treatments-xml/data/03/FC/26/03FC2669F7512F3A6130FDA9FF13FC68.xml

<document id="230FCD3FC95899CE6A6A53E43B23621E" ID-CLB-Dataset="51057" ID-DOI="10.5281/zenodo.171028" ID-GBIF-Dataset="d3f1ce59-f24d-4abf-9ebd-28af273be687" ID-ISSN="1175­5326" ID-Zenodo-Dep="171028" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459717943265" checkinUser="plazi" docAuthor="Helgen, Kristofer M." docDate="2005" docId="03FC2669F7512F3A6130FDA9FF13FC68" docLanguage="en" docName="zt00913.pdf" docOrigin="Zootaxa 913" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hydromys ziegleri Helgen, 2005, new species" docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="FFC55E11F7582F2B6038FFF5FC03FFB6" masterDocTitle="the generic status of Baiyankamys and description of a new species of Hydromys" masterLastPageNumber="20" masterPageNumber="1" pageNumber="10" updateTime="1698212093946" updateUser="ExternalLinkService">
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<mods:title id="12D0195CFBEFB29311388A8AB1438711">the generic status of Baiyankamys and description of a new species of Hydromys</mods:title>
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<mods:namePart id="251DB66381E78C332D9689F185A469E2">Helgen, Kristofer M.</mods:namePart>
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<mods:date id="D2B6124D62A04D74146CFDD7409DE5AA">2005</mods:date>
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<treatment id="03FC2669F7512F3A6130FDA9FF13FC68" ID-DOI="http://doi.org/10.5281/zenodo.5628221" ID-GBIF-Taxon="119335025" ID-Zenodo-Dep="5628221" LSID="urn:lsid:plazi:treatment:03FC2669F7512F3A6130FDA9FF13FC68" httpUri="http://treatment.plazi.org/id/03FC2669F7512F3A6130FDA9FF13FC68" lastPageId="17" lastPageNumber="18" pageId="9" pageNumber="10">
<subSubSection id="C34FC4F4F7512F226130FDA9FE21FD2F" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BEA977FF7512F226130FDA9FE79FDC3" blockId="9.[264,634,603,665]" box="[264,634,603,630]" pageId="9" pageNumber="10">
<heading id="D0A22013F7512F226130FDA9FE79FDC3" bold="true" box="[264,634,603,630]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B9214B6DF7512F226130FDA9FE79FDC3" bold="true" box="[264,634,603,630]" pageId="9" pageNumber="10">
<taxonomicName id="4C55ECFCF7512F226130FDA9FDE0FDC0" ID-CoL="3N7S8" box="[264,483,604,630]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri" status="sp. nov.">
<emphasis id="B9214B6DF7512F226130FDA9FDE0FDC0" bold="true" box="[264,483,604,630]" italics="true" pageId="9" pageNumber="10">Hydromys ziegleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212F616F7512F2261D2FDAEFE79FDC3" box="[490,634,603,629]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA977FF7512F226130FD8AFE21FD2F" blockId="9.[264,634,603,665]" box="[264,546,639,665]" pageId="9" pageNumber="10">
<figureCitation id="136E8BFAF7512F226130FD8AFD74FD2F" box="[264,375,639,665]" captionStart="FIGURE 3" captionStartId="10.[264,368,1721,1745]" captionTargetBox="[291,1296,613,1678]" captionTargetId="figure@10.[291,1296,597,1678]" captionTargetPageId="10" captionText="FIGURE 3. Dorsal and ventral views of the skins of Hydromys ziegleri (a, c; BBM­NG 101683, adult female holotype) and Hydromys hussoni (b, d; AM M 18627; adult female paratype). Scale bar = 20 mm." httpUri="https://zenodo.org/record/171031/files/figure.png" pageId="9" pageNumber="10">Figures 3</figureCitation>
, 
<figureCitation id="136E8BFAF7512F2261BCFD8AFD96FD2F" box="[388,405,639,665]" captionStart="FIGURE 4" captionStartId="11.[264,368,1454,1478]" captionTargetBox="[374,1213,729,1428]" captionTargetId="figure@11.[374,1214,729,1428]" captionTargetPageId="11" captionText="FIGURE 4. Crania and mandibles of Hydromys ziegleri (below, adult female holotype) and Hydromys hussoni (above, AM M 18627, adult female paratype)." httpUri="https://zenodo.org/record/171032/files/figure.png" pageId="9" pageNumber="10">4</figureCitation>
; 
<tableCitation id="C6D7A2C4F7512F22619BFD8AFE05FD2F" box="[419,518,639,665]" captionStart="TABLE 3" captionStartId="12.[264,352,676,700]" captionText="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." pageId="9" pageNumber="10">Tables 3</tableCitation>
, 
<tableCitation id="C6D7A2C4F7512F22622BFD8AFE21FD2F" box="[531,546,639,665]" captionStart="TABLE 4" captionStartId="13.[264,352,1209,1233]" captionTargetBox="[264,1226,1311,1770]" captionTargetPageId="13" captionText="TABLE 4. Comparative molar dimensions in specimens of Hydromys ziegleri (adult female holotype) and Hydromys hussoni (AM M 18627, adult female paratype)." httpUri="http://table.plazi.org/id/DF2AC7F7F7552F266130FB4CFFD4FB4F" pageId="9" pageNumber="10" tableUuid="DF2AC7F7F7552F266130FB4CFFD4FB4F">4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7512F226130FD3BFE22FC3F" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BEA977FF7512F226130FD3BFE22FC3F" blockId="9.[264,1323,718,1905]" pageId="9" pageNumber="10">
<typeStatus id="54EE29DDF7512F226130FD3BFD79FD5E" box="[264,378,718,744]" pageId="9" pageNumber="10" type="holotype">
<emphasis id="B9214B6DF7512F226130FD3BFD79FD5E" bold="true" box="[264,378,718,744]" pageId="9" pageNumber="10">Holotype</emphasis>
</typeStatus>
: BBM­NG 
<date id="FFEBB1BFF7512F226231FD3AFE67FD5F" box="[521,612,719,745]" pageId="9" pageNumber="10" value="83-10-16">101683</date>
, adult female, skin (
<figureCitation id="136E8BFAF7512F22636DFD3AFF9FFD5F" box="[853,924,719,745]" captionStart="FIGURE 3" captionStartId="10.[264,368,1721,1745]" captionTargetBox="[291,1296,613,1678]" captionTargetId="figure@10.[291,1296,597,1678]" captionTargetPageId="10" captionText="FIGURE 3. Dorsal and ventral views of the skins of Hydromys ziegleri (a, c; BBM­NG 101683, adult female holotype) and Hydromys hussoni (b, d; AM M 18627; adult female paratype). Scale bar = 20 mm." httpUri="https://zenodo.org/record/171031/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
) and skull (
<figureCitation id="136E8BFAF7512F226414FD3AF876FD5F" box="[1068,1141,719,745]" captionStart="FIGURE 4" captionStartId="11.[264,368,1454,1478]" captionTargetBox="[374,1213,729,1428]" captionTargetId="figure@11.[374,1214,729,1428]" captionTargetPageId="11" captionText="FIGURE 4. Crania and mandibles of Hydromys ziegleri (below, adult female holotype) and Hydromys hussoni (above, AM M 18627, adult female paratype)." httpUri="https://zenodo.org/record/171032/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
), from Bainyik (
<geoCoordinate id="EE61F1B8F7512F226129FD02FD73FCA7" box="[273,368,759,785]" direction="south" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="-3.6666667">03°40'S</geoCoordinate>
, 
<geoCoordinate id="EE61F1B8F7512F226145FD02FDEEFCA7" box="[381,493,759,785]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="143.08333">143°05'E</geoCoordinate>
), 
<emphasis id="B9214B6DF7512F22623CFD02FE3CFCA6" box="[516,575,759,784]" italics="true" pageId="9" pageNumber="10">circa</emphasis>
<quantity id="4CAD3A9AF7512F22627FFD02FE86FCA7" box="[583,645,759,785]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="km" value="5.0">5 km</quantity>
south of Maprik, 
<quantity id="4CAD3A9AF7512F226365FD02FFA0FCA7" box="[861,931,759,785]" metricMagnitude="2" metricUnit="m" metricValue="2.1336" pageId="9" pageNumber="10" unit="ft" value="700.0">700 ft</quantity>
(= 
<quantity id="4CAD3A9AF7512F2263F7FD02F81DFCA7" box="[975,1054,759,785]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="9" pageNumber="10" unit="m" value="213.0">213 m</quantity>
), East Sepik Province, 
<collectingCountry id="F342D7EFF7512F226130FCEAFDF3FC8F" box="[264,496,799,825]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua New Guinea</collectingCountry>
, collected 
<date id="FFEBB1BFF7512F226257FCEAFF3BFC8F" box="[623,824,799,825]" pageId="9" pageNumber="10" value="1972-10-29">29 October 1972</date>
by A.B. Mirza. According to its label the holotype—the only known specimen of this species—was live­trapped in a local garden (see 
<bibRefCitation id="EFC4EA8EF7512F226106FC9AFDDAFC3F" author="Ziegler" box="[318,473,879,905]" pageId="9" pageNumber="20" refString="Ziegler, A. C. (1984) A Papua New Guinea specimen of Hydromys hussoni Musser &amp; Piik, 1982 (Rodentia: Muridae). Australian Mammalogy, 7, 101 - 105." type="journal article" year="1984">Ziegler 1984</bibRefCitation>
: 101).
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7512F226100FC63FE97FBE7" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8BEA977FF7512F226100FC63FE97FBE7" blockId="9.[264,1323,718,1905]" pageId="9" pageNumber="10">
<emphasis id="B9214B6DF7512F226100FC63FDB3FC06" bold="true" box="[312,432,918,944]" pageId="9" pageNumber="10">Diagnosis</emphasis>
: 
<taxonomicName id="4C55ECFCF7512F2261FCFC62FEA3FC06" box="[452,672,919,944]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F2261FCFC62FEA3FC06" box="[452,672,919,944]" italics="true" pageId="9" pageNumber="10">Hydromys ziegleri</emphasis>
</taxonomicName>
is the smallest species of 
<taxonomicName id="4C55ECFCF7512F2263D3FC62F861FC06" box="[1003,1122,919,944]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7512F2263D3FC62F861FC06" box="[1003,1122,919,944]" italics="true" pageId="9" pageNumber="10">Hydromys</emphasis>
</taxonomicName>
. It most closely resembles 
<taxonomicName id="4C55ECFCF7512F2261BFFC4AFE0BFC6E" box="[391,520,959,984]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7512F2261BFFC4AFE0BFC6E" box="[391,520,959,984]" italics="true" pageId="9" pageNumber="10">H. hussoni</emphasis>
</taxonomicName>
, endemic to the upland Wissel Lakes of west­central New 
<collectingCountry id="F342D7EFF7512F2264ECFC4AF92AFC6F" box="[1236,1321,959,985]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
(
<figureCitation id="136E8BFAF7512F226129FC12FD81FBB7" box="[273,386,999,1025]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="10.[264,368,1721,1745]" captionStartId-1="11.[264,368,1454,1478]" captionTargetBox-0="[291,1296,613,1678]" captionTargetBox-1="[374,1213,729,1428]" captionTargetId-0="figure@10.[291,1296,597,1678]" captionTargetId-1="figure@11.[374,1214,729,1428]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 3. Dorsal and ventral views of the skins of Hydromys ziegleri (a, c; BBM­NG 101683, adult female holotype) and Hydromys hussoni (b, d; AM M 18627; adult female paratype). Scale bar = 20 mm." captionText-1="FIGURE 4. Crania and mandibles of Hydromys ziegleri (below, adult female holotype) and Hydromys hussoni (above, AM M 18627, adult female paratype)." httpUri-0="https://zenodo.org/record/171031/files/figure.png" httpUri-1="https://zenodo.org/record/171032/files/figure.png" pageId="9" pageNumber="10">Figs. 3–4</figureCitation>
; 
<tableCitation id="C6D7A2C4F7512F2261B6FC12FE06FBB7" box="[398,517,999,1025]" captionStart-0="TABLE 3" captionStart-1="TABLE 4" captionStartId-0="12.[264,352,676,700]" captionStartId-1="13.[264,352,1209,1233]" captionTargetBox-1="[264,1226,1311,1770]" captionTargetPageId-1="13" captionText-0="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." captionText-1="TABLE 4. Comparative molar dimensions in specimens of Hydromys ziegleri (adult female holotype) and Hydromys hussoni (AM M 18627, adult female paratype)." httpUri-1="http://table.plazi.org/id/DF2AC7F7F7552F266130FB4CFFD4FB4F" pageId="9" pageNumber="10" tableUuid-1="DF2AC7F7F7552F266130FB4CFFD4FB4F">Table 3–4</tableCitation>
), but differs from that species in its smaller size, broadened rostrum, shorter incisive foramina, shorter toothrow, broader first molars, lower­domed braincase, and much less densely­furred tail.
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7512F226100FBABFF6EFA57" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BEA977FF7512F226100FBABFF6EFA57" blockId="9.[264,1323,718,1905]" pageId="9" pageNumber="10">
<emphasis id="B9214B6DF7512F226100FBABFDD2FBCE" bold="true" box="[312,465,1118,1144]" pageId="9" pageNumber="10">Distribution</emphasis>
: 
<taxonomicName id="4C55ECFCF7512F2261DAFBAAFEB9FBCE" box="[482,698,1119,1144]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F2261DAFBAAFEB9FBCE" box="[482,698,1119,1144]" italics="true" pageId="9" pageNumber="10">Hydromys ziegleri</emphasis>
</taxonomicName>
is known only from the 
<typeStatus id="54EE29DDF7512F2263DAFBAAF816FBCF" box="[994,1045,1119,1145]" pageId="9" pageNumber="10">type</typeStatus>
locality, situated in the foothills of the southern slopes of the Prince Alexander Range, one of the North Coast Ranges of 
<collectingCountry id="F342D7EFF7512F2261BCFB5AFE69FB7F" box="[388,618,1199,1225]" name="Papua New Guinea" pageId="9" pageNumber="10">Papua New Guinea</collectingCountry>
. The 
<typeStatus id="54EE29DDF7512F226290FB5AFED8FB7F" box="[680,731,1199,1225]" pageId="9" pageNumber="10">type</typeStatus>
locality lies in the vicinity of the Screw and Amuk Rivers, which drain from the highlands of the Prince Alexander Range and the Torricelli Range, respectively. I suspect that future collecting efforts may show it to occur more widely (and at higher altitudes) in the North Coast Ranges (see Discussion below). Two other amphibious murines (
<taxonomicName id="4C55ECFCF7512F226269FABAFF6BFADE" box="[593,872,1359,1384]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="chrysogaster">
<emphasis id="B9214B6DF7512F226269FABAFF6BFADE" box="[593,872,1359,1384]" italics="true" pageId="9" pageNumber="10">Hydromys chrysogaster</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7512F22639DFABAF898FADE" box="[933,1179,1359,1384]" class="Mammalia" family="Muridae" genus="Parahydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="asper">
<emphasis id="B9214B6DF7512F22639DFABAF898FADE" box="[933,1179,1359,1384]" italics="true" pageId="9" pageNumber="10">Parahydromys asper</emphasis>
</taxonomicName>
) are known from other localities in the North Coast Ranges and are thus likely to occur sympatrically with 
<taxonomicName id="4C55ECFCF7512F22617CFA6AFDC3FA0E" box="[324,448,1439,1464]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F22617CFA6AFDC3FA0E" box="[324,448,1439,1464]" italics="true" pageId="9" pageNumber="10">H. ziegleri</emphasis>
</taxonomicName>
, though only 
<taxonomicName id="4C55ECFCF7512F22625EFA6AFF20FA0E" box="[614,803,1439,1464]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="chrysogaster">
<emphasis id="B9214B6DF7512F22625EFA6AFF20FA0E" box="[614,803,1439,1464]" italics="true" pageId="9" pageNumber="10">H. chrysogaster</emphasis>
</taxonomicName>
has been collected in the same general area to date (e.g. CSIRO 8346, adult male from Maprik).
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7512F226100FA1BFD81F9EF" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BEA977FF7512F226100FA1BFD81F9EF" blockId="9.[264,1323,718,1905]" pageId="9" pageNumber="10">
<emphasis id="B9214B6DF7512F226100FA1BFDBCF9BE" bold="true" box="[312,447,1518,1544]" pageId="9" pageNumber="10">Etymology</emphasis>
: For the late Dr. Alan C. Ziegler (
<date id="FFEBB1BFF7512F226367FA1AFFE0F9BF" box="[863,995,1519,1545]" pageId="9" pageNumber="10" value="1929" valueMax="2003">1929­2003</date>
), former head of the Vertebrate Zoology Division at the Bishop Museum in Honolulu and authority on New 
<collectingCountry id="F342D7EFF7512F2264EDF9E2F929F987" box="[1237,1322,1559,1585]" name="Guinea-Bissau" pageId="9" pageNumber="10">Guinea</collectingCountry>
mammals.
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7512F256100F993FD3CFAF1" lastPageId="14" lastPageNumber="15" pageId="9" pageNumber="10" type="description">
<paragraph id="8BEA977FF7512F226100F993F928F8C7" blockId="9.[264,1323,718,1905]" pageId="9" pageNumber="10">
<emphasis id="B9214B6DF7512F226100F993FDCAF936" bold="true" box="[312,457,1638,1664]" pageId="9" pageNumber="10">Description</emphasis>
: The 
<typeStatus id="54EE29DDF7512F226234F992FE70F937" box="[524,627,1639,1665]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C55ECFCF7512F2262A0F992FEF3F936" box="[664,752,1639,1664]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F2262A0F992FEF3F936" box="[664,752,1639,1664]" italics="true" pageId="9" pageNumber="10">ziegleri</emphasis>
</taxonomicName>
is smaller than most specimens of 
<taxonomicName id="4C55ECFCF7512F2264BFF992F8E2F936" box="[1159,1249,1639,1664]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7512F2264BFF992F8E2F936" box="[1159,1249,1639,1664]" italics="true" pageId="9" pageNumber="10">hussoni</emphasis>
</taxonomicName>
(
<figureCitation id="136E8BFAF7512F2264D7F992FD34F91F" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="10.[264,368,1721,1745]" captionStartId-1="11.[264,368,1454,1478]" captionTargetBox-0="[291,1296,613,1678]" captionTargetBox-1="[374,1213,729,1428]" captionTargetId-0="figure@10.[291,1296,597,1678]" captionTargetId-1="figure@11.[374,1214,729,1428]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 3. Dorsal and ventral views of the skins of Hydromys ziegleri (a, c; BBM­NG 101683, adult female holotype) and Hydromys hussoni (b, d; AM M 18627; adult female paratype). Scale bar = 20 mm." captionText-1="FIGURE 4. Crania and mandibles of Hydromys ziegleri (below, adult female holotype) and Hydromys hussoni (above, AM M 18627, adult female paratype)." httpUri-0="https://zenodo.org/record/171031/files/figure.png" httpUri-1="https://zenodo.org/record/171032/files/figure.png" pageId="9" pageNumber="10">Figs. 3–4</figureCitation>
; 
<tableCitation id="C6D7A2C4F7512F22617CF97AFD9EF91F" box="[324,413,1679,1705]" captionStart="TABLE 3" captionStartId="12.[264,352,676,700]" captionText="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." pageId="9" pageNumber="10">Table 3</tableCitation>
). The dorsal pelage of 
<taxonomicName id="4C55ECFCF7512F226288F97AFF0BF91E" box="[688,776,1679,1704]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F226288F97AFF0BF91E" box="[688,776,1679,1704]" italics="true" pageId="9" pageNumber="10">ziegleri</emphasis>
</taxonomicName>
is dark brown (somewhat darker than the rich brown of 
<taxonomicName id="4C55ECFCF7512F226143F942FDD7F966" box="[379,468,1719,1744]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7512F226143F942FDD7F966" box="[379,468,1719,1744]" italics="true" pageId="9" pageNumber="10">hussoni</emphasis>
</taxonomicName>
) tipped with paler, orange­buff hairs. The fur is considerably shorter (
<quantity id="4CAD3A9AF7512F226534F942FD4FF94F" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="9" pageNumber="10" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6– 7 mm</quantity>
on the anterior dorsum), less dense, and conspicuously more glossy than in 
<taxonomicName id="4C55ECFCF7512F2264F2F92AF927F94E" box="[1226,1316,1759,1784]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7512F2264F2F92AF927F94E" box="[1226,1316,1759,1784]" italics="true" pageId="9" pageNumber="10">hussoni</emphasis>
</taxonomicName>
. The fur of the venter is grey­based with ochraceous­buff tipping, similar to 
<taxonomicName id="4C55ECFCF7512F2264E9F8F2F928F896" box="[1233,1323,1799,1824]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7512F2264E9F8F2F928F896" box="[1233,1323,1799,1824]" italics="true" pageId="9" pageNumber="10">hussoni</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC4EA8EF7512F226128F8DAFE16F8FF" author="Musser" box="[272,533,1839,1865]" pageId="9" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik 1982</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7512F22621AF8DAFEBEF8FF" author="Ziegler" box="[546,701,1839,1865]" pageId="9" pageNumber="20" refString="Ziegler, A. C. (1984) A Papua New Guinea specimen of Hydromys hussoni Musser &amp; Piik, 1982 (Rodentia: Muridae). Australian Mammalogy, 7, 101 - 105." type="journal article" year="1984">Ziegler 1984</bibRefCitation>
), but in the 
<typeStatus id="54EE29DDF7512F22637FF8DAFFAEF8FF" box="[839,941,1839,1865]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C55ECFCF7512F2263ECF8DAF82FF8FE" box="[980,1068,1839,1864]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7512F2263ECF8DAF82FF8FE" box="[980,1068,1839,1864]" italics="true" pageId="9" pageNumber="10">ziegleri</emphasis>
</taxonomicName>
the lower cheeks and the underside of the throat and chin are dingy white, contrasting with the rest of the venter.
</paragraph>
<paragraph id="8BEA977FF7522F216130FEE9FFB3FE48" blockId="10.[264,1323,284,510]" pageId="10" pageNumber="11">
The inguinal region and ventral base of the tail are dark brown (
<figureCitation id="136E8BFAF7522F216425FEE9F884FE80" box="[1053,1159,284,310]" captionStart="FIGURE 3" captionStartId="10.[264,368,1721,1745]" captionTargetBox="[291,1296,613,1678]" captionTargetId="figure@10.[291,1296,597,1678]" captionTargetPageId="10" captionText="FIGURE 3. Dorsal and ventral views of the skins of Hydromys ziegleri (a, c; BBM­NG 101683, adult female holotype) and Hydromys hussoni (b, d; AM M 18627; adult female paratype). Scale bar = 20 mm." httpUri="https://zenodo.org/record/171031/files/figure.png" pageId="10" pageNumber="11">Figure 3</figureCitation>
). 
<collectingCountry id="F342D7EFF7522F216499FEE9F8C1FE80" box="[1185,1218,284,310]" name="American Samoa" pageId="10" pageNumber="11">As</collectingCountry>
in other 
<emphasis id="B9214B6DF7522F216130FEB1FD85FEEB" box="[264,390,324,349]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C55ECFCF7522F216130FEB1FD81FEEB" box="[264,386,324,349]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">Hydromys</taxonomicName>
,
</emphasis>
the hindfeet are long and broad and bear conspicuous webbing between the middle digits (II–IV). The dorsal surface of the pes is pigmented dark but covered with small pale hairs, and the claws are equal in size to those of 
<emphasis id="B9214B6DF7522F216384FE61F81EFE1B" box="[956,1053,404,429]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C55ECFCF7522F216384FE61F81AFE1B" box="[956,1049,404,429]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hussoni">hussoni</taxonomicName>
,
</emphasis>
and unpigmented. The skin label gives the hindfoot length as 29, probably measured with the claw, as on the dry skin this length is closer to 27 
<emphasis id="B9214B6DF7522F216255FE11FEF4FE4B" box="[621,759,484,509]" italics="true" pageId="10" pageNumber="11">sans unguis</emphasis>
(
<bibRefCitation id="EFC4EA8EF7522F21633EFE11FFA0FE48" author="Ziegler" box="[774,931,484,510]" pageId="10" pageNumber="20" refString="Ziegler, A. C. (1984) A Papua New Guinea specimen of Hydromys hussoni Musser &amp; Piik, 1982 (Rodentia: Muridae). Australian Mammalogy, 7, 101 - 105." type="journal article" year="1984">Ziegler 1984</bibRefCitation>
).
</paragraph>
<caption id="DF2AC7F7F7522F216130F94CFD73F8A3" httpUri="https://zenodo.org/record/171031/files/figure.png" pageId="10" pageNumber="11" targetBox="[291,1296,613,1678]" targetPageId="10">
<paragraph id="8BEA977FF7522F216130F94CFD73F8A3" blockId="10.[264,1322,1721,1813]" pageId="10" pageNumber="11">
<emphasis id="B9214B6DF7522F216130F94CFD8FF967" bold="true" box="[264,396,1721,1745]" pageId="10" pageNumber="11">FIGURE 3.</emphasis>
Dorsal and ventral views of the skins of 
<taxonomicName id="4C55ECFCF7522F21636BF94CF81BF966" box="[851,1048,1721,1744]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7522F21636BF94CF81BF966" box="[851,1048,1721,1744]" italics="true" pageId="10" pageNumber="11">Hydromys ziegleri</emphasis>
</taxonomicName>
(a, c; BBM­NG 101683, adult female holotype) and 
<taxonomicName id="4C55ECFCF7522F216213F929FEECF945" box="[555,751,1756,1779]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7522F216213F929FEECF945" box="[555,751,1756,1779]" italics="true" pageId="10" pageNumber="11">Hydromys hussoni</emphasis>
</taxonomicName>
(b, d; AM M18627; adult female paratype). Scale bar = 20 mm.
</paragraph>
</caption>
<paragraph id="8BEA977FF7532F206100FEE9FEA6FD28" blockId="11.[264,1323,284,670]" pageId="11" pageNumber="12">
The tail of the 
<typeStatus id="54EE29DDF7532F2061DBFEE9FE4AFE80" box="[483,585,284,310]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
is black, and lacks a white tail­tip, which is present in 19 of 24 known specimens of 
<emphasis id="B9214B6DF7532F206233FEB1FE68FEEB" box="[523,619,324,349]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C55ECFCF7532F206233FEB1FE64FEEB" box="[523,615,324,349]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">hussoni</taxonomicName>
,
</emphasis>
and always present in 
<emphasis id="B9214B6DF7532F2063B2FEB1FD3FFE33" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C55ECFCF7532F2063B2FEB1F8A0FEEB" box="[906,1187,324,349]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="chrysogaster">Hydromys chrysogaster</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7532F20648AFEB1FD3BFE33" class="Mammalia" family="Muridae" genus="Baiyankamys" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Baiyankamys</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C55ECFCF7532F206141FE99FE26FE33" box="[377,549,364,389]" class="Mammalia" family="Muridae" genus="Parahydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7532F206141FE99FE26FE33" box="[377,549,364,389]" italics="true" pageId="11" pageNumber="12">Parahydromys</emphasis>
</taxonomicName>
(absent in 
<taxonomicName id="4C55ECFCF7532F206292FE99FF29FE33" box="[682,810,364,389]" class="Mammalia" family="Muridae" genus="Crossomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7532F206292FE99FF29FE33" box="[682,810,364,389]" italics="true" pageId="11" pageNumber="12">Crossomys</emphasis>
</taxonomicName>
). There are 13–15 tail rings per centimetre in the mid­section of the tail. Probably the most conspicuous external difference between 
<taxonomicName id="4C55ECFCF7532F206130FE49FD63FE63" box="[264,352,444,469]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F206130FE49FD63FE63" box="[264,352,444,469]" italics="true" pageId="11" pageNumber="12">ziegleri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7532F2061A3FE49FDF6FE63" box="[411,501,444,469]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F2061A3FE49FDF6FE63" box="[411,501,444,469]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
is the relative hairiness of the tail. In other 
<taxonomicName id="4C55ECFCF7532F2063C2FE49F872FE63" box="[1018,1137,444,469]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7532F2063C2FE49F872FE63" box="[1018,1137,444,469]" italics="true" pageId="11" pageNumber="12">Hydromys</emphasis>
</taxonomicName>
, including 
<taxonomicName id="4C55ECFCF7532F2064C0FE49FD3AFE4B" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F2064C0FE49FD3AFE4B" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
, the tail is extensively haired, obscuring the tail rings, and the tail hairs are long, extending for the length of many tail scales. In 
<taxonomicName id="4C55ECFCF7532F20630AFDF9FF89FD93" box="[818,906,524,549]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F20630AFDF9FF89FD93" box="[818,906,524,549]" italics="true" pageId="11" pageNumber="12">ziegleri</emphasis>
</taxonomicName>
the tail is considerably less heavily furred, lending it a much more “naked” appearance, such that the tail rings are immediately evident to the naked eye. The tail hairs measure only about 2–3 tail rings in length and lie close to the body of the tail.
