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<document id="4E11258466B301A7B18D4651AA96D2D5" ID-CLB-Dataset="26748" ID-DOI="10.11646/zootaxa.4413.1.2" ID-GBIF-Dataset="e9339680-76a0-4c9d-90fd-ecab97b50d9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3066636" ID-ZooBank="D65DAEB6-0499-44F9-A70D-0720BA71D520" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558414641282" checkinUser="plazi" docAuthor="Sassi, Davide" docDate="2018" docId="03DB8796322E9836FF481020FC96FEC4" docLanguage="en" docName="zootaxa.4413.1.2.pdf" docOrigin="Zootaxa 4413 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metallactus bellatrix Sassi 2018, sp. nov." docType="treatment" docUuid="0F40ADCB-9911-4447-A095-E4084B84B03A" docUuidSource="ZooBank" docVersion="12" lastPageNumber="94" masterDocId="FFE2FFEE320B9810FFDF1107FFBCFFD9" masterDocTitle="Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae)" masterLastPageNumber="110" masterPageNumber="57" pageNumber="94" updateTime="1698731295107" updateUser="ExternalLinkService">
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<mods:title id="81DC72B2E1D6237AB6397A56864D24C3">Revision of the Metallactus kollari species-group with a new diagnosis of the genus (Coleoptera: Chrysomelidae: Cryptocephalinae)</mods:title>
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<mods:namePart id="BD09A576AC26BA67A49FE24CB991C4C8">Sassi, Davide</mods:namePart>
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<mods:date id="FDFB37F967CF83EBB925F06646E5E0EC">2018</mods:date>
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<mods:number id="BE1C4E2006474B84DDD078F1D5C24EF1">2018-04-20</mods:number>
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<mods:number id="45A3883BA3542ED3DC100DFFAD2EABFC">4413</mods:number>
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<mods:classification id="D31505C56F9AC250664BCA087FE3AA11">journal article</mods:classification>
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<mods:identifier id="28EA086E8123C866AB3CE65B3C57016B" type="DOI">10.11646/zootaxa.4413.1.2</mods:identifier>
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<treatment id="03DB8796322E9836FF481020FC96FEC4" ID-DOI="http://doi.org/10.5281/zenodo.5970582" ID-GBIF-Taxon="157232644" ID-Zenodo-Dep="5970582" LSID="urn:lsid:zoobank.org:act:0F40ADCB-9911-4447-A095-E4084B84B03A" httpUri="http://treatment.plazi.org/id/03DB8796322E9836FF481020FC96FEC4" lastPageId="38" lastPageNumber="94" pageId="37" pageNumber="94">
<subSubSection id="C368650B322E9835FF481020FE49FE9B" box="[151,501,295,322]" pageId="37" pageNumber="94" type="nomenclature">
<paragraph id="8BCD3680322E9835FF481020FE49FE9B" blockId="37.[151,501,295,355]" box="[151,501,295,322]" pageId="37" pageNumber="94">
<heading id="D08581EC322E9835FF481020FE49FE9B" bold="true" box="[151,501,295,322]" fontSize="11" level="1" pageId="37" pageNumber="94" reason="1">
<emphasis id="B906EA92322E9835FF481020FE49FE9B" bold="true" box="[151,501,295,322]" pageId="37" pageNumber="94">
<taxonomicName id="4C724D03322E9835FF481020FE2CFE98" authority="Sassi, 2018" authorityName="Sassi" authorityYear="2018" box="[151,400,295,321]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="bellatrix" status="sp. nov.">
<emphasis id="B906EA92322E9835FF481020FE2CFE98" bold="true" box="[151,400,295,321]" italics="true" pageId="37" pageNumber="94">Metallactus bellatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557E9322E9835FE47102FFE49FE9B" box="[408,501,296,322]" pageId="37" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9835FF48104DFE9BFEBA" box="[151,295,330,355]" pageId="37" pageNumber="94" type="description">
<paragraph id="8BCD3680322E9835FF48104DFE9BFEBA" blockId="37.[151,501,295,355]" box="[151,295,330,355]" pageId="37" pageNumber="94">
(
<figureCitation id="13492A05322E9835FF40104DFF4AFEBA" box="[159,246,330,355]" captionStart="FIGURES 1314" captionStartId="36.[151,264,1883,1906]" captionTargetBox="[224,1371,202,1844]" captionTargetId="figure@36.[181,1405,193,1862]" captionTargetPageId="36" captionText="FIGURES 1314. M. rileyi (13), M. bellatrix (14). 13ab: habitus; 13ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [1.0 mm]; 13f: sclerites IIIIIIV of endophallus [0.6 mm]; 13g: sclerite I and dorsal spicule of endophallus [0.9 mm]; 13h: spermatheca [0.8 mm]; 13km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]; 13n: antenna [1.5 mm]. 14ab: habitus; 14h: spermatheca [0.7 mm]; 14km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]. 13ab: holotype; 14ab: holotype. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066650/files/figure.png" pageId="37" pageNumber="94">Figs 14</figureCitation>
;
<figureCitation id="13492A05322E9835FEDD104CFEA3FEBA" box="[258,287,331,355]" captionStart="FIGURES 2028" captionStartId="50.[151,264,1944,1967]" captionTargetBox="[222,1372,226,1871]" captionTargetId="figure@50.[187,1400,195,1923]" captionTargetPageId="50" captionText="FIGURES 2028. Distribution maps for Metallactus kollari species-group. M. affinis (20); M. albifrons (21); M. albipes (22); M. bellatrix (23); M. bezoar (24); M. corruptus (25); M. dodecastictus (26); M. guarani (27); M. kollari (28). Grey squares: localities reported in a generic way." httpUri="https://zenodo.org/record/3066658/files/figure.png" pageId="37" pageNumber="94">23</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9835FF481095FD55FE2A" pageId="37" pageNumber="94" type="etymology">
<paragraph id="8BCD3680322E9835FF481095FD55FE2A" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">
<emphasis id="B906EA92322E9835FF481095FEA2FE72" bold="true" box="[151,286,402,427]" pageId="37" pageNumber="94">Etymology.</emphasis>
Bellatrix is latin for “female warrior”, stressing the fact that only females were available for the study. The word
<taxonomicName id="4C724D03322E9835FED610B1FEDBFE17" authorityName="Sassi" authorityYear="2018" box="[265,359,438,462]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="bellatrix">
<emphasis id="B906EA92322E9835FED610B1FEDBFE17" box="[265,359,438,462]" italics="true" pageId="37" pageNumber="94">bellatrix</emphasis>
</taxonomicName>
is intended here as a noun in the nominative singular, standing in apposition to the generic name (articles 11.9.1.2 and 31.2.1 of I.C.Z.N., 4th edition).
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9835FF1810F9FD6AFD7E" pageId="37" pageNumber="94" type="materials_examined">
<paragraph id="8BCD3680322E9835FF1810F9FB40FD5A" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">
<emphasis id="B906EA92322E9835FF1810F9FEC0FDCE" bold="true" box="[199,380,510,535]" pageId="37" pageNumber="94">Type material.</emphasis>
<typeStatus id="54C98822322E9835FE551305FDBCFDCC" box="[394,512,514,533]" pageId="37" pageNumber="94" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D74FD09322E9835FDC910F8FD97FDCF" box="[534,555,511,534]" pageId="37" pageNumber="94" type="female"></specimenCount>
, // “
<collectingCountry id="F3657610322E9835FDB710F9FD11FDCE" box="[616,685,510,535]" name="Brazil" pageId="37" pageNumber="94">Brasil</collectingCountry>
Santa
<taxonomicName id="4C724D03322E9835FCD810F9FC69FDCE" authority="Corupa" authorityName="Corupa" box="[775,981,510,535]" class="Polyplacophora" family="Mopaliidae" genus="Catharina" kingdom="Animalia" order="Chitonida" pageId="37" pageNumber="94" phylum="Mollusca" rank="genus">Catharina Corupá</taxonomicName>
<date id="FFCC1040322E9835FC3C10F8FB88FDCE" box="[995,1076,511,535]" pageId="37" pageNumber="94" value="1949-02">II.1949</date>
col. A. Maller” [white label, handwritten] // “F. Monrós collection 1959” [white label, printed] // “
<taxonomicName id="4C724D03322E9835FC7B1324FB33FDE2" authority="Sassi, 2018" authorityName="Sassi" authorityYear="2018" box="[932,1167,547,571]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="bellatrix" status="sp. nov.">
<emphasis id="B906EA92322E9835FC7B1324FB33FDE2" box="[932,1167,547,571]" italics="true" pageId="37" pageNumber="94">Metallactus bellatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557E9322E9835FB491324FB51FDE3" box="[1174,1261,547,570]" pageId="37" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
<typeStatus id="54C98822322E9835FB2B1321FAC5FDE0" box="[1268,1401,550,569]" pageId="37" pageNumber="94" type="holotype">HOLOTYPUS</typeStatus>
D. Sassi des.” [red label, printed] // (
<collectionCode id="ED63AE45322E9835FDD61340FD35FD86" box="[521,649,583,607]" pageId="37" pageNumber="94">USNMNH</collectionCode>
).
<typeStatus id="54C98822322E9835FD44134DFCA7FD84" box="[667,795,586,605]" pageId="37" pageNumber="94" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D74FD09322E9835FCF61340FCEEFD87" box="[809,850,583,607]" pageId="37" pageNumber="94" type="female">1 ♀</specimenCount>
, same data of the
<typeStatus id="54C98822322E9835FBC41341FB3CFD86" box="[1051,1152,582,607]" pageId="37" pageNumber="94" type="holotype">holotype</typeStatus>
, provided with additional label: // “
<taxonomicName id="4C724D03322E9835FF20136CFE55FD5A" authority="Sassi, 2018" authorityName="Sassi" authorityYear="2018" box="[255,489,619,643]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="bellatrix" status="sp. nov.">
<emphasis id="B906EA92322E9835FF20136CFE55FD5A" box="[255,489,619,643]" italics="true" pageId="37" pageNumber="94">Metallactus bellatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23557E9322E9835FE2F136CFDFBFD5B" box="[496,583,619,642]" pageId="37" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
<typeStatus id="54C98822322E9835FD921369FD6CFD58" box="[589,720,622,641]" pageId="37" pageNumber="94" type="paratype">PARATYPUS</typeStatus>
D. Sassi des.” [red label, printed] // (
<collectionCode id="ED63AE45322E9835FBAF136CFB4CFD5A" box="[1136,1264,619,643]" pageId="37" pageNumber="94">USNMNH</collectionCode>
).
</paragraph>
<paragraph id="8BCD3680322E9835FF181389FD6AFD7E" blockId="37.[151,1437,402,2011]" box="[199,726,654,679]" pageId="37" pageNumber="94">
<materialsCitation id="3B1A3CDD322E9835FF181389FD6EFD7E" ID-GBIF-Occurrence="2243028702" box="[199,722,654,679]" country="Brazil" location="Corupa" pageId="37" pageNumber="94" specimenCount="1" stateProvince="Santa Catarina" typeStatus="holotype">
<emphasis id="B906EA92322E9835FF181389FED9FD7E" bold="true" box="[199,357,654,679]" pageId="37" pageNumber="94">
<typeStatus id="54C98822322E9835FF181389FEBEFD7E" box="[199,258,654,679]" pageId="37" pageNumber="94">Type</typeStatus>
locality.
</emphasis>
<location id="8EAD605B322E9835FEB31388FE03FD7E" LSID="urn:lsid:plazi:treatment:03DB8796322E9836FF481020FC96FEC4:8EAD605B322E9835FEB31388FE03FD7E" box="[364,447,655,679]" country="Brazil" name="Corupa" pageId="37" pageNumber="94" stateProvince="Santa Catarina">Corupá</location>
(
<collectingRegion id="49B6F862322E9835FE121388FDCAFD7E" box="[461,630,654,679]" country="Brazil" name="Santa Catarina" pageId="37" pageNumber="94">Santa Catarina</collectingRegion>
,
<collectingCountry id="F3657610322E9835FD5E1389FD77FD7E" box="[641,715,654,679]" name="Brazil" pageId="37" pageNumber="94">Brazil</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9835FF1813B5FE0FFD12" box="[199,435,690,715]" pageId="37" pageNumber="94" type="distribution">
<paragraph id="8BCD3680322E9835FF1813B5FE0FFD12" blockId="37.[151,1437,402,2011]" box="[199,435,690,715]" pageId="37" pageNumber="94">
<emphasis id="B906EA92322E9835FF1813B5FEDCFD12" bold="true" box="[199,352,690,715]" pageId="37" pageNumber="94">Distribution.</emphasis>
<collectingCountry id="F3657610322E9835FEB813B5FE13FD12" box="[359,431,690,715]" name="Brazil" pageId="37" pageNumber="94">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9835FF1813D1FAEEFBD6" pageId="37" pageNumber="94" type="diagnosis">
<paragraph id="8BCD3680322E9835FF1813D1FAEEFBD6" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">
<emphasis id="B906EA92322E9835FF1813D1FEFDFD36" bold="true" box="[199,321,726,751]" pageId="37" pageNumber="94">Diagnosis.</emphasis>
A
<taxonomicName id="4C724D03322E9835FEB313D1FE4CFD37" authorityName="Suffrian" authorityYear="1866" box="[364,496,726,750]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="genus">
<emphasis id="B906EA92322E9835FEB313D1FE4CFD37" box="[364,496,726,750]" italics="true" pageId="37" pageNumber="94">Metallactus</emphasis>
</taxonomicName>
of medium size, characterized by a fairly reduced black dorsal pattern. At first glance, this species looks very similar to
<taxonomicName id="4C724D03322E9835FDA013FCFCABFCCA" authorityName="Suffrian" authorityYear="1866" box="[639,791,763,787]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="languidus">
<emphasis id="B906EA92322E9835FDA013FCFCABFCCA" box="[639,791,763,787]" italics="true" pageId="37" pageNumber="94">M. languidus</emphasis>
</taxonomicName>
, but it can be distinguished by the stronger and closer pronotal and elytral punctation and the lack of impressions on sides of pronotum. Pronotal shape is also very distinctive in having lateral margin distinctly larger than in all the other species of the group, with the exception of
<taxonomicName id="4C724D03322E9835FF481260FEAAFCA7" authorityName="Suffrian" authorityYear="1866" box="[151,278,870,894]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="spilotus">
<emphasis id="B906EA92322E9835FF481260FEAAFCA7" box="[151,278,870,894]" italics="true" pageId="37" pageNumber="94">M. spilotus</emphasis>
</taxonomicName>
which is easily distinguishable by the very different general shape and dorsal colour pattern. Light colored specimens of
<taxonomicName id="4C724D03322E9835FE4A128CFD96FC7B" authorityName="Suffrian" authorityYear="1866" box="[405,554,907,930]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="corruptus">
<emphasis id="B906EA92322E9835FE4A128CFD96FC7B" box="[405,554,907,930]" italics="true" pageId="37" pageNumber="94">M. corruptus</emphasis>
</taxonomicName>
can have a quite similar look, but they are easily diagnosed by the broadly yellow ventral side. The species looks somehow similar to
<taxonomicName id="4C724D03322E9835FCF312A8FBEAFC1E" authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[812,1110,942,967]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="verrinus">
<emphasis id="B906EA92322E9835FCF312A8FC12FC1F" box="[812,942,943,966]" italics="true" pageId="37" pageNumber="94">M. verrinus</emphasis>
<bibRefCitation id="EFE34B71322E9835FC6A12A9FBEAFC1E" author="Suffrian, E." box="[949,1110,942,967]" pageId="37" pageNumber="94" pagination="1 - 483" refId="ref45438" refString="Suffrian, E. (1866) Zur Kenntniss der Sudamerikanischen Cryptocephalen. Linnaea Entomologica, 16, 1 - 483." type="journal article" year="1866">Suffrian, 1866</bibRefCitation>
</taxonomicName>
which has not been included in
<taxonomicName id="4C724D03322E9835FF6812D4FE97FC32" authorityName="Perty" authorityYear="1832" box="[183,299,979,1003]" class="Insecta" family="Chrysomelidae" genus="Metallactus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="94" phylum="Arthropoda" rank="species" species="kollari">
<emphasis id="B906EA92322E9835FF6812D4FE97FC32" box="[183,299,979,1003]" italics="true" pageId="37" pageNumber="94">M. kollari</emphasis>
</taxonomicName>
group because of the very different aedeagal shape. The latter is also different in having a much weaker dorsal puncturation, the pronotum slightly vaulted with narrower margins, lighter ventral coloration.
</paragraph>
</subSubSection>
<subSubSection id="C368650B322E9836FF18151DFC96FEC4" lastPageId="38" lastPageNumber="95" pageId="37" pageNumber="94" type="description">
<paragraph id="8BCD3680322E9835FF18151DFC7FFB8F" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">
<emphasis id="B906EA92322E9835FF18151DFE7EFBEA" bold="true" box="[199,450,1050,1075]" italics="true" pageId="37" pageNumber="94">Description of female</emphasis>
(male unknown). Habitus in
<figureCitation id="13492A05322E9835FCD0151DFCE1FBEA" box="[783,861,1050,1075]" captionStart="FIGURES 1314" captionStartId="36.[151,264,1883,1906]" captionTargetBox="[224,1371,202,1844]" captionTargetId="figure@36.[181,1405,193,1862]" captionTargetPageId="36" captionText="FIGURES 1314. M. rileyi (13), M. bellatrix (14). 13ab: habitus; 13ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [1.0 mm]; 13f: sclerites IIIIIIV of endophallus [0.6 mm]; 13g: sclerite I and dorsal spicule of endophallus [0.9 mm]; 13h: spermatheca [0.8 mm]; 13km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]; 13n: antenna [1.5 mm]. 14ab: habitus; 14h: spermatheca [0.7 mm]; 14km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]. 13ab: holotype; 14ab: holotype. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066650/files/figure.png" pageId="37" pageNumber="94">Fig. 14</figureCitation>
ab (HT). BL =
<quantity id="4C8A9B65322E9835FBC5151DFB18FBEB" box="[1050,1188,1050,1075]" metricMagnitude="-3" metricUnit="m" metricValue="5.699999999999999" metricValueMax="5.8" metricValueMin="5.6" pageId="37" pageNumber="94" unit="mm" value="5.699999999999999" valueMax="5.8" valueMin="5.6">5.65.8 mm</quantity>
, BW =
<quantity id="4C8A9B65322E9835FB21151CFAEBFBEB" box="[1278,1367,1051,1075]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="37" pageNumber="94" unit="mm" value="3.4">3.4 mm</quantity>
, PL =
<quantity id="4C8A9B65322E9835FF481538FE9CFB8F" box="[151,288,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="37" pageNumber="94" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.71.8 mm</quantity>
, PW = 3.0
<quantity id="4C8A9B65322E9835FE791538FE41FB8F" box="[422,509,1087,1111]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="37" pageNumber="94" unit="mm" value="3.1">3.1 mm</quantity>
. Interocular distance 11.713.5% of BL.
</paragraph>
<paragraph id="8BCD3680322E9835FF181565FB77FB3E" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">Head black with a pair of faint yellow oblique “eyebrows” above the upper rim of eyes. Surface matt with scattered punctures fairly impressed on frons and clypeus, slightly smaller on vertex. Mid-cranial suture extended from mid of frons to mid part of vertex. Setosity sparse, limited to clypeus and labrum and near inner rim of eyes. Proximal antennomeres black. Four apical antennomeres missing in both the studied specimens.</paragraph>
<paragraph id="8BCD3680322E9835FF1815F4FCF8FAAE" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">Pronotum yellow with two minute rounded black spots on disc. Pronotal shape trapezoidal, rather transversally arranged. Lateral margins large with maximum width behind half of length. Posterolateral impressions almost obliterated, so that posterior margin barely swollen toward sides. Surface moderately lustrous with scattered, fairly impressed punctation, almost evenly dense on sides and disc.</paragraph>
<paragraph id="8BCD3680322E9835FF181485FC28FA66" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">Scutellum completely black, moderately raised, with truncated apex and surface moderately lustrous, smooth, bald, with only few scattered tiny punctures close to lateral margins.</paragraph>
<paragraph id="8BCD3680322E9835FF1814CDFEC2F996" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">Elytra yellow with black pattern consisting of three tiny black rounded spots, first one on humerous, second one just behind half, closer to suture than to lateral margin, third one near apex. Suture narrowly black. Surface dull with rather strong and dense punctation in particular in anterior half. Remnants of striae only on sides and apical clivus. Intervals flat.</paragraph>
<paragraph id="8BCD3680322E9835FF18175DFAF3F9AB" blockId="37.[151,1437,402,2011]" box="[199,1359,1626,1651]" pageId="37" pageNumber="94">Pygidium completely yellow with surface smooth, evenly and minutely punctate with sparse pale setae.</paragraph>
<paragraph id="8BCD3680322E9835FF181779FBE8F8FE" blockId="37.[151,1437,402,2011]" pageId="37" pageNumber="94">Inferior parts black with external part of hypomera very narrowly and lateral sides of abdominal sternites widely yellow. All ventral surface covered by scattered whitish setae, quite regularly distributed, somewhat longer on metathorax. Punctuation evenly arranged, coarser on hypomera and sides of mesothorax, minute on abdominal sternites. Prosternal process longitudinally grooved, with rounded to roughly pointed and slightly raised apex and surface covered with long dense pale setae. Legs totally black, or tibiae partly lighter.</paragraph>
<paragraph id="8BCD3680322E9836FF181635FC96FEC4" blockId="37.[151,1437,402,2011]" lastBlockId="38.[151,1436,151,285]" lastPageId="38" lastPageNumber="95" pageId="37" pageNumber="94">
Fifth abdominal ventrite with rather deep, rounded pit. Bottom of pit matt, impunctate but covered by tiny wrinkles. Spermatheca (
<figureCitation id="13492A05322E9835FE6F1651FDA8F8B6" box="[432,532,1878,1903]" captionStart="FIGURES 1314" captionStartId="36.[151,264,1883,1906]" captionTargetBox="[224,1371,202,1844]" captionTargetId="figure@36.[181,1405,193,1862]" captionTargetPageId="36" captionText="FIGURES 1314. M. rileyi (13), M. bellatrix (14). 13ab: habitus; 13ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [1.0 mm]; 13f: sclerites IIIIIIV of endophallus [0.6 mm]; 13g: sclerite I and dorsal spicule of endophallus [0.9 mm]; 13h: spermatheca [0.8 mm]; 13km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]; 13n: antenna [1.5 mm]. 14ab: habitus; 14h: spermatheca [0.7 mm]; 14km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]. 13ab: holotype; 14ab: holotype. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066650/files/figure.png" pageId="37" pageNumber="94">Fig. 14h</figureCitation>
) with lower branch of vasculum slenderer than the upper one, slightly bent backwards. Upper branch darkened, swollen on its middle part and quite abruptly tapered in a sharp, bent downward tip. Ampulla pigmented, squared, rather small, shifted on dorsal side of vasculum. Duct regularly coiled, uniform in thickness, slender. Coils only slightly loosened in the last section. Insertion on bursa copulatrix almost straight, weakly swollen and weakly pigmented. Rectal apparatus (
<figureCitation id="13492A05322D9836FBCF1190FBD9FF69" box="[1040,1125,151,176]" captionStart="FIGURES 1314" captionStartId="36.[151,264,1883,1906]" captionTargetBox="[224,1371,202,1844]" captionTargetId="figure@36.[181,1405,193,1862]" captionTargetPageId="36" captionText="FIGURES 1314. M. rileyi (13), M. bellatrix (14). 13ab: habitus; 13ce: median lobe of aedeagus in ventral (c), dorsal (d) and lateral (e) view [1.0 mm]; 13f: sclerites IIIIIIV of endophallus [0.6 mm]; 13g: sclerite I and dorsal spicule of endophallus [0.9 mm]; 13h: spermatheca [0.8 mm]; 13km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]; 13n: antenna [1.5 mm]. 14ab: habitus; 14h: spermatheca [0.7 mm]; 14km: rectal apparatus in dorsal (k) and ventral (m) view [0.9 mm]. 13ab: holotype; 14ab: holotype. Square brackets: length of scale segment." httpUri="https://zenodo.org/record/3066650/files/figure.png" pageId="38" pageNumber="95">Fig. 14</figureCitation>
km) with dorsal sclerites laterally not projected beyond rectus, abruptly tapered towards median line, with apodemes large, hyaline, bent upwards and leaning against rectum. Ventral sclerite quite deep, almost evenly pigmented. Apodemes of ventral sclerite moderately developed, slightly projecting laterally.
</paragraph>
</subSubSection>
</treatment>
</document>