297 lines
42 KiB
XML
297 lines
42 KiB
XML
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<mods:title id="16567BA750D8BAF1BAB14656C5D50DCA">Revision and phylogeny of Hypotelus Erichson: a Neotropical genus of minute rove beetles (Coleoptera, Staphylinidae, Piestinae)</mods:title>
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<mods:namePart id="7B1EED7758A0FF4E4020207190E174D3">Caron, Edilson</mods:namePart>
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<mods:namePart id="02469E5ADD90ABC7DCB8A01D90D1CEF5">Ribeiro-Costa, Cibele S.</mods:namePart>
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<treatment id="03DB878DFF8DD527CEB5FB9ED1D9FCB0" ID-DOI="http://doi.org/10.5281/zenodo.6017748" ID-GBIF-Taxon="131812446" ID-Zenodo-Dep="6017748" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0" httpUri="http://treatment.plazi.org/id/03DB878DFF8DD527CEB5FB9ED1D9FCB0" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478">
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<paragraph id="8BCD369BFF8DD520CEB5FB9ED008FB93" blockId="27.[151,609,1052,1111]" box="[151,546,1052,1078]" pageId="27" pageNumber="478">
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<heading id="D08581F7FF8DD520CEB5FB9ED008FB93" bold="true" box="[151,546,1052,1078]" fontSize="11" level="1" pageId="27" pageNumber="478" reason="1">
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<taxonomicName id="4C724D18FF8DD520CEB5FB9ED008FB93" ID-CoL="3NZDK" authority="Bierig, 1934" authorityName="Bierig" authorityYear="1934" box="[151,546,1052,1078]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">
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<emphasis id="B906EA89FF8DD520CEB5FB9ED008FB93" bold="true" box="[151,546,1052,1078]" pageId="27" pageNumber="478">
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<emphasis id="B906EA89FF8DD520CEB5FB9ED3AEFB93" bold="true" box="[151,388,1052,1078]" italics="true" pageId="27" pageNumber="478">Hypotelus testaceus</emphasis>
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<bibRefCitation id="EFE34B6AFF8DD520CFAEFB9ED008FB93" author="Bierig" box="[396,546,1052,1078]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">Bierig, 1934</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BCD369BFF8DD520CEB5FBBCD04BFBF2" blockId="27.[151,609,1052,1111]" box="[151,609,1086,1111]" pageId="27" pageNumber="478">
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(
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<figureCitation id="13492A1EFF8DD520CEBDFBBCD2D6FBF2" box="[159,252,1086,1111]" captionStart="FIGURES 9 – 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 – 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="27" pageNumber="478">Figs. 12</figureCitation>
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,
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<figureCitation id="13492A1EFF8DD520CF2AFBBCD30CFBF2" box="[264,294,1086,1111]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="27" pageNumber="478">53</figureCitation>
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,
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<figureCitation id="13492A1EFF8DD520CF10FBBCD351FBF2" box="[306,379,1086,1111]" captionStart="FIGURES 59 – 74" captionStartId="17.[151,264,1924,1946]" captionTargetBox="[290,1268,229,1830]" captionTargetId="figure@17.[237,1336,193,1893]" captionTargetPageId="17" captionText="FIGURES 59 – 74. Aedeagus, for each species with lateral view and ventral view, respectively. 59 – 60, Hypotelus pusillus; 61 – 62, H. insulanus; 63 – 64, H. marginatus; 65 – 66, H. testaceus; 67 – 68, H. praecox; 69 – 70, H. brevitarsus sp. nov., parameres removed in ventral view; 71 – 72, H. castaneus sp. nov.; 73 – 74, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803662/files/figure.png" pageId="27" pageNumber="478">65, 66</figureCitation>
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,
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<figureCitation id="13492A1EFF8DD520CFA4FBBDD38EFBF2" box="[390,420,1087,1111]" captionStart="FIGURES 75 – 82" captionStartId="18.[151,264,1403,1425]" captionTargetBox="[194,1368,254,1322]" captionTargetId="figure@18.[151,1436,193,1381]" captionTargetPageId="18" captionText="FIGURES 75 – 82. Sternite 8 of females in ventral view, right setae removed. 75, Hypotelus pusillus; 76, H. insulanus; 77, H. marginatus; 78 H. testaceus; 79, H. melanodelta sp. nov.; 80, H. brevitarsus sp. nov.; 81, H. castaneus sp. nov.; 82, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803664/files/figure.png" pageId="27" pageNumber="478">78</figureCitation>
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,
