471 lines
61 KiB
XML
471 lines
61 KiB
XML
<document id="1EA4138A039DAAF95D6F69214E5381E8" ID-DOI="10.11646/zootaxa.3941.4.6" ID-GBIF-Dataset="c87d61d6-c366-4a74-a2d8-bac34c844434" ID-ISSN="1175-5326" ID-Zenodo-Dep="244519" ID-ZooBank="CCC3BFF9-1EF6-45A6-AFE1-07ED3DFD2E41" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461028721887" checkinUser="plazi" docAuthor="Clark, Roger N. & Jewett, Stephen C." docDate="2015" docId="03DB878D6F172138F5C6C38DFCAE9E5E" docLanguage="en" docName="zt03941p584.pdf" docOrigin="Zootaxa 3941 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptasterias tatei Clark & Jewett, 2015, sp. nov." docType="treatment" docUuid="B19FC790-60F4-423B-A11E-EF2FF308FB9E" docUuidSource="ZooBank" docVersion="10" lastPageNumber="583" masterDocId="FFE2FFF56F16213CF551C351FFCB9A51" masterDocTitle="A new sea star of the genus Leptasterias (Asteroidea: Asteriidae) from the Aleutian Islands" masterLastPageNumber="584" masterPageNumber="579" pageNumber="580" updateTime="1698573018833" updateUser="plazi">
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<mods:mods id="190544C5D8D4F6FA8C39203207E32258" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EE645827E611188C28622DCE4BDD8B5F">
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<mods:title id="4BEA7A484EF6B17677FBBA5D63C8F9FA">A new sea star of the genus Leptasterias (Asteroidea: Asteriidae) from the Aleutian Islands</mods:title>
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<mods:name id="33D90F4DCDEC77F1279F0EA9C7AAE9DB" type="personal">
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<mods:roleTerm id="5C593703322A583A56C2717ED7A5FE2A">Author</mods:roleTerm>
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<mods:namePart id="301BB86A3F50C4387BC697D68CB41BE0">Clark, Roger N.</mods:namePart>
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<mods:name id="604E592F9AA309203C71B9AFB4C85716" type="personal">
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<mods:roleTerm id="CA72DCD6D3ACB7C469DCCCDAF50EAAFB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="01685385C09A0C6F50DDBBF782089AA3">Jewett, Stephen C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3FF5774B8DD487FB166C51A85EB7920F">text</mods:typeOfResource>
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<mods:title id="D6BFAC8DF60518D2DB304CFF7DF412FC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="86101BAA1789F53F5D6167BB055E8F55">
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<mods:date id="7CD6C4DD1CDC821EBD1AC6C5E12A5DB1">2015</mods:date>
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<mods:detail id="A065A4B587B5523F8DDC82A414208AF5" type="volume">
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<mods:number id="823AB027ED1B24BF05C7453D22A12B74">3941</mods:number>
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<mods:detail id="C09E94653E367AE536764709179DAEA4" type="issue">
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<mods:number id="05661EA75A0E17A4E6D9212247AFA209">4</mods:number>
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<mods:classification id="66678BC7471849E444B2858F2E8F9272">journal article</mods:classification>
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<mods:identifier id="9978B3F69C45A89A0CA94BEEBF56115E" type="DOI">10.11646/zootaxa.3941.4.6</mods:identifier>
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<treatment id="03DB878D6F172138F5C6C38DFCAE9E5E" ID-DOI="http://doi.org/10.5281/zenodo.6105211" ID-GBIF-Taxon="119607089" ID-Zenodo-Dep="6105211" LSID="urn:lsid:zoobank.org:act:B19FC790-60F4-423B-A11E-EF2FF308FB9E" httpUri="http://treatment.plazi.org/id/03DB878D6F172138F5C6C38DFCAE9E5E" lastPageId="4" lastPageNumber="583" pageId="1" pageNumber="580">
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<subSubSection id="C36865106F17213DF5C6C38DFEE19B49" pageId="1" pageNumber="580" type="nomenclature">
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<paragraph id="8BCD369B6F17213DF5C6C38DFE029AA6" blockId="1.[151,457,220,280]" box="[151,457,220,247]" pageId="1" pageNumber="580">
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<heading id="D08581F76F17213DF5C6C38DFE029AA6" bold="true" box="[151,457,220,247]" fontSize="11" level="1" pageId="1" pageNumber="580" reason="1">
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<emphasis id="B906EA896F17213DF5C6C38DFE029AA6" bold="true" box="[151,457,220,247]" pageId="1" pageNumber="580">
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<taxonomicName id="4C724D186F17213DF5C6C38DFEAF9AA7" ID-CoL="3THYJ" box="[151,356,220,246]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="1" pageNumber="580" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
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<emphasis id="B906EA896F17213DF5C6C38DFEAF9AA7" bold="true" box="[151,356,220,246]" italics="true" pageId="1" pageNumber="580">Leptasterias tatei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23557F26F17213DF43AC38CFE029AA6" box="[363,457,221,247]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCD369B6F17213DF5C6C3AEFEE19B49" blockId="1.[151,457,220,280]" box="[151,298,255,280]" pageId="1" pageNumber="580">
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<figureCitation id="13492A1E6F17213DF5C6C3AEFEE19B49" box="[151,298,255,280]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 10" captionStart-2="FIGURE 11" captionStartId-0="1.[151,264,1890,1913]" captionStartId-1="2.[151,264,1671,1694]" captionStartId-2="2.[151,250,1794,1816]" captionTargetBox-0="[151,1435,479,1866]" captionTargetBox-1="[151,1436,193,1649]" captionTargetId-0="figure@1.[151,1435,479,1866]" captionTargetId-1="figure@2.[151,1436,193,1650]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." captionText-1="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." captionText-2="FIGURE 11. Leptasterias tatei sp. nov. Live, in situ Hawadax Island, N side, 13 m (leg. R. N. Clark, 8 June, 2008), Bar = 5.0 cm." httpUri-0="https://zenodo.org/record/244520/files/figure.png" httpUri-1="https://zenodo.org/record/244521/files/figure.png" pageId="1" pageNumber="580">Figures 1–11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36865106F17213EF5C6C216FCAE9D89" lastPageId="2" lastPageNumber="581" pageId="1" pageNumber="580" type="materials_examined">
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<paragraph id="8BCD369B6F17213DF5C6C216FCBF9BD5" blockId="1.[151,1436,327,424]" pageId="1" pageNumber="580">
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<emphasis id="B906EA896F17213DF5C6C216FEF39B31" bold="true" box="[151,312,327,352]" pageId="1" pageNumber="580">
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<typeStatus id="54C988396F17213DF5C6C216FF199B31" box="[151,210,327,352]" pageId="1" pageNumber="580">Type</typeStatus>
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locality.