</paragraph>
<caption id="DF2AC7F7F7532F206130FA5BFFB8FA5E" httpUri="https://zenodo.org/record/171032/files/figure.png" pageId="11" pageNumber="12" targetBox="[374,1213,729,1428]" targetPageId="11">
<paragraph id="8BEA977FF7532F206130FA5BFFB8FA5E" blockId="11.[264,1322,1454,1513]" pageId="11" pageNumber="12">
<emphasis id="B9214B6DF7532F206130FA5BFD92FA70" bold="true" box="[264,401,1454,1478]" pageId="11" pageNumber="12">FIGURE 4.</emphasis>
Crania and mandibles of 
<taxonomicName id="4C55ECFCF7532F2062FDFA5AFF8CFA70" box="[709,911,1455,1478]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F2062FDFA5AFF8CFA70" box="[709,911,1455,1478]" italics="true" pageId="11" pageNumber="12">Hydromys ziegleri</emphasis>
</taxonomicName>
(below, adult female holotype) and 
<taxonomicName id="4C55ECFCF7532F206130FA27FDCFFA5F" box="[264,460,1490,1513]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F206130FA27FDCFFA5F" box="[264,460,1490,1513]" italics="true" pageId="11" pageNumber="12">Hydromys hussoni</emphasis>
</taxonomicName>
(above, AM M18627, adult female paratype).
</paragraph>
</caption>
<paragraph id="8BEA977FF7532F276100F9E9FDBDFDC2" blockId="11.[264,1324,1564,1868]" lastBlockId="12.[264,1323,284,628]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
In many ways the cranial resemblance between 
<taxonomicName id="4C55ECFCF7532F20634CF9E9FFCEF983" box="[884,973,1564,1589]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F20634CF9E9FFCEF983" box="[884,973,1564,1589]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7532F206432F9E9F861F983" box="[1034,1122,1564,1589]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F206432F9E9F861F983" box="[1034,1122,1564,1589]" italics="true" pageId="11" pageNumber="12">ziegleri</emphasis>
</taxonomicName>
is close (
<bibRefCitation id="EFC4EA8EF7532F2064ECF9E9FD44F9E8" author="Ziegler" pageId="11" pageNumber="20" refString="Ziegler, A. C. (1984) A Papua New Guinea specimen of Hydromys hussoni Musser &amp; Piik, 1982 (Rodentia: Muridae). Australian Mammalogy, 7, 101 - 105." type="journal article" year="1984">Ziegler 1984</bibRefCitation>
). However, judging from direct comparisons between the 
<typeStatus id="54EE29DDF7532F2063C1F9B1F85CF9E8" box="[1017,1119,1604,1630]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C55ECFCF7532F2064B2F9B1F8E1F9EB" box="[1162,1250,1604,1629]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F2064B2F9B1F8E1F9EB" box="[1162,1250,1604,1629]" italics="true" pageId="11" pageNumber="12">ziegleri</emphasis>
</taxonomicName>
and a 
<typeStatus id="54EE29DDF7532F206130F999FD6EF930" box="[264,365,1644,1670]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
<emphasis id="B9214B6DF7532F20614EF999FDECF933" box="[374,495,1644,1669]" italics="true" pageId="11" pageNumber="12">
of 
<taxonomicName id="4C55ECFCF7532F2061AEF999FDECF933" box="[406,495,1644,1669]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">hussoni</taxonomicName>
</emphasis>
(AM M18627, formerly RMNH 29162, also adult female), my notes on the 
<typeStatus id="54EE29DDF7532F20616EF966FD8AF91B" box="[342,393,1683,1709]" pageId="11" pageNumber="12">type</typeStatus>
series of 
<taxonomicName id="4C55ECFCF7532F2061C0F961FE52F91B" box="[504,593,1684,1709]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F2061C0F961FE52F91B" box="[504,593,1684,1709]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
at RMNH, comparisons with the verbal and mensural descriptions of 
<taxonomicName id="4C55ECFCF7532F206151F94EFDC0F962" box="[361,451,1723,1748]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F206151F94EFDC0F962" box="[361,451,1723,1748]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFC4EA8EF7532F2061C8F94EFF05F963" author="Musser" box="[496,774,1723,1749]" pageId="11" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik (1982)</bibRefCitation>
and Voss (1988), and illustrations of the 
<typeStatus id="54EE29DDF7532F2064D6F94EFD39F94B" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C55ECFCF7532F20615FF916FDC2F94A" box="[359,449,1763,1788]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F20615FF916FDC2F94A" box="[359,449,1763,1788]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
figured by 
<bibRefCitation id="EFC4EA8EF7532F206268F916FF73F94B" author="Musser" box="[592,880,1763,1789]" pageId="11" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik (1982)</bibRefCitation>
, many qualitative cranial differences between the two species are immediately apparent. The skull of 
<taxonomicName id="4C55ECFCF7532F206414F8FEF887F892" box="[1068,1156,1803,1828]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7532F206414F8FEF887F892" box="[1068,1156,1803,1828]" italics="true" pageId="11" pageNumber="12">ziegleri</emphasis>
</taxonomicName>
is noticeably shorter and stouter than that of 
<taxonomicName id="4C55ECFCF7532F2062BEF8C7FEE3F8FD" box="[646,736,1842,1867]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7532F2062BEF8C7FEE3F8FD" box="[646,736,1842,1867]" italics="true" pageId="11" pageNumber="12">hussoni</emphasis>
</taxonomicName>
(
<figureCitation id="136E8BFAF7532F2062CBF8C7FF3DF8FA" box="[755,830,1842,1868]" captionStart="FIGURE 4" captionStartId="11.[264,368,1454,1478]" captionTargetBox="[374,1213,729,1428]" captionTargetId="figure@11.[374,1214,729,1428]" captionTargetPageId="11" captionText="FIGURE 4. Crania and mandibles of Hydromys ziegleri (below, adult female holotype) and Hydromys hussoni (above, AM M 18627, adult female paratype)." httpUri="https://zenodo.org/record/171032/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
). The rostrum is short and broad in both species, but the nasolacrimal capsules are laterally expanded to a much greater degree in 
<taxonomicName id="4C55ECFCF7542F276130FEB1FD63FEEB" box="[264,352,324,349]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7542F276130FEB1FD63FEEB" box="[264,352,324,349]" italics="true" pageId="12" pageNumber="13">ziegleri</emphasis>
</taxonomicName>
, as reflected in its relatively greater rostral breadth, and the nasals are correspondingly broader (
<tableCitation id="C6D7A2C4F7542F27618FFE9EFE11FE33" box="[439,530,363,389]" captionStart="TABLE 3" captionStartId="12.[264,352,676,700]" captionText="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." pageId="12" pageNumber="13">Table 3</tableCitation>
) and also more strongly retracted. The incisive foramina are more truncate in 
<taxonomicName id="4C55ECFCF7542F2761A9FE66FDEAFE1A" box="[401,489,403,428]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7542F2761A9FE66FDEAFE1A" box="[401,489,403,428]" italics="true" pageId="12" pageNumber="13">ziegleri</emphasis>
</taxonomicName>
, only 31% of the length of the diastema, versus 37% (in 
<taxonomicName id="4C55ECFCF7542F27649DFE66F8FDFE1A" box="[1189,1278,403,428]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7542F27649DFE66F8FDFE1A" box="[1189,1278,403,428]" italics="true" pageId="12" pageNumber="13">hussoni</emphasis>
</taxonomicName>
) or greater in all other species of 
<taxonomicName id="4C55ECFCF7542F276248FE4EFEE4FE62" box="[624,743,443,468]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7542F276248FE4EFEE4FE62" box="[624,743,443,468]" italics="true" pageId="12" pageNumber="13">Hydromys</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7542F27631DFE4EF925FE63" authority="Musser and Piik 1982: 164" authorityName="Musser and Piik" authorityPageNumber="164" authorityYear="1982" box="[805,1318,443,469]" class="Mammalia" family="Muridae" genus="Baiyankamys" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7542F27631DFE4EFFC2FE62" box="[805,961,443,468]" italics="true" pageId="12" pageNumber="13">Baiyankamys</emphasis>
(
<bibRefCitation id="EFC4EA8EF7542F2763EBFE4EF91EFE63" author="Musser" box="[979,1309,443,469]" pageId="12" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik 1982: 164</bibRefCitation>
)
</taxonomicName>
. The zygomatic plate is more excavated in 
<taxonomicName id="4C55ECFCF7542F2762CFFE16FF4CFE4A" box="[759,847,483,508]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7542F2762CFFE16FF4CFE4A" box="[759,847,483,508]" italics="true" pageId="12" pageNumber="13">ziegleri</emphasis>
</taxonomicName>
than 
<taxonomicName id="4C55ECFCF7542F2763B7FE16FFEAFE4A" box="[911,1001,483,508]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7542F2763B7FE16FFEAFE4A" box="[911,1001,483,508]" italics="true" pageId="12" pageNumber="13">hussoni</emphasis>
</taxonomicName>
, and the braincase is markedly less expanded dorso­ventrally (
<tableCitation id="C6D7A2C4F7542F276288FDFFFF0FFD92" box="[688,780,522,548]" captionStart="TABLE 3" captionStartId="12.[264,352,676,700]" captionText="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." pageId="12" pageNumber="13">Table 3</tableCitation>
; braincase height is 64% of braincase breadth in 
<taxonomicName id="4C55ECFCF7542F276112FDC7FD81FDFD" box="[298,386,562,587]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7542F276112FDC7FD81FDFD" box="[298,386,562,587]" italics="true" pageId="12" pageNumber="13">ziegleri</emphasis>
</taxonomicName>
, versus 69% or greater in all other 
<taxonomicName id="4C55ECFCF7542F276301FDC7FFB2FDFD" box="[825,945,562,587]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7542F276301FDC7FFB2FDFD" box="[825,945,562,587]" italics="true" pageId="12" pageNumber="13">Hydromys</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7542F2763C8FDC7F88FFDFD" box="[1008,1164,562,587]" class="Mammalia" family="Muridae" genus="Baiyankamys" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7542F2763C8FDC7F88FFDFD" box="[1008,1164,562,587]" italics="true" pageId="12" pageNumber="13">Baiyankamys</emphasis>
</taxonomicName>
; 
<bibRefCitation id="EFC4EA8EF7542F2764A7FDC7FD7DFDC2" author="Musser" pageId="12" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik 1982</bibRefCitation>
:164).
</paragraph>
<caption id="DF2AC7F7F7542F276130FD51FF2DFD52" pageId="12" pageNumber="13">
<paragraph id="8BEA977FF7542F276130FD51FF2DFD52" blockId="12.[264,1322,676,740]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276130FD51FD7EFD0A" bold="true" box="[264,381,676,700]" pageId="12" pageNumber="13">TABLE 3.</emphasis>
Craniodental metrics for the adult female holotype of 
<taxonomicName id="4C55ECFCF7542F2763F4FD51F892FD0D" box="[972,1169,676,699]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7542F2763F4FD51F892FD0D" box="[972,1169,676,699]" italics="true" pageId="12" pageNumber="13">Hydromys ziegleri</emphasis>
</taxonomicName>
and for series of adult female and mixed adult 
<taxonomicName id="4C55ECFCF7542F27625BFD3EFF24FD54" box="[611,807,715,738]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7542F27625BFD3EFF24FD54" box="[611,807,715,738]" italics="true" pageId="12" pageNumber="13">Hydromys hussoni</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BEA977FF7542F27616EFCF9F910FCA8" blockId="12.[283,1308,779,1573]" box="[342,1299,779,810]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F27616EFCF9FFA3FC9C" bold="true" box="[342,928,780,810]" pageId="12" pageNumber="13">
Measurements 
<emphasis id="B9214B6DF7542F27625BFCF8FFA3FC9C" bold="true" box="[611,928,781,810]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C55ECFCF7542F27625BFCF8FED7FC93" box="[611,724,781,805]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="ziegleri">H. ziegleri</taxonomicName>
<taxonomicName id="4C55ECFCF7542F276310FCE7FFA3FC9C" box="[808,928,786,810]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hussoni">H. hussoni</taxonomicName>
</emphasis>
</emphasis>
ΨΨ 
<emphasis id="B9214B6DF7542F2763E9FCFEFFDFFCA8" bold="true" box="[977,988,779,798]" pageId="12" pageNumber="13">1</emphasis>
<taxonomicName id="4C55ECFCF7542F276416FCE7F8A5FC9C" box="[1070,1190,786,810]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7542F276416FCE7F8A5FC9C" bold="true" box="[1070,1190,786,810]" italics="true" pageId="12" pageNumber="13">H. hussoni</emphasis>
</taxonomicName>
ΨΨ, ♂♂ 
<emphasis id="B9214B6DF7542F276530FCFEF910FCA8" bold="true" box="[1288,1299,779,798]" pageId="12" pageNumber="13">2</emphasis>
</paragraph>
<paragraph id="8BEA977FF7542F276363FCB0F8C0FCEA" blockId="12.[283,1308,779,1573]" box="[859,1219,837,860]" pageId="12" pageNumber="13">n = 6­9 n = 20</paragraph>
<paragraph id="8BEA977FF7542F276123FC8DF91FFC39" blockId="12.[283,1308,779,1573]" box="[283,1308,888,912]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276123FC8DFE34FC26" bold="true" box="[283,567,888,912]" pageId="12" pageNumber="13">Length of head and body</emphasis>
132 150.1 ± 9.4 (138­164) 147.9 ± 15.4 (122­171)
</paragraph>
<paragraph id="8BEA977FF7542F276164FC5EF91FFC74" blockId="12.[283,1308,779,1573]" box="[348,1308,939,963]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276164FC5EFDF4FC75" bold="true" box="[348,503,939,963]" pageId="12" pageNumber="13">Length of tail</emphasis>
118 125.5 ± 14.3 (103­144) 126.4 ± 13.8 (103­152)
</paragraph>
<paragraph id="8BEA977FF7542F276107FC2BF901FC43" blockId="12.[283,1308,779,1573]" box="[319,1282,990,1014]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276107FC2BFE17FC40" bold="true" box="[319,532,990,1014]" pageId="12" pageNumber="13">Length of hindfoot</emphasis>
29 27.4 ± 1.1 (26­29) 29.8 ± 2.0 (27­33)
</paragraph>
<paragraph id="8BEA977FF7542F276164FBE4F902FB9E" blockId="12.[283,1308,779,1573]" box="[348,1281,1041,1065]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276164FBE4FDF5FB9F" bold="true" box="[348,502,1041,1065]" pageId="12" pageNumber="13">Length of ear</emphasis>
13 12.4 ± 0.8 (12­14) 12.1 ± 0.8 (11­14)
</paragraph>
<paragraph id="8BEA977FF7542F27611DFBB6F8ADFBE3" blockId="12.[283,1308,779,1573]" box="[293,1198,1086,1115]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F27611DFBB6FE2DFBED" bold="true" box="[293,558,1091,1115]" pageId="12" pageNumber="13">Greatest length of skull</emphasis>
29.01 31.54 ± 0.87 (30.1­32.8) —
</paragraph>
<paragraph id="8BEA977FF7542F276110FB83F918FB3B" blockId="12.[283,1308,779,1573]" box="[296,1307,1137,1166]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276110FB83FE29FB38" bold="true" box="[296,554,1142,1166]" pageId="12" pageNumber="13">Condyloincisive length</emphasis>
28.50 — 30.5 ± 1.41 (27.9­32.9)
</paragraph>
<paragraph id="8BEA977FF7542F276104FB5CF91FFB76" blockId="12.[283,1308,779,1573]" box="[316,1308,1193,1217]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276104FB5CFE14FB77" bold="true" box="[316,535,1193,1217]" pageId="12" pageNumber="13">Zygomatic breadth</emphasis>
14.98 15.60 ± 0.50 (14.8­16.3) 15.5 ± 0.87 (14.0­17.3)
</paragraph>
<paragraph id="8BEA977FF7542F276103FB29F8ADFB5B" blockId="12.[283,1308,779,1573]" box="[315,1198,1238,1268]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276103FB29FE1BFB42" bold="true" box="[315,536,1244,1268]" pageId="12" pageNumber="13">Breadth of rostrum</emphasis>
6.23 5.82 ± 0.21 (5.6 ­ 6.1) —
</paragraph>
<paragraph id="8BEA977FF7542F276174FAFAF90CFA90" blockId="12.[283,1308,779,1573]" box="[332,1295,1295,1319]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276174FAFAFE05FA91" bold="true" box="[332,518,1295,1319]" pageId="12" pageNumber="13">Length of nasals</emphasis>
8.73 10.17 ± 0.43 (9.4­10.6) 9.8 ± 0.68 (8.6­11.2)
</paragraph>
<paragraph id="8BEA977FF7542F27617EFAB7F913FAEF" blockId="12.[283,1308,779,1573]" box="[326,1296,1340,1370]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F27617EFAB7FE0EFAEC" bold="true" box="[326,525,1346,1370]" pageId="12" pageNumber="13">Breadth of nasals</emphasis>
3.39 — 3.10 ± 0.18 (2.8­3.4)
</paragraph>
<paragraph id="8BEA977FF7542F276123FA81F90AFA3A" blockId="12.[283,1308,779,1573]" box="[283,1289,1396,1420]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276123FA81FE34FA3A" bold="true" box="[283,567,1396,1420]" pageId="12" pageNumber="13">Length incisive foramina</emphasis>
2.48 3.08 ± 0.13 (2.9­3.3) 3.0 ± 0.33 (2.0­3.3)
</paragraph>
<paragraph id="8BEA977FF7542F276102FA52F90AFA09" blockId="12.[283,1308,779,1573]" box="[314,1289,1447,1471]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276102FA52FE1AFA09" bold="true" box="[314,537,1447,1471]" pageId="12" pageNumber="13">Maxillary toothrow</emphasis>
4.13 4.71 ± 0.15 (4.5­5.0) 4.6 ± 0.12 (4.4­4.8)
</paragraph>
<paragraph id="8BEA977FF7542F27610BFA2FF91FFA47" blockId="12.[283,1308,779,1573]" box="[307,1308,1498,1522]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F27610BFA2FFE23FA44" bold="true" box="[307,544,1498,1522]" pageId="12" pageNumber="13">Breadth of braincase</emphasis>
13.09 13.44 ± 0.28 (13.1­13.8) 13.7 ± 0.36 (13.0­14.4)
</paragraph>
<paragraph id="8BEA977FF7542F276103F9F8F8ADF9A8" blockId="12.[283,1308,779,1573]" box="[315,1198,1543,1573]" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276103F9F8FE1AF993" bold="true" box="[315,537,1549,1573]" pageId="12" pageNumber="13">Height of braincase</emphasis>
8.39 9.21 ± 0.19 (8.9­9.5) —
</paragraph>
<paragraph id="8BEA977FF7542F27611CF9B5F90AF90B" pageId="12" pageNumber="13">
<table id="F95565DFF754D0D46123F9B5F91FF90B" box="[283,1308,1600,1725]" gridcols="4" gridrows="3" pageId="12" pageNumber="13">
<tr id="3565953DF754D0D46123F9B5F91FF9EE" box="[283,1308,1600,1624]" gridrow="0" pageId="12" pageNumber="13">
<th id="76B4FC41F754D0D46123F9B5FE2CF9EE" box="[283,559,1600,1624]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F27611CF9B5FE2CF9EE" bold="true" box="[292,559,1600,1624]" pageId="12" pageNumber="13">Postorbital constriction</emphasis>
</th>
<th id="76B4FC41F754D0D462BCF9B5FEB0F9EE" box="[644,691,1600,1624]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">5.09</th>
<th id="76B4FC41F754D0D4632BF9B5FFF3F9EE" box="[787,1008,1600,1624]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">5.28 ± 0.14 (5.1­5.5)</th>
<th id="76B4FC41F754D0D46401F9B5F91FF9EE" box="[1081,1308,1600,1624]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">5.3 ± 0.14 (5.0­5.6)</th>
</tr>
<tr id="3565953DF754D0D46123F986F91FF93D" box="[283,1308,1651,1675]" gridrow="1" pageId="12" pageNumber="13">
<th id="76B4FC41F754D0D46123F986FE2CF93D" box="[283,559,1651,1675]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276113F986FE2BF93D" bold="true" box="[299,552,1651,1675]" pageId="12" pageNumber="13">Breadth of incisor tips</emphasis>
</th>
<td id="76B4FC41F754D0D462BCF986FEB0F93D" box="[644,691,1651,1675]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">1.62</td>
<td id="76B4FC41F754D0D4632BF986FFF3F93D" box="[787,1008,1651,1675]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">1.67 ± 0.13 (1.5­1.9)</td>
<td id="76B4FC41F754D0D46401F986F91FF93D" box="[1081,1308,1651,1675]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">1.6 ± 0.15 (1.4­1.9)</td>
</tr>
<tr id="3565953DF754D0D46123F950F91FF90B" box="[283,1308,1701,1725]" gridrow="2" pageId="12" pageNumber="13">
<th id="76B4FC41F754D0D46123F950FE2CF90B" box="[283,559,1701,1725]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">
<emphasis id="B9214B6DF7542F276104F950FE14F90B" bold="true" box="[316,535,1701,1725]" pageId="12" pageNumber="13">Length of diastema</emphasis>
</th>
<td id="76B4FC41F754D0D462BCF950FEB0F90B" box="[644,691,1701,1725]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">7.99</td>
<td id="76B4FC41F754D0D4632BF950FFF3F90B" box="[787,1008,1701,1725]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">8.39 ± 0.36 (7.9­9.0)</td>
<td id="76B4FC41F754D0D46401F950F91FF90B" box="[1081,1308,1701,1725]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">8.5 ± 0.54 (7.6­9.3)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEA977FF7542F276130F928FE91F94D" blockId="12.[264,710,1757,1829]" box="[264,658,1757,1787]" pageId="12" pageNumber="13">
1 From 
<bibRefCitation id="EFC4EA8EF7542F276162F911FE8DF94D" author="Musser" box="[346,654,1764,1787]" pageId="12" pageNumber="20" refString="Musser, G. G. &amp; Piik, E. (1982) A new species of Hydromys (Muridae) from western New Guinea (Irian Jaya). Zoologische Mededelingen, 56, 153 - 166." type="journal article" year="1982">Musser and Piik (1982: 159)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEA977FF7542F276130F8F2FEC5F893" blockId="12.[264,710,1757,1829]" box="[264,710,1799,1829]" pageId="12" pageNumber="13">2 From Voss (1988: 467): 12ΨΨ and 8♂♂.</paragraph>
<paragraph id="8BEA977FF7552F266100FEE9FE69FC38" blockId="13.[264,1324,284,1156]" pageId="13" pageNumber="14">
The faces of the upper and lower incisors are pigmented yellowish­orange in 
<taxonomicName id="4C55ECFCF7552F2664EBFEE9F928FE83" box="[1235,1323,284,309]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F2664EBFEE9F928FE83" box="[1235,1323,284,309]" italics="true" pageId="13" pageNumber="14">ziegleri</emphasis>
</taxonomicName>
(with pigmentation more obvious in the upper incisors), while those of 
<taxonomicName id="4C55ECFCF7552F26645DFEB1F8BDFEEB" box="[1125,1214,324,349]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F26645DFEB1F8BDFEEB" box="[1125,1214,324,349]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
are very weakly pigmented or almost unpigmented. In 
<taxonomicName id="4C55ECFCF7552F26630CFE99FF8FFE33" box="[820,908,364,389]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F26630CFE99FF8FFE33" box="[820,908,364,389]" italics="true" pageId="13" pageNumber="14">ziegleri</emphasis>
</taxonomicName>
the first upper molars are broader relative to their length compared to 
<taxonomicName id="4C55ECFCF7552F266282FE61FF10FE1B" box="[698,787,404,429]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F266282FE61FF10FE1B" box="[698,787,404,429]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
, and the second upper and lower molars are more reduced (e.g. 
<tableCitation id="C6D7A2C4F7552F2661CAFE49FE4CFE60" box="[498,591,444,470]" captionStart="TABLE 4" captionStartId="13.[264,352,1209,1233]" captionTargetBox="[264,1226,1311,1770]" captionTargetPageId="13" captionText="TABLE 4. Comparative molar dimensions in specimens of Hydromys ziegleri (adult female holotype) and Hydromys hussoni (AM M 18627, adult female paratype)." httpUri="http://table.plazi.org/id/DF2AC7F7F7552F266130FB4CFFD4FB4F" pageId="13" pageNumber="14" tableUuid="DF2AC7F7F7552F266130FB4CFFD4FB4F">Table 4</tableCitation>
). The posterior palatine foramina are lengthened into a more extensive narrow groove in the 
<taxonomicName id="4C55ECFCF7552F266244FE11FED5FE4B" box="[636,726,484,509]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F266244FE11FED5FE4B" box="[636,726,484,509]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
<typeStatus id="54EE29DDF7552F2662E6FE11FF40FE48" box="[734,835,484,510]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
than in the 
<typeStatus id="54EE29DDF7552F2663F7FE11F835FE48" box="[975,1078,484,510]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C55ECFCF7552F266465FE11F8B6FE4B" box="[1117,1205,484,509]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F266465FE11F8B6FE4B" box="[1117,1205,484,509]" italics="true" pageId="13" pageNumber="14">ziegleri</emphasis>
</taxonomicName>
, in which these foramina are essentially small ovate holes. The posterior palatal spine is extremely well­developed. Relative to 
<taxonomicName id="4C55ECFCF7552F266269FDC1FEA8FDFB" box="[593,683,564,589]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F266269FDC1FEA8FDFB" box="[593,683,564,589]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
the auditory bullae are less flattened and less elongate. The foramen ovale is distinctly larger in 
<taxonomicName id="4C55ECFCF7552F2662DAFDA9FF39FDC3" box="[738,826,604,629]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F2662DAFDA9FF39FDC3" box="[738,826,604,629]" italics="true" pageId="13" pageNumber="14">ziegleri</emphasis>
</taxonomicName>
than 
<taxonomicName id="4C55ECFCF7552F266342FDA9FFD7FDC3" box="[890,980,604,629]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F266342FDA9FFD7FDC3" box="[890,980,604,629]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
. In both species the stapedial foramen is well­developed and the foramen ovale bears a conspicuous groove for the passage of the infraorbital branch of the stapedial artery, suggesting that both possess the primitive murine cephalic arterial configuration typical of other New Guinean water­rats of the genera 
<emphasis id="B9214B6DF7552F266187FD09FFB1FCA3" box="[447,946,764,789]" italics="true" pageId="13" pageNumber="14">
<taxonomicName id="4C55ECFCF7552F266187FD09FE5DFCA3" box="[447,606,764,789]" class="Mammalia" family="Muridae" genus="Baiyankamys" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">Baiyankamys</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7552F26624AFD09FEEFFCA3" box="[626,748,764,789]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">Hydromys</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7552F266338FD09FFACFCA3" box="[768,943,764,789]" class="Mammalia" family="Muridae" genus="Parahydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">Parahydromys</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C55ECFCF7552F2663C2FD09F879FCA3" box="[1018,1146,764,789]" class="Mammalia" family="Muridae" genus="Crossomys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7552F2663C2FD09F879FCA3" box="[1018,1146,764,789]" italics="true" pageId="13" pageNumber="14">Crossomys</emphasis>
</taxonomicName>
(and the less closely­related lowland false water­rat, 
<taxonomicName id="4C55ECFCF7552F2662E5FCD1FFAFFC8B" box="[733,940,804,829]" class="Mammalia" family="Muridae" genus="Xeromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="myoides">
<emphasis id="B9214B6DF7552F2662E5FCD1FFAFFC8B" box="[733,940,804,829]" italics="true" pageId="13" pageNumber="14">Xeromys myoides</emphasis>
</taxonomicName>
, as stated by Musser and Carleton [2005] and confirmed by my examination of specimens in the Australian Museum; cf. 
<bibRefCitation id="EFC4EA8EF7552F266130FC81FE3AFC38" author="Musser" box="[264,569,884,910]" pageId="13" pageNumber="20" refString="Musser, G. G. &amp; Heaney, L. R. (1992) Philippine rodents: definitions of Tarsomys and Limnomys plus a preliminary assessment of phylogenetic patterns among native Philippine murines (Murinae, Muridae). Bulletin of the American Museum of Natural History, 211, 1 - 138." type="journal article" year="1992">Musser and Heaney 1992</bibRefCitation>
:87).
</paragraph>
<paragraph id="8BEA977FF7552F266100FC69F825FB32" blockId="13.[264,1324,284,1156]" pageId="13" pageNumber="14">
Additional measurements of the 
<typeStatus id="54EE29DDF7552F2662F7FC69FF35FC00" box="[719,822,924,950]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(other than those presented in 
<tableCitation id="C6D7A2C4F7552F2664FCFC69F921FC00" box="[1220,1314,924,950]" captionStart="TABLE 3" captionStartId="12.[264,352,676,700]" captionText="TABLE 3. Craniodental metrics for the adult female holotype of Hydromys ziegleri and for series of adult female and mixed adult Hydromys hussoni." pageId="13" pageNumber="14">Table 3</tableCitation>
) include: palatal length 13.08; breadth of mesopterygoid fossa 2.13; greatest breadth of incisive foramina 1.67; breadth of the palatal bridge between M2–M2 2.60; breadth of the zygomatic plate 1.54; height of upper incisor 4.58; depth of upper incisor 1.55; breadth across occipital condyles 8.37; coronoid height of mandible 7.99; alveolar length of M1–2 4.35; and mandible length minus incisor 14.85 (with incisor, 18.38).
</paragraph>
<caption id="DF2AC7F7F7552F266130FB4CFFD4FB4F" ID-Table-UUID="DF2AC7F7F7552F266130FB4CFFD4FB4F" httpUri="http://table.plazi.org/id/DF2AC7F7F7552F266130FB4CFFD4FB4F" pageId="13" pageNumber="14" targetBox="[264,1226,1311,1770]" targetIsTable="true" targetPageId="13">
<paragraph id="8BEA977FF7552F266130FB4CFFD4FB4F" blockId="13.[264,1322,1209,1273]" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F266130FB4CFD7FFB67" bold="true" box="[264,380,1209,1233]" pageId="13" pageNumber="14">TABLE 4.</emphasis>
Comparative molar dimensions in specimens of 
<taxonomicName id="4C55ECFCF7552F2663B7FB4FF850FB67" box="[911,1107,1210,1233]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F2663B7FB4FF850FB67" box="[911,1107,1210,1233]" italics="true" pageId="13" pageNumber="14">Hydromys ziegleri</emphasis>
</taxonomicName>
(adult female holotype) and 
<taxonomicName id="4C55ECFCF7552F26614BFB14FE34FB4E" box="[371,567,1249,1272]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F26614BFB14FE34FB4E" box="[371,567,1249,1272]" italics="true" pageId="13" pageNumber="14">Hydromys hussoni</emphasis>
</taxonomicName>
(AM M18627, adult female paratype).
</paragraph>
</caption>
<paragraph id="8BEA977FF7552F266314FAEAFDBFF95C" pageId="13" pageNumber="14">
<table id="F95565DFF755D0D46130FAEAF8C9F95C" box="[264,1226,1311,1770]" gridcols="3" gridrows="8" pageId="13" pageNumber="14">
<tr id="3565953DF755D0D46130FAEAF8C9FA81" box="[264,1226,1311,1335]" gridrow="0" pageId="13" pageNumber="14" rowspan-0="1">
<th id="76B4FC41F755D0D46320FAEAFF93FA81" box="[792,912,1311,1335]" gridcol="1" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="4C55ECFCF7552F266314FAEAFF78FA81" box="[812,891,1311,1335]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7552F266314FAEAFF78FA81" bold="true" box="[812,891,1311,1335]" italics="true" pageId="13" pageNumber="14">ziegleri</emphasis>
</taxonomicName>
</th>
<th id="76B4FC41F755D0D4646BFAEAF8C9FA81" box="[1107,1226,1311,1335]" gridcol="2" gridrow="0" pageId="13" pageNumber="14">
<taxonomicName id="4C55ECFCF7552F26645CFAEAF8B9FA81" box="[1124,1210,1311,1335]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7552F26645CFAEAF8B9FA81" bold="true" box="[1124,1210,1311,1335]" italics="true" pageId="13" pageNumber="14">hussoni</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3565953DF755D0D46130FABBF8C9FADB" box="[264,1226,1358,1389]" gridrow="1" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130FABBFE58FADB" box="[264,603,1358,1389]" gridcol="0" gridrow="1" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F266141FAA0FD91FADB" bold="true" box="[377,402,1365,1389]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F2661AAFABBFD9EFAD7" bold="true" box="[402,413,1358,1377]" pageId="13" pageNumber="14">1</emphasis>
<emphasis id="B9214B6DF7552F26619CFAA0FE45FADB" bold="true" box="[420,582,1365,1389]" pageId="13" pageNumber="14">length x width</emphasis>
</th>
<td id="76B4FC41F755D0D46320FABBFF93FADB" box="[792,912,1358,1389]" gridcol="1" gridrow="1" pageId="13" pageNumber="14">2.98 x 1.54</td>
<td id="76B4FC41F755D0D4646BFABBF8C9FADB" box="[1107,1226,1358,1389]" gridcol="2" gridrow="1" pageId="13" pageNumber="14">3.24 x 1.49</td>
</tr>
<tr id="3565953DF755D0D46130FA74F8C9FA16" box="[264,1226,1409,1440]" gridrow="2" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130FA74FE58FA16" box="[264,603,1409,1440]" gridcol="0" gridrow="2" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F266141FA7DFD91FA16" bold="true" box="[377,402,1416,1440]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F2661AAFA74FD9EFA22" bold="true" box="[402,413,1409,1428]" pageId="13" pageNumber="14">2</emphasis>
<emphasis id="B9214B6DF7552F26619CFA7DFE45FA16" bold="true" box="[420,582,1416,1440]" pageId="13" pageNumber="14">length x width</emphasis>
</th>
<td id="76B4FC41F755D0D46320FA74FF93FA16" box="[792,912,1409,1440]" gridcol="1" gridrow="2" pageId="13" pageNumber="14">1.22 x 1.07</td>
<td id="76B4FC41F755D0D4646BFA74F8C9FA16" box="[1107,1226,1409,1440]" gridcol="2" gridrow="2" pageId="13" pageNumber="14">1.52 x 1.34</td>
</tr>
<tr id="3565953DF755D0D46130FA40F8C9FA62" box="[264,1226,1461,1492]" gridrow="3" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130FA40FE58FA62" box="[264,603,1461,1492]" gridcol="0" gridrow="3" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F26615CFA49FD7EFA62" bold="true" box="[356,381,1468,1492]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F266145FA40FD8BFA7E" bold="true" box="[381,392,1461,1480]" pageId="13" pageNumber="14">2</emphasis>
<emphasis id="B9214B6DF7552F2661B6FA49FE00FA62" bold="true" box="[398,515,1468,1492]" pageId="13" pageNumber="14">length / M</emphasis>
<emphasis id="B9214B6DF7552F26623BFA40FE0DFA7E" bold="true" box="[515,526,1461,1480]" pageId="13" pageNumber="14">1</emphasis>
<emphasis id="B9214B6DF7552F26622CFA49FE58FA62" bold="true" box="[532,603,1468,1492]" pageId="13" pageNumber="14">length</emphasis>
</th>
<td id="76B4FC41F755D0D46320FA40FF93FA62" box="[792,912,1461,1492]" gridcol="1" gridrow="3" pageId="13" pageNumber="14">0.41</td>
<td id="76B4FC41F755D0D4646BFA40F8C9FA62" box="[1107,1226,1461,1492]" gridcol="2" gridrow="3" pageId="13" pageNumber="14">0.47</td>
</tr>
<tr id="3565953DF755D0D46130FA1DF8C9F9B1" box="[264,1226,1512,1543]" gridrow="4" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130FA1DFE58F9B1" box="[264,603,1512,1543]" gridcol="0" gridrow="4" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F26615FFA1AFD83F9B1" bold="true" box="[359,384,1519,1543]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F2661B8FA1DFD88FA4D" bold="true" box="[384,395,1512,1531]" pageId="13" pageNumber="14">1</emphasis>
<emphasis id="B9214B6DF7552F2661A9FA1AFE03F9B1" bold="true" box="[401,512,1519,1543]" pageId="13" pageNumber="14">width / M</emphasis>
<emphasis id="B9214B6DF7552F266238FA1DFE08FA4D" bold="true" box="[512,523,1512,1531]" pageId="13" pageNumber="14">1</emphasis>
<emphasis id="B9214B6DF7552F26622AFA1AFE5AF9B1" bold="true" box="[530,601,1519,1543]" pageId="13" pageNumber="14">length</emphasis>
</th>
<td id="76B4FC41F755D0D46320FA1DFF93F9B1" box="[792,912,1512,1543]" gridcol="1" gridrow="4" pageId="13" pageNumber="14">0.52</td>
<td id="76B4FC41F755D0D4646BFA1DF8C9F9B1" box="[1107,1226,1512,1543]" gridcol="2" gridrow="4" pageId="13" pageNumber="14">0.46</td>
</tr>
<tr id="3565953DF755D0D46130F9E8F8C9F98C" box="[264,1226,1565,1594]" gridrow="5" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130F9E8FE58F98C" box="[264,603,1565,1594]" gridcol="0" gridrow="5" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F266141F9E8FD91F983" bold="true" box="[377,402,1565,1589]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F2661AAF9D2FD9EF98C" bold="true" box="[402,413,1575,1594]" pageId="13" pageNumber="14">1</emphasis>
<emphasis id="B9214B6DF7552F26619CF9E8FE45F983" bold="true" box="[420,582,1565,1589]" pageId="13" pageNumber="14">length x width</emphasis>
</th>
<td id="76B4FC41F755D0D46320F9E8FF93F98C" box="[792,912,1565,1594]" gridcol="1" gridrow="5" pageId="13" pageNumber="14">2.59 x 1.28</td>
<td id="76B4FC41F755D0D4646BF9E8F8C9F98C" box="[1107,1226,1565,1594]" gridcol="2" gridrow="5" pageId="13" pageNumber="14">2.84 x 1.59</td>
</tr>
<tr id="3565953DF755D0D46130F9A5F8C9F9DB" box="[264,1226,1616,1645]" gridrow="6" pageId="13" pageNumber="14">
<th id="76B4FC41F755D0D46130F9A5FE58F9DB" box="[264,603,1616,1645]" gridcol="0" gridrow="6" pageId="13" pageNumber="14">
<emphasis id="B9214B6DF7552F266141F9A5FD91F9DE" bold="true" box="[377,402,1616,1640]" pageId="13" pageNumber="14">M</emphasis>
<emphasis id="B9214B6DF7552F2661AAF9AFFD9EF9DB" bold="true" box="[402,413,1626,1645]" pageId="13" pageNumber="14">2</emphasis>
<emphasis id="B9214B6DF7552F26619CF9A5FE45F9DE" bold="true" box="[420,582,1616,1640]" pageId="13" pageNumber="14">length x width</emphasis>
</th>
<td id="76B4FC41F755D0D46320F9A5FF93F9DB" box="[792,912,1616,1645]" gridcol="1" gridrow="6" pageId="13" pageNumber="14">1.67 x 1.18</td>
<td id="76B4FC41F755D0D4646BF9A5F8C9F9DB" box="[1107,1226,1616,1645]" gridcol="2" gridrow="6" pageId="13" pageNumber="14">1.81 x 1.34</td>
</tr>
<tr id="3565953DF755D0D46130F925F8C9F95C" box="[264,1226,1744,1770]" gridrow="7" pageId="13" pageNumber="14" rowspan-1="1" rowspan-2="1">
<th id="76B4FC41F755D0D46130F925FE58F95C" box="[264,603,1744,1770]" gridcol="0" gridrow="7" pageId="13" pageNumber="14">Zoogeography</th>
</tr>
</table>
</paragraph>
<paragraph id="8BEA977FF7552F256130F8D5FF17FE18" blockId="13.[264,1322,1824,1890]" lastBlockId="14.[264,1324,284,430]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Several isolated mountain ranges are situated along the northern coast of New 
<collectingCountry id="F342D7EFF7552F2664F4F8D5F927F88C" box="[1228,1316,1824,1850]" name="Guinea-Bissau" pageId="13" pageNumber="14">Guinea</collectingCountry>
, including the Adelbert, Prince Alexander, Torricelli, Bewani, Cyclops, Foya (= Gauttier), and Van Rees Mountains, collectively referred to hereafter as the North Coast Ranges. These mountain ranges are well­separated from the Central Dividing Ranges of New 
<collectingCountry id="F342D7EFF7562F256130FE99FD5EFE30" box="[264,349,364,390]" name="Guinea-Bissau" pageId="14" pageNumber="15">Guinea</collectingCountry>
by large expanses of lowland forest, have distinctive geological histories (
<bibRefCitation id="EFC4EA8EF7562F2564F8FE99FD45FE18" author="Flannery" pageId="14" pageNumber="19" refString="Flannery, T. F. (1995) Mammals of New Guinea, revised edition, Reed Books: Ithaca, New York, 568 pp." type="book" year="1995">Flannery 1995</bibRefCitation>
), and support highly distinctive faunas.
</paragraph>
<caption id="DF2AC7F7F7562F256130FDFEFFAFFDDE" pageId="14" pageNumber="15">
<paragraph id="8BEA977FF7562F256130FDFEFFAFFDDE" blockId="14.[264,1322,523,616]" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256130FDFEFD78FD95" bold="true" box="[264,379,523,547]" pageId="14" pageNumber="15">TABLE 5.</emphasis>
North Coast endemic mammal species recorded from the various North Coast Ranges of northern New Guinea. PA = Prince Alexander Range; T = Torricelli Range; B = Bewani Range (including Mt. Menawa); C = Cyclops Range; F = Foya Range.
