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<document id="9E0874E5FD696563EDD7C23CABC3E9EC" ID-CLB-Dataset="8111" ID-DOI="10.37520/aemnp.2020.26" ID-GBIF-Dataset="1a61f2ea-5825-49cd-afeb-bb9d9f9be24a" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489332" ID-ZooBank="7CF8BFF6-034F-475E-9A3A-1FA8AF862824" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221451257" checkinUser="felipe" docAuthor="Zettel, Herbert &amp; Laciny, Alice" docDate="2020" docId="03FC501AFFBB6853FE9B4D8EFA06FA06" docLanguage="en" docName="ActaEntMusNatPrag.60.2.419-425.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Rhagovelia madli Zettel &amp; Laciny 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="422" masterDocId="FFC52862FFBA6850FFA24F03FF83FFD5" masterDocTitle="Rhagovelia species from Nosy Boraha east of Madagascar (Hemiptera: Heteroptera: Veliidae)" masterLastPageNumber="425" masterPageNumber="419" pageNumber="420" updateTime="1698926184911" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="838ABD6CFAAABC50487AA67376AF7146">Rhagovelia species from Nosy Boraha east of Madagascar (Hemiptera: Heteroptera: Veliidae)</mods:title>
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<mods:name id="E8F2F8916EB794F83BCD43D917F5C1EF" type="personal">
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<mods:namePart id="42A8758CD509F0DFD1B2CA3B94AAC621">Zettel, Herbert</mods:namePart>
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<mods:namePart id="15B09F1117A5A69EFDF7E2F40FAFB338">Laciny, Alice</mods:namePart>
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<mods:title id="D5D77D232EB2A8D6FE68CCFE64266AAE">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="23FB367D511DB90D9EE2ACAB294FCB38">2020</mods:date>
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<treatment id="03FC501AFFBB6853FE9B4D8EFA06FA06" ID-DOI="http://doi.org/10.5281/zenodo.4550761" ID-GBIF-Taxon="177426401" ID-Zenodo-Dep="4550761" LSID="urn:lsid:plazi:treatment:03FC501AFFBB6853FE9B4D8EFA06FA06" httpUri="http://treatment.plazi.org/id/03FC501AFFBB6853FE9B4D8EFA06FA06" lastPageId="3" lastPageNumber="422" pageId="1" pageNumber="420">
<subSubSection id="C34FB287FFBB6851FE9B4D8EFDD5FD70" box="[313,598,653,677]" pageId="1" pageNumber="420" type="nomenclature">
<paragraph id="8BEAE10CFFBB6851FE9B4D8EFDD5FD70" blockId="1.[313,598,653,677]" box="[313,598,653,677]" pageId="1" pageNumber="420">
<heading id="D0A25660FFBB6851FE9B4D8EFDD5FD70" box="[313,598,653,677]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="420" reason="3">
<emphasis id="B9213D1EFFBB6851FE9B4D8EFDD5FD70" bold="true" box="[313,598,653,677]" pageId="1" pageNumber="420">
<taxonomicName id="4C559A8FFFBB6851FE9B4D8EFE79FD70" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[313,506,653,677]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="420" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFBB6851FE9B4D8EFE79FD70" bold="true" box="[313,506,653,677]" italics="true" pageId="1" pageNumber="420">Rhagovelia madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFBB6851FDA34D8DFDD5FD70" box="[513,598,654,677]" pageId="1" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34FB287FFBB6851FEFE4DAFFDB0FD6A" box="[348,563,684,703]" pageId="1" pageNumber="420" type="description">
<paragraph id="8BEAE10CFFBB6851FEFE4DAFFDB0FD6A" blockId="1.[151,760,684,798]" box="[348,563,684,703]" pageId="1" pageNumber="420">
(
<figureCitation id="136EFD89FFBB6851FEC04DAFFE2DFD6A" box="[354,430,684,703]" captionStart="Figs 14. 12" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. 12 Rhagovelia madli sp. nov.: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3 R. kalalaoensis sp. nov., holotype, apterous male; 4 R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="1" pageNumber="420">Figs 1, 2</figureCitation>
,
<figureCitation id="136EFD89FFBB6851FE154DAFFD8EFD6A" box="[439,525,684,703]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">5, 7, 9, 11</figureCitation>
,
<figureCitation id="136EFD89FFBB6851FDB54DAFFDAEFD6A" box="[535,557,684,703]" captionStart="Figs 1318" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 1318.Characteristics of females: 1316 apex of abdomen, lateral view: 13 Rhagovelia madli sp. nov., 14 R. kalalaoensis sp. nov., 15 R. ambra Polhemus &amp; Andersen, 2015, 16 R. aquacola Polhemus &amp; Andersen, 2015; 1718 tergites 7 and 8, dorsal view: 17 R. kalalaoensis sp. nov., 18 R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="1" pageNumber="420">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34FB287FFBB6851FF354DD5FE02FCCB" pageId="1" pageNumber="420" type="materials_examined">
<paragraph id="8BEAE10CFFBB6851FF354DD5FE02FCCB" blockId="1.[151,760,684,798]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FF354DD5FE9EFD3C" bold="true" box="[151,285,726,745]" pageId="1" pageNumber="420">
<typeStatus id="54EE5FAEFFBB6851FF354DD5FF47FD3C" box="[151,196,726,745]" pageId="1" pageNumber="420">Type</typeStatus>
material.