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</emphasis>
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Alaska, Aleutian Islands, Adak Island, Thumb Bay entrance, W side (51°
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<geoCoordinate id="EE46505C6F17213DF1D8C216FB269B0E" box="[1161,1261,327,352]" direction="north" orientation="latitude" pageId="1" pageNumber="580" precision="555" value="50.08">50.08 N</geoCoordinate>
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, 176°
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<geoCoordinate id="EE46505C6F17213DF07FC216FA5F9B31" box="[1326,1428,327,352]" direction="west" orientation="longitude" pageId="1" pageNumber="580" precision="555" value="-32.31">32.31 W</geoCoordinate>
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) (
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<emphasis id="B906EA896F17213DF5F1C23DFF019BD5" box="[160,202,364,388]" italics="true" pageId="1" pageNumber="580">leg.</emphasis>
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Roger N. Clark, scuba
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<quantity id="4C8A9B7E6F17213DF480C23DFDC79BD5" box="[465,524,364,388]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="580" unit="m" value="11.0">11 m</quantity>
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;
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<date id="FFCC105B6F17213DF746C23DFDA09BD4" box="[535,619,364,389]" pageId="1" pageNumber="580" value="1920-07">20 July</date>
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, 2011; R/V
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<emphasis id="B906EA896F17213DF7A0C23CFCAF9BD5" box="[753,868,365,388]" italics="true" pageId="1" pageNumber="580">Norseman</emphasis>
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).
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</paragraph>
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<paragraph id="8BCD369B6F17213DF596C2DEFDFE9BF9" blockId="1.[151,1436,327,424]" box="[199,565,399,424]" pageId="1" pageNumber="580">
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<emphasis id="B906EA896F17213DF596C2DEFEF09BF9" bold="true" box="[199,315,399,424]" pageId="1" pageNumber="580">
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<typeStatus id="54C988396F17213DF596C2DEFEFD9BF9" box="[199,310,399,424]" pageId="1" pageNumber="580" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED63AE5E6F17213DF413C2C1FE589BF9" LSID="urn:lsid:biocol.org:col:34802" box="[322,403,400,424]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="580">LACM</collectionCode>
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2011-158.001
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</paragraph>
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<caption id="DF0D66136F17213DF5C6C433FDA49DA2" httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="1" pageNumber="580" targetBox="[151,1435,479,1866]" targetPageId="1">
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<paragraph id="8BCD369B6F17213DF5C6C433FDA49DA2" blockId="1.[151,1436,1890,2035]" pageId="1" pageNumber="580">
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<emphasis id="B906EA896F17213DF5C6C433FEF19D29" bold="true" box="[151,314,1890,1913]" pageId="1" pageNumber="580">FIGURES 1–6.</emphasis>
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<taxonomicName id="4C724D186F17213DF411C432FE3A9D29" box="[320,497,1891,1912]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="1" pageNumber="580" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
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||
<emphasis id="B906EA896F17213DF411C432FE3A9D29" box="[320,497,1891,1912]" italics="true" pageId="1" pageNumber="580">Leptasterias tatei</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B906EA896F17213DF4A6C432FD839D29" bold="true" box="[503,584,1891,1912]" pageId="1" pageNumber="580">
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||
<taxonomicNameLabel id="A23557F26F17213DF4A6C432FD839D29" box="[503,584,1891,1912]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Holotype, 2011-158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm.
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</paragraph>
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</caption>
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<caption id="DF0D66136F14213EF5C6C5D6FDBD9CAB" httpUri="https://zenodo.org/record/244521/files/figure.png" pageId="2" pageNumber="581" targetBox="[151,1436,193,1649]" targetPageId="2">
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<paragraph id="8BCD369B6F14213EF5C6C5D6FDBD9CAB" blockId="2.[151,1436,1671,1908]" pageId="2" pageNumber="581">
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<emphasis id="B906EA896F14213EF5C6C5D6FE819CCC" bold="true" box="[151,330,1671,1694]" pageId="2" pageNumber="581">FIGURES 7–10.</emphasis>
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<taxonomicName id="4C724D186F14213EF402C5D8FDCC9CCF" box="[339,519,1673,1694]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="2" pageNumber="581" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
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<emphasis id="B906EA896F14213EF402C5D8FDCC9CCF" box="[339,519,1673,1694]" italics="true" pageId="2" pageNumber="581">Leptasterias tatei</emphasis>
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</taxonomicName>
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<emphasis id="B906EA896F14213EF741C5D9FDA89CCC" bold="true" box="[528,611,1672,1693]" pageId="2" pageNumber="581">
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<taxonomicNameLabel id="A23557F26F14213EF741C5D9FDA89CCC" box="[528,611,1672,1693]" pageId="2" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, Holotype, 2011-158.001.Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live,
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<emphasis id="B906EA896F14213EF4E2C597FE3E9C8A" box="[435,501,1734,1755]" italics="true" pageId="2" pageNumber="581">in situ</emphasis>
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Adak Island, Thumb Bay, 13 m (
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<emphasis id="B906EA896F14213EF60DC594FC4A9C8B" box="[860,897,1733,1754]" italics="true" pageId="2" pageNumber="581">leg.</emphasis>
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R.N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011-159.001. Oral side, Bar = 4 cm.
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</paragraph>
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</caption>
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<caption id="DF0D66136F14213EF5C6C453FF769D66" pageId="2" pageNumber="581">
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<paragraph id="8BCD369B6F14213EF5C6C453FF769D66" blockId="2.[151,1436,1671,1908]" pageId="2" pageNumber="581">
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||
<emphasis id="B906EA896F14213EF5C6C453FED59D46" bold="true" box="[151,286,1794,1816]" pageId="2" pageNumber="581">FIGURE 11.</emphasis>
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<taxonomicName id="4C724D186F14213EF474C452FE1D9D49" box="[293,470,1795,1816]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="2" pageNumber="581" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
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<emphasis id="B906EA896F14213EF474C452FE1D9D49" box="[293,470,1795,1816]" italics="true" pageId="2" pageNumber="581">Leptasterias tatei</emphasis>
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</taxonomicName>
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<emphasis id="B906EA896F14213EF48CC452FDE59D49" bold="true" box="[477,558,1795,1816]" pageId="2" pageNumber="581">
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<taxonomicNameLabel id="A23557F26F14213EF48CC452FDE59D49" box="[477,558,1795,1816]" pageId="2" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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Live,
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<emphasis id="B906EA896F14213EF720C452FD7A9D49" box="[625,689,1795,1816]" italics="true" pageId="2" pageNumber="581">in situ</emphasis>
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Hawadax Island, N side, 13 m (
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<emphasis id="B906EA896F14213EF6ADC452FBEA9D49" box="[1020,1057,1795,1816]" italics="true" pageId="2" pageNumber="581">leg.</emphasis>
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R.N. Clark, 8 June, 2008), Bar = 5.0 cm.
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</paragraph>
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||
</caption>
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||
<caption id="DF0D66136F14213EF5C6C411FEE99D25" pageId="2" pageNumber="581">
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||
<paragraph id="8BCD369B6F14213EF5C6C411FEE99D25" blockId="2.[151,1436,1671,1908]" pageId="2" pageNumber="581">
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<emphasis id="B906EA896F14213EF5C6C411FED49D04" bold="true" box="[151,287,1856,1878]" pageId="2" pageNumber="581">FIGURE 12.</emphasis>
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<taxonomicName id="4C724D186F14213EF474C410FD6E9D07" authority="Fisher, 1930" authorityName="Fisher" authorityYear="1930" box="[293,677,1856,1878]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="2" pageNumber="581" phylum="Echinodermata" rank="species" species="stolacantha">
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<emphasis id="B906EA896F14213EF474C410FDD59D04" box="[293,542,1856,1878]" italics="true" pageId="2" pageNumber="581">Leptasterias stolacantha</emphasis>
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Fisher, 1930
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</taxonomicName>
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. Live,
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<emphasis id="B906EA896F14213EF7B8C410FCE39D07" box="[745,808,1857,1878]" italics="true" pageId="2" pageNumber="581">in situ</emphasis>
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Kagamil Island, E side, 11 m (
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<emphasis id="B906EA896F14213EF131C411FB4D9D04" box="[1120,1158,1856,1877]" italics="true" pageId="2" pageNumber="581">leg.</emphasis>
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R.N. Clark, 19 July, 2007), Bar = 2.0 cm.
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</paragraph>
|
||
</caption>
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||
<paragraph id="8BCD369B6F14213EF596C4CDFCAE9D89" blockId="2.[151,1436,1948,2008]" pageId="2" pageNumber="581">
|
||
<emphasis id="B906EA896F14213EF596C4CDFEF09DE4" bold="true" box="[199,315,1948,1973]" pageId="2" pageNumber="581">
|
||
<typeStatus id="54C988396F14213EF596C4CDFEFD9DE4" box="[199,310,1948,1973]" pageId="2" pageNumber="581" type="paratype">Paratype</typeStatus>
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||
.
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</emphasis>
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<collectionCode id="ED63AE5E6F14213EF414C4CDFE5D9DE5" LSID="urn:lsid:biocol.org:col:34802" box="[325,406,1948,1972]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="2" pageNumber="581">LACM</collectionCode>
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||
2011-159.001; genetic sample
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<collectionCode id="ED63AE5E6F14213EF653C4CDFC989DE5" LSID="urn:lsid:biocol.org:col:34802" box="[770,851,1948,1972]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="2" pageNumber="581">LACM</collectionCode>
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||
2011-159.002 Alaska, Aleutian Islands, Amchitka Island, SE side, W of Makarius Bay (51°
|
||
<geoCoordinate id="EE46505C6F14213EF70DC4EEFD759D86" box="[604,702,1983,2008]" direction="north" orientation="latitude" pageId="2" pageNumber="581" precision="555" value="23.16">23.16 N</geoCoordinate>
|
||
, 179°
|
||
<geoCoordinate id="EE46505C6F14213EF7ADC491FC939D89" box="[764,856,1984,2008]" direction="east" orientation="longitude" pageId="2" pageNumber="581" precision="555" value="11.02">11.02 E</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36865106F15213FF596C3C6FDAC9B11" pageId="3" pageNumber="582" type="diagnosis">
|
||
<paragraph id="8BCD369B6F15213FF596C3C6FDAC9B11" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
<emphasis id="B906EA896F15213FF596C3C6FEF19AE1" bold="true" box="[199,314,151,176]" pageId="3" pageNumber="582">Diagnosis</emphasis>
|
||
.Star of moderate size; R to
|
||
<quantity id="4C8A9B7E6F15213FF727C3C9FD7A9AFE" box="[630,689,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="582" unit="cm" value="9.0">9 cm</quantity>
|
||
, R:r 6–9 (
|
||
<figureCitation id="13492A1E6F15213FF673C3C6FCAF9AE1" box="[802,868,151,176]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 1</figureCitation>
|
||
). Five rays, relatively long; abactinal ossicles 2–4 lobed, with single spines; spines relatively short, stout, truncated bearing numerous fine vertical ridges; superomarginal and inferomarginal ossicles bear a single thick, blunt spine, similar to abactinal spines; all spines wreathed with numerous crossed pedicellariae. Tube feet, in four rows. Color in life reddish with lighter mottlings abactinally, actinal surface straw colored.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36865106F15213FF596C21DFBFC99BC" pageId="3" pageNumber="582" type="description">
|
||
<paragraph id="8BCD369B6F15213FF596C21DFE4898D4" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
<emphasis id="B906EA896F15213FF596C21DFE929B34" bold="true" box="[199,345,332,357]" pageId="3" pageNumber="582">Description.</emphasis>
|
||
Moderate sized star R to
|
||
<quantity id="4C8A9B7E6F15213FF72CC21DFD729B35" box="[637,697,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="582" unit="cm" value="9.0">9 cm</quantity>
|
||
, r =
|
||
<quantity id="4C8A9B7E6F15213FF7BFC21DFCE29B35" box="[750,809,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="582" unit="cm" value="1.0">1 cm</quantity>
|
||
, R:r 9.1 (
|
||
<typeStatus id="54C988396F15213FF6C2C21DFBCB9B34" box="[915,1024,332,357]" pageId="3" pageNumber="582" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<figureCitation id="13492A1E6F15213FF15DC21DFB859B35" box="[1036,1102,332,357]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 1</figureCitation>
|
||
). Disc small, five rays, fairly long, tapering, relatively soft. Skeleton relatively open; papular areas relatively large, bearing 3–6 papulae, obscured by wreaths of pedicellariae on spines in live individuals; bearing scattered large straight pedicellaria to
|
||
<quantity id="4C8A9B7E6F15213FF5C6C2E6FF369B9E" box="[151,253,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="3" pageNumber="582" unit="mm" value="1.25">1.25 mm</quantity>
|
||
long (
|
||
<figureCitation id="13492A1E6F15213FF418C2E6FE7E9B81" box="[329,437,439,464]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Figs. 2–3</figureCitation>
|
||
) on papular areas; pedicellariae broadened distally, and bearing four “teeth” per valve. Carinal ossicles 2–4 lobed, bearing single spines, dorsolateral ossicles single spined, also 2–4 lobed, but forming a more open meshwork (
|
||
<figureCitation id="13492A1E6F15213FF4C6C2AEFE109849" box="[407,475,511,536]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 4</figureCitation>
|
||
), and bearing small, irregular accessory plates; spines relatively short, thick truncated, with strong vertical ridges continuing onto the dorsal surface; encircled by a fleshy wreath of small, crossed pedicellariae up to
|
||
<quantity id="4C8A9B7E6F15213FF421C116FE1D980E" box="[368,470,583,608]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="582" unit="mm" value="0.25">0.25 mm</quantity>
|
||
in length (
|
||
<figureCitation id="13492A1E6F15213FF703C116FD0A9831" box="[594,705,583,608]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Figs. 4–5</figureCitation>
|
||
). Madreporite small, about
|
||
<quantity id="4C8A9B7E6F15213FF6ABC116FB9A980E" box="[1018,1105,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="582" unit="mm" value="3.5">3.5 mm</quantity>
|
||
in diameter, roughly circular with irregular ridges.
|
||
</paragraph>
|
||
<paragraph id="8BCD369B6F15213FF596C1DEFA93989C" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
Superomarginal ossicles 4 lobed and inferomarginal ossicles 3–4 lobed (
|
||
<figureCitation id="13492A1E6F15213FF6BDC1DEFBFB98F9" box="[1004,1072,655,680]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 6</figureCitation>
|
||
), typically bearing single (rarely two) pedicellariae-wreathed spines, longer and more slender than those of the carinal and dorsolateral series.
|
||
</paragraph>
|
||
<paragraph id="8BCD369B6F15213FF596C186FAB499D1" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
Actinal ossicles (
|
||
<figureCitation id="13492A1E6F15213FF4D8C186FE0698A1" box="[393,461,727,752]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 6</figureCitation>
|
||
) in a single nearly complete series, and bearing a single spine. Adambulacral ossicles (
|
||
<figureCitation id="13492A1E6F15213FF5CEC1ADFEC09945" box="[159,267,764,789]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1671,1694]" captionTargetBox="[151,1436,193,1649]" captionTargetId="figure@2.[151,1436,193,1650]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." httpUri="https://zenodo.org/record/244521/files/figure.png" pageId="3" pageNumber="582">Figs. 7–8</figureCitation>
|
||
) bearing single spines on proximal five, then alternating one and two, (two spines in a vertical series), the proximal spine about ¾ as long and thick as the distal spine; actinal and adambulacral spines with a partial wreath on distal side of variously sized straight pedicellariae
|
||
<quantity id="4C8A9B7E6F15213FF603C015FBD9990D" box="[850,1042,836,861]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="7.5" metricValueMin="2.5" pageId="3" pageNumber="582" unit="mm" value="0.5" valueMax="0.75" valueMin="0.25">0.25 to 0.75 mm</quantity>
|
||
. Oral ossicles narrow, bearing two rather slender spines with partial wreaths of pedicellariae similar to those of the actinal and adambulacral series.
|
||
</paragraph>
|
||
<paragraph id="8BCD369B6F15213FF596C0DDFBFC99BC" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
Tube feet in four rows. Color in life: dorsal surface reddish with dark and pale mottlings; spines and madreporite white; oral surface straw colored (
|
||
<figureCitation id="13492A1E6F15213FF793C0FEFCFD9999" box="[706,822,943,968]" captionStart-0="FIGURES 7 – 10" captionStart-1="FIGURE 11" captionStartId-0="2.[151,264,1671,1694]" captionStartId-1="2.[151,250,1794,1816]" captionTargetBox-0="[151,1436,193,1649]" captionTargetId-0="figure@2.[151,1436,193,1650]" captionTargetPageId-0="2" captionText-0="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." captionText-1="FIGURE 11. Leptasterias tatei sp. nov. Live, in situ Hawadax Island, N side, 13 m (leg. R. N. Clark, 8 June, 2008), Bar = 5.0 cm." httpUri-0="https://zenodo.org/record/244521/files/figure.png" pageId="3" pageNumber="582">Fig. 9–11</figureCitation>
|
||
).
|
||
<typeStatus id="54C988396F15213FF61EC0E1FC7A9999" box="[847,945,944,968]" pageId="3" pageNumber="582" type="paratype">Paratype</typeStatus>
|
||
(
|
||
<figureCitation id="13492A1E6F15213FF694C0FEFBD79999" box="[965,1052,943,968]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1671,1694]" captionTargetBox="[151,1436,193,1649]" captionTargetId="figure@2.[151,1436,193,1650]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." httpUri="https://zenodo.org/record/244521/files/figure.png" pageId="3" pageNumber="582">Fig. 10</figureCitation>
|
||
) agrees with the
|
||
<typeStatus id="54C988396F15213FF1A1C0FEFA999999" box="[1264,1362,943,968]" pageId="3" pageNumber="582" type="holotype">holotype</typeStatus>
|
||
in all respects, but is smaller, R =
|
||
<quantity id="4C8A9B7E6F15213FF482C085FDC699BD" box="[467,525,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="582" unit="cm" value="6.0">6 cm</quantity>
|
||
, r =
|
||
<quantity id="4C8A9B7E6F15213FF76EC085FDB199BD" box="[575,634,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="582" unit="cm" value="1.0">1 cm</quantity>
|
||
, R:r 6.1 and has somewhat stouter rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36865106F152138F596C0A6FC6E9A84" lastPageId="4" lastPageNumber="583" pageId="3" pageNumber="582" type="distribution">
|
||
<paragraph id="8BCD369B6F15213FF596C0A6FAD89E2C" blockId="3.[151,1437,151,1149]" pageId="3" pageNumber="582">
|
||
<emphasis id="B906EA896F15213FF596C0A6FEAB9E41" bold="true" box="[199,352,1015,1040]" pageId="3" pageNumber="582">Distribution.</emphasis>
|
||
The distribution of
|
||
<taxonomicName id="4C724D186F15213FF719C0A8FD529E5E" box="[584,665,1016,1040]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="3" pageNumber="582" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F15213FF719C0A8FD529E5E" box="[584,665,1016,1040]" italics="true" pageId="3" pageNumber="582">L. tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F15213FF7F3C0A6FCCE9E41" bold="true" box="[674,773,1015,1040]" pageId="3" pageNumber="582">
|
||
<taxonomicNameLabel id="A23557F26F15213FF7F3C0A6FD349E41" box="[674,767,1015,1040]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
as well as
|
||
<taxonomicName id="4C724D186F15213FF6D6C0A8FBE19E41" box="[903,1066,1016,1040]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="3" pageNumber="582" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F15213FF6D6C0A8FBE19E41" box="[903,1066,1016,1040]" italics="true" pageId="3" pageNumber="582">L. stolacantha</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13492A1E6F15213FF16AC0A6FB459E41" box="[1083,1166,1015,1040]" captionStart="FIGURE 12" captionStartId="2.[151,250,1856,1878]" captionText="FIGURE 12. Leptasterias stolacantha Fisher, 1930. Live, in situ Kagamil Island, E side, 11 m (leg. R. N. Clark, 19 July, 2007), Bar = 2.0 cm." pageId="3" pageNumber="582">Fig. 12</figureCitation>
|
||
) is the central Aleutian Islands, Andreanof and Rat Islands (
|
||
<emphasis id="B906EA896F15213FF764C74CFD409E65" box="[565,651,1053,1076]" italics="true" pageId="3" pageNumber="582">approx.</emphasis>
|
||
176° W to 178° E) (
|
||
<figureCitation id="13492A1E6F15213FF620C74DFC0D9E65" box="[881,966,1052,1077]" captionStart="FIGURE 13" captionStartId="3.[151,250,1948,1970]" captionTargetBox="[363,1218,1219,1899]" captionTargetId="figure@3.[313,1273,1204,1924]" captionTargetPageId="3" captionText="FIGURE 13. Map of Aleutian Islands, showing known geographic distributions of Leptasterias tatei sp. nov. (●) and Leptasterias stolacantha Fisher, 1930 (▲)." httpUri="https://zenodo.org/record/244522/files/figure.png" pageId="3" pageNumber="582">Fig. 13</figureCitation>
|
||
).
|
||
<taxonomicName id="4C724D186F15213FF68AC74CFB6B9E65" box="[987,1184,1053,1076]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="3" pageNumber="582" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F15213FF68AC74CFB6B9E65" box="[987,1184,1053,1076]" italics="true" pageId="3" pageNumber="582">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23557F26F15213FF1F9C74DFAC99E64" box="[1192,1282,1052,1077]" pageId="3" pageNumber="582" rank="species">
|
||
<emphasis id="B906EA896F15213FF1F9C74DFAC99E64" bold="true" box="[1192,1282,1052,1077]" pageId="3" pageNumber="582">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found at Adak (
|
||
<typeStatus id="54C988396F15213FF5B2C76EFE9B9E09" box="[227,336,1087,1112]" pageId="3" pageNumber="582" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<figureCitation id="13492A1E6F15213FF40CC76EFE6B9E09" box="[349,416,1087,1112]" captionStart="FIGURES 1 – 6" captionStartId="1.[151,264,1890,1913]" captionTargetBox="[151,1435,479,1866]" captionTargetId="figure@1.[151,1435,479,1866]" captionTargetPageId="1" captionText="FIGURES 1 – 6. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 1, whole animal, aboral, Bar = 3.0 cm. Fig. 2, aboral surface, Bar = 3.0 mm. Fig. 3, aboral spines & pedicellariae, Bar = 1.0 mm. Fig. 4, aboral surface cleaned of tissue, showing carinal [C] and dorsolateral [DL] ossicles & spines, Bar = 3.0 mm. Fig. 5, aboral spine with pedicellariae, Bar = 1.0 mm. Fig. 6, marginal area of ray, cleaned of tissue, showing arrangement of ossicles (superomarginal [SM], inferomarginal [IM], actinal [AC], and adambulacral [AD]), Bar = 3.0 mm." httpUri="https://zenodo.org/record/244520/files/figure.png" pageId="3" pageNumber="582">Fig. 1</figureCitation>
|
||
;
|
||
<emphasis id="B906EA896F15213FF4FEC711FE3E9E06" box="[431,501,1088,1111]" italics="true" pageId="3" pageNumber="582">in situ</emphasis>
|
||
,
|
||
<figureCitation id="13492A1E6F15213FF755C76EFD8C9E09" box="[516,583,1087,1112]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1671,1694]" captionTargetBox="[151,1436,193,1649]" captionTargetId="figure@2.[151,1436,193,1650]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." httpUri="https://zenodo.org/record/244521/files/figure.png" pageId="3" pageNumber="582">Fig. 9</figureCitation>
|
||
), Amchitka (
|
||
<typeStatus id="54C988396F15213FF7B7C711FC869E09" box="[742,845,1088,1112]" pageId="3" pageNumber="582" type="paratype">Paratype</typeStatus>
|
||
,
|
||
<figureCitation id="13492A1E6F15213FF608C76EFC669E09" box="[857,941,1087,1112]" captionStart="FIGURES 7 – 10" captionStartId="2.[151,264,1671,1694]" captionTargetBox="[151,1436,193,1649]" captionTargetId="figure@2.[151,1436,193,1650]" captionTargetPageId="2" captionText="FIGURES 7 – 10. Leptasterias tatei sp. nov., Holotype, 2011 - 158.001. Fig. 7, cleaned adambulacral region (actinal [AC] & adambulacral [AD] spines), Bar = 3.0 mm. Fig. 8, uncleaned adambulacral region, showing wreaths of straight & crossed pedicellariae. Fig. 9, Live, in situ Adak Island, Thumb Bay, 13 m (leg. R. N. Clark, 20 July, 2011), Bar = 8.0 cm. Fig. 10, Paratype, 2011 - 159.001. Oral side, Bar = 4 cm." httpUri="https://zenodo.org/record/244521/files/figure.png" pageId="3" pageNumber="582">Fig. 10</figureCitation>
|
||
), and Hawadax (formerly Rat) (
|
||
<emphasis id="B906EA896F15213FF04BC711FAAB9E06" box="[1306,1376,1088,1111]" italics="true" pageId="3" pageNumber="582">in situ</emphasis>
|
||
,
|
||
<figureCitation id="13492A1E6F15213FF021C76EFF7E9E2D" captionStart="FIGURE 11" captionStartId="2.[151,250,1794,1816]" captionText="FIGURE 11. Leptasterias tatei sp. nov. Live, in situ Hawadax Island, N side, 13 m (leg. R. N. Clark, 8 June, 2008), Bar = 5.0 cm." pageId="3" pageNumber="582">Fig. 11</figureCitation>
|
||
; not collected) islands. Additional specimens were photographed at Adak as well, but not collected.
|
||
</paragraph>
|
||
<caption id="DF0D66136F15213FF5C6C4CDFD809D81" httpUri="https://zenodo.org/record/244522/files/figure.png" pageId="3" pageNumber="582" targetBox="[363,1218,1219,1899]" targetPageId="3">
|
||
<paragraph id="8BCD369B6F15213FF5C6C4CDFD809D81" blockId="3.[151,1436,1948,2000]" pageId="3" pageNumber="582">
|
||
<emphasis id="B906EA896F15213FF5C6C4CDFEEE9DE0" bold="true" box="[151,293,1948,1970]" pageId="3" pageNumber="582">FIGURE 13.</emphasis>
|
||
Map of Aleutian Islands, showing known geographic distributions of
|
||
<taxonomicName id="4C724D186F15213FF177C4CDFB179DE0" box="[1062,1244,1948,1969]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="3" pageNumber="582" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F15213FF177C4CDFB179DE0" box="[1062,1244,1948,1969]" italics="true" pageId="3" pageNumber="582">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F15213FF1B9C4CDFAF59DE0" bold="true" box="[1256,1342,1948,1969]" pageId="3" pageNumber="582">
|
||
<taxonomicNameLabel id="A23557F26F15213FF1B9C4CDFAF59DE0" box="[1256,1342,1948,1969]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(●) and
|
||
<taxonomicName id="4C724D186F15213FF5C6C4EAFDDE9D81" authority="Fisher, 1930" authorityName="Fisher" authorityYear="1930" box="[151,533,1978,2000]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="3" pageNumber="582" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F15213FF5C6C4EAFE5B9D81" box="[151,400,1979,2000]" italics="true" pageId="3" pageNumber="582">Leptasterias stolacantha</emphasis>
|
||
Fisher, 1930
|
||
</taxonomicName>
|
||
(▲).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCD369B6F122138F596C3C6FC6E9A84" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="583">
|
||
<emphasis id="B906EA896F122138F596C3C6FEE19AE1" bold="true" box="[199,298,151,176]" pageId="4" pageNumber="583">Habitat.</emphasis>
|
||
Found on cobble, boulder and bedrock substrate encrusted with the coralline algae
|
||
<taxonomicName id="4C724D186F122138F18AC3C9FEE09A85" class="Florideophyceae" family="Corallinaceae" genus="Clathromorphum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Corallinales" pageId="4" pageNumber="583" phylum="Rhodophyta" rank="species" species="nereostratum">
|
||
<emphasis id="B906EA896F122138F18AC3C9FEE09A85" italics="true" pageId="4" pageNumber="583">Clathromorphum nereostratum</emphasis>
|
||
</taxonomicName>
|
||
, at depths of
|
||
<quantity id="4C8A9B7E6F122138F493C3EDFDD29A85" box="[450,537,188,213]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="1.6" metricValueMin="0.8" pageId="4" pageNumber="583" unit="m" value="12.0" valueMax="16.0" valueMin="8.0">8–16 m</quantity>
|
||
, and temperatures of 3.8° to 5.0°C.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36865106F122138F596C38EFE0A9B6E" pageId="4" pageNumber="583" type="etymology">
|
||
<paragraph id="8BCD369B6F122138F596C38EFE0A9B6E" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="583">
|
||
<emphasis id="B906EA896F122138F596C38EFE859AA9" bold="true" box="[199,334,223,248]" pageId="4" pageNumber="583">Etymology.</emphasis>
|
||
The name honors Mr. Paul Tate. For more than 40 years he worked throughout the Aleutian Islands, first as a biologist for the Alaska Department of Fish and Game in Dutch Harbor, later as the Captain of the research vessel
|
||
<emphasis id="B906EA896F122138F417C278FE729B11" box="[326,441,297,320]" italics="true" pageId="4" pageNumber="583">Norseman</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36865106F122138F596C21DFCAE9E5E" pageId="4" pageNumber="583" type="discussion">
|
||
<paragraph id="8BCD369B6F122138F596C21DFC379849" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="583">
|
||
<emphasis id="B906EA896F122138F596C21DFEF09B34" bold="true" box="[199,315,332,357]" pageId="4" pageNumber="583">Remarks.</emphasis>
|
||
<taxonomicName id="4C724D186F122138F410C21CFDCF9B35" box="[321,516,333,356]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F410C21CFDCF9B35" box="[321,516,333,356]" italics="true" pageId="4" pageNumber="583">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F75BC21DFDA89B34" bold="true" box="[522,611,332,357]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F75BC21DFDA89B34" box="[522,611,332,357]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C724D186F122138F7B1C21CFBDA9B35" authority="Fisher, 1930" authorityName="Fisher" authorityYear="1930" box="[736,1041,332,357]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F122138F7B1C21CFC4B9B35" box="[736,896,332,356]" italics="true" pageId="4" pageNumber="583">L. stolacantha</emphasis>
|
||
<bibRefCitation id="EFE34B6A6F122138F6D6C21DFBDA9B35" author="Fisher" box="[903,1041,332,357]" pageId="4" pageNumber="583" refString="Fisher, W. K. (1930) Asteroidea of the North Pacific and adjacent waters. Part 3. Forcipulata (concluded). Smithsonian Institution, Washington, D. C., U. S. National Museum Bulletin 76 (Part 3), 1 - 356." type="journal article" year="1930">Fisher, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13492A1E6F122138F171C21DFBB99B35" box="[1056,1138,332,357]" captionStart="FIGURE 12" captionStartId="2.[151,250,1856,1878]" captionText="FIGURE 12. Leptasterias stolacantha Fisher, 1930. Live, in situ Kagamil Island, E side, 11 m (leg. R. N. Clark, 19 July, 2007), Bar = 2.0 cm." pageId="4" pageNumber="583">Fig. 12</figureCitation>
|
||
), but differs in 1) its larger size, 2) having thicker spines, 3) wreaths of fewer and larger crossed pedicellariae, to
|
||
<quantity id="4C8A9B7E6F122138F1DBC23EFB399BD6" box="[1162,1266,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="583" unit="mm" value="0.25">0.25 mm</quantity>
|
||
in length, and scattered large straight, pedicellariae to
|
||
<quantity id="4C8A9B7E6F122138F725C2C5FD2B9BFD" box="[628,736,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="583" unit="mm" value="1.25">1.25 mm</quantity>
|
||
, as opposed to
|
||
<quantity id="4C8A9B7E6F122138F6F5C2C5FBC59BFD" box="[932,1038,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="583" unit="mm" value="0.13">0.13 mm</quantity>
|
||
and
|
||
<quantity id="4C8A9B7E6F122138F101C2C5FB779BFD" box="[1104,1212,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="583" unit="mm" value="0.37">0.37 mm</quantity>
|
||
, respectively in
|
||
<emphasis id="B906EA896F122138F0D4C2C4FEEB9B81" italics="true" pageId="4" pageNumber="583">
|
||
<taxonomicName id="4C724D186F122138F0D4C2C4FED69B81" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="stolacantha">L. stolacantha</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
4) the superomarginal and inferomarginal ossicles bear single spines in
|
||
<taxonomicName id="4C724D186F122138F116C2E8FB5D9B9E" box="[1095,1174,440,464]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F116C2E8FB5D9B9E" box="[1095,1174,440,464]" italics="true" pageId="4" pageNumber="583">L. tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F1CCC2E6FB3D9B81" bold="true" box="[1181,1270,439,464]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F1CCC2E6FB3D9B81" box="[1181,1270,439,464]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, as opposed to two (sometimes three) in
|
||
<taxonomicName id="4C724D186F122138F4E5C28CFD9E9BA5" box="[436,597,476,500]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F122138F4E5C28CFD9E9BA5" box="[436,597,476,500]" italics="true" pageId="4" pageNumber="583">L. stolacantha</emphasis>
|
||
</taxonomicName>
|
||
. Additionally, the spines of
|
||
<taxonomicName id="4C724D186F122138F6DEC28CFBFB9BA5" box="[911,1072,476,500]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F122138F6DEC28CFBFB9BA5" box="[911,1072,476,500]" italics="true" pageId="4" pageNumber="583">L. stolacantha</emphasis>
|
||
</taxonomicName>
|
||
are slender and needle-like. The color of live
|
||
<taxonomicName id="4C724D186F122138F476C150FE0C9849" box="[295,455,512,536]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="stolacantha">
|
||
<emphasis id="B906EA896F122138F476C150FE0C9849" box="[295,455,512,536]" italics="true" pageId="4" pageNumber="583">L. stolacantha</emphasis>
|
||
</taxonomicName>
|
||
also differs in being uniformly tan or light brown.
|
||
</paragraph>
|
||
<paragraph id="8BCD369B6F122138F596C175FCAE9E5E" blockId="4.[151,1437,151,1040]" pageId="4" pageNumber="583">
|
||
Although its thick, truncated abactinal spines and large, straight pedicellariae clearly distinguish
|
||
<taxonomicName id="4C724D186F122138F040C174FF02980E" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F040C174FF02980E" italics="true" pageId="4" pageNumber="583">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F581C116FEE29831" bold="true" box="[208,297,583,608]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F581C116FEE29831" box="[208,297,583,608]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from all other Aleutian species, it bears some resemblance to four congeners described by
|
||
<bibRefCitation id="EFE34B6A6F122138F079C116FF2A98D4" author="Dyakonov" pageId="4" pageNumber="583" refString="Dyakonov, A. M. (1950) Keys to the Fauna of the USSR. No. 34. Sea Stars (Asteroids) of the USSR Seas. Zoological Institute of the Academy of Sciences of the USSR (translated 1968 by Israel Program for Scientific Translations, Jerusalem), USSR / Jerusalem, 183 pp." type="book" year="1950">Dyakonov (1950)</bibRefCitation>
|
||
from the western part of the Okhotsk Sea, chiefly near Sakhalin Island.
|
||
<taxonomicName id="4C724D186F122138F155C13CFA5798D5" authority="Dyakonov, 1929" authorityName="Dyakonov" authorityYear="1929" box="[1028,1436,620,645]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="fisheri">
|
||
<emphasis id="B906EA896F122138F155C13CFB1798D5" box="[1028,1244,620,644]" italics="true" pageId="4" pageNumber="583">Leptasterias fisheri</emphasis>
|
||
Dyakonov, 1929
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C724D186F122138F472C1C1FEA098F6" box="[291,363,656,679]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F472C1C1FEA098F6" box="[291,363,656,679]" italics="true" pageId="4" pageNumber="583">L.tatei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23557F26F122138F423C1DEFE0198F6" box="[370,458,655,680]" pageId="4" pageNumber="583" rank="species">
|
||
<emphasis id="B906EA896F122138F423C1DEFE0F98F9" bold="true" box="[370,452,655,680]" pageId="4" pageNumber="583">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by having the aboral spines arranged in more or less longitudinal rows;
|
||
<taxonomicName id="4C724D186F122138F1BCC1C0FAF698F6" box="[1261,1341,656,680]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F1BCC1C0FAF698F6" box="[1261,1341,656,680]" italics="true" pageId="4" pageNumber="583">L. tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F012C1DEFA5798F9" bold="true" box="[1347,1436,655,680]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F012C1DEFA5798F9" box="[1347,1436,655,680]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has random arrangement.
|
||
<taxonomicName id="4C724D186F122138F4EBC1E4FDB6989D" box="[442,637,693,716]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F4EBC1E4FDB6989D" box="[442,637,693,716]" italics="true" pageId="4" pageNumber="583">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F7D5C1E5FD15989C" bold="true" box="[644,734,692,717]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F7D5C1E5FD15989C" box="[644,734,692,717]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="4C724D186F122138F689C1E4FAB2989D" authority="Dyakonov, 1938" authorityName="Dyakonov" authorityYear="1938" box="[984,1401,692,717]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="hirsuta">
|
||
<emphasis id="B906EA896F122138F689C1E4FB72989D" box="[984,1209,692,716]" italics="true" pageId="4" pageNumber="583">Leptasterias hirsuta</emphasis>
|
||
Dyakonov, 1938
|
||
</taxonomicName>
|
||
by the truncate tips of the aboral spines, and single spines on the inferomarginals. The spines of
|
||
<taxonomicName id="4C724D186F122138F1BAC188FA9098A1" box="[1259,1371,728,752]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="hirsuta">
|
||
<emphasis id="B906EA896F122138F1BAC188FA9098A1" box="[1259,1371,728,752]" italics="true" pageId="4" pageNumber="583">L. hirsuta</emphasis>
|
||
</taxonomicName>
|
||
have rounded tips. Also,
|
||
<taxonomicName id="4C724D186F122138F42DC1ACFE209945" box="[380,491,764,788]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="hirsuta">
|
||
<emphasis id="B906EA896F122138F42DC1ACFE209945" box="[380,491,764,788]" italics="true" pageId="4" pageNumber="583">L. hirsuta</emphasis>
|
||
</taxonomicName>
|
||
has diverse
|
||
<typeStatus id="54C988396F122138F72EC1ACFD719944" box="[639,698,765,789]" pageId="4" pageNumber="583">types</typeStatus>
|
||
of straight pedicellariae scattered aborally that are smaller than those of
|
||
<taxonomicName id="4C724D186F122138F5A9C070FE839966" box="[248,328,800,824]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F5A9C070FE839966" box="[248,328,800,824]" italics="true" pageId="4" pageNumber="583">L. tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F401C04EFE619969" bold="true" box="[336,426,799,824]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F401C04EFE619969" box="[336,426,799,824]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="4C724D186F122138F4E0C071FDBD9966" box="[433,630,800,823]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F4E0C071FDBD9966" box="[433,630,800,823]" italics="true" pageId="4" pageNumber="583">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23557F26F122138F72CC04EFD1C9969" box="[637,727,799,824]" pageId="4" pageNumber="583" rank="species">
|
||
<emphasis id="B906EA896F122138F72CC04EFD1C9969" bold="true" box="[637,727,799,824]" pageId="4" pageNumber="583">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C724D186F122138F63DC071FAE09969" authority="Dyakonov, 1929" authorityName="Dyakonov" authorityYear="1929" box="[876,1323,799,824]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="orientalis">
|
||
<emphasis id="B906EA896F122138F63DC071FBA19969" box="[876,1130,800,824]" italics="true" pageId="4" pageNumber="583">Leptasterias orientalis</emphasis>
|
||
Dyakonov,
|
||
<quantity id="4C8A9B7E6F122138F1A2C071FAE09969" box="[1267,1323,800,824]" metricMagnitude="1" metricUnit="m" metricValue="4.89966" pageId="4" pageNumber="583" unit="in" value="1929.0">1929</quantity>
|
||
</taxonomicName>
|
||
in having only a single row of actinal ossicles.
|
||
<taxonomicName id="4C724D186F122138F702C014FC9E990D" box="[595,853,836,860]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="orientalis">
|
||
<emphasis id="B906EA896F122138F702C014FC9E990D" box="[595,853,836,860]" italics="true" pageId="4" pageNumber="583">Leptasterias orientalis</emphasis>
|
||
</taxonomicName>
|
||
has two rows of actinal ossicles (ventolaterals in Dyakonov), and the spines on the inferomarginals change from one to two at about mid-ray. Also, the aboral spines of
|
||
<taxonomicName id="4C724D186F122138F5E6C0DCFE8999F5" box="[183,322,908,932]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="orientalis">
|
||
<emphasis id="B906EA896F122138F5E6C0DCFE8999F5" box="[183,322,908,932]" italics="true" pageId="4" pageNumber="583">L. orientalis</emphasis>
|
||
</taxonomicName>
|
||
are long and very fine (acicular).
|
||
<taxonomicName id="4C724D186F122138F79FC0DCFC5F99F5" box="[718,916,909,932]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F79FC0DCFC5F99F5" box="[718,916,909,932]" italics="true" pageId="4" pageNumber="583">Leptasterias tatei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23557F26F122138F6CCC0DDFC3299F5" box="[925,1017,908,933]" pageId="4" pageNumber="583" rank="species">
|
||
<emphasis id="B906EA896F122138F6CCC0DDFC3999F4" bold="true" box="[925,1010,908,933]" pageId="4" pageNumber="583">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C724D186F122138F1C2C0DCFE999999" authority="Dyakonov, 1938" authorityName="Dyakonov" authorityYear="1938" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="subarctica">
|
||
<emphasis id="B906EA896F122138F1C2C0DCFA5099F5" box="[1171,1435,908,932]" italics="true" pageId="4" pageNumber="583">Leptasterias subarctica</emphasis>
|
||
Dyakonov,
|
||
<quantity id="4C8A9B7E6F122138F44BC0E1FE999999" box="[282,338,944,968]" metricMagnitude="1" metricUnit="m" metricValue="4.92252" pageId="4" pageNumber="583" unit="in" value="1938.0">1938</quantity>
|
||
</taxonomicName>
|
||
in having smaller actinal ossicles than inferomarginal ossicles; the actinals of
|
||
<taxonomicName id="4C724D186F122138F18AC0E0FABB9999" box="[1243,1392,944,968]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="subarctica">
|
||
<emphasis id="B906EA896F122138F18AC0E0FABB9999" box="[1243,1392,944,968]" italics="true" pageId="4" pageNumber="583">L. subarctica</emphasis>
|
||
</taxonomicName>
|
||
are notably larger than the inferomarginals. Also,
|
||
<taxonomicName id="4C724D186F122138F7F2C084FD3F99BD" box="[675,756,981,1004]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="tatei" status="sp. nov.">
|
||
<emphasis id="B906EA896F122138F7F2C084FD3F99BD" box="[675,756,981,1004]" italics="true" pageId="4" pageNumber="583">L. tatei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B906EA896F122138F7ADC085FC9D99BC" bold="true" box="[764,854,980,1005]" pageId="4" pageNumber="583">
|
||
<taxonomicNameLabel id="A23557F26F122138F7ADC085FC9D99BC" box="[764,854,980,1005]" pageId="4" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has profuse wreaths of crossed pedicellaria around the aboral spines; the wreaths of
|
||
<taxonomicName id="4C724D186F122138F756C0A8FD519E41" box="[519,666,1016,1040]" class="Asteroidea" family="Asteriidae" genus="Leptasterias" kingdom="Animalia" order="Forcipulatida" pageId="4" pageNumber="583" phylum="Echinodermata" rank="species" species="subarctica">
|
||
<emphasis id="B906EA896F122138F756C0A8FD519E41" box="[519,666,1016,1040]" italics="true" pageId="4" pageNumber="583">L. subarctica</emphasis>
|
||
</taxonomicName>
|
||
are much sparser.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |