196 lines
20 KiB
XML
196 lines
20 KiB
XML
<document id="B7911496E65DAEF508CD934F5CF84A16" ID="10.11646/zootaxa.4231.3.7SLASH10134" ID-CLB-Dataset="36563" ID-DOI="10.11646/zootaxa.4231.3.7" ID-GBIF-Dataset="b00e90b8-95b1-4d04-96b0-7e1b2abd5bf8" ID-ISSN="1175-5326" ID-Zenodo-Dep="291315" ID-ZooBank="C228937C-DC62-40F9-A42E-0FB2575447E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1486975824547" checkinUser="plazi" docAuthor="Rifkind, Jacques" docDate="2017" docId="03DB87C2CD42B678AFE83343FE9DC6ED" docLanguage="en" docName="zootaxa.4231.3.7SLASH10134.pdf" docOrigin="Zootaxa 4231 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enoclerus incanus Rifkind, n. sp." docType="treatment" docVersion="9" lastPageNumber="413" masterDocId="FFE2FFBACD41B67CAF7F3429FFEDC051" masterDocTitle="New species of Mutillid Mimicking Enoclerus Gahan (Coleoptera: Cleridae: Clerinae) from Mexico and Central America" masterLastPageNumber="420" masterPageNumber="409" pageNumber="412" updateTime="1698436846717" updateUser="ExternalLinkService">
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<mods:titleInfo id="41B3AAC41A722D83BA2FC1B6685B5A46">
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<mods:title id="5B8E4941473998E48913D92A5615E382">New species of Mutillid Mimicking Enoclerus Gahan (Coleoptera: Cleridae: Clerinae) from Mexico and Central America</mods:title>
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<mods:name id="3DAB6E1D5C8B4814D49ADEE407643420" type="personal">
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<mods:namePart id="A4FF62C03F5C384EF0AEE88EF690B2C8">Rifkind, Jacques</mods:namePart>
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<mods:title id="97444CBFD76F7B31C8DB63F603DC93DC">Zootaxa</mods:title>
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<mods:part id="BDA5960DCB12C4398076730C7D8DAD26">
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<mods:date id="4B118B20678F2A9ACFBAC34462AA2F4D">2017</mods:date>
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<mods:number id="D6887DECC3C6F687D3DFCD1E203D0498">4231</mods:number>
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<mods:number id="B1B4A0B97A2FFAB4033D5D92EF62437A">3</mods:number>
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<mods:classification id="B1E0E8E964C22FC7E6DFD22E76D37138">journal article</mods:classification>
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<mods:identifier id="7BDD41C1D62CCA2E9287B66B941B6281" type="CLB-Dataset">36563</mods:identifier>
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<mods:identifier id="8383B5855848E4141E5E2E34932F1E81" type="DOI">10.11646/zootaxa.4231.3.7</mods:identifier>
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<mods:identifier id="31C040689E6058FF31DD85C4897ACE69" type="GBIF-Dataset">b00e90b8-95b1-4d04-96b0-7e1b2abd5bf8</mods:identifier>
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<mods:identifier id="10B0FE4658B3CB1E5FECE4AA14574B6B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="034E1DF9D519AD107C1797EA3A88BBCF" type="Zenodo-Dep">291315</mods:identifier>
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<mods:identifier id="D7AA91E09521E1A4BBF787B0CED2AC28" type="ZooBank">C228937C-DC62-40F9-A42E-0FB2575447E6</mods:identifier>
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<treatment id="03DB87C2CD42B678AFE83343FE9DC6ED" ID-DOI="http://doi.org/10.5281/zenodo.6030409" ID-GBIF-Taxon="127694952" ID-Zenodo-Dep="6030409" LSID="urn:lsid:plazi:treatment:03DB87C2CD42B678AFE83343FE9DC6ED" httpUri="http://treatment.plazi.org/id/03DB87C2CD42B678AFE83343FE9DC6ED" lastPageId="4" lastPageNumber="413" pageId="3" pageNumber="412">
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<subSubSection id="C368655FCD42B67FAFE83343FF06C7F7" pageId="3" pageNumber="412" type="nomenclature">
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<paragraph id="8BCD36D4CD42B67FAFE83343FDDFC7D4" blockId="3.[151,562,1898,1958]" box="[151,562,1898,1925]" pageId="3" pageNumber="412">
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<heading id="D08581B8CD42B67FAFE83343FDDFC7D4" bold="true" box="[151,562,1898,1925]" fontSize="11" level="1" pageId="3" pageNumber="412" reason="1">
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<emphasis id="B906EAC6CD42B67FAFE83343FDDFC7D4" bold="true" box="[151,562,1898,1925]" pageId="3" pageNumber="412">
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<taxonomicName id="4C724D57CD42B67FAFE83343FE0EC7D4" authority="Rifkind" authorityName="Rifkind" box="[151,483,1898,1925]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="incanus" status="sp. nov.">
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<emphasis id="B906EAC6CD42B67FAFE83343FE97C7D5" bold="true" box="[151,378,1898,1924]" italics="true" pageId="3" pageNumber="412">Enoclerus incanus</emphasis>
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Rifkind
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23557BDCD42B67FAE8F3342FDDFC7D4" box="[496,562,1899,1925]" pageId="3" pageNumber="412" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCD36D4CD42B67FAFE833A4FF06C7F7" blockId="3.[151,562,1898,1958]" box="[151,235,1933,1958]" pageId="3" pageNumber="412">(Fig. 4)</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD42B678AFE833FCFB8EC0A9" lastPageId="4" lastPageNumber="413" pageId="3" pageNumber="412" type="diagnosis">
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<paragraph id="8BCD36D4CD42B678AFE833FCFB8EC0A9" blockId="3.[151,1437,2005,2030]" lastBlockId="4.[151,1437,151,1725]" lastPageId="4" lastPageNumber="413" pageId="3" pageNumber="412">
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<emphasis id="B906EAC6CD42B67FAFE833FCFEFCC7BF" bold="true" box="[151,273,2005,2030]" pageId="3" pageNumber="412">Diagnosis.</emphasis>
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Distinguishable from other, similar congeners by a unique combination of the following characteristics: large size (longer than 13.0 mm); pronotum and base of elytra with a preponderance of very long, pale setae; pronotal pale setae forming a complete ring around disk, which appears as a large black macula; each pale median elytral fascia rather large, constricted at middle on both anterior and posterior margins.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD45B678AFB8352DFB2BC4B9" pageId="4" pageNumber="413" type="description">
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<paragraph id="8BCD36D4CD45B678AFB8352DFF17C42C" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="413">
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<emphasis id="B906EAC6CD45B678AFB8352DFDC3C14C" bold="true" box="[199,558,260,285]" pageId="4" pageNumber="413">
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Description (
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<typeStatus id="54C98876CD45B678AE20352DFE2AC14C" box="[351,455,260,285]" pageId="4" pageNumber="413" type="holotype">holotype</typeStatus>
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female).
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</emphasis>
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Length: 16.5 mm. Color: black; antennal scape testaceous in part; each elytron with a rather large pinkish–yellow fascia (Fig. 4) broadly rounded internally (interrupted before suture), constricted at about mid–elytron, then broadening laterally, where it is complete to lateral margins. Head: antennae elongate, club moderately expanded; surface rather finely, shallowly granulate–punctate, densely set with mostly short, suberect, whitish setae. Pronotum: about as broad as long, subflattened above; transverse impression obscured by vestiture; integument densely granulate–punctate; disk thickly set at middle with erect and suberect black setae of various lengths, this appearing as a large scutiform macula, circumscribed on base, sides and anterior margin by a continuous ring of thickly arrayed, robust, suberect white setae, including many of inordinate length. Elytra: elongate (ratio of length to width 37:20); base subquadrate; umbones well developed; sides subparallel, slightly, gradually narrowed behind anterior 1/5, broadly expanded behind middle; posterior 1/3 broadly, arcuately convergent to separately rounded, slightly dehiscent apices; subbasal tumescences obsolete; disk subflattened above; posterior slope moderate; integument moderately densely, deeply but rather finely and irregularly punctate and somewhat roughened, each elytron with a couple of shallow, indistinct longitudinal costae, these broadly separated and obsolete before apices; vestiture rather conspicuous: base and suture to anterior 1/3 bearing a roughly T–shaped covering of suberect, robust whitish setae of moderate length, interspersed with quite elongate robust, erect whitish setae, these angled haphazardly; black integumental surfaces thickly covered with short, suberect black setae, intermingled with longer, more robust black setae, most conspicuously on anterior 1/3; pale median fascia rather less conspicuously clothed, setae yellowish, mostly short, suberect and rather fine, interspersed with an occasional longer, more robust, erect yellowish seta; apices densely covered with moderately long, whitish adpressed setae, interspersed with longer, robust, suberect whitish setae (pale apical band eclipsed on each elytron by a large black macula). Mesosternum: posterior median process shallow. Metasternum: rather flattened, surface granulate and rugulose, densely set with rather long, whitish setae. Abdomen: sculpted as on metasternum, though vestiture somewhat less dense; visible sternite 5 with sides tapered posteriorly, hind margin rather broadly and feebly arcuate–emarginate; visible sternite 6 and visible tergite 6 with hind margins conjointly, semicircularly rounded.
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</paragraph>
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<paragraph id="8BCD36D4CD45B678AFB830AEFB2BC4B9" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="413">
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<emphasis id="B906EAC6CD45B678AFB830AEFED1C4F1" bold="true" box="[199,316,1159,1184]" pageId="4" pageNumber="413">Variation.</emphasis>
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The male
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<typeStatus id="54C98876CD45B678AECA30A0FDFBC4F0" box="[437,534,1161,1185]" pageId="4" pageNumber="413" type="paratype">paratype</typeStatus>
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measures 13.0 mm in length. The shape of visible sternite 6 is ambiguous in this specimen, but it appears to be distinctly shorter than visible tergite 6, which has its posterior margin rather broadly subtruncate in dorsal aspect, and its underside concave. The male pygidium lacks setal daggers.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD45B678AFB830DDFE54C504" pageId="4" pageNumber="413" type="materials_examined">
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<paragraph id="8BCD36D4CD45B678AFB830DDFE54C504" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="413">
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<materialsCitation id="3B1A3C89CD45B678AFB830DDFEA1C561" ID-GBIF-Occurrence="1434499453" collectingDate="1992-07-11" collectionCode="CSCA" collectorName="Muhle & Kifernwald & Eichen" country="Mexico" location="Type" pageId="4" pageNumber="413" specimenCount="2" stateProvince="Oaxaca" typeStatus="holotype">
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<emphasis id="B906EAC6CD45B678AFB830DDFE67C55C" bold="true" box="[199,394,1268,1293]" pageId="4" pageNumber="413">
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<location id="8EAD600FCD45B678AFB830DDFEEFC55C" LSID="urn:lsid:plazi:treatment:03DB87C2CD42B678AFE83343FE9DC6ED:8EAD600FCD45B678AFB830DDFEEFC55C" box="[199,258,1268,1293]" country="Mexico" name="Type" pageId="4" pageNumber="413" stateProvince="Oaxaca">
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<typeStatus id="54C98876CD45B678AFB830DDFEEFC55C" box="[199,258,1268,1293]" pageId="4" pageNumber="413">Type</typeStatus>
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</location>
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specimens.
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</emphasis>
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<typeStatus id="54C98876CD45B678AEED30DDFE12C55C" box="[402,511,1268,1293]" pageId="4" pageNumber="413" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F3657644CD45B678AD7530DDFD8FC55C" box="[522,610,1268,1293]" name="Mexico" pageId="4" pageNumber="413">México</collectingCountry>
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,
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<collectingRegion id="49B6F836CD45B678AD1130DDFD28C55D" box="[622,709,1268,1292]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="413">Oaxaca</collectingRegion>
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,
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<locationDeviation id="9A7F2FF6CD45B678ADAE30DDFC20C55C" box="[721,973,1268,1293]" pageId="4" pageNumber="413">
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||
<quantity id="4C8A9B31CD45B678ADAE30DDFCF4C55C" box="[721,793,1268,1293]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="413" unit="km" value="10.0">10 km</quantity>
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W Tequisistlán
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</locationDeviation>
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,
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<collectingDate id="EF88E9FCCD45B678ACA630DDFBB7C55D" box="[985,1114,1268,1292]" pageId="4" pageNumber="413" value="1992-07-11">
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<date id="FFCC1014CD45B678ACA630DDFBB7C55D" box="[985,1114,1268,1292]" pageId="4" pageNumber="413" value="1992-07-11">11–7–1992</date>
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</collectingDate>
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; leg.
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<collectorName id="26875302CD45B678ABE830DDFB0EC55C" box="[1175,1251,1268,1293]" pageId="4" pageNumber="413">Mühle</collectorName>
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,
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<collectorName id="26875302CD45B678AB9030DDFA83C55C" box="[1263,1390,1268,1293]" pageId="4" pageNumber="413">Kifernwald</collectorName>
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mit
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<collectorName id="26875302CD45B678AFE8313EFF08C561" box="[151,229,1303,1328]" pageId="4" pageNumber="413">Eichen</collectorName>
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(
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<collectionCode id="ED63AE11CD45B678AF8A3131FEA9C561" box="[245,324,1304,1328]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="4" pageNumber="413">CSCA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B1A3C89CD45B678AE273130FE54C504" ID-GBIF-Occurrence="1434499455" collectingDate="1992-07-11" collectionCode="JNRC" collectorName="C. L. Bellamy" country="Mexico" elevation="1280" location="Hwy" pageId="4" pageNumber="413" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
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<typeStatus id="54C98876CD45B678AE273130FE52C560" box="[344,447,1305,1329]" pageId="4" pageNumber="413" type="paratype">Paratype</typeStatus>
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: 1 (
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<collectionCode id="ED63AE11CD45B678AE963131FDDEC561" box="[489,563,1304,1328]" pageId="4" pageNumber="413">JNRC</collectionCode>
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),
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<collectingCountry id="F3657644CD45B678AD38313EFD72C561" box="[583,671,1303,1328]" name="Mexico" pageId="4" pageNumber="413">México</collectingCountry>
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,
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<collectingRegion id="49B6F836CD45B678ADD33131FCEEC561" box="[684,771,1304,1328]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="413">Oaxaca</collectingRegion>
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,
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<location id="8EAD600FCD45B678AC703130FCABC560" LSID="urn:lsid:plazi:treatment:03DB87C2CD42B678AFE83343FE9DC6ED:8EAD600FCD45B678AC703130FCABC560" box="[783,838,1305,1329]" country="Mexico" name="Hwy" pageId="4" pageNumber="413" stateProvince="Oaxaca">Hwy</location>
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<locationDeviation id="9A7F2FF6CD45B678AC303131FB3CC561" box="[847,1233,1303,1328]" pageId="4" pageNumber="413">
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||
<quantity id="4C8A9B31CD45B678AC303131FC22C561" box="[847,975,1303,1328]" metricMagnitude="5" metricUnit="m" metricValue="1.9011" pageId="4" pageNumber="413" unit="km" value="190.11">190, 11 km</quantity>
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WSW Las Margaritas
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</locationDeviation>
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,
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<quantity id="4C8A9B31CD45B678ABA23131FADBC561" box="[1245,1334,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="413" unit="m" value="1280.0">
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||
<elevation id="005FD1E7CD45B678ABA23131FADBC561" box="[1245,1334,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="413" unit="m" value="1280.0">1280 m</elevation>
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</quantity>
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,
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<collectingDate id="EF88E9FCCD45B678AA3C313EFF3FC505" pageId="4" pageNumber="413" value="1992-07-11">
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<date id="FFCC1014CD45B678AA3C313EFF3FC505" pageId="4" pageNumber="413" value="1992-07-11">July 11, 1992</date>
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</collectingDate>
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,
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<collectorName id="26875302CD45B678AFA23115FE91C504" box="[221,380,1340,1365]" pageId="4" pageNumber="413">C. L. Bellamy</collectorName>
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, coll.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C368655FCD45B678AFB83176FCD2C529" box="[199,831,1375,1400]" pageId="4" pageNumber="413" type="distribution">
|
||
<paragraph id="8BCD36D4CD45B678AFB83176FCD2C529" blockId="4.[151,1437,151,1725]" box="[199,831,1375,1400]" pageId="4" pageNumber="413">
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<emphasis id="B906EAC6CD45B678AFB83176FE8DC529" bold="true" box="[199,352,1375,1400]" pageId="4" pageNumber="413">Distribution.</emphasis>
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Isthmus of Tehuantepec,
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<collectingRegion id="49B6F836CD45B678ADFF3149FD3AC529" box="[640,727,1376,1400]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="413">Oaxaca</collectingRegion>
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,
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<collectingRegion id="49B6F836CD45B678AD9D3176FCD6C529" box="[738,827,1375,1400]" country="Mexico" name="Mexico" pageId="4" pageNumber="413">
|
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<collectingCountry id="F3657644CD45B678AD9D3176FCD6C529" box="[738,827,1375,1400]" name="Mexico" pageId="4" pageNumber="413">México</collectingCountry>
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</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD45B678AFB831ADFF12C591" pageId="4" pageNumber="413" type="etymology">
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<paragraph id="8BCD36D4CD45B678AFB831ADFF12C591" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="413">
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<emphasis id="B906EAC6CD45B678AFB831ADFEA3C5CC" bold="true" box="[199,334,1412,1437]" pageId="4" pageNumber="413">Etymology.</emphasis>
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The specific name “
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<taxonomicName id="4C724D57CD45B678AD4631ADFD78C5CC" box="[569,661,1412,1437]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="incanus">incanus</taxonomicName>
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,” Latin for “hoary,” refers to this species’ distinctive elongate white vestiture.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD45B678AFB831E5FB83C5B4" box="[199,1134,1484,1509]" pageId="4" pageNumber="413" type="biology_ecology">
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||
<paragraph id="8BCD36D4CD45B678AFB831E5FB83C5B4" blockId="4.[151,1437,151,1725]" box="[199,1134,1484,1509]" pageId="4" pageNumber="413">
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||
<emphasis id="B906EAC6CD45B678AFB831E5FECAC5B4" bold="true" box="[199,295,1484,1509]" pageId="4" pageNumber="413">Biology.</emphasis>
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Based on label information, the
|
||
<typeStatus id="54C98876CD45B678ADEA31E5FD1AC5B4" box="[661,759,1484,1509]" pageId="4" pageNumber="413" type="holotype">holotype</typeStatus>
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was collected in pine–oak forest.
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</paragraph>
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</subSubSection>
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<subSubSection id="C368655FCD45B678AFB831C6FE9DC6ED" pageId="4" pageNumber="413" type="discussion">
|
||
<paragraph id="8BCD36D4CD45B678AFB831C6FE9DC6ED" blockId="4.[151,1437,151,1725]" pageId="4" pageNumber="413">
|
||
<emphasis id="B906EAC6CD45B678AFB831C6FEA1C659" bold="true" box="[199,332,1519,1544]" pageId="4" pageNumber="413">Discussion.</emphasis>
|
||
I examined two specimens of an undescribed species of
|
||
<taxonomicName id="4C724D57CD45B678AC8731C6FB87C659" box="[1016,1130,1519,1544]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B906EAC6CD45B678AC8731C6FB87C659" box="[1016,1130,1519,1544]" italics="true" pageId="4" pageNumber="413">Enoclerus</emphasis>
|
||
</taxonomicName>
|
||
from Chamela in
|
||
<collectingRegion id="49B6F836CD45B678AA3631C6FA75C659" box="[1353,1432,1519,1544]" country="Mexico" name="Jalisco" pageId="4" pageNumber="413">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B6F836CD45B678AFE8323DFF01C67C" box="[151,236,1556,1581]" country="Mexico" name="Mexico" pageId="4" pageNumber="413">
|
||
<collectingCountry id="F3657644CD45B678AFE8323DFF01C67C" box="[151,236,1556,1581]" name="Mexico" pageId="4" pageNumber="413">México</collectingCountry>
|
||
</collectingRegion>
|
||
that are very similar to
|
||
<taxonomicName id="4C724D57CD45B678AE89323CFD81C67D" box="[502,620,1557,1580]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="incanus">
|
||
<emphasis id="B906EAC6CD45B678AE89323CFD81C67D" box="[502,620,1557,1580]" italics="true" pageId="4" pageNumber="413">E. incanus</emphasis>
|
||
</taxonomicName>
|
||
. They differ primarily by the shape of the median elytral fascia, which is constricted only on the anterior margin, rather than both anteriorly and posteriorly. I have not included them in the
|
||
<typeStatus id="54C98876CD45B678AFE83274FF2AC624" box="[151,199,1629,1653]" pageId="4" pageNumber="413">type</typeStatus>
|
||
series because this feature, combined with these specimens’ disjunct distribution in coastal
|
||
<collectingRegion id="49B6F836CD45B678ABA13275FAC0C624" box="[1246,1325,1628,1653]" country="Mexico" name="Jalisco" pageId="4" pageNumber="413">Jalisco</collectingRegion>
|
||
, seems to militate against their conspecificity with
|
||
<taxonomicName id="4C724D57CD45B678AD0A32A8FD02C6C9" box="[629,751,1665,1688]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="incanus">
|
||
<emphasis id="B906EAC6CD45B678AD0A32A8FD02C6C9" box="[629,751,1665,1688]" italics="true" pageId="4" pageNumber="413">E. incanus</emphasis>
|
||
</taxonomicName>
|
||
. Further collecting and molecular analysis may eventually resolve their status.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |