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<document id="43320923D845544DB0DE93EE5D2B96EA" ID-CLB-Dataset="50957" ID-DOI="10.5281/zenodo.169855" ID-GBIF-Dataset="09004b07-2510-4306-8979-c24c4ccbd9bb" ID-ISSN="1175­5326" ID-Zenodo-Dep="169855" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459747142710" checkinUser="plazi" docAuthor="Tumanov, Denis V." docDate="2005" docId="03DA87E1D14EFF888736FBCAFC1DEDE9" docLanguage="en" docName="zt01043p046.pdf" docOrigin="Zootaxa 1043" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrobiotus barabanovi Tumanov, 2005, sp. nov." docType="treatment" docVersion="10" lastPageNumber="38" masterDocId="FFE3FF99D14FFF8D863EFFF6FF90E927" masterDocTitle="Two new species of Macrobiotus (Eutardigrada, Macrobiotidae) from Tien Shan (Kirghizia), with notes on Macrobiotus tenuis group" masterLastPageNumber="46" masterPageNumber="33" pageNumber="34" updateTime="1698214190570" updateUser="ExternalLinkService">
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<mods:title id="3A9FA03E19A9D388C206063B066DFC4B">Two new species of Macrobiotus (Eutardigrada, Macrobiotidae) from Tien Shan (Kirghizia), with notes on Macrobiotus tenuis group</mods:title>
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<mods:namePart id="A05175658EB5EAFF0A203D93929B5BF1">Tumanov, Denis V.</mods:namePart>
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<mods:date id="9674295E1802032534CC59C5F20EB203">2005</mods:date>
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<mods:classification id="56AD1B0260CE694D98B5F2E249A05B3C">journal article</mods:classification>
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<treatment id="03DA87E1D14EFF888736FBCAFC1DEDE9" ID-DOI="http://doi.org/10.5281/zenodo.5629673" ID-GBIF-Taxon="119336669" ID-Zenodo-Dep="5629673" LSID="urn:lsid:plazi:treatment:03DA87E1D14EFF888736FBCAFC1DEDE9" httpUri="http://treatment.plazi.org/id/03DA87E1D14EFF888736FBCAFC1DEDE9" lastPageId="5" lastPageNumber="38" pageId="1" pageNumber="34">
<subSubSection id="C369657CD14EFF8C8736FBCAFC08ED71" box="[264,920,1084,1110]" pageId="1" pageNumber="34" type="nomenclature">
<paragraph id="8BCC36F7D14EFF8C8736FBCAFC08ED71" blockId="1.[264,920,1084,1110]" box="[264,920,1084,1110]" pageId="1" pageNumber="34">
<heading id="D084819BD14EFF8C8736FBCAFC08ED71" bold="true" box="[264,920,1084,1110]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
<emphasis id="B907EAE5D14EFF8C8736FBCAFC08ED71" bold="true" box="[264,920,1084,1110]" pageId="1" pageNumber="34">
<taxonomicName id="4C734D74D14EFF8C8736FBCAFDA1ED71" ID-CoL="3X2R3" box="[264,561,1084,1110]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="34" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14EFF8C8736FBCAFDA1ED71" bold="true" box="[264,561,1084,1110]" italics="true" pageId="1" pageNumber="34">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14EFF8C8407FBCAFD06ED71" box="[569,662,1084,1110]" pageId="1" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13482A72D14EFF8C8498FBCAFCB5ED71" box="[678,805,1084,1110]" captionStart-0="FIGURES 1 – 8" captionStart-1="FIGURES 9 – 17" captionStartId-0="3.[264,383,1720,1744]" captionStartId-1="4.[264,382,1753,1777]" captionTargetBox-0="[276,1286,323,1652]" captionTargetBox-1="[324,1261,283,1752]" captionTargetId-0="figure@3.[264,1323,302,1694]" captionTargetId-1="figure@4.[319,1282,281,1759]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." captionText-1="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri-0="https://zenodo.org/record/169856/files/figure.png" httpUri-1="https://zenodo.org/record/169857/files/figure.png" pageId="1" pageNumber="34">Figs. 1–17</figureCitation>
,
<tableCitation id="C6F1034CD14EFF8C850DFBCAFC1EED71" box="[819,910,1084,1110]" captionStart="TABLE 1" captionStartId="2.[264,352,686,710]" captionTargetBox="[264,1323,740,1766]" captionTargetPageId="2" captionText="TABLE 1. Summary of morphometric data for Macrobiotus barabanovi sp. nov. (n = 16)." httpUri="http://table.plazi.org/id/DF0C667FD14DFF8F8736FD58FB2EEBE1" pageId="1" pageNumber="34" tableUuid="DF0C667FD14DFF8F8736FD58FB2EEBE1">Table 1</tableCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369657CD14EFF8C8736FB70FEC4EF2E" pageId="1" pageNumber="34" type="materials_examined">
<paragraph id="8BCC36F7D14EFF8C8736FB70FEC4EF2E" blockId="1.[264,1323,1158,1905]" pageId="1" pageNumber="34">
<emphasis id="B907EAE5D14EFF8C8736FB70FE49ED87" bold="true" box="[264,473,1158,1184]" pageId="1" pageNumber="34">
<typeStatus id="54C88855D14EFF8C8736FB70FED5ED87" box="[264,325,1158,1184]" pageId="1" pageNumber="34">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="54C88855D14EFF8C87D9FB71FDC9ED86" box="[487,601,1159,1185]" pageId="1" pageNumber="34" type="holotype">Holotype</typeStatus>
: slide number 197(17) (sex indet.), collected by Dr. A.V. Barabanov (Zoological Institute Russian Academy of Sciences, St. Petersburg)
<date id="FFCD1037D14EFF8C8736FB21FE1EEDD6" box="[264,398,1239,1265]" pageId="1" pageNumber="34" value="1998-08-17">17.08.1998</date>
,
<collectingCountry id="F3647667D14EFF8C87A5FB21FD9CEDD6" box="[411,524,1239,1265]" name="Kyrgyzstan" pageId="1" pageNumber="34">Kirghizia</collectingCountry>
, Chui Area.
<typeStatus id="54C88855D14EFF8C849FFB21FC88EDD6" box="[673,792,1239,1265]" pageId="1" pageNumber="34" type="paratype">Paratypes</typeStatus>
: slide numbers 195(1, 2, 4, 5, 6), 197(16, 18, 20) from the same locality.
<typeStatus id="54C88855D14EFF8C8453FB09FD4AEC3E" box="[621,730,1279,1305]" pageId="1" pageNumber="34" type="holotype">Holotype</typeStatus>
and
<typeStatus id="54C88855D14EFF8C851CFB09FC02EC3E" box="[802,914,1279,1305]" pageId="1" pageNumber="34" type="paratype">paratypes</typeStatus>
are preserved at the Zoological Institute of the Russian Academy of Sciences, St. Petersburg,
<collectingCountry id="F3647667D14EFF8C85DAFAD1FBA6EC66" box="[996,1078,1319,1345]" name="Russia" pageId="1" pageNumber="34">Russia</collectingCountry>
. One
<typeStatus id="54C88855D14EFF8C8245FAD1FB70EC66" box="[1147,1248,1319,1345]" pageId="1" pageNumber="34" type="paratype">paratype</typeStatus>
(slide number 195(1)) preserved in the collection of Binda and Pilato (Department of Animal Biology, University of Catania,
<collectingCountry id="F3647667D14EFF8C84BFFA81FD2BECB6" box="[641,699,1399,1425]" name="Italy" pageId="1" pageNumber="34">Italy</collectingCountry>
).
<typeStatus id="54C88855D14EFF8C84F1FA81FC99ECB6" box="[719,777,1399,1425]" pageId="1" pageNumber="34">Type</typeStatus>
locality:
<collectingCountry id="F3647667D14EFF8C8547FA81FC7AECB6" box="[889,1002,1399,1425]" name="Kyrgyzstan" pageId="1" pageNumber="34">Kirghizia</collectingCountry>
, Chui Area, northern slope of Kirghizian ridge, Turakaiyn canyon, vicinity of Turakaiyn village, near
<quantity id="4C8B9B12D14EFF8C82B8FA69FB71EC9E" box="[1158,1249,1439,1465]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="34" unit="m" value="2000.0">2000 m</quantity>
a.s.l., moss from soil. Twenty adult and juveniles and
<specimenCount id="9D75FD7ED14EFF8C8502FA31FC08ECC6" box="[828,920,1479,1505]" pageId="1" pageNumber="34" type="egg">15 eggs</specimenCount>
(2 with developed embryos) were found.
</paragraph>
</subSubSection>
<subSubSection id="C369657CD14EFF8C8706F9E0FDC8EF7E" pageId="1" pageNumber="34" type="etymology">
<paragraph id="8BCC36F7D14EFF8C8706F9E0FDC8EF7E" blockId="1.[264,1323,1158,1905]" pageId="1" pageNumber="34">
<emphasis id="B907EAE5D14EFF8C8706F9E0FE54EF17" bold="true" box="[312,452,1558,1584]" pageId="1" pageNumber="34">Etymology.</emphasis>
This species is dedicated to Dr. A.V. Barabanov, who kindly collected the material for me in
<collectingCountry id="F3647667D14EFF8C87DCF9C9FDC4EF7E" box="[482,596,1599,1625]" name="Kyrgyzstan" pageId="1" pageNumber="34">Kirghizia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C369657CD14EFF888706F990FCDDEAA9" lastPageId="5" lastPageNumber="38" pageId="1" pageNumber="34" type="description">
<paragraph id="8BCC36F7D14EFF8F8706F990FC17EB5E" blockId="1.[264,1323,1158,1905]" lastBlockId="2.[264,1323,284,633]" lastPageId="2" lastPageNumber="35" pageId="1" pageNumber="34">
<emphasis id="B907EAE5D14EFF8C8706F990FE5FEFA7" bold="true" box="[312,463,1638,1664]" pageId="1" pageNumber="34">Description.</emphasis>
Body long and relatively plump (body length of adult animals
<quantity id="4C8B9B12D14EFF8C82E4F991FEEDEF8E" metricMagnitude="2" metricUnit="m" metricValue="5.2204999999999995" metricValueMax="5.919" metricValueMin="4.522" pageId="1" pageNumber="34" unit="m" value="522.05" valueMax="591.9" valueMin="452.2">452.2– 591.9 m</quantity>
) (
<figureCitation id="13482A72D14EFF8C87A6F979FE7CEF8E" box="[408,492,1679,1705]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="1" pageNumber="34">Figs. 1</figureCitation>
,
<figureCitation id="13482A72D14EFF8C87C3F979FD9EEF8E" box="[509,526,1679,1705]" captionStart="FIGURES 9 – 17" captionStartId="4.[264,382,1753,1777]" captionTargetBox="[324,1261,283,1752]" captionTargetId="figure@4.[319,1282,281,1759]" captionTargetPageId="4" captionText="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="1" pageNumber="34">9</figureCitation>
). Color white (colourless). With little granular eye spots (in some specimens eyespots dissolved during the slide preparation). Cuticle smooth, only with well defined small areas of granulation on the ventral side, similar to those described by
<bibRefCitation id="EFE24B06D14EFF8C82EDF929FEC7EE06" author="Maucci" pageId="1" pageNumber="45" refString="Maucci, W. (1973) Macrobiotus spallanzanii sp. nov. and redescription of Macrobiotus tenuis Binda and Pilato (Tardigrada, Macrobiotidae). Bolletino di Zoologia, 40, 261 - 267." type="journal article" year="1973">Maucci (1973)</bibRefCitation>
in
<taxonomicName id="4C734D74D14EFF8C8742F8F1FDCEEE07" box="[380,606,1799,1824]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="34" phylum="Tardigrada" rank="species" species="tenuis">
<emphasis id="B907EAE5D14EFF8C8742F8F1FDCEEE07" box="[380,606,1799,1824]" italics="true" pageId="1" pageNumber="34">Macrobiotus tenuis</emphasis>
</taxonomicName>
. Typical distribution of these areas is the following: one area between legs I, two areas aligned in a transverse row between legs II and between legs III, three areas aligned transversally between first three pairs of legs, two rows of three areas between the 3rd and the 4th pairs of legs and a pair of areas laterally to anus. Fine dot­like sculpture on legs IV. Bucco­pharyngeal apparatus of
<taxonomicName id="4C734D74D14DFF8F854FFEB1FB96E847" box="[881,1030,327,352]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="35" phylum="Tardigrada" rank="genus">
<emphasis id="B907EAE5D14DFF8F854FFEB1FB96E847" box="[881,1030,327,352]" italics="true" pageId="2" pageNumber="35">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="54C88855D14DFF8F8232FEB1FBAFE846" box="[1036,1087,327,353]" pageId="2" pageNumber="35">type</typeStatus>
(
<figureCitation id="13482A72D14DFF8F8270FEB1FB31E846" box="[1102,1185,327,353]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="2" pageNumber="35">Figs. 2</figureCitation>
,
<figureCitation id="13482A72D14DFF8F8293FEB1FB5BE846" box="[1197,1227,327,353]" captionStart="FIGURES 9 – 17" captionStartId="4.[264,382,1753,1777]" captionTargetBox="[324,1261,283,1752]" captionTargetId="figure@4.[319,1282,281,1759]" captionTargetPageId="4" captionText="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="2" pageNumber="35">10</figureCitation>
) (for all dimensions see
<tableCitation id="C6F1034CD14DFF8F87FDFE99FD8AE8AE" box="[451,538,367,393]" captionStart="TABLE 1" captionStartId="2.[264,352,686,710]" captionTargetBox="[264,1323,740,1766]" captionTargetPageId="2" captionText="TABLE 1. Summary of morphometric data for Macrobiotus barabanovi sp. nov. (n = 16)." httpUri="http://table.plazi.org/id/DF0C667FD14DFF8F8736FD58FB2EEBE1" pageId="2" pageNumber="35" tableUuid="DF0C667FD14DFF8F8736FD58FB2EEBE1">Table 1</tableCitation>
). Buccal cavity with ten peribuccal lamellae, a posterior crown of triangular teeth and a system of three dorsal and three ventral transverse ridges dorsally and ventrally, consisting of medial and paired lateral crests (
<figureCitation id="13482A72D14DFF8F85CDFE49FBFDE8FE" box="[1011,1133,447,473]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="2" pageNumber="35">Figs. 3, 4</figureCitation>
,
<figureCitation id="13482A72D14DFF8F8243FE49FB5EE8FE" box="[1149,1230,447,473]" captionStart="FIGURES 9 – 17" captionStartId="4.[264,382,1753,1777]" captionTargetBox="[324,1261,283,1752]" captionTargetId="figure@4.[319,1282,281,1759]" captionTargetPageId="4" captionText="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="2" pageNumber="35">11, 12</figureCitation>
). Large specimens with anterior band of minute dots. Ventral strengthening bar relatively long exceeding half of the length of the buccal tube. Pharyngeal bulb suboval, with well developed apophyses, three macroplacoids and elongated microplacoid. Third macroplacoid with a distinct pre­terminal constriction.
</paragraph>
<caption id="DF0C667FD14DFF8F8736FD58FB2EEBE1" ID-Table-UUID="DF0C667FD14DFF8F8736FD58FB2EEBE1" box="[264,1214,686,710]" httpUri="http://table.plazi.org/id/DF0C667FD14DFF8F8736FD58FB2EEBE1" pageId="2" pageNumber="35" targetBox="[264,1323,740,1766]" targetIsTable="true" targetPageId="2">
<paragraph id="8BCC36F7D14DFF8F8736FD58FB2EEBE1" blockId="2.[264,1214,686,710]" box="[264,1214,686,710]" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8736FD58FEEBEBE1" bold="true" box="[264,379,686,710]" pageId="2" pageNumber="35">TABLE 1.</emphasis>
Summary of morphometric data for
<taxonomicName id="4C734D74D14DFF8F853AFD59FB9DEBE1" box="[772,1037,687,710]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="35" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14DFF8F853AFD59FB9DEBE1" box="[772,1037,687,710]" italics="true" pageId="2" pageNumber="35">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14DFF8F822DFD58FBF9EBE1" box="[1043,1129,686,710]" pageId="2" pageNumber="35" rank="species">
<emphasis id="B907EAE5D14DFF8F822DFD58FBF9EBE1" bold="true" box="[1043,1129,686,710]" pageId="2" pageNumber="35">sp. nov.</emphasis>
</taxonomicNameLabel>
(n=16).
</paragraph>
</caption>
<paragraph id="8BCC36F7D14DFF8F8724FD12FA99EFC7" pageId="2" pageNumber="35">
<table id="F973C457D14D00728736FD12FABBEFC1" box="[264,1323,740,1766]" gridcols="6" gridrows="11" pageId="2" pageNumber="35">
<tr id="354334B5D14D00728736FD12FABBEA39" box="[264,1323,740,798]" gridrow="0" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FD12FDE5EA39" box="[264,629,740,798]" gridcol="0" gridrow="0" pageId="2" pageNumber="35">Character</th>
<th id="76925DC9D14D007284B7FD12FD7CEA39" box="[649,748,740,798]" gridcol="1" gridrow="0" pageId="2" pageNumber="35">Holotype</th>
<th id="76925DC9D14D007284C3FD12FCF2EA39" box="[765,866,740,798]" gridcol="2" gridrow="0" pageId="2" pageNumber="35">Smallest specimen</th>
<th id="76925DC9D14D00728549FD12FC4CEA39" box="[887,988,740,798]" gridcol="3" gridrow="0" pageId="2" pageNumber="35">Largest specimen</th>
<th id="76925DC9D14D007285D1FD12FB23EA39" box="[1007,1203,740,798]" gridcol="4" gridrow="0" pageId="2" pageNumber="35">Mean (Range)</th>
<th id="76925DC9D14D007282FBFD12FABBEA39" box="[1221,1323,740,798]" gridcol="5" gridrow="0" pageId="2" pageNumber="35">Standard error</th>
</tr>
<tr id="354334B5D14D00728736FCD9FABBEA52" box="[264,1323,815,885]" gridrow="1" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FCD9FDE5EA52" box="[264,629,815,885]" gridcol="0" gridrow="1" pageId="2" pageNumber="35">Body length (m) Buccal tube length (m)</th>
<td id="76925DC9D14D007284B7FCD9FD7CEA52" box="[649,748,815,885]" gridcol="1" gridrow="1" pageId="2" pageNumber="35">591.9 51.1</td>
<td id="76925DC9D14D007284C3FCD9FCF2EA52" box="[765,866,815,885]" gridcol="2" gridrow="1" pageId="2" pageNumber="35">452.2 44.4</td>
<td id="76925DC9D14D00728549FCD9FC4CEA52" box="[887,988,815,885]" gridcol="3" gridrow="1" pageId="2" pageNumber="35">591.9 51.1</td>
<td id="76925DC9D14D007285D1FCD9FB23EA52" box="[1007,1203,815,885]" gridcol="4" gridrow="1" pageId="2" pageNumber="35">514.3 (452.2–591.9) 47.9 (39.2–54.8)</td>
<td id="76925DC9D14D007282FBFCD9FABBEA52" box="[1221,1323,815,885]" gridcol="5" gridrow="1" pageId="2" pageNumber="35">10.42 1.09</td>
</tr>
<tr id="354334B5D14D00728736FC70FABBEA87" box="[264,1323,902,928]" gridrow="2" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FC70FDE5EA87" box="[264,629,902,928]" gridcol="0" gridrow="2" pageId="2" pageNumber="35">Ss (m)</th>
<td id="76925DC9D14D007284B7FC70FD7CEA87" box="[649,748,902,928]" gridcol="1" gridrow="2" pageId="2" pageNumber="35">39.2</td>
<td id="76925DC9D14D007284C3FC70FCF2EA87" box="[765,866,902,928]" gridcol="2" gridrow="2" pageId="2" pageNumber="35">33.7</td>
<td id="76925DC9D14D00728549FC70FC4CEA87" box="[887,988,902,928]" gridcol="3" gridrow="2" pageId="2" pageNumber="35">39.2</td>
<td id="76925DC9D14D007285D1FC70FB23EA87" box="[1007,1203,902,928]" gridcol="4" gridrow="2" pageId="2" pageNumber="35">36.4 (29.6–42.2)</td>
<td id="76925DC9D14D007282FBFC70FABBEA87" box="[1221,1323,902,928]" gridcol="5" gridrow="2" pageId="2" pageNumber="35">0.87</td>
</tr>
<tr id="354334B5D14D00728736FC47FABBED3B" box="[264,1323,945,1052]" gridrow="3" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FC47FDE5ED3B" box="[264,629,945,1052]" gridcol="0" gridrow="3" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735FC44FEB0EAEE" box="[267,288,946,969]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
Ss Outer buccal tube width (m)
<emphasis id="B907EAE5D14DFF8F8735FBF3FEB0ED3B" box="[267,288,1029,1052]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of outer buccal tube width
</th>
<td id="76925DC9D14D007284B7FC47FD7CED3B" box="[649,748,945,1052]" gridcol="1" gridrow="3" pageId="2" pageNumber="35">76.8 8.1 15.9</td>
<td id="76925DC9D14D007284C3FC47FCF2ED3B" box="[765,866,945,1052]" gridcol="2" gridrow="3" pageId="2" pageNumber="35">75.8 6.8 15.3</td>
<td id="76925DC9D14D00728549FC47FC4CED3B" box="[887,988,945,1052]" gridcol="3" gridrow="3" pageId="2" pageNumber="35">76.8 8.1 15.9</td>
<td id="76925DC9D14D007285D1FC47FB23ED3B" box="[1007,1203,945,1052]" gridcol="4" gridrow="3" pageId="2" pageNumber="35">76.1 (75.0–77.1) 7.4 (5.9–9.6) 15.5 (13.6–17.6)</td>
<td id="76925DC9D14D007282FBFC47FABBED3B" box="[1221,1323,945,1052]" gridcol="5" gridrow="3" pageId="2" pageNumber="35">0.16 0.27 0.27</td>
</tr>
<tr id="354334B5D14D00728736FBDBFABBED60" box="[264,1323,1069,1095]" gridrow="4" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FBDBFDE5ED60" box="[264,629,1069,1095]" gridcol="0" gridrow="4" pageId="2" pageNumber="35">Placoid row length (m)</th>
<td id="76925DC9D14D007284B7FBDBFD7CED60" box="[649,748,1069,1095]" gridcol="1" gridrow="4" pageId="2" pageNumber="35">26.6</td>
<td id="76925DC9D14D007284C3FBDBFCF2ED60" box="[765,866,1069,1095]" gridcol="2" gridrow="4" pageId="2" pageNumber="35">25.2</td>
<td id="76925DC9D14D00728549FBDBFC4CED60" box="[887,988,1069,1095]" gridcol="3" gridrow="4" pageId="2" pageNumber="35">26.6</td>
<td id="76925DC9D14D007285D1FBDBFB23ED60" box="[1007,1203,1069,1095]" gridcol="4" gridrow="4" pageId="2" pageNumber="35">27.4 (22.2–32.6)</td>
<td id="76925DC9D14D007282FBFBDBFABBED60" box="[1221,1323,1069,1095]" gridcol="5" gridrow="4" pageId="2" pageNumber="35">0.86</td>
</tr>
<tr id="354334B5D14D00728736FBAEFABBEDE4" box="[264,1323,1112,1219]" gridrow="5" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FBAEFDE5EDE4" box="[264,629,1112,1219]" gridcol="0" gridrow="5" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735FBAFFEB0ED57" box="[267,288,1113,1136]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of placoid row length Macroplacoid row length (m)
<emphasis id="B907EAE5D14DFF8F8735FB5AFEB0EDE4" box="[267,288,1196,1219]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of macroplacoid row length
</th>
<td id="76925DC9D14D007284B7FBAEFD7CEDE4" box="[649,748,1112,1219]" gridcol="1" gridrow="5" pageId="2" pageNumber="35">52.2 22.2 43.5</td>
<td id="76925DC9D14D007284C3FBAEFCF2EDE4" box="[765,866,1112,1219]" gridcol="2" gridrow="5" pageId="2" pageNumber="35">56.7 19.6 44.2</td>
<td id="76925DC9D14D00728549FBAEFC4CEDE4" box="[887,988,1112,1219]" gridcol="3" gridrow="5" pageId="2" pageNumber="35">52.2 22.2 43.5</td>
<td id="76925DC9D14D007285D1FBAEFB23EDE4" box="[1007,1203,1112,1219]" gridcol="4" gridrow="5" pageId="2" pageNumber="35">57.1 (52.2–61.4) 21.9 (17.0–26.6) 45.7 (42.6–50.0)</td>
<td id="76925DC9D14D007282FBFBAEFABBEDE4" box="[1221,1323,1112,1219]" gridcol="5" gridrow="5" pageId="2" pageNumber="35">0.66 0.71 0.59</td>
</tr>
<tr id="354334B5D14D00728736FB22FABBEDC9" box="[264,1323,1236,1262]" gridrow="6" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FB22FDE5EDC9" box="[264,629,1236,1262]" gridcol="0" gridrow="6" pageId="2" pageNumber="35">1st macroplacoid length (m)</th>
<td id="76925DC9D14D007284B7FB22FD7CEDC9" box="[649,748,1236,1262]" gridcol="1" gridrow="6" pageId="2" pageNumber="35">7.0</td>
<td id="76925DC9D14D007284C3FB22FCF2EDC9" box="[765,866,1236,1262]" gridcol="2" gridrow="6" pageId="2" pageNumber="35">5.9</td>
<td id="76925DC9D14D00728549FB22FC4CEDC9" box="[887,988,1236,1262]" gridcol="3" gridrow="6" pageId="2" pageNumber="35">7.0</td>
<td id="76925DC9D14D007285D1FB22FB23EDC9" box="[1007,1203,1236,1262]" gridcol="4" gridrow="6" pageId="2" pageNumber="35">6.9 (4.4–8.9)</td>
<td id="76925DC9D14D007282FBFB22FABBEDC9" box="[1221,1323,1236,1262]" gridcol="5" gridrow="6" pageId="2" pageNumber="35">0.31</td>
</tr>
<tr id="354334B5D14D00728736FB09FABBEC4D" box="[264,1323,1279,1386]" gridrow="7" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FB09FDE5EC4D" box="[264,629,1279,1386]" gridcol="0" gridrow="7" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735FAF6FEB0EC30" box="[267,288,1280,1303]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of 1st macroplacoid length 2nd macroplacoid length (m)
<emphasis id="B907EAE5D14DFF8F8735FAA5FEB0EC4D" box="[267,288,1363,1386]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of 2nd macroplacoid length
</th>
<td id="76925DC9D14D007284B7FB09FD7CEC4D" box="[649,748,1279,1386]" gridcol="1" gridrow="7" pageId="2" pageNumber="35">13.8 5.2 10.1</td>
<td id="76925DC9D14D007284C3FB09FCF2EC4D" box="[765,866,1279,1386]" gridcol="2" gridrow="7" pageId="2" pageNumber="35">13.3 4.8 10.8</td>
<td id="76925DC9D14D00728549FB09FC4CEC4D" box="[887,988,1279,1386]" gridcol="3" gridrow="7" pageId="2" pageNumber="35">13.8 5.2 10.1</td>
<td id="76925DC9D14D007285D1FB09FB23EC4D" box="[1007,1203,1279,1386]" gridcol="4" gridrow="7" pageId="2" pageNumber="35">14.4 (11.3–16.4) 5.6 (4.4–6.7) 11.6 (9.8–13.6)</td>
<td id="76925DC9D14D007282FBFB09FABBEC4D" box="[1221,1323,1279,1386]" gridcol="5" gridrow="7" pageId="2" pageNumber="35">0.37 0.21 0.30</td>
</tr>
<tr id="354334B5D14D00728736FA8DFABBECB2" box="[264,1323,1403,1429]" gridrow="8" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FA8DFDE5ECB2" box="[264,629,1403,1429]" gridcol="0" gridrow="8" pageId="2" pageNumber="35">3rd macroplacoid length (m)</th>
<td id="76925DC9D14D007284B7FA8DFD7CECB2" box="[649,748,1403,1429]" gridcol="1" gridrow="8" pageId="2" pageNumber="35">7.4</td>
<td id="76925DC9D14D007284C3FA8DFCF2ECB2" box="[765,866,1403,1429]" gridcol="2" gridrow="8" pageId="2" pageNumber="35">6.7</td>
<td id="76925DC9D14D00728549FA8DFC4CECB2" box="[887,988,1403,1429]" gridcol="3" gridrow="8" pageId="2" pageNumber="35">7.4</td>
<td id="76925DC9D14D007285D1FA8DFB23ECB2" box="[1007,1203,1403,1429]" gridcol="4" gridrow="8" pageId="2" pageNumber="35">6.9 (4.8–8.1)</td>
<td id="76925DC9D14D007282FBFA8DFABBECB2" box="[1221,1323,1403,1429]" gridcol="5" gridrow="8" pageId="2" pageNumber="35">0.22</td>
</tr>
<tr id="354334B5D14D00728736FA50FABBEF1B" box="[264,1323,1446,1596]" gridrow="9" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736FA50FDE5EF1B" box="[264,629,1446,1596]" gridcol="0" gridrow="9" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735FA51FEB0EC99" box="[267,288,1447,1470]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of 3rd macroplacoid length Microplacoid length (m)
<emphasis id="B907EAE5D14DFF8F8735FA0CFEB0EF36" box="[267,288,1530,1553]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of microplacoid length External claw length, leg I (m)
</th>
<td id="76925DC9D14D007284B7FA50FD7CEF1B" box="[649,748,1446,1596]" gridcol="1" gridrow="9" pageId="2" pageNumber="35">14.5 3.7 7.2 17.0</td>
<td id="76925DC9D14D007284C3FA50FCF2EF1B" box="[765,866,1446,1596]" gridcol="2" gridrow="9" pageId="2" pageNumber="35">15.0 4.4 10.0 14.8</td>
<td id="76925DC9D14D00728549FA50FC4CEF1B" box="[887,988,1446,1596]" gridcol="3" gridrow="9" pageId="2" pageNumber="35">14.5 3.7 7.2 17.0</td>
<td id="76925DC9D14D007285D1FA50FB23EF1B" box="[1007,1203,1446,1596]" gridcol="4" gridrow="9" pageId="2" pageNumber="35">14.3 (12.3–15.7) 4.2 (3.0–5.2) 8.8 (7.2–10.3) 18.5 (14.8–22.2)</td>
<td id="76925DC9D14D007282FBFA50FABBEF1B" box="[1221,1323,1446,1596]" gridcol="5" gridrow="9" pageId="2" pageNumber="35">0.21 0.16 0.27 0.57</td>
</tr>
<tr id="354334B5D14D00728736F9BBFABBEFC1" box="[264,1323,1613,1766]" gridrow="10" pageId="2" pageNumber="35">
<th id="76925DC9D14D00728736F9BBFDE5EFC1" box="[264,629,1613,1766]" gridcol="0" gridrow="10" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735F9B8FEB0EF42" box="[267,288,1614,1637]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of external claw length, leg I External claw length, leg III (m)
<emphasis id="B907EAE5D14DFF8F8735F957FEB0EF9F" box="[267,288,1697,1720]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of external claw length, leg III External claw length, leg IV (m)
</th>
<td id="76925DC9D14D007284B7F9BBFD7CEFC1" box="[649,748,1613,1766]" gridcol="1" gridrow="10" pageId="2" pageNumber="35">33.3 – – 30.3</td>
<td id="76925DC9D14D007284C3F9BBFCF2EFC1" box="[765,866,1613,1766]" gridcol="2" gridrow="10" pageId="2" pageNumber="35">33.3 16.3 36.7 22.9</td>
<td id="76925DC9D14D00728549F9BBFC4CEFC1" box="[887,988,1613,1766]" gridcol="3" gridrow="10" pageId="2" pageNumber="35">33.3 – – 30.3</td>
<td id="76925DC9D14D007285D1F9BBFB23EFC1" box="[1007,1203,1613,1766]" gridcol="4" gridrow="10" pageId="2" pageNumber="35">38.8 (32.3–44.1) 20.5 (16.3–25.2) 43.5 (36.7–51.7) 28.7 (22.9–32.9)</td>
<td id="76925DC9D14D007282FBF9BBFABBEFC1" box="[1221,1323,1613,1766]" gridcol="5" gridrow="10" pageId="2" pageNumber="35">0.99 0.79 1.45 0.75</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCC36F7D14DFF8F8735F90EFC5CEE08" blockId="2.[264,1323,1783,1893]" pageId="2" pageNumber="35">
<emphasis id="B907EAE5D14DFF8F8735F90EFEB0EE28" box="[267,288,1784,1807]" italics="true" pageId="2" pageNumber="35">pt</emphasis>
of external claw length, leg IV 59.4 51.7 59.4 60.1 (51.7–66.7) 1.22 Ss = distance from cephalic end of buccal tube to insertion point of stylet supports.
</paragraph>
<paragraph id="8BCC36F7D14DFF8F8736F8C1FDD2EE42" blockId="2.[264,1323,1783,1893]" pageId="2" pageNumber="35">
Supplementary measurements for
<typeStatus id="54C88855D14DFF8F8412F8C1FDEAEE6D" box="[556,634,1847,1866]" pageId="2" pageNumber="35" type="holotype">holotype</typeStatus>
: length of strengthening bar on buccal tube: above
<quantity id="4C8B9B12D14DFF8F820FF8C1FBECEE6D" box="[1073,1148,1847,1866]" metricMagnitude="1" metricUnit="m" metricValue="3.26" pageId="2" pageNumber="35" unit="m" value="32.6">32.6 m</quantity>
, lunules on claws of the fourth pair of legs: 5.9 x
<quantity id="4C8B9B12D14DFF8F87C2F8A4FDD2EE42" box="[508,578,1874,1893]" metricMagnitude="1" metricUnit="m" metricValue="1.1099999999999999" pageId="2" pageNumber="35" unit="m" value="11.1">11.1 m</quantity>
</paragraph>
<caption id="DF0C667FD14CFF8E8736F94EFC30EE10" httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="3" pageNumber="36" targetBox="[276,1286,323,1652]" targetPageId="3">
<paragraph id="8BCC36F7D14CFF8E8736F94EFC30EE10" blockId="3.[264,1323,1720,1847]" pageId="3" pageNumber="36">
<emphasis id="B907EAE5D14CFF8E8736F94EFE20EFF7" bold="true" box="[264,432,1720,1744]" pageId="3" pageNumber="36">FIGURES 1–8</emphasis>
.
<taxonomicName id="4C734D74D14CFF8E87FFF94FFD5EEFF7" box="[449,718,1721,1744]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="36" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14CFF8E87FFF94FFD5EEFF7" box="[449,718,1721,1744]" italics="true" pageId="3" pageNumber="36">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14CFF8E84E9F94EFCA0EFF7" box="[727,816,1720,1744]" pageId="3" pageNumber="36" rank="species">
<emphasis id="B907EAE5D14CFF8E84E9F94EFCA0EFF7" bold="true" box="[727,816,1720,1744]" pageId="3" pageNumber="36">sp. nov.</emphasis>
</taxonomicNameLabel>
1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series).
</paragraph>
</caption>
<caption id="DF0C667FD14BFF898736F92FFE18EE57" httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="4" pageNumber="37" targetBox="[324,1261,283,1752]" targetPageId="4">
<paragraph id="8BCC36F7D14BFF898736F92FFE18EE57" blockId="4.[264,1323,1753,1904]" pageId="4" pageNumber="37">
<emphasis id="B907EAE5D14BFF898736F92FFE2BEFD6" bold="true" box="[264,443,1753,1777]" pageId="4" pageNumber="37">FIGURES 9–17</emphasis>
.
<taxonomicName id="4C734D74D14BFF8987F4F92CFD44EFD6" box="[458,724,1754,1777]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="37" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14BFF8987F4F92CFD44EFD6" box="[458,724,1754,1777]" italics="true" pageId="4" pageNumber="37">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14BFF8984E2F92FFCA2EFD6" box="[732,818,1753,1777]" pageId="4" pageNumber="37" rank="species">
<emphasis id="B907EAE5D14BFF8984E2F92FFCA2EFD6" bold="true" box="[732,818,1753,1777]" pageId="4" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9–12, 14 – holotype, 13 – paratype).
</paragraph>
</caption>
<paragraph id="8BCC36F7D14AFF888706FEEAFB88EB51" blockId="5.[264,1325,284,1230]" pageId="5" pageNumber="38">
Claws long, thin, with flexible main branches (degree of bending of the distal part of main branches could vary much even within the same leg), minute stalk, poorly marked distal part of the basal portion and short common tract (
<figureCitation id="13482A72D14AFF888590FE9AFBB4E8A1" box="[942,1060,364,390]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="5" pageNumber="38">Figs. 5, 6</figureCitation>
,
<figureCitation id="13482A72D14AFF88820DFE9AFB12E8A1" box="[1075,1154,364,390]" captionStart="FIGURES 9 – 17" captionStartId="4.[264,382,1753,1777]" captionTargetBox="[324,1261,283,1752]" captionTargetId="figure@4.[319,1282,281,1759]" captionTargetPageId="4" captionText="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="5" pageNumber="38">13, 14</figureCitation>
). Primary and secondary branches diverge near 1/3 of claw length, main branches with well developed accessory points. Claws of fourth pair of legs longer than claws of first three pairs of legs. All claws with large lunules, especially on legs IV. Lunules of legs IV with distinct marginal teeth, lunules of legs I–III with slightly waved edges. Lunules of legs IV with clearly darkened medial area, the same area is scarcely visible in lunules of legs I–III. Legs I–III with long curved band of thickened cuticle below claw bases.
</paragraph>
<paragraph id="8BCC36F7D14AFF888706FD72FCDDEAA9" blockId="5.[264,1325,284,1230]" pageId="5" pageNumber="38">
Eggs laid free, spherical in shape, white, and ornamented with bulbous processes with attenuated apical parts clearly septate and sometimes bent downward (
<figureCitation id="13482A72D14AFF88826BFD5AFB58EBE1" box="[1109,1224,684,710]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – bucco­pharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="5" pageNumber="38">Figs. 7, 8</figureCitation>
,
<figureCitation id="13482A72D14AFF8882E6FD5AFEB8EBC9" captionStart="FIGURES 9 – 17" captionStartId="4.[264,382,1753,1777]" captionTargetBox="[324,1261,283,1752]" captionTargetId="figure@4.[319,1282,281,1759]" captionTargetPageId="4" captionText="FIGURES 9 – 17. Macrobiotus barabanovi sp. nov. 9 – habitus, 10 – bucco­pharyngeal apparatus, ventral, 11 – buccal armature, dorsal, 12 – buccal armature, ventral, 13 – claws of the II pair of legs (arrow points to the cuticular thickening), 14 – claws of the IV pair of legs, 15 – general appearance of the egg, 16 – part of the egg surface, 17 – egg processes (phase contrast; 9 – 12, 14 – holotype, 13 – paratype)." httpUri="https://zenodo.org/record/169857/files/figure.png" pageId="5" pageNumber="38">15, 16, 17</figureCitation>
). Processes with developed polygonal to roundish reticulation. Processes bases with small projections. Egg shell between processes with scarcely visible radial ridges. Usually egg surface poorly visible, covered with debris particles. Diameter of eggs (without processes):
<quantity id="4C8B9B12D14AFF8887ABFCBAFDD0EA41" box="[405,576,844,870]" metricMagnitude="1" metricUnit="m" metricValue="8.625" metricValueMax="9.129999999999999" metricValueMin="8.12" pageId="5" pageNumber="38" unit="m" value="86.25" valueMax="91.3" valueMin="81.2">81.2–91.3 m</quantity>
; height of processes (about 20 on the circumference of egg): 11.0–
<quantity id="4C8B9B12D14AFF888774FC82FE3EEAA9" box="[330,430,884,910]" metricMagnitude="1" metricUnit="m" metricValue="1.72" pageId="5" pageNumber="38" unit="m" value="17.2">17.2 m</quantity>
, their basal diameter:
<quantity id="4C8B9B12D14AFF88848FFC82FCD9EAA9" box="[689,841,884,910]" metricMagnitude="1" metricUnit="m" metricValue="1.06" metricValueMax="1.27" metricValueMin="0.85" pageId="5" pageNumber="38" unit="m" value="10.6" valueMax="12.7" valueMin="8.5">8.5–12.7 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C369657CD14AFF888706FC6AFC1DEDE9" pageId="5" pageNumber="38" type="discussion">
<paragraph id="8BCC36F7D14AFF888706FC6AFC1DEDE9" blockId="5.[264,1325,284,1230]" pageId="5" pageNumber="38">
<emphasis id="B907EAE5D14AFF888706FC6AFE21EA91" bold="true" box="[312,433,924,950]" pageId="5" pageNumber="38">Remarks.</emphasis>
<taxonomicName id="4C734D74D14AFF888783FC6AFD76EA92" box="[445,742,924,949]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14AFF888783FC6AFD76EA92" box="[445,742,924,949]" italics="true" pageId="5" pageNumber="38">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14AFF8884CDFC6AFCC5EA91" box="[755,853,924,950]" pageId="5" pageNumber="38" rank="species">
<emphasis id="B907EAE5D14AFF8884CDFC6AFCC5EA91" bold="true" box="[755,853,924,950]" pageId="5" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the
<emphasis id="B907EAE5D14AFF888220FC6AFB24EA92" box="[1054,1204,924,949]" italics="true" pageId="5" pageNumber="38">harmsworthi</emphasis>
group. It could be easily distinguished from the majority of species of this group in having noticeably longer claws. Within the
<emphasis id="B907EAE5D14AFF888490FC1AFCD4ED22" box="[686,836,1004,1029]" italics="true" pageId="5" pageNumber="38">harmsworthi</emphasis>
group, the only species with similar claw morphology is
<taxonomicName id="4C734D74D14AFF8887FFFBE2FC19ED09" authority="Biserov, 1997" authorityName="Biserov" authorityYear="1997" box="[449,905,1044,1070]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="altitudinalis">
<emphasis id="B907EAE5D14AFF8887FFFBE2FD16ED0A" box="[449,646,1044,1069]" italics="true" pageId="5" pageNumber="38">M. altitudinalis</emphasis>
<bibRefCitation id="EFE24B06D14AFF8884AEFBE2FCAAED09" author="Biserov" box="[656,826,1044,1070]" pageId="5" pageNumber="45" refString="Biserov, V. (1997 (1998 )) Tardigrades of the Caucasus with a Taxonomic Analysis of the Genus Ramazzottius (Parachela: Hypsibiidae). Zoologischer Anzeiger, 236, 139 - 159." type="journal article" year="1997">Biserov, 1997</bibRefCitation>
(1998)
</taxonomicName>
.
<taxonomicName id="4C734D74D14AFF8885A9FBE2FB2EED0A" box="[919,1214,1044,1069]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14AFF8885A9FBE2FB2EED0A" box="[919,1214,1044,1069]" italics="true" pageId="5" pageNumber="38">Macrobiotus barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14AFF8882F7FBE2FABAED09" box="[1225,1322,1044,1070]" pageId="5" pageNumber="38" rank="species">
<emphasis id="B907EAE5D14AFF8882F7FBE2FABAED09" bold="true" box="[1225,1322,1044,1070]" pageId="5" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from this species in having longer claws of the hind legs (
<emphasis id="B907EAE5D14AFF888237FBCAFBB0ED72" box="[1033,1056,1084,1109]" italics="true" pageId="5" pageNumber="38">pt</emphasis>
value
<quantity id="4C8B9B12D14AFF888251FBCAFA93ED71" box="[1135,1283,1084,1110]" metricMagnitude="0" metricUnit="m" metricValue="1.51384" metricValueMax="1.69418" metricValueMin="1.3335" pageId="5" pageNumber="38" unit="in" value="59.6" valueMax="66.7" valueMin="52.5">52.5–66.7 in</quantity>
<taxonomicName id="4C734D74D14AFF888335FBCAFE1FED5A" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
<emphasis id="B907EAE5D14AFF888335FBCAFE1FED5A" italics="true" pageId="5" pageNumber="38">M. barabanovi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A234579ED14AFF8887A7FB92FE68ED59" box="[409,504,1124,1150]" pageId="5" pageNumber="38" rank="species">
<emphasis id="B907EAE5D14AFF8887A7FB92FE68ED59" bold="true" box="[409,504,1124,1150]" pageId="5" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
and 47.2 (mean
<emphasis id="B907EAE5D14AFF8884FBFB92FD4CED5A" box="[709,732,1124,1149]" italics="true" pageId="5" pageNumber="38">pt</emphasis>
) in
<taxonomicName id="4C734D74D14AFF888531FB92FC56ED5A" box="[783,966,1124,1149]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="altitudinalis">
<emphasis id="B907EAE5D14AFF888531FB92FC56ED5A" box="[783,966,1124,1149]" italics="true" pageId="5" pageNumber="38">M. altitudinalis</emphasis>
</taxonomicName>
), relatively shorter secondary branches of claws of the hind legs, lunules of the legs IV with marginal teeth and darkened zone, eggs with smaller and more numerous processes.
</paragraph>
</subSubSection>
</treatment>
</document>
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