation id="136ECBD7DC4B2C78FDFCFB1E6958FAF8" box="[530,620,1268,1295]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5D</figureCitation>
). Maxilla and mandibula as in
<figureCitation id="136ECBD7DC4B2C78FC28FB1E6F0CFAF8" box="[966,1080,1268,1295]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5E, F</figureCitation>
.
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29FAF36A20FA35" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Antenna. As in
<figureCitation id="136ECBD7DC4B2C78FE9DFAF26AFEFAC4" box="[371,458,1304,1331]" captionStart="FIGURE 5" captionStartId="7.[151,250,1437,1461]" captionTargetBox="[156,1432,186,1408]" captionTargetId="figure-19@7.[151,1436,181,1413]" captionTargetPageId="7" captionText="FIGURE 5. Megalothorax boerneri sp. nov., (A) Head; (B) labium, bmf=basomedian field, blf=basolateral field; (C) labrum; (D) maxillary outer lobe (mol) and oral fold (of); (E) maxilla; (F) mandibula; (G) Antenna." figureDoi="http://doi.org/10.5281/zenodo.8181862" httpUri="https://zenodo.org/record/8181862/files/figure.png" pageId="8" pageNumber="482">Fig. 5G</figureCitation>
. Ant. I with one chaeta. Ant. II with four chaetae, the anterior one stronger than the three others. Ant. III with 9 chaetae and the four S-chaetae
<emphasis id="B9210B40DC4B2C78FD19FAD66809FAA1" box="[759,829,1340,1367]" italics="true" pageId="8" pageNumber="482">S1–S4</emphasis>
from the sensory organ;
<emphasis id="B9210B40DC4B2C78FBB4FAD66F42FAA1" box="[1114,1142,1340,1366]" italics="true" pageId="8" pageNumber="482">S2</emphasis>
and
<emphasis id="B9210B40DC4B2C78FB40FAD66FFEFAA1" box="[1198,1226,1340,1366]" italics="true" pageId="8" pageNumber="482">S3</emphasis>
clearly protruding from the cupule; all
<emphasis id="B9210B40DC4B2C78FE97FA8A6AF4FA8D" box="[377,448,1376,1403]" italics="true" pageId="8" pageNumber="482">S1–S4</emphasis>
rather indistinct in our preparations. Ant. IV with six ordinary chaetae (including
<emphasis id="B9210B40DC4B2C78FABDFA8B6E50FA8D" box="[1363,1380,1377,1402]" italics="true" pageId="8" pageNumber="482">X</emphasis>
) and 10 + 2 S-chaetae (
<emphasis id="B9210B40DC4B2C78FE8BFA6E6A9FFA68" box="[357,427,1412,1439]" italics="true" pageId="8" pageNumber="482">Sb1–5</emphasis>
,
<emphasis id="B9210B40DC4B2C78FE54FA6E6934FA68" box="[442,512,1412,1439]" italics="true" pageId="8" pageNumber="482">Sa1–5</emphasis>
,
<emphasis id="B9210B40DC4B2C78FDE1FA6F691DFA68" box="[527,553,1413,1439]" italics="true" pageId="8" pageNumber="482">Sx</emphasis>
and
<emphasis id="B9210B40DC4B2C78FD8FFA6F694FFA68" box="[609,635,1413,1439]" italics="true" pageId="8" pageNumber="482">Sy</emphasis>
),
<emphasis id="B9210B40DC4B2C78FD7AFA6E698AFA69" box="[660,702,1412,1438]" italics="true" pageId="8" pageNumber="482">Sb3</emphasis>
implanted basally, slightly above
<emphasis id="B9210B40DC4B2C78FBAFFA6E6F69FA69" box="[1089,1117,1412,1438]" italics="true" pageId="8" pageNumber="482">S2</emphasis>
and
<emphasis id="B9210B40DC4B2C78FB7BFA6E6F85FA69" box="[1173,1201,1412,1438]" italics="true" pageId="8" pageNumber="482">S3</emphasis>
of Ant. III. Organite (
<emphasis id="B9210B40DC4B2C78FF4FFA436BF4FA34" box="[161,192,1449,1475]" italics="true" pageId="8" pageNumber="482">Or</emphasis>
) small.
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29FA266ACFF9D9" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Thoracic tergites. As in
<figureCitation id="136ECBD7DC4B2C78FE37FA26692AFA11" box="[473,542,1484,1510]" captionStart="FIGURE 6" captionStartId="8.[151,250,1018,1042]" captionTargetBox="[156,1431,186,989]" captionTargetId="figure-17@8.[151,1436,181,994]" captionTargetPageId="8" captionText="FIGURE 6. Megalothorax boerneri sp. nov., trunk tergites chaetotaxy (Th. I–Abd. V)." figureDoi="http://doi.org/10.5281/zenodo.8181864" httpUri="https://zenodo.org/record/8181864/files/figure.png" pageId="8" pageNumber="482">Fig. 6</figureCitation>
. Pseudopores and τ-chaetae indistinct, not studied. Th. II with 12 + 12 chaetae and 1 + 1 s-chaetae
<emphasis id="B9210B40DC4B2C78FE90FA1A6AAAF9FD" box="[382,414,1520,1546]" italics="true" pageId="8" pageNumber="482">s1.</emphasis>
Th. III with 10 + 10 chaetae,
<emphasis id="B9210B40DC4B2C78FD15FA1B6823F9FC" box="[763,791,1521,1547]" italics="true" pageId="8" pageNumber="482">a5</emphasis>
slightly stronger than
<emphasis id="B9210B40DC4B2C78FBF3FA1A6F0DF9FD" box="[1053,1081,1520,1546]" italics="true" pageId="8" pageNumber="482">a6</emphasis>
,
<emphasis id="B9210B40DC4B2C78FBA4FA1A6F52F9FD" box="[1098,1126,1520,1546]" italics="true" pageId="8" pageNumber="482">p4</emphasis>
close to
<emphasis id="B9210B40DC4B2C78FB3CFA1A6E3DF9FD" box="[1234,1289,1520,1546]" italics="true" pageId="8" pageNumber="482">wrc2</emphasis>
. Diagram of chaetotaxy provided in
<figureCitation id="136ECBD7DC4B2C78FE71F9FE6ACCF9D9" box="[415,504,1556,1582]" captionStart="FIGURE 8" captionStartId="10.[151,250,1251,1275]" captionTargetBox="[156,1431,186,1223]" captionTargetId="figure-17@10.[151,1436,181,1228]" captionTargetPageId="10" captionText="FIGURE 8. Chaetotaxy diagrams: M. incertus sensu nov. (A) head, (B) trunk, M. boerneri sp. nov. (A) head, (B) trunk." figureDoi="http://doi.org/10.5281/zenodo.8181868" httpUri="https://zenodo.org/record/8181868/files/figure.png" pageId="8" pageNumber="482">Fig. 8D</figureCitation>
.
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29F9D2692EF981" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Abd. I–V tergites. With 19 + 19 ordinary chaetae, 2 + 2 globular s-chaetae (
<emphasis id="B9210B40DC4B2C78FBF7F9D26F06F9A5" box="[1049,1074,1592,1618]" italics="true" pageId="8" pageNumber="482">s2</emphasis>
,
<emphasis id="B9210B40DC4B2C78FBAEF9D26F6DF9A5" box="[1088,1113,1592,1618]" italics="true" pageId="8" pageNumber="482">s3</emphasis>
) with
<emphasis id="B9210B40DC4B2C78FB4CF9D26F8FF9A5" box="[1186,1211,1592,1618]" italics="true" pageId="8" pageNumber="482">s3</emphasis>
slightly bigger than
<emphasis id="B9210B40DC4B2C78FF79F9B66B84F981" box="[151,176,1628,1654]" italics="true" pageId="8" pageNumber="482">s2</emphasis>
; chaetae
<emphasis id="B9210B40DC4B2C78FEF4F9B66A02F981" box="[282,310,1628,1654]" italics="true" pageId="8" pageNumber="482">β2</emphasis>
absent (
<figureCitation id="136ECBD7DC4B2C78FE7DF9B66AE8F981" box="[403,476,1628,1654]" captionStart="FIGURE 6" captionStartId="8.[151,250,1018,1042]" captionTargetBox="[156,1431,186,989]" captionTargetId="figure-17@8.[151,1436,181,994]" captionTargetPageId="8" captionText="FIGURE 6. Megalothorax boerneri sp. nov., trunk tergites chaetotaxy (Th. I–Abd. V)." figureDoi="http://doi.org/10.5281/zenodo.8181864" httpUri="https://zenodo.org/record/8181864/files/figure.png" pageId="8" pageNumber="482">Figs 6</figureCitation>
,
<figureCitation id="136ECBD7DC4B2C78FE06F9B6693AF981" box="[488,526,1628,1654]" captionStart="FIGURE 8" captionStartId="10.[151,250,1251,1275]" captionTargetBox="[156,1431,186,1223]" captionTargetId="figure-17@10.[151,1436,181,1228]" captionTargetPageId="10" captionText="FIGURE 8. Chaetotaxy diagrams: M. incertus sensu nov. (A) head, (B) trunk, M. boerneri sp. nov. (A) head, (B) trunk." figureDoi="http://doi.org/10.5281/zenodo.8181868" httpUri="https://zenodo.org/record/8181868/files/figure.png" pageId="8" pageNumber="482">8D</figureCitation>
).
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29F96A68D4F949" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Abd. VI. Tergite with 4 + 4 and 1 unpaired chaetae (
<figureCitation id="136ECBD7DC4B2C78FCDEF96A68B9F96D" box="[816,909,1664,1690]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7A</figureCitation>
). Each anal valve with one chaeta (
<figureCitation id="136ECBD7DC4B2C78FADCF96A6EBBF96D" box="[1330,1423,1664,1690]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7A</figureCitation>
). Sternite with 7 + 7 chaetae, the two axial pairs strongly curvated (
<figureCitation id="136ECBD7DC4B2C78FC96F94E68E6F949" box="[888,978,1700,1726]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29F922688EF915" blockId="8.[151,1437,1124,1943]" box="[199,954,1736,1763]" pageId="8" pageNumber="482">
Genital plate. Female with 3 + 3 chaetae (
<figureCitation id="136ECBD7DC4B2C78FD75F92269C1F915" box="[667,757,1736,1762]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7A</figureCitation>
). Male unknown.
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29F9066BFEF8DD" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Abd. IV sternite. With 2 + 2 usual neosminthuroid chaetae, 3 + 3 ordinary chaetae and 1 + 1 small lobes (
<figureCitation id="136ECBD7DC4B2C78FA81F9066B89F8DD" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="8BEAD752DC4B2C78FF29F8DE699EF861" blockId="8.[151,1437,1124,1943]" pageId="8" pageNumber="482">
Abdominal appendages. Manubrium with 1 + 1 posterior chaetae (
<figureCitation id="136ECBD7DC4B2C78FC49F8DE68CAF8B9" box="[935,1022,1844,1870]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7B</figureCitation>
). Dens ordinary, as in
<figureCitation id="136ECBD7DC4B2C78FB1CF8DE6E73F8B8" box="[1266,1351,1844,1871]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7B</figureCitation>
. Mucro with thin lamellae, with ~10–12 teeth on each posterior lamellae (
<figureCitation id="136ECBD7DC4B2C78FC6CF8B268E8F885" box="[898,988,1880,1906]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7B</figureCitation>
). Ventral tube with 2 + 2 apical chaetae (
<figureCitation id="136ECBD7DC4B2C78FF71F8966BCCF861" box="[159,248,1916,1942]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="8" pageNumber="482">Fig. 7C</figureCitation>
), retinaculum with 4 + 4 distinct teeth.
</paragraph>
<caption id="DF2A87DADC4A2C79FF79F9256831F8FD" ID-DOI="http://doi.org/10.5281/zenodo.8181866" ID-Zenodo-Dep="8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="9" pageNumber="483" startId="9.[151,250,1743,1767]" targetBox="[156,1432,186,1714]" targetPageId="9" targetType="figure">
<paragraph id="8BEAD752DC4A2C79FF79F9256831F8FD" blockId="9.[151,1436,1743,1803]" pageId="9" pageNumber="483">
<emphasis id="B9210B40DC4A2C79FF79F9256A26F910" bold="true" box="[151,274,1743,1767]" pageId="9" pageNumber="483">FIGURE 7.</emphasis>
<taxonomicName id="4C55ACD1DC4A2C79FEF6F9256931F910" authority="Schneider &amp; Panina, 2023" authorityName="Schneider &amp; Panina" authorityYear="2023" box="[280,517,1743,1767]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="9" pageNumber="483" phylum="Arthropoda" rank="species" species="boerneri" status="sp. nov.">
<emphasis id="B9210B40DC4A2C79FEF6F9256931F910" box="[280,517,1743,1767]" italics="true" pageId="9" pageNumber="483">Megalothorax boerneri</emphasis>
</taxonomicName>
<emphasis id="B9210B40DC4A2C79FDE4F925696EF910" bold="true" box="[522,602,1743,1767]" pageId="9" pageNumber="483">
<taxonomicNameLabel id="A212B63BDC4A2C79FDE4F925696EF910" box="[522,602,1743,1767]" pageId="9" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III.
</paragraph>
</caption>
<paragraph id="8BEAD752DC4A2C79FF29F8B46F6DF814" blockId="9.[151,1436,1885,2020]" pageId="9" pageNumber="483">
Legs. Chaetal composition on each legs subcoxa 1, 2, coxa, trochanter, femur and tibiotarsus: leg I—1, 0, 1, 3, 7, 12 chaeta(e) (
<figureCitation id="136ECBD7DC4A2C79FE82F86B6AF3F86C" box="[364,455,1921,1947]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="9" pageNumber="483">Fig. 7D</figureCitation>
); leg II—1, 1, 1, 3, 8 and 12 chaeta(e) (
<figureCitation id="136ECBD7DC4A2C79FC67F86B68D5F86C" box="[905,993,1921,1947]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="9" pageNumber="483">Fig. 7E</figureCitation>
); leg III—2, 1, 1, 4, 8 and 11 chaeta(e) (
<figureCitation id="136ECBD7DC4A2C79FF71F84F6BC6F848" box="[159,242,1957,1983]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="9" pageNumber="483">Fig. 7F</figureCitation>
). Claw as in
<figureCitation id="136ECBD7DC4A2C79FE62F84F6AC9F848" box="[396,509,1957,1983]" captionStart="FIGURE 7" captionStartId="9.[151,250,1743,1767]" captionTargetBox="[156,1432,186,1714]" captionTargetId="figure-19@9.[151,1436,181,1719]" captionTargetPageId="9" captionText="FIGURE 7. Megalothorax boerneri sp. nov., (A) Abd. VI, genital plate and Abd. IV sternite, (B) furca, (C) ventral tube, (D) leg I, (E) leg II, (F) leg III, (G) claw I, (H, I) claw II, (J) claw III." figureDoi="http://doi.org/10.5281/zenodo.8181866" httpUri="https://zenodo.org/record/8181866/files/figure.png" pageId="9" pageNumber="483">Fig. 7G–J</figureCitation>
, ordinary, subequal in length. Basal and posterior auxiliary lamellae of unguis well developed, anterior crest indistinct. Basal lobe of unguiculus not or feebly protruding.
</paragraph>
<caption id="DF2A87DADC492C7AFF79FB096E6EFB0B" ID-DOI="http://doi.org/10.5281/zenodo.8181868" ID-Zenodo-Dep="8181868" box="[151,1370,1251,1276]" httpUri="https://zenodo.org/record/8181868/files/figure.png" pageId="10" pageNumber="484" startId="10.[151,250,1251,1275]" targetBox="[156,1431,186,1223]" targetPageId="10" targetType="figure">
<paragraph id="8BEAD752DC492C7AFF79FB096E6EFB0B" blockId="10.[151,1370,1251,1276]" box="[151,1370,1251,1276]" pageId="10" pageNumber="484">
<emphasis id="B9210B40DC492C7AFF79FB096A27FB0B" bold="true" box="[151,275,1251,1276]" pageId="10" pageNumber="484">FIGURE 8.</emphasis>
Chaetotaxy diagrams:
<taxonomicName id="4C55ACD1DC492C7AFE13FB0E6945FB0C" authorityName="Borner" authorityYear="1903" box="[509,625,1252,1275]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="10" pageNumber="484" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="B9210B40DC492C7AFE13FB0E6945FB0C" box="[509,625,1252,1275]" italics="true" pageId="10" pageNumber="484">M. incertus</emphasis>
</taxonomicName>
<emphasis id="B9210B40DC492C7AFD99FB0E69D0FB0B" bold="true" box="[631,740,1252,1276]" pageId="10" pageNumber="484">sensu nov.</emphasis>
(A) head, (B) trunk,
<taxonomicName id="4C55ACD1DC492C7AFC55FB0E6F00FB0B" authority="Schneider &amp; Panina, 2023" authorityName="Schneider &amp; Panina" authorityYear="2023" box="[955,1076,1252,1276]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="10" pageNumber="484" phylum="Arthropoda" rank="species" species="boerneri" status="sp. nov.">
<emphasis id="B9210B40DC492C7AFC55FB0E6F00FB0B" box="[955,1076,1252,1276]" italics="true" pageId="10" pageNumber="484">M. boerneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212B63BDC492C7AFBD5FB0E6FBEFB0C" box="[1083,1162,1252,1275]" pageId="10" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
(A) head, (B) trunk.
</paragraph>
</caption>
<paragraph id="8BEAD752DC492C7AFF29FAA46A54FA47" blockId="10.[151,1436,1358,1456]" pageId="10" pageNumber="484">
<emphasis id="B9210B40DC492C7AFF29FAA46A09FA9F" bold="true" box="[199,317,1358,1384]" pageId="10" pageNumber="484">Affinities.</emphasis>
The most similar species are
<taxonomicName id="4C55ACD1DC492C7AFD69FAA5680CFA9F" authorityName="Schneider &amp; D'Haese" authorityYear="2013" box="[647,824,1358,1384]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="10" pageNumber="484" phylum="Arthropoda" rank="species" species="perspicillum">
<emphasis id="B9210B40DC492C7AFD69FAA5680CFA9F" box="[647,824,1358,1384]" italics="true" pageId="10" pageNumber="484">M. perspicillum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C55ACD1DC492C7AFCA8FAA568F1FA9F" authorityName="Borner" authorityYear="1903" box="[838,965,1359,1384]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="10" pageNumber="484" phylum="Arthropoda" rank="species" species="incertus">
<emphasis id="B9210B40DC492C7AFCA8FAA568F1FA9F" box="[838,965,1359,1384]" italics="true" pageId="10" pageNumber="484">M. incertus</emphasis>
</taxonomicName>
<emphasis id="B9210B40DC492C7AFC22FAA46F71FA9F" bold="true" box="[972,1093,1358,1384]" pageId="10" pageNumber="484">sensu nov.</emphasis>
and
<taxonomicName id="4C55ACD1DC492C7AFB94FAA56FD5FA9F" authorityName="Denis" authorityYear="1948" box="[1146,1249,1358,1384]" class="Collembola" family="Neelidae" genus="Megalothorax" kingdom="Animalia" order="Neelipleona" pageId="10" pageNumber="484" phylum="Arthropoda" rank="species" species="laevis">
<emphasis id="B9210B40DC492C7AFB94FAA56FD5FA9F" box="[1146,1249,1358,1384]" italics="true" pageId="10" pageNumber="484">M. laevis</emphasis>
</taxonomicName>
. The differential diagnosis is provided in
<tableCitation id="C6D7E2E9DC492C7AFE5CFA98693EFA7B" box="[434,522,1394,1420]" captionStart="TABLE 2" captionStartId="6.[151,239,548,572]" captionText="TABLE 2. Differential diagnosis of the closest relatives of M. incertus sensu nov." pageId="10" pageNumber="484">Table 2</tableCitation>
. Also see the updated interactive key of the
<taxonomicName id="4C55ACD1DC492C7AFBE2FA986E31FA7B" box="[1036,1285,1394,1420]" pageId="10" pageNumber="484">
<emphasis id="B9210B40DC492C7AFBE2FA986F9FFA7B" box="[1036,1195,1394,1420]" italics="true" pageId="10" pageNumber="484">Megalothorax</emphasis>
species
</taxonomicName>
of the world (
<bibRefCitation id="EFC4AAA3DC492C7AFF71FA7C6A60FA47" author="Schneider, C." box="[159,340,1430,1456]" pageId="10" pageNumber="484" refId="ref7591" refString="Schneider, C. (2022) Open key of the Megalothorax species of the world (Collembola: Neelidae) (v 1.0.0). Zenodo. https: // doi. org / 10.5281 / zenodo. 7341417" type="book" year="2022">Schneider 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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