</emphasis>
HƟΓƟΤΥΡΕ (apterous ♁) and ΡΑRΑΤΥΡΕඌ (1 apterous ♁, 6 apterous
<specimenCount id="9D532A85FFBB6851FF4D4DEEFE8DFCD1" box="[239,270,749,772]" pageId="1" pageNumber="420" type="female">♀♀</specimenCount>
):
<collectingCountry id="F342A19CFFBB6851FEBD4DF2FE06FCD1" box="[287,389,753,772]" name="Madagascar" pageId="1" pageNumber="420">Madagascar</collectingCountry>
,Nosy Boraha:Forêt de Kalalao,
<date id="FFEBC7CCFFBB6851FD314DF2FD77FCD1" box="[659,756,753,772]" pageId="1" pageNumber="420" value="1993-11-18">18.XI.1993</date>
, leg. M. Madl (all
<collectionCode id="ED4479C9FFBB6851FE8C4C08FEFBFCCB" box="[302,376,779,798]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="420" type="Museum">NHMW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34FB287FFBB6853FF354C34FB7CFA66" lastPageId="3" lastPageNumber="422" pageId="1" pageNumber="420" type="description">
<paragraph id="8BEAE10CFFBB6851FF354C34FD01FBBB" blockId="1.[151,761,822,2062]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FF354C34FE62FC9B" bold="true" box="[151,481,822,846]" pageId="1" pageNumber="420">
Description.
<emphasis id="B9213D1EFFBB6851FE8B4C35FE59FC9B" bold="true" box="[297,474,822,846]" italics="true" pageId="1" pageNumber="420">Male (apterous)</emphasis>
.
</emphasis>
<emphasis id="B9213D1EFFBB6851FE4A4C34FD00FC9B" box="[488,643,823,846]" italics="true" pageId="1" pageNumber="420">Measurements</emphasis>
(
<typeStatus id="54EE5FAEFFBB6851FD304C34FD77FC9B" box="[658,756,823,846]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
,
<typeStatus id="54EE5FAEFFBB6851FF354C54FF7BFCBB" box="[151,248,855,878]" pageId="1" pageNumber="420" type="paratype">paratype</typeStatus>
). Body length 2.66,\ 2.69; maximum body width (at metapleura) 1.06, 1.08. Head length 0.36, 0.38, width 0.75, 0.77; minimum eye distance 0.10, 0.12. Pronotum length 0.75, 0.73, width 0.97, 0.97. Lengths of antennomeres (
<typeStatus id="54EE5FAEFFBB6851FF414CD4FEC6FC3B" box="[227,325,983,1006]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
), I 0.53, II 0.36, III 0.42, IV 0.47. Lengths of leg segments (
<typeStatus id="54EE5FAEFFBB6851FEEF4CF4FE32FBDB" box="[333,433,1015,1038]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
): profemur 0.78, protibia 0.88, protarsus 0.01 + 0.02 + 0.20, mesofemur 1.44, mesotibia 1.19, mesotarsus 0.07 + 0.41 + 0.66, metafemur 1.09, metatibia 1.20, metatarsus 0.03 + 0.04 + 0.23.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FF174B74FE3BFA3B" blockId="1.[151,761,822,2062]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FF174B74FE82FB5B" box="[181,257,1143,1166]" italics="true" pageId="1" pageNumber="420">Colour</emphasis>
(
<figureCitation id="136EFD89FFBB6851FEB04B74FED0FB5B" box="[274,339,1143,1166]" captionStart="Figs 14. 12" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. 12 Rhagovelia madli sp. nov.: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3 R. kalalaoensis sp. nov., holotype, apterous male; 4 R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="1" pageNumber="420">Fig. 1</figureCitation>
). Trunk chiefly black. Pronotum anteriorly with transverse yellow stripe, medially interrupted by narrow blackish line. Tergite 6 with small yellowish brown spot, tergite 7 with large yellow spot. Sternites 36 with narrow brownish or yellowish brown lateral margins. Propleuron, mesopleuron, and all acetabula yellow. Venter bicoloured: mesosternum to sternite 6 black, other parts yellow (segment 8 slightly darker). Antenna black, basal third of antennomere 1 yellow. Legs chiefly black; all coxae, pro- and metatrochanter, basal third of profemur, and base of metafemur yellow; mesotrochanter black, with small yellow patch at base.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FF174AF4FED5F8DB" blockId="1.[151,761,822,2062]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FF174AF4FE93F9DB" box="[181,272,1527,1550]" italics="true" pageId="1" pageNumber="420">Pilosity.</emphasis>
Body dorsally with short, grey, appressed pubescence; abdomen additionally with inconspicuous, dispersed, dark grey, subdecumbent, short setae; only head, sides of pronotum, and posterior margin of segment 8 with some long black setae. Protrochanter without conspicuous pilosity. Profemur (
<figureCitation id="136EFD89FFBB6851FEC34994FE1CF97B" box="[353,415,1687,1710]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 5</figureCitation>