</paragraph>
</caption>
<paragraph id="8BEA977FF7562F256368FD7AF91DFD11" blockId="14.[269,1310,655,774]" box="[848,1310,655,679]" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256368FD7AF91DFD11" bold="true" box="[848,1310,655,679]" pageId="14" pageNumber="15">PA T B C F Recorded elevation</emphasis>
</paragraph>
<paragraph id="8BEA977FF7562F256135FD4AFD9FFD61" blockId="14.[269,1310,655,774]" box="[269,412,703,727]" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256135FD4AFD9FFD61" bold="true" box="[269,412,703,727]" pageId="14" pageNumber="15">Monotremes</emphasis>
</paragraph>
<paragraph id="8BEA977FF7562F256135FD1AFD3CFAF1" blockId="14.[269,1310,655,774]" lastBlockId="14.[269,1236,1327,1398]" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FD1AFF2FFCB0" authority="Flannery &amp; Groves, 1998" authorityName="Flannery &amp; Groves" authorityYear="1998" box="[269,812,751,774]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="attenboroughi">
<emphasis id="B9214B6DF7562F256135FD1AFE16FCB0" box="[269,533,751,774]" italics="true" pageId="14" pageNumber="15">Zaglossus attenboroughi</emphasis>
<bibRefCitation id="EFC4EA8EF7562F256224FD1AFF2FFCB0" author="Flannery" box="[540,812,751,774]" pageId="14" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998" yearSuffix="x">Flannery &amp; Groves, 1998</bibRefCitation>
</taxonomicName>
x 
<quantity id="4CAD3A9AF7562F2564B1FD1AF8D9FCB0" box="[1161,1242,751,774]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="1600.0">1600 m</quantity>
<emphasis id="B9214B6DF7562F256135FADAFD3CFAF1" bold="true" box="[269,319,1327,1351]" pageId="14" pageNumber="15">Bats</emphasis>
</paragraph>
</subSubSection>
<paragraph id="8BEA977FF7562F256135FCBAF8FCFC40" pageId="14" pageNumber="15">
<table id="F95565DFF756D0D46135FCBAF8FCFC40" box="[269,1279,847,1014]" gridcols="4" gridrows="4" pageId="14" pageNumber="15">
<tr id="3565953DF756D0D46135FCBAF8FCFCD1" box="[269,1279,847,871]" gridrow="0" pageId="14" pageNumber="15" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76B4FC41F756D0D46135FCBAFEDCFCD1" box="[269,735,847,871]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256135FCBAFD8DFCD1" bold="true" box="[269,398,847,871]" pageId="14" pageNumber="15">Marsupials</emphasis>
</th>
</tr>
<tr id="3565953DF756D0D46135FC8AF8FCFC20" box="[269,1279,895,918]" gridrow="1" pageId="14" pageNumber="15">
<th id="76B4FC41F756D0D46135FC8AFEDCFC20" box="[269,735,895,918]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FC8AFED0FC20" authority="Flannery, 1993" authorityName="Flannery" authorityYear="1993" box="[269,723,895,918]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="B9214B6DF7562F256135FC8AFE2FFC20" box="[269,556,895,918]" italics="true" pageId="14" pageNumber="15">Dendrolagus pulcherrimus</emphasis>
Flannery, 1993
</taxonomicName>
</th>
<td id="76B4FC41F756D0D46363FC8AFFC4FC20" box="[859,967,895,918]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">? x</td>
<td id="76B4FC41F756D0D46422FC8AF825FC20" box="[1050,1062,895,918]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">?</td>
<td id="76B4FC41F756D0D4645CFC8AF8FCFC20" box="[1124,1279,895,918]" gridcol="3" gridrow="1" pageId="14" pageNumber="15">680 to 1120 m</td>
</tr>
<tr id="3565953DF756D0D46135FC5AF8FCFC70" box="[269,1279,943,966]" gridrow="2" pageId="14" pageNumber="15" rowspan-2="1">
<th id="76B4FC41F756D0D46135FC5AFEDCFC70" box="[269,735,943,966]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FC5AFEDCFC70" authority="Flannery &amp; Seri, 1990" authorityName="Flannery &amp; Seri" authorityYear="1990" box="[269,735,943,966]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="scottae">
<emphasis id="B9214B6DF7562F256135FC5AFDE9FC70" box="[269,490,943,966]" italics="true" pageId="14" pageNumber="15">Dendrolagus scottae</emphasis>
Flannery &amp; Seri, 1990
</taxonomicName>
</th>
<td id="76B4FC41F756D0D46363FC5AFFC4FC70" box="[859,967,943,966]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">x x</td>
<td id="76B4FC41F756D0D4645CFC5AF8FCFC70" box="[1124,1279,943,966]" gridcol="3" gridrow="2" pageId="14" pageNumber="15">900 to 2000 m</td>
</tr>
<tr id="3565953DF756D0D46135FC2AF8FCFC40" box="[269,1279,991,1014]" gridrow="3" pageId="14" pageNumber="15" rowspan-2="1">
<th id="76B4FC41F756D0D46135FC2AFEDCFC40" box="[269,735,991,1014]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FC2AFE43FC40" authority="Ziegler, 1981" authorityName="Ziegler" authorityYear="1981" box="[269,576,991,1014]" class="Mammalia" family="Petauridae" genus="Petaurus" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="abidi">
<emphasis id="B9214B6DF7562F256135FC2AFD6EFC40" box="[269,365,991,1014]" italics="true" pageId="14" pageNumber="15">Petaurus</emphasis>
<emphasis id="B9214B6DF7562F25614CFC2AFDA8FC40" box="[372,427,991,1014]" italics="true" pageId="14" pageNumber="15">abidi</emphasis>
Ziegler, 1981
</taxonomicName>
</th>
<td id="76B4FC41F756D0D46363FC2AFFC4FC40" box="[859,967,991,1014]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">x</td>
<td id="76B4FC41F756D0D4645CFC2AF8FCFC40" box="[1124,1279,991,1014]" gridcol="3" gridrow="3" pageId="14" pageNumber="15">800 to 1220 m</td>
</tr>
</table>
</paragraph>
<paragraph id="8BEA977FF7562F256135FBCAF905FB50" pageId="14" pageNumber="15">
<table id="F95565DFF756D0D46135FBCAF905FB50" box="[269,1286,1087,1254]" gridcols="3" gridrows="4" pageId="14" pageNumber="15">
<tr id="3565953DF756D0D46135FBCAF905FBE1" box="[269,1286,1087,1111]" gridrow="0" pageId="14" pageNumber="15" rowspan-1="1" rowspan-2="1">
<th id="76B4FC41F756D0D46135FBCAFEC5FBE1" box="[269,710,1087,1111]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256135FBCAFD69FBE1" bold="true" box="[269,362,1087,1111]" pageId="14" pageNumber="15">Rodents</emphasis>
</th>
</tr>
<tr id="3565953DF756D0D46135FB9AF905FB30" box="[269,1286,1135,1158]" gridrow="1" pageId="14" pageNumber="15">
<th id="76B4FC41F756D0D46135FB9AFEC5FB30" box="[269,710,1135,1158]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FB9AFDD2FB30" box="[269,465,1135,1158]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="ziegleri" status="sp. nov.">
<emphasis id="B9214B6DF7562F256135FB9AFDD2FB30" box="[269,465,1135,1158]" italics="true" pageId="14" pageNumber="15">Hydromys ziegleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A212F616F7562F2561EFFB9AFE12FB30" box="[471,529,1135,1158]" pageId="14" pageNumber="15" rank="species">n. sp.</taxonomicNameLabel>
</th>
<td id="76B4FC41F756D0D46362FB9AFFF5FB30" box="[858,1014,1135,1158]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">x</td>
<td id="76B4FC41F756D0D46465FB9AF905FB30" box="[1117,1286,1135,1158]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">213 m</td>
</tr>
<tr id="3565953DF756D0D46135FB6AF905FB00" box="[269,1286,1183,1206]" gridrow="2" pageId="14" pageNumber="15">
<th id="76B4FC41F756D0D46135FB6AFEC5FB00" box="[269,710,1183,1206]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256135FB6AFE1CFB00" box="[269,543,1183,1206]" italics="true" pageId="14" pageNumber="15">‘Microhydromys’ musseri</emphasis>
Flannery, 1989
</th>
<td id="76B4FC41F756D0D46362FB6AFFF5FB00" box="[858,1014,1183,1206]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">x</td>
<td id="76B4FC41F756D0D46465FB6AF905FB00" box="[1117,1286,1183,1206]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">1350 m</td>
</tr>
<tr id="3565953DF756D0D46135FB3AF905FB50" box="[269,1286,1231,1254]" gridrow="3" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FB3AFF94FB50" authority="Osgood, 1945" authorityName="Osgood" authorityYear="1945" box="[269,919,1231,1254]" class="Mammalia" family="Muridae" genus="Paraleptomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rufilatus">
<th id="76B4FC41F756D0D46135FB3AFEC5FB50" box="[269,710,1231,1254]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">
<emphasis id="B9214B6DF7562F256135FB3AFE01FB50" box="[269,514,1231,1254]" italics="true" pageId="14" pageNumber="15">Paraleptomys rufilatus</emphasis>
Osgood, 1945
</th>
<td id="76B4FC41F756D0D46362FB3AFFF5FB50" box="[858,1014,1231,1254]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">x x x</td>
</taxonomicName>
<td id="76B4FC41F756D0D46465FB3AF905FB50" box="[1117,1286,1231,1254]" gridcol="2" gridrow="3" pageId="14" pageNumber="15">1200 to 1700 m</td>
</tr>
</table>
</paragraph>
<subSubSection id="C34FC4F4F7562F246135FAAAFEF3FB98" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BEA977FF7562F246135FAAAF866FDC0" blockId="14.[269,1236,1327,1398]" lastBlockId="15.[264,1324,284,1870]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<taxonomicName id="4C55ECFCF7562F256135FAAAFF3DFAC0" ID-CoL="3M36V" authority="Flannery &amp; Colgan, 1993" authorityName="Flannery &amp; Colgan" authorityYear="1993" box="[269,830,1375,1398]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="edwardshilli">
<emphasis id="B9214B6DF7562F256135FAAAFE2BFAC0" box="[269,552,1375,1398]" italics="true" pageId="14" pageNumber="15">Hipposideros edwardshilli</emphasis>
<bibRefCitation id="EFC4EA8EF7562F256216FAAAFF3DFAC0" author="Flannery" box="[558,830,1375,1398]" pageId="14" pageNumber="19" refString="Flannery, T. F. &amp; Colgan, D. J. (1993) A new species and two new subspecies of Hipposideros (Chiroptera) from western Papua New Guinea. Records of the Australian Museum, 45, 43 - 57." type="journal article" year="1993" yearSuffix="x">Flannery &amp; Colgan, 1993</bibRefCitation>
</taxonomicName>
x 
<quantity id="4CAD3A9AF7562F2564A8FAAAF8D7FAC0" box="[1168,1236,1375,1398]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="15" unit="m" value="240.0">240 m</quantity>
Many endemic mammals occur in these mountain ranges (
<tableCitation id="C6D7A2C4F7562F256427FA2EF882FA43" box="[1055,1153,1499,1525]" captionStart="TABLE 5" captionStartId="14.[264,352,523,547]" captionText="TABLE 5. North Coast endemic mammal species recorded from the various North Coast Ranges of northern New Guinea. PA = Prince Alexander Range; T = Torricelli Range; B = Bewani Range (including Mt. Menawa); C = Cyclops Range; F = Foya Range." pageId="14" pageNumber="15">Table 5</tableCitation>
). A recentlydescribed monotreme, the dwarf long­beaked echidna (
<taxonomicName id="4C55ECFCF7562F2563F2F9F6F8F6F9AA" ID-CoL="7GBTK" box="[970,1269,1539,1564]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="attenboroughi">
<emphasis id="B9214B6DF7562F2563F2F9F6F8F6F9AA" box="[970,1269,1539,1564]" italics="true" pageId="14" pageNumber="15">Zaglossus attenboroughi</emphasis>
</taxonomicName>
), is known only by a single specimen collected in 
<quantity id="4CAD3A9AF7562F256308F9DEFF88F9F3" box="[816,907,1579,1605]" metricMagnitude="1" metricUnit="m" metricValue="4.98094" pageId="14" pageNumber="15" unit="in" value="1961.0">1961 in</quantity>
upper montane forest (
<quantity id="4CAD3A9AF7562F2564A7F9DEF8FFF9F3" box="[1183,1276,1579,1605]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="15" unit="m" value="1600.0">1600 m</quantity>
) on Mt. Rara in the Cyclops Range, behind Jayapura (
<bibRefCitation id="EFC4EA8EF7562F256367F9A6F8A9F9DB" author="Flannery" box="[863,1194,1619,1645]" pageId="14" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998">Flannery and Groves, 1998</bibRefCitation>
). The treekangaroo 
<taxonomicName id="4C55ECFCF7562F256145F98EFE73F922" ID-CoL="34QBX" box="[381,624,1659,1684]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="scottae">
<emphasis id="B9214B6DF7562F256145F98EFE73F922" box="[381,624,1659,1684]" italics="true" pageId="14" pageNumber="15">Dendrolagus scottae</emphasis>
</taxonomicName>
(the “tenkile”) is known from the western Torricelli Range and Mt. Menawa in the Bewanis, where it is primarily restricted to narrow bands of montane forest situated at about 
<quantity id="4CAD3A9AF7562F256276F93EFEECF953" box="[590,751,1739,1765]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="14" pageNumber="15" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">1200–1500 m</quantity>
in the Torricellis and about 
<quantity id="4CAD3A9AF7562F25640DF93EF8D4F953" box="[1077,1239,1739,1765]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="14" pageNumber="15" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">1500–2000 m</quantity>
on Mt. Menawa (
<bibRefCitation id="EFC4EA8EF7562F256144F906FEA1F8BB" author="Flannery" box="[380,674,1779,1805]" pageId="14" pageNumber="19" refString="Flannery, T. F. &amp; Seri, L. (1990 b) Dendrolagus scottae n. sp. (Marsupialia: Macropodidae): a new tree-kangaroo from Papua New Guinea. Records of the Australian Museum, 42, 237 - 245." type="journal article" year="1990" yearSuffix="b">Flannery and Seri 1990b</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7562F256296F906FF9DF8BB" author="Flannery" box="[686,926,1779,1805]" pageId="14" pageNumber="19" refString="Flannery, T. F., Martin, R., &amp; Szalay, A. (1996) Tree Kangaroos: A Curious Natural History, Reed Books: Melbourne, 202 pp." type="book" year="1996">
Flannery 
<emphasis id="B9214B6DF7562F256325F906FF5BF8BA" box="[797,856,1779,1804]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
1996
</bibRefCitation>
). Another tree kangaroo, 
<taxonomicName id="4C55ECFCF7562F2564F0F906FDE8F882" ID-CoL="34QBW" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="B9214B6DF7562F2564F0F906FDE8F882" italics="true" pageId="14" pageNumber="15">Dendrolagus pulcherrimus</emphasis>
</taxonomicName>
, is recorded as a living animal from the eastern Torricellis and (on the basis of a sight record) from the Foya Range (Flannery 1993; 
<bibRefCitation id="EFC4EA8EF7562F2563E7F8B6F8D3F8EB" author="Flannery" box="[991,1232,1859,1885]" pageId="14" pageNumber="19" refString="Flannery, T. F., Martin, R., &amp; Szalay, A. (1996) Tree Kangaroos: A Curious Natural History, Reed Books: Melbourne, 202 pp." type="book" year="1996">
Flannery 
<emphasis id="B9214B6DF7562F256477F8B6F888F8EA" box="[1103,1163,1859,1884]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
1996
</bibRefCitation>
), and is rumoured to occur in the Prince Alexander Range. The latter species (originally described as a subspecies of 
<taxonomicName id="4C55ECFCF7572F2461D6FEB1FE9CFEEB" ID-CoL="34QBR" box="[494,671,324,349]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="goodfellowi">
<emphasis id="B9214B6DF7572F2461D6FEB1FE9CFEEB" box="[494,671,324,349]" italics="true" pageId="15" pageNumber="16">D. goodfellowi</emphasis>
</taxonomicName>
by Flannery 1993) has disappeared from much of its former range in the Torricellis in recent decades, probably as a result of unsustainable hunting practices (
<bibRefCitation id="EFC4EA8EF7572F2461D8FE61FED1FE18" author="Flannery" box="[480,722,404,430]" pageId="15" pageNumber="19" refString="Flannery, T. F., Martin, R., &amp; Szalay, A. (1996) Tree Kangaroos: A Curious Natural History, Reed Books: Melbourne, 202 pp." type="book" year="1996">
Flannery 
<emphasis id="B9214B6DF7572F246269FE61FE88FE1B" box="[593,651,404,429]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1996
</bibRefCitation>
). In the past, 
<taxonomicName id="4C55ECFCF7572F246355FE61F82EFE1B" ID-CoL="34QBW" box="[877,1069,404,429]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="B9214B6DF7572F246355FE61F82EFE1B" box="[877,1069,404,429]" italics="true" pageId="15" pageNumber="16">D. pulcherrimus</emphasis>
</taxonomicName>
probably had a much more extensive distribution, as it or a very closely­related taxon is also represented in Quaternary subfossil deposits from the Vogelkop Peninsula of north­west New 
<collectingCountry id="F342D7EFF7572F2464B9FE11F8D5FE48" box="[1153,1238,484,510]" name="Guinea-Bissau" pageId="15" pageNumber="16">Guinea</collectingCountry>
(
<bibRefCitation id="EFC4EA8EF7572F2464DEFE11FD8BFD90" author="Aplin" lastPageId="15" lastPageNumber="19" pageId="15" pageNumber="18" refString="Aplin, K. P., Pasveer, J. M. &amp; Boles, W. E. (1999) Late Quaternary vertebrates from the Birds Head Peninsula, Irian Jaya, Indonesia, including descriptions of two previously unknown marsupial species. Records of the Western Australian Museum, Supplement 57, 351 - 387." type="book chapter" year="1999">
Aplin 
<emphasis id="B9214B6DF7572F246130FDF9FD41FD93" box="[264,322,524,549]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1999
</bibRefCitation>
). A third marsupial found only in the North Coast Ranges is the Northern glider (
<taxonomicName id="4C55ECFCF7572F24612AFDC1FDBDFDFB" ID-CoL="4FD5F" box="[274,446,564,589]" class="Mammalia" family="Petauridae" genus="Petaurus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="abidi">
<emphasis id="B9214B6DF7572F24612AFDC1FDBDFDFB" box="[274,446,564,589]" italics="true" pageId="15" pageNumber="16">Petaurus abidi</emphasis>
</taxonomicName>
), recorded only from mid­ and upper montane forest in the vicinity of Mts. Somoro and Sapau in the Torricelli Range (
<bibRefCitation id="EFC4EA8EF7572F24633BFDA9FF9DFDC0" author="Ziegler" box="[771,926,604,630]" pageId="15" pageNumber="20" refString="Ziegler, A. C. (1981) Petaurus abidi, a new species of glider (Marsupialia: Petauridae) from Papua New Guinea. Australian Mammalogy, 4, 81 - 88." type="journal article" year="1981">Ziegler 1981</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7572F246392FDA9F85AFDC0" author="Flannery" box="[938,1113,604,630]" pageId="15" pageNumber="19" refString="Flannery, T. F. (1995) Mammals of New Guinea, revised edition, Reed Books: Ithaca, New York, 568 pp." type="book" year="1995">Flannery 1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA977FF7572F246100FD71FEF3FB98" blockId="15.[264,1324,284,1870]" pageId="15" pageNumber="16">
Among murine rodents, three endemic hydromyins are known from the North Coast Ranges. These are 
<emphasis id="B9214B6DF7572F2461D2FD59FEFCFD73" box="[490,767,684,709]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C55ECFCF7572F2461D2FD59FEF8FD73" ID-CoL="4CYLR" box="[490,763,684,709]" class="Mammalia" family="Muridae" genus="Paraleptomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="rufilatus">Paraleptomys rufilatus</taxonomicName>
,
</emphasis>
a medium­sized terrestrial rat which occurs in mountaintop forests in the Cyclops, Bewani, and Torricelli Ranges (
<bibRefCitation id="EFC4EA8EF7572F246402FD21F8DCFD58" author="Osgood" box="[1082,1247,724,750]" pageId="15" pageNumber="20" refString="Osgood, W. H. (1945) A new rodent from Dutch New Guinea. Fieldiana (Zoology), 31, 1 - 2." type="journal article" year="1945">Osgood 1945</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7572F2464D5FD21FD81FCA0" author="Flannery" pageId="15" pageNumber="19" refString="Flannery, T. F. (1995) Mammals of New Guinea, revised edition, Reed Books: Ithaca, New York, 568 pp." type="book" year="1995">Flannery 1995</bibRefCitation>
; Musser and Carleton 2005); ‘ 
<emphasis id="B9214B6DF7572F2462D1FD09F80EFCA3" box="[745,1037,764,789]" italics="true" pageId="15" pageNumber="16">Microhydromys’ musseri</emphasis>
, a small moss­mouse of uncertain phylogenetic affinity, known by a single specimen collected in 
<quantity id="4CAD3A9AF7572F2464B9FCD1F8DDFC88" box="[1153,1246,804,830]" metricMagnitude="1" metricUnit="m" metricValue="5.00888" pageId="15" pageNumber="16" unit="in" value="1972.0">1972 in</quantity>
upper montane forest (
<quantity id="4CAD3A9AF7572F2461F5FCB9FE2FFCD0" box="[461,556,844,870]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="15" pageNumber="16" unit="m" value="1350.0">1350 m</quantity>
) on Mt. Somoro in the Torricellis (Flannery 1989; Musser and Carleton, 2005); and now 
<taxonomicName id="4C55ECFCF7572F246202FC81FF12FC3B" ID-CoL="3N7S8" box="[570,785,884,909]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7572F246202FC81FF12FC3B" box="[570,785,884,909]" italics="true" pageId="15" pageNumber="16">Hydromys ziegleri</emphasis>
</taxonomicName>
, known by a single specimen collected on the southern margin of the Prince Alexander Range. Finally, one species and one distinctive subspecies of leaf­nosed bat (
<taxonomicName id="4C55ECFCF7572F2462B8FC31FFBEFC6B" ID-CoL="3M36V" box="[640,957,964,989]" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="edwardshilli">
<emphasis id="B9214B6DF7572F2462B8FC31FFBEFC6B" box="[640,957,964,989]" italics="true" pageId="15" pageNumber="16">Hipposideros edwardshilli</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C55ECFCF7572F24643DFC31FD67FBB3" ID-CoL="5HYN3" class="Mammalia" family="Hipposideridae" genus="Hipposideros" kingdom="Animalia" order="Chiroptera" pageId="15" pageNumber="16" phylum="Chordata" rank="subSpecies" species="wollastoni" subSpecies="fasensis">
<emphasis id="B9214B6DF7572F24643DFC31FD67FBB3" italics="true" pageId="15" pageNumber="16">Hipposideros wollastoni fasensis</emphasis>
</taxonomicName>
) are known only from the northern and southern foothills of the Bewani range, respectively (
<bibRefCitation id="EFC4EA8EF7572F24619DFBE1FEE0FB98" author="Flannery" box="[421,739,1044,1070]" pageId="15" pageNumber="19" refString="Flannery, T. F. &amp; Colgan, D. J. (1993) A new species and two new subspecies of Hipposideros (Chiroptera) from western Papua New Guinea. Records of the Australian Museum, 45, 43 - 57." type="journal article" year="1993">Flannery and Colgan 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34FC4F4F7572F3A6100FBC9FF13FC68" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEA977FF7572F246100FBC9F87CFAD8" blockId="15.[264,1324,284,1870]" pageId="15" pageNumber="16">
During cooler climatic periods during the Late Pleistocene, montane forests extended to much lower elevations in New 
<collectingCountry id="F342D7EFF7572F2462B7FB91FEE7FBC8" box="[655,740,1124,1150]" name="Guinea-Bissau" pageId="15" pageNumber="16">Guinea</collectingCountry>
than they do today, and the montane­adapted species of the North Coast Ranges likely had considerably more expansive or even contiguous distributions across certain of these mountain ranges. I suspect that the seemingly haphazard distribution of endemic species recorded from the various North Coast ranges today (
<tableCitation id="C6D7A2C4F7572F246128FAF1FD68FAA8" box="[272,363,1284,1310]" captionStart="TABLE 5" captionStartId="14.[264,352,523,547]" captionText="TABLE 5. North Coast endemic mammal species recorded from the various North Coast Ranges of northern New Guinea. PA = Prince Alexander Range; T = Torricelli Range; B = Bewani Range (including Mt. Menawa); C = Cyclops Range; F = Foya Range." pageId="15" pageNumber="16">Table 5</tableCitation>
) reflects a combination of two factors: artefacts of geographically uneven sampling effort and, at least for some of the larger­bodied species (
<emphasis id="B9214B6DF7572F2463C4FAD9F91AFAF3" box="[1020,1305,1324,1349]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C55ECFCF7572F2463C4FAD9F877FAF3" box="[1020,1140,1324,1349]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Zaglossus</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7572F2464BAFAD9F91AFAF3" box="[1154,1305,1324,1349]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">Dendrolagus</taxonomicName>
</emphasis>
), historical extinctions (see 
<bibRefCitation id="EFC4EA8EF7572F246203FAA1FF29FAD8" author="Flannery" box="[571,810,1364,1390]" pageId="15" pageNumber="19" refString="Flannery, T. F., Martin, R., &amp; Szalay, A. (1996) Tree Kangaroos: A Curious Natural History, Reed Books: Melbourne, 202 pp." type="book" year="1996">
Flannery 
<emphasis id="B9214B6DF7572F246292FAA1FEE6FADB" box="[682,741,1364,1389]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1996
</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7572F24630EFAA1F871FAD8" author="Flannery" box="[822,1138,1364,1390]" pageId="15" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998">Flannery and Groves 1998</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA977FF7572F3B6100FA89FD57FE30" blockId="15.[264,1324,284,1870]" lastBlockId="16.[264,1325,284,1870]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Some of the larger endemic mammals of the North Coast Ranges, especially 
<emphasis id="B9214B6DF7572F24648DFA89FEB5FA0B" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C55ECFCF7572F24648DFA89FDB3FA0B" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="attenboroughi">Zaglossus attenboroughi</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7572F246184FA51FEB1FA0B" box="[444,690,1444,1469]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="scottae">Dendrolagus scottae</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C55ECFCF7572F2462C8FA51F82FFA0B" box="[752,1068,1444,1469]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pulcherrimus">
<emphasis id="B9214B6DF7572F2462C8FA51F82FFA0B" box="[752,1068,1444,1469]" italics="true" pageId="15" pageNumber="16">Dendrolagus pulcherrimus</emphasis>
</taxonomicName>
, have minuscule global geographic ranges and are actively hunted for food (
<bibRefCitation id="EFC4EA8EF7572F2463ABFA39F886FA50" author="Flannery" box="[915,1157,1484,1510]" pageId="15" pageNumber="19" refString="Flannery, T. F., Martin, R., &amp; Szalay, A. (1996) Tree Kangaroos: A Curious Natural History, Reed Books: Melbourne, 202 pp." type="book" year="1996">
Flannery 
<emphasis id="B9214B6DF7572F24643EFA39F843FA53" box="[1030,1088,1484,1509]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1996
</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7572F2464A9FA39FDA1F9B8" author="Flannery" pageId="15" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998">Flannery and Groves 1998</bibRefCitation>
), and are probably among the most endangered of all Australasian mammals (indeed, 
<taxonomicName id="4C55ECFCF7572F246148F9E9FE35F983" box="[368,566,1564,1589]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="attenboroughi">
<emphasis id="B9214B6DF7572F246148F9E9FE35F983" box="[368,566,1564,1589]" italics="true" pageId="15" pageNumber="16">Z. attenboroughi</emphasis>
</taxonomicName>
may already be extinct; 
<bibRefCitation id="EFC4EA8EF7572F246358F9E9F8A2F980" author="Flannery" box="[864,1185,1564,1590]" pageId="15" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998">Flannery and Groves 1998</bibRefCitation>
). All North Coast Range endemic mammals, even the smallest (such as ‘ 
<emphasis id="B9214B6DF7572F2463ECF9B1F8FAF9EB" box="[980,1273,1604,1629]" italics="true" pageId="15" pageNumber="16">Microhydromys’ musseri</emphasis>
and 
<taxonomicName id="4C55ECFCF7572F246130F999FDE5F933" box="[264,486,1644,1669]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7572F246130F999FDE5F933" box="[264,486,1644,1669]" italics="true" pageId="15" pageNumber="16">Hydromys ziegleri</emphasis>
</taxonomicName>
), should be considered conservation priorities on account of their extremely limited geographic and altitudinal ranges, which in most cases are centred on small tracts of upper montane forest, highly susceptible to threatening processes including logging and climate change. In addition to supporting a markedly unique mammal fauna, the North Coast Ranges are also an area of considerable geographically­restricted endemism for frogs, birds, and other biotic groups (
<bibRefCitation id="EFC4EA8EF7572F246308F8C1F84BF8F8" author="Stattersfield" box="[816,1096,1844,1870]" pageId="15" pageNumber="20" refString="Stattersfield, A. J., Corsby, M. J., Long, A. J. &amp; Wege, D. C. (1998) Global Directory of Endemic Bird Areas, BirdLife International: Cambridge, U. K., 846 pp." type="book" year="1998">
Stattersfield 
<emphasis id="B9214B6DF7572F2463FFF8C1F801F8FB" box="[967,1026,1844,1869]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
1998
</bibRefCitation>
; Kraus and Allison 2000; Allison and Kraus 2000, 2003), ranking these mountain ranges among the most critical priorities for conservation efforts in the Melanesian region (
<bibRefCitation id="EFC4EA8EF7482F3B6427FEB1FD4BFE30" author="Wikramanayake" pageId="16" pageNumber="20" refString="Wikramanayake, E., Dinerstein, E. Loucks, C. J., et al. (2002) Terrestrial Ecoregions of the Indo- Pacific: A Conservation Assessment, Island Press: Washington, D. C., 643 pp." type="book" year="2002">
Wikramanayake 
<emphasis id="B9214B6DF7482F3B64D3FEB1F929FEEB" box="[1259,1322,324,349]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2002
</bibRefCitation>
).
</paragraph>
<paragraph id="8BEA977FF7482F3B6100FE61F872FC68" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
Of the eight endemic mammal species currently known from the North Coast Ranges, seven were described as new to science only within the last 15 years, and all eight were both discovered and described within the last 60 years (
<tableCitation id="C6D7A2C4F7482F3B6390FE11F805FE48" box="[936,1030,484,510]" captionStart="TABLE 5" captionStartId="14.[264,352,523,547]" captionText="TABLE 5. North Coast endemic mammal species recorded from the various North Coast Ranges of northern New Guinea. PA = Prince Alexander Range; T = Torricelli Range; B = Bewani Range (including Mt. Menawa); C = Cyclops Range; F = Foya Range." pageId="16" pageNumber="17">Table 5</tableCitation>
). Further, three of these remain known by a single museum specimen (
<emphasis id="B9214B6DF7482F3B6316FDF9FD69FDFB" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6316FDF9F850FD93" box="[814,1107,524,549]" class="Mammalia" family="Tachyglossidae" genus="Zaglossus" kingdom="Animalia" order="Monotremata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="attenboroughi">Zaglossus attenboroughi</taxonomicName>
, ‘Microhydromys’ musseri,
</emphasis>
and 
<taxonomicName id="4C55ECFCF7482F3B619EFDC1FE7DFDFB" box="[422,638,564,589]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7482F3B619EFDC1FE7DFDFB" box="[422,638,564,589]" italics="true" pageId="16" pageNumber="17">Hydromys ziegleri</emphasis>
</taxonomicName>
). These cursory but remarkable statistics for North Coast Range mammals aptly demonstrate how incompletely catalogued the biota of these mountains remains. While the mammal faunas of some areas in the North Coast Ranges (including the western Torricellis, Cyclops, and Bewanis) have become much better understood through recent survey efforts (e.g. Flannery 1993, 1995), others, such as the Foya and Van Rees Ranges of western New 
<collectingCountry id="F342D7EFF7482F3B624CFD09FECFFCA0" box="[628,716,764,790]" name="Guinea-Bissau" pageId="16" pageNumber="17">Guinea</collectingCountry>
, remain almost wholly unknown mammalogically. For example, until recently the ornithologist Jared Diamond was the only westerner known to have visited the relatively expansive Foya Range, which is mostly uninhabited and largely unexplored (see 
<bibRefCitation id="EFC4EA8EF7482F3B6216FC81FEE5FC38" author="Diamond" box="[558,742,884,910]" pageId="16" pageNumber="19" refString="Diamond, J. M. (1985) New distributional records and taxa from the outlying mountain ranges of New Guinea. Emu, 85, 65 - 91." type="journal article" year="1985">Diamond 1985</bibRefCitation>
). Given the startling rate of new discoveries in North Coast mountain ranges in recent decades, there is no doubt that future survey efforts in this region will continue to encounter vertebrate species new to science.
</paragraph>
<paragraph id="8BEA977FF7482F3B6100FC19F8B8F9B8" blockId="16.[264,1325,284,1870]" pageId="16" pageNumber="17">
The mammal fauna of the Prince Alexander Mountains is moderately well­known. Apart from 
<emphasis id="B9214B6DF7482F3B61AAFBE1FE73FB9B" box="[402,624,1044,1069]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B61AAFBE1FE6FFB9B" box="[402,620,1044,1069]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="ziegleri">Hydromys ziegleri</taxonomicName>
,
</emphasis>
at least 23 other species are recorded from these mountains (specimens at AM, AMNH, and BBM). These include the cuscuses 
<emphasis id="B9214B6DF7482F3B6417FBC9FD95FBCB" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6417FBC9F925FBE3" box="[1071,1318,1084,1109]" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="orientalis">Phalanger orientalis</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B6130FB91FD91FBCB" box="[264,402,1124,1149]" class="Mammalia" family="Phalangeridae" genus="Phalanger" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="gymnotis">P. gymnotis</taxonomicName>
,
</emphasis>
and 
<emphasis id="B9214B6DF7482F3B61E9FB91FEE1FBCB" box="[465,738,1124,1149]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B61E9FB91FEDDFBCB" box="[465,734,1124,1149]" class="Mammalia" family="Phalangeridae" genus="Spilocuscus" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="maculatus">Spilocuscus maculatus</taxonomicName>
,
</emphasis>
the tree­kangaroo 
<emphasis id="B9214B6DF7482F3B63F9FB91F8B8FBCB" box="[961,1211,1124,1149]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B63F9FB91F8B5FBCB" box="[961,1206,1124,1149]" class="Mammalia" family="Macropodidae" genus="Dendrolagus" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="inustus">Dendrolagus inustus</taxonomicName>
,
</emphasis>
the wallabies 
<taxonomicName id="4C55ECFCF7482F3B616CFB79FE29FB13" box="[340,554,1164,1189]" class="Mammalia" family="Macropodidae" genus="Thylogale" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="browni">
<emphasis id="B9214B6DF7482F3B616CFB79FE29FB13" box="[340,554,1164,1189]" italics="true" pageId="16" pageNumber="17">Thylogale browni</emphasis>
</taxonomicName>
and 
<emphasis id="B9214B6DF7482F3B624AFB79FF53FB13" box="[626,848,1164,1189]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B624AFB79FF4FFB13" box="[626,844,1164,1189]" class="Mammalia" family="Macropodidae" genus="Dorcopsis" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hageni">Dorcopsis hageni</taxonomicName>
,
</emphasis>
the bandicoots 
<emphasis id="B9214B6DF7482F3B6418FB79FD5CFB7B" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6418FB79F8FDFB13" box="[1056,1278,1164,1189]" class="Mammalia" family="Peramelidae" genus="Echymipera" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="clara">Echymipera clara</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B652AFB79FD58FB7B" class="Mammalia" family="Peramelidae" genus="Echymipera" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="kalubu">E. kalubu</taxonomicName>
,
</emphasis>
and 
<emphasis id="B9214B6DF7482F3B61A4FB41FE37FB7B" box="[412,564,1204,1229]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B61A4FB41FE33FB7B" box="[412,560,1204,1229]" class="Mammalia" family="Peramelidae" genus="Echymipera" kingdom="Animalia" order="Peramelemorphia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="rufescens">E. rufescens</taxonomicName>
,
</emphasis>
the rodents 
<emphasis id="B9214B6DF7482F3B62F1FB41FD67FB43" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B62F1FB41FFE3FB7B" box="[713,992,1204,1229]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chrysogaster">Hydromys chrysogaster</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B63D6FB41F8D0FB7B" box="[1006,1235,1204,1229]" class="Mammalia" family="Muridae" genus="Melomys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="rufescens">Melomys rufescens</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B64D8FB41FD63FB43" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="praetor">Rattus praetor</taxonomicName>
,
</emphasis>
and 
<emphasis id="B9214B6DF7482F3B61A4FB29FE1EFB43" box="[412,541,1244,1269]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B61A4FB29FE1AFB43" box="[412,537,1244,1269]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="exulans">R. exulans</taxonomicName>
,
</emphasis>
the fruit­bats 
<emphasis id="B9214B6DF7482F3B62F9FB29F89EFAAB" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B62F9FB29F854FB43" authority="Dobsonia" authorityName="Dobsonia" box="[705,1111,1244,1269]" class="Mammalia" family="Pteropodidae" genus="Pteropus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="neohibernicus">Pteropus neohibernicus, Dobsonia</taxonomicName>
minor, 
<taxonomicName id="4C55ECFCF7482F3B6497FB29FD45FAAB" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="aello">Nyctimene aello</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B616AFAF1FDE8FAAB" box="[338,491,1284,1309]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="albiventer">N. albiventer</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B61CFFAF1FF3FFAAB" authority="Paranyctimene" authorityName="Paranyctimene" box="[503,828,1284,1309]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cyclotis">N. cyclotis, Paranyctimene</taxonomicName>
tenax, Syconycteris australis,
</emphasis>
and 
<emphasis id="B9214B6DF7482F3B64ECFAF1FDD2FAF3" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B64ECFAF1FDCEFAF3" class="Mammalia" family="Pteropodidae" genus="Macroglossus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="minimus">Macroglossus minimus</taxonomicName>
,
</emphasis>
the vespertilionid bat 
<emphasis id="B9214B6DF7482F3B62E6FAD9FFE5FAF3" box="[734,998,1324,1349]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B62E6FAD9FFE1FAF3" box="[734,994,1324,1349]" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="papuanus">Pipistrellus papuanus</taxonomicName>
,
</emphasis>
and the molossid bat 
<taxonomicName id="4C55ECFCF7482F3B64D6FAD9FDDFFADB" class="Mammalia" family="Molossidae" genus="Mormopterus" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="beccarii">
<emphasis id="B9214B6DF7482F3B64D6FAD9FDDFFADB" italics="true" pageId="16" pageNumber="17">Mormopterus beccarii</emphasis>
</taxonomicName>
. However, essentially all of these species (excepting 
<emphasis id="B9214B6DF7482F3B647FFAA1F928FADB" box="[1095,1323,1364,1389]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B647FFAA1F924FADB" box="[1095,1319,1364,1389]" class="Mammalia" family="Pteropodidae" genus="Nyctimene" kingdom="Animalia" order="Chiroptera" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cyclotis">Nyctimene cyclotis</taxonomicName>
,
</emphasis>
a widespread inhabitant of New Guinean montane forests) are characteristic inhabitants of lowland habitats and are widespread in northern New 
<collectingCountry id="F342D7EFF7482F3B6347FA51FFDBFA08" box="[895,984,1444,1470]" name="Guinea-Bissau" pageId="16" pageNumber="17">Guinea</collectingCountry>
. Future surveys in the Prince Alexanders should thus focus on the very highest peaks in the range, which rise to 
<quantity id="4CAD3A9AF7482F3B64EAFA39F928FA50" box="[1234,1323,1484,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.24" pageId="16" pageNumber="17" unit="m" value="1240.0">1240 m</quantity>
and are likely to support unique animal assemblages unknown at lower altitudes.
</paragraph>
<paragraph id="8BEA977FF7482F3A6100F9E9FE16FDC0" blockId="16.[264,1325,284,1870]" lastBlockId="17.[264,1325,284,990]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<collectingCountry id="F342D7EFF7482F3B6100F9E9FD5AF980" box="[312,345,1564,1590]" name="American Samoa" pageId="16" pageNumber="17">As</collectingCountry>
discussed above, rodent­collecting efforts in the Melanesian region have so far yielded five species of 
<emphasis id="B9214B6DF7482F3B622CF9B1FEA6F9EB" box="[532,677,1604,1629]" italics="true" pageId="16" pageNumber="17">Hydromys—</emphasis>
the widespread 
<taxonomicName id="4C55ECFCF7482F3B6364F9B1F816F9EB" box="[860,1045,1604,1629]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chrysogaster">
<emphasis id="B9214B6DF7482F3B6364F9B1F816F9EB" box="[860,1045,1604,1629]" italics="true" pageId="16" pageNumber="17">H. chrysogaster</emphasis>
</taxonomicName>
, the insular endemic 
<emphasis id="B9214B6DF7482F3B6536F9B1FDBBF933" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6536F9B1FDB7F933" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="neobrittanicus">H. neobrittanicus</taxonomicName>
,
</emphasis>
the mid­montane Wissel Lakes endemic 
<emphasis id="B9214B6DF7482F3B6393F999F832F933" box="[939,1073,1644,1669]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6393F999F82EF933" box="[939,1069,1644,1669]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hussoni">H. hussoni</taxonomicName>
,
</emphasis>
the newly­described 
<emphasis id="B9214B6DF7482F3B6130F961FD8DF91B" box="[264,398,1684,1709]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6130F961FD8AF91B" box="[264,393,1684,1709]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="ziegleri">H. ziegleri</taxonomicName>
,
</emphasis>
and an unnamed species from the Moluccan island of Obi. The three larger species (
<emphasis id="B9214B6DF7482F3B6157F949FF0EF963" box="[367,781,1724,1749]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C55ECFCF7482F3B6157F949FE28F963" box="[367,555,1724,1749]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="chrysogaster">H. chrysogaster</taxonomicName>
, 
<taxonomicName id="4C55ECFCF7482F3B6200F949FF0AF963" box="[568,777,1724,1749]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="neobrittanicus">H. neobrittanicus</taxonomicName>
,
</emphasis>
and the Obi taxon) have all been collected in lowland rainforest, including in forest at sea level, while one of the smaller species (
<taxonomicName id="4C55ECFCF7482F3B6536F911FD61F893" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7482F3B6536F911FD61F893" italics="true" pageId="16" pageNumber="17">H. hussoni</emphasis>
</taxonomicName>
) is known only from forest at much higher elevations. The similarly small­bodied 
<taxonomicName id="4C55ECFCF7482F3B6130F8C1FDDCF8FB" box="[264,479,1844,1869]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7482F3B6130F8C1FDDCF8FB" box="[264,479,1844,1869]" italics="true" pageId="16" pageNumber="17">Hydromys ziegleri</emphasis>
</taxonomicName>
is thus far known only by a single specimen taken in relatively low elevation forest (
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); given that extensive collecting efforts in other areas of lowland northern New 
<collectingCountry id="F342D7EFF7492F3A6188FEB1FE06FEE8" box="[432,517,324,350]" name="Guinea-Bissau" pageId="17" pageNumber="18">Guinea</collectingCountry>
have failed to record it elsewhere, I strongly suspect that it is a legitimate endemic of the North Coast Ranges. That the other small­bodied 
<taxonomicName id="4C55ECFCF7492F3A6474FE99F8C7FE33" box="[1100,1220,364,389]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B9214B6DF7492F3A6474FE99F8C7FE33" box="[1100,1220,364,389]" italics="true" pageId="17" pageNumber="18">Hydromys</emphasis>
</taxonomicName>
(
<taxonomicName id="4C55ECFCF7492F3A64ECFE99FD3AFE1B" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="hussoni">
<emphasis id="B9214B6DF7492F3A64ECFE99FD3AFE1B" italics="true" pageId="17" pageNumber="18">H. hussoni</emphasis>
</taxonomicName>
) and most other North Coast Range endemic mammals are restricted to higherelevation forests suggests to me that 
<taxonomicName id="4C55ECFCF7492F3A6287FE49FF3EFE63" box="[703,829,444,469]" class="Mammalia" family="Muridae" genus="Hydromys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B9214B6DF7492F3A6287FE49FF3EFE63" box="[703,829,444,469]" italics="true" pageId="17" pageNumber="18">H. ziegleri</emphasis>
</taxonomicName>
may be an inhabitant of montane forests, and that thus far it may be known only from the very lowest altitudinal bound of its range. Future trapping regimes targeted at streams along elevational gradients in the Prince Alexander and eastern Torricelli Mountains should serve to illuminate its real geographic and altitudinal distribution.
</paragraph>
<paragraph id="8BEA977FF7492F3A6100FD71FF13FC68" blockId="17.[264,1325,284,990]" pageId="17" pageNumber="18">
The ongoing discovery and description of new mammal species from throughout mainland New 
<collectingCountry id="F342D7EFF7492F3A6187FD59FE17FD70" box="[447,532,684,710]" name="Guinea-Bissau" pageId="17" pageNumber="18">Guinea</collectingCountry>
(e.g. 
<bibRefCitation id="EFC4EA8EF7492F3A6262FD59FF98FD70" author="Flannery" box="[602,923,684,710]" pageId="17" pageNumber="19" refString="Flannery, T. F. &amp; Groves, C. P. (1998) A revision of the genus Zaglossus (Monotremata, Tachyglossidae), with description of new species and subspecies. Mammalia, 62, 367 - 396." type="journal article" year="1998">Flannery and Groves 1998</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7492F3A6391FD59F8E9FD70" author="Helgen" box="[937,1258,684,710]" pageId="17" pageNumber="19" refString="Helgen, K. M. &amp; Flannery, T. F. (2004) A new species of bandicoot, Microperoryctes aplini, from western New Guinea. Journal of Zoology (London), 264, 117 - 124." type="journal article" year="2004">Helgen and Flannery 2004</bibRefCitation>
; 
<bibRefCitation id="EFC4EA8EF7492F3A64C0FD59FD78FD58" author="Helgen" pageId="17" pageNumber="19" refString="Helgen, K. M. (2005) A new species of murid rodent (genus Mayermys) from south-eastern New Guinea. Mammalian Biology, 70, 61 - 67." type="journal article" year="2005">Helgen 2005</bibRefCitation>
), highlights how poorly­studied the biota of many areas of the island remains. Amphibious murines, which can be difficult to trap, are often especially overlooked in rapid faunal inventories. I strongly suspect that future survey efforts will identify additional small­bodied amphibious murines in outlying areas of montane New Guinea—per­haps, for example, in the mountain ranges of the Vogelkop Peninsula (the Arfaks and Tamraus), in the expansive Foya Range, or on other large mountainous islands adjacent to New 
<collectingCountry id="F342D7EFF7492F3A617EFC31FD9DFC68" box="[326,414,964,990]" name="Guinea-Bissau" pageId="17" pageNumber="18">Guinea</collectingCountry>
, such as New 
<collectingCountry id="F342D7EFF7492F3A627DFC31FE95FC68" box="[581,662,964,990]" name="United Kingdom" pageId="17" pageNumber="18">Britain</collectingCountry>
or Seram.
</paragraph>
</subSubSection>
</treatment>
</document>