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<figureCitation id="13492A1EFF8DD520CF92FBBDD009FBF2" box="[432,547,1087,1111]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="27" pageNumber="478">88, 89, 90</figureCitation>
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,
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<figureCitation id="13492A1EFF8DD520CC0CFBBDD072FBF2" box="[558,600,1087,1111]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="27" pageNumber="478">114</figureCitation>
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)
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</paragraph>
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<materialsCitation id="3B1A3CC6FF8DD520CEB5FB04D0A5FB1E" ID-GBIF-Occurrence="1563407715" country="Panama" county="Herman" location="Zona" municipality="Canal" pageId="27" pageNumber="478" specimenCount="1" typeStatus="holotype">
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<treatmentCitationGroup id="AB6211B5FF8DD520CEB5FB04D0A5FB1E" pageId="27" pageNumber="478">
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<taxonomicName id="4C724D18FF8DD520CEB5FB04D004FB39" ID-CoL="3NZDK" authority="Bierig, 1934: 342" authorityName="Bierig" authorityPageNumber="342" authorityYear="1934" box="[151,558,1158,1181]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">
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<emphasis id="B906EA89FF8DD520CEB5FB04D34CFB38" box="[151,358,1158,1181]" italics="true" pageId="27" pageNumber="478">Hypotelus testaceus</emphasis>
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<treatmentCitation id="0AD3108AFF8DD520CF53FB04D004FB39" author="Bierig" box="[369,558,1158,1180]" page="342" pageId="27" pageNumber="478" year="1934">
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<bibRefCitation id="EFE34B6AFF8DD520CF53FB04D3D3FB39" author="Bierig" box="[369,505,1158,1180]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">Bierig, 1934</bibRefCitation>
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: 342
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</treatmentCitation>
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</taxonomicName>
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(original description,
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<typeStatus id="54C98839FF8DD520CD3CFB05D160FB39" box="[798,842,1159,1180]" pageId="27" pageNumber="478">type</typeStatus>
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locality: “
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<collectingCountry id="F365760BFF8DD520CD9CFB04D638FB39" box="[958,1042,1158,1180]" name="Panama" pageId="27" pageNumber="478">Panamá</collectingCountry>
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, cercanía de
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<collectingCountry id="F365760BFF8DD520CA87FB05D6C0FB39" box="[1189,1258,1159,1180]" name="France" pageId="27" pageNumber="478">France</collectingCountry>
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Field (
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<location id="8EAD6040FF8DD520CB19FB04D75BFB39" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CB19FB04D75BFB39" box="[1339,1393,1158,1180]" country="Panama" county="Herman" municipality="Canal" name="Zona" pageId="27" pageNumber="478">Zona</location>
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del
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<collectingMunicipality id="6BA9ACE1FF8DD520CEE5FB27D322FB1E" box="[199,264,1189,1211]" pageId="27" pageNumber="478">Canal</collectingMunicipality>
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)”);
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<treatmentCitation id="0AD3108AFF8DD520CF09FB24D3D7FB1E" author="Herman" box="[299,509,1189,1211]" page="1786" pageId="27" pageNumber="478" year="2001">
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<bibRefCitation id="EFE34B6AFF8DD520CF09FB24D395FB1E" author="Herman" box="[299,447,1189,1211]" pageId="27" pageNumber="478" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera) 1758 to the end of the second millennium, parts I - VII. Bulletin of the American Museum of Natural History, 167, 1 - 4218." type="journal article" year="2001">
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<collectingCounty id="62AC4E17FF8DD520CF09FB24D355FB1E" box="[299,383,1190,1211]" pageId="27" pageNumber="478">Herman</collectingCounty>
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, 2001
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</bibRefCitation>
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: 1786
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</treatmentCitation>
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(distribution).
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</materialsCitation>
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</paragraph>
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<subSubSection id="C3686510FF8DD520CEB5FB65D711F915" pageId="27" pageNumber="478" type="materials_examined">
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<paragraph id="8BCD369BFF8DD520CEB5FB65D2CEF9CD" blockId="27.[151,1437,1255,2000]" pageId="27" pageNumber="478">
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<materialsCitation id="3B1A3CC6FF8DD520CEB5FB65D6BBFB5A" ID-GBIF-Occurrence="1563407718" box="[151,1169,1255,1280]" collectionCode="FMNH" location="Four" pageId="27" pageNumber="478" specimenCount="2" typeStatus="syntype">
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<emphasis id="B906EA89FF8DD520CEB5FB65D36FFAA5" bold="true" box="[151,325,1255,1280]" pageId="27" pageNumber="478">
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<typeStatus id="54C98839FF8DD520CEB5FB65D2F8FAA5" box="[151,210,1255,1280]" pageId="27" pageNumber="478">Type</typeStatus>
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material.
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</emphasis>
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<taxonomicName id="4C724D18FF8DD520CF6EFB65D0D1FAA5" authority="Bierig, 1934: 342" authorityName="Bierig" authorityPageNumber="342" authorityYear="1934" box="[332,763,1255,1280]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">
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<emphasis id="B906EA89FF8DD520CF6EFB65D001FAA5" box="[332,555,1255,1280]" italics="true" pageId="27" pageNumber="478">Hypotelus testaceus</emphasis>
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<bibRefCitation id="EFE34B6AFF8DD520CC11FB65D0E9FAA5" author="Bierig" box="[563,707,1255,1280]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">Bierig, 1934</bibRefCitation>
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: 342
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</taxonomicName>
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.
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<location id="8EAD6040FF8DD520CD25FB6AD116FB5A" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CD25FB6AD116FB5A" box="[775,828,1256,1279]" name="Four" pageId="27" pageNumber="478">Four</location>
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<typeStatus id="54C98839FF8DD520CD61FB6AD18FFAA5" box="[835,933,1256,1280]" pageId="27" pageNumber="478" type="syntype">syntypes</typeStatus>
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deposited in
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<collectionCode id="ED63AE5EFF8DD520CA1EFB6AD6BBFB5A" box="[1084,1169,1256,1279]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="27" pageNumber="478">FMNH</collectionCode>
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</materialsCitation>
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.
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<materialsCitation id="3B1A3CC6FF8DD520CABEFB6AD7B3FAC8" ID-GBIF-Occurrence="1563407713" collectingDate="1930-06" collectorName="Brg. & Field Mus. Nat. Hist. & A. Bierig Colln. & Acc" country="Canada" location="One" pageId="27" pageNumber="478" specimenCount="1" typeStatus="holotype">
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<location id="8EAD6040FF8DD520CABEFB6AD6E0FAA5" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CABEFB6AD6E0FAA5" box="[1180,1226,1256,1280]" country="Canada" name="One" pageId="27" pageNumber="478">One</location>
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female [dissected, abdominal segments 8 to 10 were fixed on acetate plastic card and covered with
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||
<collectingCountry id="F365760BFF8DD520CA0EFA8ED6AAFA80" box="[1068,1152,1292,1317]" name="Canada" pageId="27" pageNumber="478">Canada</collectingCountry>
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||
balsam], with labels: (1) “
|
||
<collectingCountry id="F365760BFF8DD520CE83FAB2D2DAFAED" box="[161,240,1328,1352]" name="France" pageId="27" pageNumber="478">France</collectingCountry>
|
||
Field/
|
||
<collectingDate id="EF88E9B3FF8DD520CF64FAADD3B4FAED" box="[326,414,1327,1352]" pageId="27" pageNumber="478" value="1930-06">
|
||
<date id="FFCC105BFF8DD520CF64FAADD3B4FAED" box="[326,414,1327,1352]" pageId="27" pageNumber="478" value="1930-06">VI.1930</date>
|
||
</collectingDate>
|
||
/
|
||
<collectingCountry id="F365760BFF8DD520CF85FAADD029FAED" box="[423,515,1327,1352]" name="Panama" pageId="27" pageNumber="478">Panamá</collectingCountry>
|
||
” [white label, handwritten]; (2) “
|
||
<typeStatus id="54C98839FF8DD520CD92FAB2D627FAED" box="[944,1037,1328,1352]" pageId="27" pageNumber="478">TYPUS</typeStatus>
|
||
” [black label, handwritten]; (3) “
|
||
<taxonomicName id="4C724D18FF8DD520CE83FAD6D33DFAC8" box="[161,279,1364,1389]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">Hypotelus</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C724D18FF8DD520CF02FAD6D3A2FAC8" box="[288,392,1364,1389]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">testaceus</taxonomicName>
|
||
/
|
||
<collectorName id="2687534DFF8DD520CFB3FAD6D395FAC8" box="[401,447,1364,1389]" pageId="27" pageNumber="478">Brg.</collectorName>
|
||
” [white label, handwritten]; (4) “
|
||
<collectorName id="2687534DFF8DD520CD71FAD6D67DFAC8" box="[851,1111,1364,1389]" pageId="27" pageNumber="478">Field Mus. Nat. Hist.</collectorName>
|
||
/1966/
|
||
<collectorName id="2687534DFF8DD520CA83FAD6D74AFAC8" box="[1185,1376,1364,1389]" pageId="27" pageNumber="478">A. Bierig Colln.</collectorName>
|
||
/
|
||
<collectorName id="2687534DFF8DD520CB4BFAD6D7B3FAC8" box="[1385,1433,1364,1389]" pageId="27" pageNumber="478">Acc</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B1A3CC6FF8DD520CEB5FAFAD2CEF9CD" ID-GBIF-Occurrence="1563407725" collectingDate="1930-06" collectorName="Field Mus. Nat. Hist. & A. Bierig Colln. & Acc. & Bierig" country="France" county="Photographed" determinerName="S. Bortoluzzi" location="Emu Catalog" municipality="Kelsey Keaton" pageId="27" pageNumber="478" specimenCount="2" typeStatus="syntype">
|
||
<accessionNumber id="9421AB78FF8DD520CEB5FAFAD2DAFA35" box="[151,240,1400,1424]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/Z13812" pageId="27" pageNumber="478" type="EnaNcbi">Z13812</accessionNumber>
|
||
” [white label, printed in black]; (5) “FMNHINS/0000 131 009” [white label, printed in black]; (6) “
|
||
<collectingCounty id="62AC4E17FF8DD520CE82FA1ED314FA10" box="[160,318,1436,1461]" pageId="27" pageNumber="478">Photographed</collectingCounty>
|
||
/
|
||
<collectingMunicipality id="6BA9ACE1FF8DD520CF65FA1ED3DFFA11" box="[327,501,1436,1461]" pageId="27" pageNumber="478">Kelsey Keaton</collectingMunicipality>
|
||
2014/
|
||
<location id="8EAD6040FF8DD520CC67FA1ED0C9FA10" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CC67FA1ED0C9FA10" box="[581,739,1436,1461]" country="France" county="Photographed" municipality="Kelsey Keaton" name="Emu Catalog" pageId="27" pageNumber="478">Emu Catalog</location>
|
||
”[blue label, printed in black]; (7) “
|
||
<typeStatus id="54C98839FF8DD520CA83FA1ED70BFA10" box="[1185,1313,1436,1461]" pageId="27" pageNumber="478" type="syntype">SYNTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C724D18FF8DD520CB0BFA1ED2D4FA7D" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">Hypotelus testaceus</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EFE34B6AFF8DD520CF24FA3DD3B3FA7D" author="Bierig" box="[262,409,1471,1496]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">
|
||
<location id="8EAD6040FF8DD520CF24FA3DD367FA7D" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CF24FA3DD367FA7D" box="[262,333,1471,1496]" country="France" county="Photographed" municipality="Kelsey Keaton" name="Bierig" pageId="27" pageNumber="478">Bierig</location>
|
||
, 1934
|
||
</bibRefCitation>
|
||
/det.
|
||
<determinerName id="7E11DA5FFF8DD520CFF8FA42D050FA7D" box="[474,634,1471,1496]" pageId="27" pageNumber="478">S. Bortoluzzi</determinerName>
|
||
, 2016” [red label, printed in black].
|
||
<location id="8EAD6040FF8DD520CA72FA3DD6BBFA7D" LSID="urn:lsid:plazi:treatment:03DB878DFF8DD527CEB5FB9ED1D9FCB0:8EAD6040FF8DD520CA72FA3DD6BBFA7D" box="[1104,1169,1471,1496]" country="France" county="Photographed" municipality="Kelsey Keaton" name="Three" pageId="27" pageNumber="478">Three</location>
|
||
specimens, one male [dissected], one female [dissected] and one sex undetermined, whit same labels: (1) “
|
||
<collectingCountry id="F365760BFF8DD520CAB8FA66D6C3FA59" box="[1178,1257,1508,1532]" name="France" pageId="27" pageNumber="478">France</collectingCountry>
|
||
Field/
|
||
<collectingDate id="EF88E9B3FF8DD520CB19FA66D7B9FA58" box="[1339,1427,1508,1533]" pageId="27" pageNumber="478" value="1930-06">
|
||
<date id="FFCC105BFF8DD520CB19FA66D7B9FA58" box="[1339,1427,1508,1533]" pageId="27" pageNumber="478" value="1930-06">VI.1930</date>
|
||
</collectingDate>
|
||
/
|
||
<collectingCountry id="F365760BFF8DD520CEB5F98AD2D8F985" box="[151,242,1544,1568]" name="Panama" pageId="27" pageNumber="478">Panamá</collectingCountry>
|
||
” [white label, handwritten]; (2) “
|
||
<collectorName id="2687534DFF8DD520CC4BF985D170F985" box="[617,858,1543,1568]" pageId="27" pageNumber="478">Field Mus. Nat. Hist.</collectorName>
|
||
/1966/
|
||
<collectorName id="2687534DFF8DD520CD86F985D67DF985" box="[932,1111,1543,1568]" pageId="27" pageNumber="478">A. Bierig Colln.</collectorName>
|
||
/
|
||
<collectorName id="2687534DFF8DD520CA42F985D6BEF985" box="[1120,1172,1543,1568]" pageId="27" pageNumber="478">Acc.</collectorName>
|
||
Z-13812” [white label, printed in black]; (3) “
|
||
<typeStatus id="54C98839FF8DD520CFB0F9AED038F9E0" box="[402,530,1580,1605]" pageId="27" pageNumber="478" type="syntype">SYNTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C724D18FF8DD520CC39F9AED0D6F9E0" box="[539,764,1580,1605]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">Hypotelus testaceus</taxonomicName>
|
||
/
|
||
<bibRefCitation id="EFE34B6AFF8DD520CD26F9AED1A5F9E0" author="Bierig" box="[772,911,1580,1605]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">
|
||
<collectorName id="2687534DFF8DD520CD26F9AED161F9E0" box="[772,843,1580,1605]" pageId="27" pageNumber="478">Bierig</collectorName>
|
||
, 1934
|
||
</bibRefCitation>
|
||
/det.
|
||
<determinerName id="7E11DA5FFF8DD520CDE5F9AED674F9E0" box="[967,1118,1580,1605]" pageId="27" pageNumber="478">S. Bortoluzzi</determinerName>
|
||
, 2016” [red label, printed in black].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8DD520CEE5F9F7D711F915" blockId="27.[151,1437,1255,2000]" pageId="27" pageNumber="478">
|
||
<emphasis id="B906EA89FF8DD520CEE5F9F7D2D6F929" box="[199,252,1653,1676]" italics="true" pageId="27" pageNumber="478">Note</emphasis>
|
||
: In the original description,
|
||
<bibRefCitation id="EFE34B6AFF8DD520CC18F9F6D0F8F929" author="Bierig" box="[570,722,1652,1677]" pageId="27" pageNumber="478" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">Bierig (1934)</bibRefCitation>
|
||
did not specify how many specimens he observed. We received from FMNH four specimens, one specimen with label “TYPUS”, but we consider all of them as syntypes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3686510FF8DD520CEE5F93ED6B8F8B9" pageId="27" pageNumber="478" type="diagnosis">
|
||
<paragraph id="8BCD369BFF8DD520CEE5F93ED6B8F8B9" blockId="27.[151,1437,1255,2000]" pageId="27" pageNumber="478">
|
||
<emphasis id="B906EA89FF8DD520CEE5F93ED36BF970" bold="true" box="[199,321,1724,1749]" pageId="27" pageNumber="478">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C724D18FF8DD520CF6BF93ED000F971" box="[329,554,1724,1749]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="species" species="testaceus">
|
||
<emphasis id="B906EA89FF8DD520CF6BF93ED000F971" box="[329,554,1724,1749]" italics="true" pageId="27" pageNumber="478">Hypotelus testaceus</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from other species of
|
||
<taxonomicName id="4C724D18FF8DD520CA06F93ED6BCF970" box="[1060,1174,1724,1749]" class="Insecta" family="Staphylinidae" genus="Hypotelus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B906EA89FF8DD520CA06F93ED6BCF970" box="[1060,1174,1724,1749]" italics="true" pageId="27" pageNumber="478">Hypotelus</emphasis>
|
||
</taxonomicName>
|
||
by the body color (see Redescription); male with sternite 8 at posterior margin with short tricuspidate projection medially (
|
||
<figureCitation id="13492A1EFF8DD520CB14F95DD7A4F95D" box="[1334,1422,1759,1784]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="27" pageNumber="478">Fig. 53</figureCitation>
|
||
); sternite 8 of female with posterior margin sinuous and shape of spermatheca (
|
||
<figureCitation id="13492A1EFF8DD520CDDEF886D6ACF8B9" box="[1020,1158,1796,1821]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="27" pageNumber="478">Figs. 88–90</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3686510FF8DD527CEE5F8A5D3DAFD68" lastPageId="28" lastPageNumber="479" pageId="27" pageNumber="478" type="description">
|
||
<paragraph id="8BCD369BFF8DD520CEE5F8A5D2C9F875" blockId="27.[151,1437,1255,2000]" pageId="27" pageNumber="478">
|
||
<emphasis id="B906EA89FF8DD520CEE5F8A5D35EF8E5" bold="true" box="[199,372,1831,1856]" pageId="27" pageNumber="478">Redescription.</emphasis>
|
||
BL: 2.6–3.0 mm, BW:
|
||
<quantity id="4C8A9B7EFF8DD520CCA8F8A5D13CF89A" box="[650,790,1831,1856]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="27" pageNumber="478" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 mm</quantity>
|
||
. Body somewhat slightly convex; dorsal surface glossy; brown; basal two-thirds of elytra, apical thirds of abdominal segments 3–6 and appendages lighter than body, except mandibles (
|
||
<figureCitation id="13492A1EFF8DD520CF4DF8EDD3ECF82D" box="[367,454,1903,1928]" captionStart="FIGURES 9 – 19" captionStartId="9.[151,264,1913,1935]" captionTargetBox="[216,1355,230,1832]" captionTargetId="figure@9.[151,1436,157,1945]" captionTargetPageId="9" captionText="FIGURES 9 – 19. Habitus, dorsal view. 9, Hypotelus insulanus, syntype, male; 10, H. praecox, lectotype, female; 11, H. scheerpeltzi sp. nov., holotype, male; 12, H. testaceus, syntype, male; 13, H. melanodelta sp. nov., holotype, female; 14, H. castaneus sp. nov., holotype, male; 15, H. brevitarsus sp. nov., holotype, male; 16, H. corniculatus sp. nov., holotype, female; 17, H. marginatus, syntype, male; 18, H. laevis, male; 19, H. andinus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/803652/files/figure.png" pageId="27" pageNumber="478">Fig. 12</figureCitation>
|
||
). Dorsal integument of head and pronotum entirely with fine punctures and undulate microstriae; elytra with dispersed fine punctures and only one longitudinal finely punctate stria close to elytral suture.
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FF15D6C6FF70" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FF15D361FF15" box="[199,331,151,176]" italics="true" pageId="28" pageNumber="479">Male. Head</emphasis>
|
||
. Supra-antennal area slightly prominent. Antennae reaching humeral angle; antennomeres 2 and 3 of equal length, 5–11 gradually increasing in length toward antennal apex. Mandibles symmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FF5DD0A8FEC1" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FF5DD332FF5D" box="[199,280,223,248]" italics="true" pageId="28" pageNumber="479">Thorax</emphasis>
|
||
. Pronotum wider than long (PW/PL=1.4); anterior angles rounded and slightly prominent; apical half with somewhat parallel sides and basal half gradually narrowing toward the base; with complete internal midlongitudinal ridge and slight longitudinal median sulcus only on disc. Elytra somewhat longer than wide (EL/ BW=1.2), not covering abdominal tergite 3.