) on posterior face with basal row of three short spiny setae; posterior row of flexor side with short stout setae. Other black setae on legs normal, slender, of varying length.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FF174814FB84FD40" blockId="1.[151,761,822,2062]" lastBlockId="1.[828,1437,190,2069]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FF174814FEAAF8FB" box="[181,297,1815,1838]" italics="true" pageId="1" pageNumber="420">Structures.</emphasis>
Jugum not flattened, narrow, with few black spicula posteriorly. Proepisternum anteriorly without black spicula. Pronotum more than twice as long as head, covering mesonotum. Protrochanter (
<figureCitation id="136EFD89FFBB6851FE4D4874FDB3F85B" box="[495,560,1911,1934]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 5</figureCitation>
) without tubercle. Profemur (
<figureCitation id="136EFD89FFBB6851FEAA4894FEC4F87B" box="[264,327,1943,1966]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 5</figureCitation>
) with distinct concavity on extensor side. Metafemur (
<figureCitation id="136EFD89FFBB6851FE8148B4FEE4F81B" box="[291,359,1975,1998]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 7</figureCitation>
) moderately incrassate, ca. 3.53.6 times as long as wide, on flexor side with rich dentition: basal row consisting of 1215 densely set short spines only weakly increasing in length distally, their distances mostly smaller than their length; posterodistal row consisting of 1214 teeth strongly decreasing in size distally, the first one considerably longer than the second; anterodistal row consisting of 1012 minute spines. Metatibia (
<figureCitation id="136EFD89FFBB6851FAED4E3DFA17FE80" box="[1359,1428,318,341]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 7</figureCitation>
) straight, on flexor side with fine tooth-like structures all over length, with very short apical spine. Pregenital segments of abdomen moderately slender. Tergites relatively narrow; tergites 15 matt, 6 with a medial shiny spot, 7 almost entirely shiny (except margins). In the
<typeStatus id="54EE5FAEFFBB6851FA9A4EDDFA1BFE20" box="[1336,1432,478,501]" pageId="1" pageNumber="420" type="holotype">holotype</typeStatus>
, fifth tergite 2.8 times as wide as long; seventh tergite 1.8 times as long as sixth and 1.0 times as long as wide. Sternites almost unmodified; sternite 7 posteriorly with very shallow semicircular depressed area; hind margin slightly concave in middle.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FCF84D9DFA1EFCE0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="420">
Genital segments small and weakly modified. Pygophore slender, subovate. Proctiger (
<figureCitation id="136EFD89FFBB6851FB1A4DBDFB7AFD00" box="[1208,1273,702,725]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 9</figureCitation>
) slender, basolateral lobes moderately protruding and posteriorly angled. Paramere (
<figureCitation id="136EFD89FFBB6851FC0E4DFDFC75FCC0" box="[940,1014,766,789]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="1" pageNumber="420">Fig. 11</figureCitation>
) small, curved, but almost parallel-sided over its length; apex hardly widened and evenly rounded.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FCF84C3EFC72FC00" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FCF84C3EFBBEFC80" bold="true" box="[858,1085,829,853]" pageId="1" pageNumber="420">
<emphasis id="B9213D1EFFBB6851FCF84C3EFBB7FC80" bold="true" box="[858,1076,829,853]" italics="true" pageId="1" pageNumber="420">Female (apterous)</emphasis>
.
</emphasis>
Measurements (n = 6). Body length 2.632.94; maximum body width (at metapleura) 1.131.20. Head length 0.360.42, width 0.750.81; minimum eye distance 0.100.13. Pronotum length 0.750.83, width 1.031.14.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FCF84CDDFBBFFB00" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="420">
Colour (
<figureCitation id="136EFD89FFBB6851FC174CDDFC75FC20" box="[949,1014,990,1013]" captionStart="Figs 14. 12" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. 12 Rhagovelia madli sp. nov.: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3 R. kalalaoensis sp. nov., holotype, apterous male; 4 R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="1" pageNumber="420">Fig. 2</figureCitation>
) of head, thorax and appendages as in male. Tergite 6 more often without, rarely with, very small yellowish dot; spot on tergite 7 very large, almost filling its entire surface. Laterotergites (2)36 with a continuous, posteriorly narrowed, yellow stripe over middle parts. Sternites with broad yellow submarginal stripe near connexiva 37; on sternite 7 strongly narrowed and absent from connexival corner.
</paragraph>
<paragraph id="8BEAE10CFFBB6851FCF84BDDFBC0F9A0" blockId="1.[828,1437,190,2069]" pageId="1" pageNumber="420">Pilosity of head and thorax similar to that of male, but strongly reduced on tergites 13, completely absent from tergites 46; tergite 7 with sparse pubescence and some longer black setae at hind margin; tergite 8 with sparse pubescence medially, but with stripes of comparatively long, black setae at sides. Laterotergites 26 with stripes of short, inconspicuous pilosity at lateral margins; laterotergite 7 with long, black, posteriorly directed pilosity all over surface. Proctiger with short pilosity; gonocoxa 1 dorsally with black pilosity, other areas only inconspicuously hirsute. Profemur with 3 setae at base of posterior face and the posterior row of flexor side longer and more slender when compared to male.</paragraph>
<paragraph id="8BEAE10CFFBB6852FCF8497DFE0DF84F" blockId="1.[828,1437,190,2069]" lastBlockId="2.[151,760,1827,2074]" lastPageId="2" lastPageNumber="421" pageId="1" pageNumber="420">
Structural characteristics. Head and thorax as in male. Mesofemur flattened. Metafemur more slender than that of male, ca. 4.14.4 times as long as wide, on flexor side with distal row consisting of 710 teeth. Metatibia without dentition on flexor side. Abdomen (
<figureCitation id="136EFD89FFBB6851FB1549FDFB74F8C0" box="[1207,1271,1790,1813]" captionStart="Figs 14. 12" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. 12 Rhagovelia madli sp. nov.: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3 R. kalalaoensis sp. nov., holotype, apterous male; 4 R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="1" pageNumber="420">Fig. 2</figureCitation>
) strongly modified. Connexiva convergent on segment 23, strongly convergent on segments 45, and subparallel on segments 67; sternites 37 narrowly visible in dorsal aspect of specimen. Connexiva on segments 47 slightly thicker than on segments 23, but without distinct swellings. Tergites 13 convex, 48 flat. Fifth tergite 1.9 times as wide as long; seventh tergite 1.6 times as long as sixth and 1.2 times as long as wide; tergite 6 without or with small shiny spot at middle; tergite 7 almost entirely shiny. Tergite 8 either directed posteriorly or ventroposteriorly. In lateral view (
<figureCitation id="136EFD89FFB86852FF3D4860FF6AF8AF" box="[159,233,1891,1914]" captionStart="Figs 1318" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 1318.Characteristics of females: 1316 apex of abdomen, lateral view: 13 Rhagovelia madli sp. nov., 14 R. kalalaoensis sp. nov., 15 R. ambra Polhemus &amp; Andersen, 2015, 16 R. aquacola Polhemus &amp; Andersen, 2015; 1718 tergites 7 and 8, dorsal view: 17 R. kalalaoensis sp. nov., 18 R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="2" pageNumber="421">Fig. 13</figureCitation>
) posterior corner of sternite 7 blunt; ventral outline of gonocoxa 1 straight.
</paragraph>
<caption id="DF2AB184FFB86852FF3549B0FC62F934" ID-DOI="http://doi.org/10.5281/zenodo.4489336" ID-Zenodo-Dep="4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="2" pageNumber="421" startId="2.[151,188,1715,1734]" targetBox="[151,1436,189,1697]" targetPageId="2">
<paragraph id="8BEAE10CFFB86852FF3549B0FC62F934" blockId="2.[151,1437,1715,1761]" pageId="2" pageNumber="421">
Figs 14. 12
<taxonomicName id="4C559A8FFFB86852FE8249B0FE34F913" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[288,439,1715,1734]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB86852FE8249B0FE34F913" box="[288,439,1715,1734]" italics="true" pageId="2" pageNumber="421">Rhagovelia madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB86852FE1949B0FD82F913" box="[443,513,1715,1734]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3
<taxonomicName id="4C559A8FFFB86852FBD849B0FB7FF913" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[1146,1276,1715,1734]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB86852FBD849B0FB7FF913" box="[1146,1276,1715,1734]" italics="true" pageId="2" pageNumber="421">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB86852FAA349B0FAC6F913" box="[1281,1349,1715,1734]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, apterous male; 4
<taxonomicName id="4C559A8FFFB86852FE9B49CDFDA4F934" authority="Hoberlandt, 1951" authorityName="Hoberlandt" authorityYear="1951" box="[313,551,1742,1761]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="reuteri">
<emphasis id="B9213D1EFFB86852FE9B49CDFE09F934" box="[313,394,1742,1761]" italics="true" pageId="2" pageNumber="421">R. reuteri</emphasis>
<bibRefCitation id="EFC49CFDFFB86852FE3249CDFDA4F934" author="HOBERLANDT L." box="[400,551,1742,1761]" pageId="2" pageNumber="421" pagination="1 - 12" refId="ref6147" refString="HOBERLANDT L. 1951: Madagascan species of Rhagovelia. Societas Scientiarum Fennica, Commentationes Biologicae 12 (5): 1 - 12." type="journal article" year="1951">Hoberlandt, 1951</bibRefCitation>
</taxonomicName>
, small apterous male with reduced metatibial tooth.
</paragraph>
</caption>
<paragraph id="8BEAE10CFFB86852FF1748A1FDA4F86F" blockId="2.[151,760,1827,2074]" box="[181,551,1954,1978]" pageId="2" pageNumber="421">
<emphasis id="B9213D1EFFB86852FF1748A1FE33F86F" bold="true" box="[181,432,1954,1978]" italics="true" pageId="2" pageNumber="421">Macropterous morphs.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BEAE10CFFB86853FF3548C0FE33F96A" blockId="2.[151,760,1827,2074]" lastBlockId="3.[151,760,1672,2079]" lastPageId="3" lastPageNumber="422" pageId="2" pageNumber="421">
<emphasis id="B9213D1EFFB86852FF3548C0FEF7F80F" bold="true" box="[151,372,1987,2010]" pageId="2" pageNumber="421">Comparative notes.</emphasis>
<taxonomicName id="4C559A8FFFB86852FED848C0FDB6F80F" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[378,565,1987,2010]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB86852FED848C0FDB6F80F" box="[378,565,1987,2010]" italics="true" pageId="2" pageNumber="421">Rhagovelia madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB86852FD9948C0FD0FF80F" box="[571,652,1987,2010]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
is a member of the
<taxonomicName id="4C559A8FFFB86852FEAF48E0FEE9F82F" authorityName="Hoberlandt" authorityYear="1941" box="[269,362,2019,2042]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="tesari">
<emphasis id="B9213D1EFFB86852FEAF48E0FEE9F82F" box="[269,362,2019,2042]" italics="true" pageId="2" pageNumber="421">R. tesari</emphasis>
</taxonomicName>
group. Within this group it belongs to an assemblage of species (the
<taxonomicName id="4C559A8FFFB86852FDB74700FDFDF7CF" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[533,638,2051,2074]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB86852FDB74700FDFDF7CF" box="[533,638,2051,2074]" italics="true" pageId="2" pageNumber="421">R. ambra</emphasis>
</taxonomicName>
subgroup) characterized by spiny setae on the profemora of males (
<figureCitation id="136EFD89FFB86852FCE64840FC00F88F" box="[836,899,1859,1882]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="2" pageNumber="421">Fig. 5</figureCitation>
) and the flattened mesofemora of females (PƟΓΗΕΜUඌ &amp; AΝĐΕRඌΕΝ 2015: key couplet 4). An approximate identification of the new species by the key of PƟΓΗΕΜUඌ &amp; AΝĐΕRඌΕΝ 2015) already fails in couplet 5, because the characters are contradictory (yellow pronotal spots separated by a narrow black line, but ventral parts of thorax and abdomen partly yellow). Moreover,
<taxonomicName id="4C559A8FFFB86852FB484700FAC7F7CF" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[1258,1348,2051,2074]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB86852FB484700FAC7F7CF" box="[1258,1348,2051,2074]" italics="true" pageId="2" pageNumber="421">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB86852FAE84700FA1EF7CF" box="[1354,1437,2051,2074]" pageId="2" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
is distinctly smaller than any previously described species of the
<taxonomicName id="4C559A8FFFB96853FF7949ABFEBCF96A" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[219,319,1704,1727]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FF7949ABFEBCF96A" box="[219,319,1704,1727]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
subgroup.
</paragraph>
<caption id="DF2AB184FFB96853FF354C8BFB04FC04" ID-DOI="http://doi.org/10.5281/zenodo.4489340" ID-Zenodo-Dep="4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="3" pageNumber="422" startId="3.[151,188,904,923]" targetBox="[283,1301,192,879]" targetPageId="3">
<paragraph id="8BEAE10CFFB96853FF354C8BFB04FC04" blockId="3.[151,1437,904,977]" pageId="3" pageNumber="422">
Figs 512. Characteristics of males: 56 right foreleg: 5
<taxonomicName id="4C559A8FFFB96853FD3F4C8BFCB6FC4E" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[669,821,904,923]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FD3F4C8BFCB6FC4E" box="[669,821,904,923]" italics="true" pageId="3" pageNumber="422">Rhagovelia madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FC984C8BFCFCFC4E" box="[826,895,904,923]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
, 6
<taxonomicName id="4C559A8FFFB96853FC0A4C8BFBA8FC4E" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[936,1067,904,923]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FC0A4C8BFBA8FC4E" box="[936,1067,904,923]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FB924C8BFBF5FC4E" box="[1072,1142,904,923]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
; 78 right hind leg: 7
<taxonomicName id="4C559A8FFFB96853FAF64C8BFA1EFC4E" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[1364,1437,904,923]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FAF64C8BFA1EFC4E" box="[1364,1437,904,923]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FF354CA0FF59FC63" box="[151,218,931,950]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
8
<taxonomicName id="4C559A8FFFB96853FF5D4CA0FE01FC63" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[255,386,931,950]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FF5D4CA0FE01FC63" box="[255,386,931,950]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FE254CA0FE4EFC63" box="[391,461,931,950]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
; 910 pygophore, dorsal view: 9
<taxonomicName id="4C559A8FFFB96853FCA84CA0FCD0FC63" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[778,851,931,950]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FCA84CA0FCD0FC63" box="[778,851,931,950]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FCFA4CA0FC1FFC63" box="[856,924,931,950]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
10
<taxonomicName id="4C559A8FFFB96853FC694CA0FBCDFC63" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[971,1102,931,950]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FC694CA0FBCDFC63" box="[971,1102,931,950]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FBF14CA0FB1BFC63" box="[1107,1176,931,950]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
; 1112 left paramere, lateral view: 11
<taxonomicName id="4C559A8FFFB96853FF544CBDFEBDFC04" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[246,318,958,977]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FF544CBDFEBDFC04" box="[246,318,958,977]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FEE14CBDFE05FC04" box="[323,390,958,977]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
12
<taxonomicName id="4C559A8FFFB96853FE174CBDFDBBFC04" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[437,568,958,977]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FE174CBDFDBBFC04" box="[437,568,958,977]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FD9F4CBDFD03FC04" box="[573,640,958,977]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
Pilosity in Figs 56 partly, in Figs 710 completely omitted.
</paragraph>
</caption>
<caption id="DF2AB184FFB96853FF344AAFFDC7F9FF" ID-DOI="http://doi.org/10.5281/zenodo.4489344" ID-Zenodo-Dep="4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="3" pageNumber="422" startId="3.[150,187,1452,1471]" targetBox="[151,756,1021,1417]" targetPageId="3">
<paragraph id="8BEAE10CFFB96853FF344AAFFDC7F9FF" blockId="3.[150,760,1452,1578]" pageId="3" pageNumber="422">
Figs 1318.Characteristics of females: 1316 apex of abdomen, lateral view: 13
<taxonomicName id="4C559A8FFFB96853FF514AC4FE09FA0F" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[243,394,1479,1498]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FF514AC4FE09FA0F" box="[243,394,1479,1498]" italics="true" pageId="3" pageNumber="422">Rhagovelia madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FE2C4AC4FE51FA0F" box="[398,466,1479,1498]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
, 14
<taxonomicName id="4C559A8FFFB96853FDA04AC4FD07FA0F" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[514,644,1479,1498]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FDA04AC4FD07FA0F" box="[514,644,1479,1498]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FD2A4AC4FD4FFA0F" box="[648,716,1479,1498]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
, 15
<taxonomicName id="4C559A8FFFB96853FF344AE1FE6BFA20" authority="Polhemus &amp; Andersen, 2015" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[150,488,1506,1525]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FF344AE1FF64FA20" box="[150,231,1506,1525]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
<bibRefCitation id="EFC49CFDFFB96853FF4F4AE1FE6BFA20" author="POLHEMUS D. A. &amp; ANDERSEN N. M." box="[237,488,1506,1525]" pageId="3" pageNumber="422" pagination="115 - 220" refId="ref6614" refString="POLHEMUS D. A. &amp; ANDERSEN N. M. 2015: Rhagovelia of Madagascar and nearby Indian Ocean Islands (Hemiptera: Veliidae), Part 2: Revision of the tesari species group. Insect Systematics and Evolution 46: 115 - 220." type="journal article" year="2015">Polhemus &amp; Andersen, 2015</bibRefCitation>
</taxonomicName>
, 16
<taxonomicName id="4C559A8FFFB96853FDBC4AE1FE98F9DA" authority="Polhemus &amp; Andersen, 2015" authorityName="Polhemus &amp; Andersen" authorityYear="2015" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="aquacola">
<emphasis id="B9213D1EFFB96853FDBC4AE1FD05FA20" box="[542,646,1506,1525]" italics="true" pageId="3" pageNumber="422">R. aquacola</emphasis>
<bibRefCitation id="EFC49CFDFFB96853FD2E4AE1FE98F9DA" author="POLHEMUS D. A. &amp; ANDERSEN N. M." pageId="3" pageNumber="422" pagination="115 - 220" refId="ref6614" refString="POLHEMUS D. A. &amp; ANDERSEN N. M. 2015: Rhagovelia of Madagascar and nearby Indian Ocean Islands (Hemiptera: Veliidae), Part 2: Revision of the tesari species group. Insect Systematics and Evolution 46: 115 - 220." type="journal article" year="2015">Polhemus &amp; Andersen, 2015</bibRefCitation>
</taxonomicName>
; 1718 tergites 7 and 8, dorsal view: 17
<taxonomicName id="4C559A8FFFB96853FDD84AFFFD7BF9DA" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[634,760,1532,1551]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="kalalaoensis" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FDD84AFFFD7BF9DA" box="[634,760,1532,1551]" italics="true" pageId="3" pageNumber="422">R. kalalaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FF344914FF58F9FF" box="[150,219,1559,1578]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
, 18
<taxonomicName id="4C559A8FFFB96853FEAF4914FEF7F9FF" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[269,372,1559,1578]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="aquacola">
<emphasis id="B9213D1EFFB96853FEAF4914FEF7F9FF" box="[269,372,1559,1578]" italics="true" pageId="3" pageNumber="422">R. aquacola</emphasis>
</taxonomicName>
; pilosity partly omitted.
</paragraph>
</caption>
<paragraph id="8BEAE10CFFB96853FF1749CBFB7CFA66" blockId="3.[151,760,1672,2079]" lastBlockId="3.[828,1437,1020,1491]" pageId="3" pageNumber="422">
Based on the structure of the females abdomen,
<taxonomicName id="4C559A8FFFB96853FD0349CBFD7BF90A" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[673,760,1736,1759]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FD0349CBFD7BF90A" box="[673,760,1736,1759]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FF3549EBFF69F92A" box="[151,234,1768,1791]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
appears closely related to
<taxonomicName id="4C559A8FFFB96853FDAA49EBFEBCF8CA" authority="Polhemus &amp; Andersen, 2015" authorityName="Polhemus &amp; Andersen" authorityYear="2015" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FDAA49EBFDEFF92A" box="[520,620,1768,1791]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
<bibRefCitation id="EFC49CFDFFB96853FDD149EBFEBCF8CA" author="POLHEMUS D. A. &amp; ANDERSEN N. M." pageId="3" pageNumber="422" pagination="115 - 220" refId="ref6614" refString="POLHEMUS D. A. &amp; ANDERSEN N. M. 2015: Rhagovelia of Madagascar and nearby Indian Ocean Islands (Hemiptera: Veliidae), Part 2: Revision of the tesari species group. Insect Systematics and Evolution 46: 115 - 220." type="journal article" year="2015">Polhemus &amp; Andersen, 2015</bibRefCitation>
</taxonomicName>
, a species endemic to the northernmost region of
<collectingCountry id="F342A19CFFB96853FF4A482BFEEBF8EA" box="[232,360,1832,1855]" name="Madagascar" pageId="3" pageNumber="422">Madagascar</collectingCountry>
. However, there are several characters separating these two species. For comparison, we used an apterous female of
<taxonomicName id="4C559A8FFFB96853FEC5486BFE48F8AA" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[359,459,1896,1919]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FEC5486BFE48F8AA" box="[359,459,1896,1919]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
from the
<typeStatus id="54EE5FAEFFB96853FD9A486BFDE6F8AA" box="[568,613,1896,1919]" pageId="3" pageNumber="422">type</typeStatus>
area, and the original description for the differences in males. In colouration
<taxonomicName id="4C559A8FFFB96853FF7348ABFEA9F86A" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[209,298,1960,1983]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FF7348ABFEA9F86A" box="[209,298,1960,1983]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FE8D48ABFE03F86A" box="[303,384,1960,1983]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="136EFD89FFB96853FE2E48ABFE6BF86A" box="[396,488,1960,1983]" captionStart="Figs 14. 12" captionStartId="2.[151,188,1715,1734]" captionTargetBox="[151,1436,189,1697]" captionTargetId="figure-13@2.[151,1436,189,1698]" captionTargetPageId="2" captionText="Figs 14. 12 Rhagovelia madli sp. nov.: 1 habitus of holotype, apterous male; 2 paratype, apterous female; 3 R. kalalaoensis sp. nov., holotype, apterous male; 4 R. reuteri Hoberlandt, 1951, small apterous male with reduced metatibial tooth." figureDoi="http://doi.org/10.5281/zenodo.4489336" httpUri="https://zenodo.org/record/4489336/files/figure.png" pageId="3" pageNumber="422">Figs 1, 2</figureCitation>
) differs from
<taxonomicName id="4C559A8FFFB96853FDD448ABFD5BF86A" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[630,728,1960,1983]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FDD448ABFD5BF86A" box="[630,728,1960,1983]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
by having the yellow pronotal spots separated by a black line (vs. confluent), and by sternites 26 being black except for the connexival margins (vs. yellow except for a black lateral stripe). Differences in apterous females are conspicuous: in
<taxonomicName id="4C559A8FFFB96853FC9E4B1FFC14FBE6" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[828,919,1052,1075]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FC9E4B1FFC14FBE6" box="[828,919,1052,1075]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FC3D4B1FFC70FBE6" box="[927,1011,1052,1075]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
the connexiva are almost parallel-sided over segments 6 and
<quantity id="4CAD4CE9FFB96853FBB54B3FFBBCFB86" box="[1047,1087,1084,1107]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="3" pageNumber="422" unit="in" value="7.0">7 in</quantity>
dorsal view (
<figureCitation id="136EFD89FFB96853FB6E4B3FFA8FFB86" box="[1228,1292,1084,1107]" captionStart="Figs 512" captionStartId="3.[151,188,904,923]" captionTargetBox="[283,1301,192,879]" captionTargetId="figure-441@3.[281,1304,189,884]" captionTargetPageId="3" captionText="Figs 512. Characteristics of males: 56 right foreleg: 5 Rhagovelia madli sp. nov., 6 R. kalalaoensis sp. nov.; 78 right hind leg: 7 R. madli sp. nov. 8 R. kalalaoensis sp. nov.; 910 pygophore, dorsal view: 9 R. madli sp. nov. 10 R. kalalaoensis sp. nov.; 1112 left paramere, lateral view: 11 R. madli sp. nov. 12 R. kalalaoensis sp. nov. Pilosity in Figs 56 partly, in Figs 710 completely omitted." figureDoi="http://doi.org/10.5281/zenodo.4489340" httpUri="https://zenodo.org/record/4489340/files/figure.png" pageId="3" pageNumber="422">Fig. 7</figureCitation>
; vs. distinctly convex in
<taxonomicName id="4C559A8FFFB96853FC0A4B5FFB8AFBA6" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[936,1033,1116,1139]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FC0A4B5FFB8AFBA6" box="[936,1033,1116,1139]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
), the laterotergites 7 bear long pilosity all over their surface (vs. long pilosity only posteriorly present in
<taxonomicName id="4C559A8FFFB96853FC044B9FFB84FB66" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[934,1031,1180,1203]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FC044B9FFB84FB66" box="[934,1031,1180,1203]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
), the posterolateral connexival angle is bluntly angular in lateral aspect (vs. broadly rounded in
<taxonomicName id="4C559A8FFFB96853FA274BBFFC03FB26" authorityName="Polhemus &amp; Andersen" authorityYear="2015" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FA274BBFFC03FB26" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
; comp.
<figureCitation id="136EFD89FFB96853FC714BDFFBEEFB26" box="[979,1133,1244,1267]" captionStart="Figs 1318" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 1318.Characteristics of females: 1316 apex of abdomen, lateral view: 13 Rhagovelia madli sp. nov., 14 R. kalalaoensis sp. nov., 15 R. ambra Polhemus &amp; Andersen, 2015, 16 R. aquacola Polhemus &amp; Andersen, 2015; 1718 tergites 7 and 8, dorsal view: 17 R. kalalaoensis sp. nov., 18 R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="3" pageNumber="422">Figs 13 and 15</figureCitation>
), and the apex of gonocoxa 1 is lacking long black setae (present in
<taxonomicName id="4C559A8FFFB96853FB1C4BFFFA9EFAC6" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[1214,1309,1276,1299]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FB1C4BFFFA9EFAC6" box="[1214,1309,1276,1299]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
; comp.
<figureCitation id="136EFD89FFB96853FACD4BFFFC25FAE6" captionStart="Figs 1318" captionStartId="3.[150,187,1452,1471]" captionTargetBox="[151,756,1021,1417]" captionTargetId="figure-527@3.[151,756,1019,1417]" captionTargetPageId="3" captionText="Figs 1318.Characteristics of females: 1316 apex of abdomen, lateral view: 13 Rhagovelia madli sp. nov., 14 R. kalalaoensis sp. nov., 15 R. ambra Polhemus &amp; Andersen, 2015, 16 R. aquacola Polhemus &amp; Andersen, 2015; 1718 tergites 7 and 8, dorsal view: 17 R. kalalaoensis sp. nov., 18 R. aquacola; pilosity partly omitted." figureDoi="http://doi.org/10.5281/zenodo.4489344" httpUri="https://zenodo.org/record/4489344/files/figure.png" pageId="3" pageNumber="422">Figs 13 and 15</figureCitation>
). Males differ by the absence of a hairy tubercle on the protrochanter that is present in
<taxonomicName id="4C559A8FFFB96853FB6B4A3FFAA9FA86" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[1225,1322,1340,1363]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FB6B4A3FFAA9FA86" box="[1225,1322,1340,1363]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
, but not in
<taxonomicName id="4C559A8FFFB96853FC9E4A5FFC16FAA6" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[828,917,1372,1395]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FC9E4A5FFC16FAA6" box="[828,917,1372,1395]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FC384A5FFC6EFAA6" box="[922,1005,1372,1395]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
, and by the shape of the paramere, which is much shorter in
<taxonomicName id="4C559A8FFFB96853FC544A7FFBD5FA46" authorityName="Polhemus &amp; Andersen" authorityYear="2015" box="[1014,1110,1404,1427]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="ambra">
<emphasis id="B9213D1EFFB96853FC544A7FFBD5FA46" box="[1014,1110,1404,1427]" italics="true" pageId="3" pageNumber="422">R. ambra</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C559A8FFFB96853FB074A7FFB7FFA46" authority="Zettel &amp; Laciny, 2020" authorityName="Zettel &amp; Laciny" authorityYear="2020" box="[1189,1276,1404,1427]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="422" phylum="Arthropoda" rank="species" species="madli" status="sp. nov.">
<emphasis id="B9213D1EFFB96853FB074A7FFB7FFA46" box="[1189,1276,1404,1427]" italics="true" pageId="3" pageNumber="422">R. madli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2128065FFB96853FAA24A7FFACCFA46" box="[1280,1359,1404,1427]" pageId="3" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
(comp. PƟΓΗΕΜUඌ &amp; AΝĐΕRඌΕΝ 2015: Figs 61, 62).
</paragraph>
</subSubSection>
<subSubSection id="C34FB287FFB96853FC9E4ABFFA06FA06" box="[828,1413,1468,1491]" pageId="3" pageNumber="422" type="etymology">
<paragraph id="8BEAE10CFFB96853FC9E4ABFFA06FA06" blockId="3.[828,1437,1020,1491]" box="[828,1413,1468,1491]" pageId="3" pageNumber="422">
<emphasis id="B9213D1EFFB96853FC9E4ABFFC3FFA06" bold="true" box="[828,956,1468,1491]" pageId="3" pageNumber="422">Etymology.</emphasis>
Dedicated to our colleague Michael Madl.
</paragraph>
</subSubSection>
</treatment>
</document>