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FEEDD6E1FE50" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FEEDD31BFE2D" box="[199,305,367,392]" italics="true" pageId="28" pageNumber="479">Abdomen</emphasis>
|
||
. Tergite 8 with posterior margin rounded; sternite 8 with posterior margin with short tricuspidate projection at middle region and with short setae (
|
||
<figureCitation id="13492A1EFF8AD527CCC7FE16D114FE08" box="[741,830,404,429]" captionStart="FIGURES 48 – 58" captionStartId="16.[151,264,1844,1866]" captionTargetBox="[200,1385,250,1781]" captionTargetId="figure@16.[151,1436,193,1823]" captionTargetPageId="16" captionText="FIGURES 48 – 58. Tergites and sternites of males in dorsal and ventral view, respectively and right setae removed. 48, Hypotelus pusillus, tergite 8; 49, H. scheerpeltzi sp. nov., tergite 8; 50, H. castaneus sp. nov., tergite 8; 51, H. pusillus, sternite 8; 52, H. insulanus, sternite 8; 53, H. testaceus, sternite 8; 54, H. castaneus sp. nov., sternite 8; 55, H. pusillus, tergites 9 and 10; 56, H. insulanus, tergite 10; 57, H. castaneus sp. nov., tergite 10; 58, H. brevitarsus sp. nov., tergite 10. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803660/files/figure.png" pageId="28" pageNumber="479">Fig. 53</figureCitation>
|
||
); tergite 9 with short ventral struts; sternite 9 with posterior margin truncate and with two pairs of long setae. Median lobe of aedeagus with bulbous base and apex triangular in ventral view and curved shape in lateral, with apex curved as a hook (
|
||
<figureCitation id="13492A1EFF8AD527CA17FE5ED695FE50" box="[1077,1215,476,501]" captionStart="FIGURES 59 – 74" captionStartId="17.[151,264,1924,1946]" captionTargetBox="[290,1268,229,1830]" captionTargetId="figure@17.[237,1336,193,1893]" captionTargetPageId="17" captionText="FIGURES 59 – 74. Aedeagus, for each species with lateral view and ventral view, respectively. 59 – 60, Hypotelus pusillus; 61 – 62, H. insulanus; 63 – 64, H. marginatus; 65 – 66, H. testaceus; 67 – 68, H. praecox; 69 – 70, H. brevitarsus sp. nov., parameres removed in ventral view; 71 – 72, H. castaneus sp. nov.; 73 – 74, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803662/files/figure.png" pageId="28" pageNumber="479">Figs. 65–66</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FE7DD0D1FD21" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FE7DD331FDBD" box="[199,283,511,536]" italics="true" pageId="28" pageNumber="479">Female</emphasis>
|
||
. Similar to male except for: abdominal sternite 8 with posterior margin sinuous (
|
||
<figureCitation id="13492A1EFF8AD527CAECFE7DD70FFDBD" box="[1230,1317,511,536]" captionStart="FIGURES 75 – 82" captionStartId="18.[151,264,1403,1425]" captionTargetBox="[194,1368,254,1322]" captionTargetId="figure@18.[151,1436,193,1381]" captionTargetPageId="18" captionText="FIGURES 75 – 82. Sternite 8 of females in ventral view, right setae removed. 75, Hypotelus pusillus; 76, H. insulanus; 77, H. marginatus; 78 H. testaceus; 79, H. melanodelta sp. nov.; 80, H. brevitarsus sp. nov.; 81, H. castaneus sp. nov.; 82, H. scheerpeltzi sp. nov. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803664/files/figure.png" pageId="28" pageNumber="479">Fig. 78</figureCitation>
|
||
); tergite 9 without ventral struts; tergite 10 with lateral and posterior margin weakly pigmented (
|
||
<figureCitation id="13492A1EFF8AD527CA7CFDA6D698FD99" box="[1118,1202,548,573]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="28" pageNumber="479">Fig. 89</figureCitation>
|
||
); bursa copulatrix as
|
||
<figureCitation id="13492A1EFF8AD527CEB5FDC5D2C1FDC5" box="[151,235,583,608]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="28" pageNumber="479">Fig. 88</figureCitation>
|
||
; ovipositor consisting of a pair of weakly pigmented hemisternites and a pair of more apical coxites, and with many long setae on apex; spermatheca as
|
||
<figureCitation id="13492A1EFF8AD527CC87FDEED0D2FD21" box="[677,760,620,645]" captionStart="FIGURES 83 – 108" captionStartId="19.[151,264,1784,1806]" captionTargetBox="[245,1323,259,1717]" captionTargetId="figure@19.[207,1375,193,1777]" captionTargetPageId="19" captionText="FIGURES 83 – 108. Tergites in dorsal view, right setae removed, and genitalia of female. 83 – 84, Hypotelus pusillus. 83, tergite 9, 10 and bursa copulatrix; 84, spermatheca. 85 – 86, H. insulanus. 85, tergite 10; 86, spermatheca. 87, H. marginatus, spermatheca. 88 – 90, H. testaceus. 88, bursa copulatrix; 89, tergite 10; 90, spermatheca. 91, H. melanodelta sp. nov., spermatheca. 92 – 94, H. praecox. 92, bursa copulatrix; 93, tergite 10; 94, spermatheca, 95 – 97, H. castaneus sp. nov. 95, bursa copulatrix; 96, tergite 10; 97, spermatheca. 98 – 99, H. brevitarsus sp. nov. 98, tergite 10; 99, spermatheca. 100 – 102, H. corniculatus sp. nov. 100, bursa copulatrix; 101, tergite 10; 102, spermatheca. 103 – 105, H. laevis. 103, bursa copulatrix; 104, tergite 10; 105, spermatheca. 106 – 108, H. scheerpeltzi sp. nov. 106, bursa copulatrix; 107, tergite 10; 108, spermatheca. Scale bars = 0.125 mm." httpUri="https://zenodo.org/record/803666/files/figure.png" pageId="28" pageNumber="479">Fig. 90</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FD0DD121FD0D" blockId="28.[151,1437,151,789]" box="[199,779,655,680]" pageId="28" pageNumber="479">
|
||
<materialsCitation id="3B1A3CC6FF8AD527CEE5FD0DD121FD0D" ID-GBIF-Occurrence="1563407712" box="[199,779,655,680]" country="Panama" location="Colon" pageId="28" pageNumber="479" specimenCount="1" stateProvince="Colon">
|
||
<emphasis id="B906EA89FF8AD527CEE5FD0DD3FAFD0D" bold="true" box="[199,464,655,680]" pageId="28" pageNumber="479">Geographical records.</emphasis>
|
||
<collectingCountry id="F365760BFF8AD527CFF5FD12D01AFD02" box="[471,560,656,679]" name="Panama" pageId="28" pageNumber="479">Panama</collectingCountry>
|
||
(
|
||
<collectingRegion id="49B6F879FF8AD527CC1DFD0DD0ADFD0D" box="[575,647,655,680]" country="Panama" name="Colon" pageId="28" pageNumber="479">Colon</collectingRegion>
|
||
) (
|
||
<figureCitation id="13492A1EFF8AD527CCBFFD0DD0D5FD0D" box="[669,767,655,680]" captionStart="FIGURE 114" captionStartId="31.[151,250,1701,1723]" captionTargetBox="[190,1375,215,1655]" captionTargetId="figure@31.[150,1437,153,1679]" captionTargetPageId="31" captionText="FIGURE 114. Geographical records of Hypotelus. Hypotelus andinus, Colombia, Ecuador and Bolivia; H. insulanus, Caribbean islands; H. marginatus, Colombia, Venezuela and Guatemala; H. testaceus, Panama; H. brevitarsus sp. nov., Panama. Countries without dots but colored represent the occurrence but no locality given." httpUri="https://zenodo.org/record/803674/files/figure.png" pageId="28" pageNumber="479">Fig. 114</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCD369BFF8AD527CEE5FD36D3DAFD68" blockId="28.[151,1437,151,789]" box="[199,496,692,717]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FD36D3A3FD68" bold="true" box="[199,393,692,717]" pageId="28" pageNumber="479">Biological notes.</emphasis>
|
||
No data.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3686510FF8AD527CEE5FD55D1D9FCB0" pageId="28" pageNumber="479" type="discussion">
|
||
<paragraph id="8BCD369BFF8AD527CEE5FD55D1D9FCB0" blockId="28.[151,1437,151,789]" pageId="28" pageNumber="479">
|
||
<emphasis id="B906EA89FF8AD527CEE5FD55D311FD55" bold="true" box="[199,315,727,752]" pageId="28" pageNumber="479">Remarks.</emphasis>
|
||
In the original description,
|
||
<bibRefCitation id="EFE34B6AFF8AD527CC5BFD55D138FD55" author="Bierig" box="[633,786,727,752]" pageId="28" pageNumber="479" refString="Bierig, A. (1934) Dos nuevas especies del genero Hypotelus Er. (Col.). Revistade Entomologia, 4, 342 - 344." type="journal article" year="1934">Bierig (1934)</bibRefCitation>
|
||
reported that the male of this species is unknown. In the
|
||
<typeStatus id="54C98839FF8AD527CEB5FD7FD2EDFCB0" box="[151,199,765,789]" pageId="28" pageNumber="479">type</typeStatus>
|
||
material, we found one male and it is described it here for the first time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |