612 lines
78 KiB
XML
612 lines
78 KiB
XML
<document id="0376F55FE2CBA2135C2E6F49ECF87B43" ID-CLB-Dataset="50957" ID-DOI="10.5281/zenodo.169855" ID-GBIF-Dataset="09004b07-2510-4306-8979-c24c4ccbd9bb" ID-ISSN="11755326" ID-Zenodo-Dep="169855" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459747142710" checkinUser="plazi" docAuthor="Tumanov, Denis V." docDate="2005" docId="03DA87E1D14AFF818736FADAFD17EBE1" docLanguage="en" docName="zt01043p046.pdf" docOrigin="Zootaxa 1043" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrobiotus kirghizicus Tumanov, 2005, sp. nov." docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="FFE3FF99D14FFF8D863EFFF6FF90E927" masterDocTitle="Two new species of Macrobiotus (Eutardigrada, Macrobiotidae) from Tien Shan (Kirghizia), with notes on Macrobiotus tenuis group" masterLastPageNumber="46" masterPageNumber="33" pageNumber="38" updateTime="1698214190570" updateUser="ExternalLinkService">
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<mods:titleInfo id="4B0CA6DF5BFCAC43FB52B157F5BBA3A5">
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<mods:title id="A817BE57A1A19BA129D6C88C3DF823B8">Two new species of Macrobiotus (Eutardigrada, Macrobiotidae) from Tien Shan (Kirghizia), with notes on Macrobiotus tenuis group</mods:title>
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<mods:name id="E06BC489C33F7754E3AD1AF3376ED906" type="personal">
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<mods:roleTerm id="380795A9D8DE1618623320CFD40C40B4">Author</mods:roleTerm>
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<mods:namePart id="15614473FFFB55786FF4ED53283F3C73">Tumanov, Denis V.</mods:namePart>
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<mods:typeOfResource id="97000FB5284BD881D4536CE36E6D37F6">text</mods:typeOfResource>
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<mods:title id="7A81C6CCBCA2DF6F9D925E8F2A98B9A5">Zootaxa</mods:title>
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<mods:part id="0E334C4C2D62028D92BA734F9FFA4241">
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<mods:date id="AB7C0A2E2ED33276466AC051326694D6">2005</mods:date>
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<mods:number id="4F275F3BCF1381E97A4152BC338EACD4">1043</mods:number>
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<mods:start id="EA1679C693E167EC3D0BB5C8ECFAA415">33</mods:start>
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<mods:classification id="B02291BF854FEBCFCF66BB26A3015F30">journal article</mods:classification>
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<mods:identifier id="9119950926BEB45781E7296E4A1456CE" type="CLB-Dataset">50957</mods:identifier>
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<mods:identifier id="91980226E48841F5EFC575F17BAE3AA6" type="DOI">10.5281/zenodo.169855</mods:identifier>
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<mods:identifier id="2D36CE477641A5AEF9AED0DAB93794D8" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="C4BA4DEBFA2EF365BCC0B21DB6CD351E" type="Zenodo-Dep">169855</mods:identifier>
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<treatment id="03DA87E1D14AFF818736FADAFD17EBE1" ID-DOI="http://doi.org/10.5281/zenodo.5629675" ID-GBIF-Taxon="119336670" ID-Zenodo-Dep="5629675" LSID="urn:lsid:plazi:treatment:03DA87E1D14AFF818736FADAFD17EBE1" httpUri="http://treatment.plazi.org/id/03DA87E1D14AFF818736FADAFD17EBE1" lastPageId="12" lastPageNumber="45" pageId="5" pageNumber="38">
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<subSubSection id="C369657CD14AFF888736FADAFBC3EC61" box="[264,1107,1324,1350]" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph id="8BCC36F7D14AFF888736FADAFBC3EC61" blockId="5.[264,1107,1324,1350]" box="[264,1107,1324,1350]" pageId="5" pageNumber="38">
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<heading id="D084819BD14AFF888736FADAFBC3EC61" bold="true" box="[264,1107,1324,1350]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<emphasis id="B907EAE5D14AFF888736FADAFBC3EC61" bold="true" box="[264,1107,1324,1350]" pageId="5" pageNumber="38">
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<taxonomicName id="4C734D74D14AFF888736FADAFDBDEC61" ID-CoL="3X2TJ" box="[264,557,1324,1350]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="38" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
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<emphasis id="B907EAE5D14AFF888736FADAFDBDEC61" bold="true" box="[264,557,1324,1350]" italics="true" pageId="5" pageNumber="38">Macrobiotus kirghizicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A234579ED14AFF88840BFADAFD03EC61" box="[565,659,1324,1350]" pageId="5" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="13482A72D14AFF88849DFADAFCA1EC61" box="[675,817,1324,1350]" captionStart="FIGURES 18 – 24" captionStartId="7.[264,382,1676,1700]" captionTargetBox="[271,1302,322,1594]" captionTargetId="figure@7.[264,1323,304,1650]" captionTargetPageId="7" captionText="FIGURES 18 – 24. Macrobiotus kirghizicus sp. nov. 18 — habitus, 19 — buccopharyngeal apparatus, dorsal, 20 — part of the egg surface, 21 — claws of the II pair of legs, 22 — claw of the III pair of legs, lateral view, 23 — claws of the IV pair of legs (18, 19, 21, 23 — paratypes, 22 — specimen from the type series). Macrobiotus ariekammensis 24 — claws of the IV pair of legs." httpUri="https://zenodo.org/record/169858/files/figure.png" pageId="5" pageNumber="38">Figs. 18–23</figureCitation>
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,
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<figureCitation id="13482A72D14AFF888501FADAFC1AEC61" box="[831,906,1324,1350]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="5" pageNumber="38">25–30</figureCitation>
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,
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<figureCitation id="13482A72D14AFF8885A9FADAFC72EC61" box="[919,994,1324,1350]" captionStart="FIGURES 34 – 39" captionStartId="10.[264,383,1713,1737]" captionTargetBox="[285,1300,310,1687]" captionTargetId="figure@10.[264,1323,304,1709]" captionTargetPageId="10" captionText="FIGURES 34 – 39. Macrobiotus kirghizicus sp. nov. 34 — general appearance of the egg, 35 — part of the egg surface, 36 — egg processes. Macrobiotus ariekammensis 37 — general appearance of the egg, 38 — part of the egg surface, 39 — egg processes (phase contrast)." httpUri="https://zenodo.org/record/169860/files/figure.png" pageId="5" pageNumber="38">34–36</figureCitation>
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,
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<tableCitation id="C6F1034CD14AFF8885D0FADAFBDBEC61" box="[1006,1099,1324,1350]" captionStart="TABLE 2" captionStartId="6.[264,352,654,678]" captionTargetBox="[264,1323,714,1699]" captionTargetPageId="6" captionText="TABLE 2. Summary of morphometric data for Macrobiotus kirghizicus sp. nov. (n = 9)." httpUri="http://table.plazi.org/id/DF0C667FD149FF8B8736FD78FB3AEB81" pageId="5" pageNumber="38" tableUuid="DF0C667FD149FF8B8736FD78FB3AEB81">Table 2</tableCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369657CD14AFF888736FA80FA90EFF6" pageId="5" pageNumber="38" type="materials_examined">
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<paragraph id="8BCC36F7D14AFF888736FA80FA90EFF6" blockId="5.[264,1323,1398,1905]" pageId="5" pageNumber="38">
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<emphasis id="B907EAE5D14AFF888736FA80FE49ECB7" bold="true" box="[264,473,1398,1424]" pageId="5" pageNumber="38">
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<typeStatus id="54C88855D14AFF888736FA80FED5ECB7" box="[264,325,1398,1424]" pageId="5" pageNumber="38">Type</typeStatus>
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specimens.
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</emphasis>
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<typeStatus id="54C88855D14AFF8887D9FA81FDC9ECB6" box="[487,601,1399,1425]" pageId="5" pageNumber="38" type="holotype">Holotype</typeStatus>
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: slide number 195(10) (sex indet.), collected by Dr. A.V. Barabanov (Zoological Institute Russian Academy of Sciences, St. Petersburg)
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<date id="FFCD1037D14AFF888736FA31FE1EECC6" box="[264,398,1479,1505]" pageId="5" pageNumber="38" value="1998-08-17">17.08.1998</date>
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,
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<collectingCountry id="F3647667D14AFF8887A2FA31FD9DECC6" box="[412,525,1479,1505]" name="Kyrgyzstan" pageId="5" pageNumber="38">Kirghizia</collectingCountry>
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, Chui Area.
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<typeStatus id="54C88855D14AFF88849DFA31FC8AECC6" box="[675,794,1479,1505]" pageId="5" pageNumber="38" type="paratype">Paratypes</typeStatus>
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: slides 194(5, 6, 7), 198(4, 7) from the same locality.
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<typeStatus id="54C88855D14AFF888751FA19FE4CEF2E" box="[367,476,1519,1545]" pageId="5" pageNumber="38" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="54C88855D14AFF888424FA19FD1AEF2E" box="[538,650,1519,1545]" pageId="5" pageNumber="38" type="paratype">paratypes</typeStatus>
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are preserved at the Zoological Institute of the Russian Academy of Sciences, St. Petersburg,
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<collectingCountry id="F3647667D14AFF8884C9F9E1FCD9EF16" box="[759,841,1559,1585]" name="Russia" pageId="5" pageNumber="38">Russia</collectingCountry>
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. One
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<typeStatus id="54C88855D14AFF88859EF9E1FB95EF16" box="[928,1029,1559,1585]" pageId="5" pageNumber="38" type="paratype">paratype</typeStatus>
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(slide number 198(7)) preserved in the collection of Binda and Pilato (Department of Animal Biology, University of Catania,
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<collectingCountry id="F3647667D14AFF8887AEF991FE5BEFA6" box="[400,459,1639,1665]" name="Italy" pageId="5" pageNumber="38">Italy</collectingCountry>
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).
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<typeStatus id="54C88855D14AFF8887E0F991FD88EFA6" box="[478,536,1639,1665]" pageId="5" pageNumber="38">Type</typeStatus>
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locality:
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<collectingCountry id="F3647667D14AFF8884B9F991FD68EFA6" box="[647,760,1639,1665]" name="Kyrgyzstan" pageId="5" pageNumber="38">Kirghizia</collectingCountry>
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, Chui Area., northern slope of Kirghizian ridge, Turakaiyn canyon, vicinity of Turakaiyn village, near
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<quantity id="4C8B9B12D14AFF888591F979FB9EEF8E" box="[943,1038,1679,1705]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="38" unit="m" value="2000.0">2000 m</quantity>
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a.s.l., moss from soil. Twentyeight adult and juveniles and
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<specimenCount id="9D75FD7ED14AFF8884FFF941FC8BEFF6" box="[705,795,1719,1745]" pageId="5" pageNumber="38" type="egg">41 eggs</specimenCount>
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(3 with developed embryos) were found.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369657CD14AFF888706F928FABBEFDE" box="[312,1323,1758,1785]" pageId="5" pageNumber="38" type="etymology">
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<paragraph id="8BCC36F7D14AFF888706F928FABBEFDE" blockId="5.[264,1323,1398,1905]" box="[312,1323,1758,1785]" pageId="5" pageNumber="38">
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<emphasis id="B907EAE5D14AFF888706F928FE54EFDF" bold="true" box="[312,452,1758,1784]" pageId="5" pageNumber="38">Etymology.</emphasis>
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The name of the new species was derived from
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<collectingCountry id="F3647667D14AFF8885CBF929FBF6EFDE" box="[1013,1126,1759,1785]" name="Kyrgyzstan" pageId="5" pageNumber="38">Kirghizia</collectingCountry>
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, its
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<typeStatus id="54C88855D14AFF8882AAF929FB57EFDE" box="[1172,1223,1759,1785]" pageId="5" pageNumber="38">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369657CD14AFF848706F8F0FC02EDD1" lastPageId="9" lastPageNumber="42" pageId="5" pageNumber="38" type="description">
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<paragraph id="8BCC36F7D14AFF8B8706F8F0FB8DEB09" blockId="5.[264,1323,1398,1905]" lastBlockId="6.[264,1323,284,558]" lastPageId="6" lastPageNumber="39" pageId="5" pageNumber="38">
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<emphasis id="B907EAE5D14AFF888706F8F0FE40EE07" bold="true" box="[312,464,1798,1824]" pageId="5" pageNumber="38">Description.</emphasis>
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Body long and relatively plump (
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<figureCitation id="13482A72D14AFF888544F8F1FC70EE06" box="[890,992,1799,1825]" captionStart="FIGURES 18 – 24" captionStartId="7.[264,382,1676,1700]" captionTargetBox="[271,1302,322,1594]" captionTargetId="figure@7.[264,1323,304,1650]" captionTargetPageId="7" captionText="FIGURES 18 – 24. Macrobiotus kirghizicus sp. nov. 18 — habitus, 19 — buccopharyngeal apparatus, dorsal, 20 — part of the egg surface, 21 — claws of the II pair of legs, 22 — claw of the III pair of legs, lateral view, 23 — claws of the IV pair of legs (18, 19, 21, 23 — paratypes, 22 — specimen from the type series). Macrobiotus ariekammensis 24 — claws of the IV pair of legs." httpUri="https://zenodo.org/record/169858/files/figure.png" pageId="5" pageNumber="38">Figs. 18</figureCitation>
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,
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<figureCitation id="13482A72D14AFF8885CFF8F1FB9FEE06" box="[1009,1039,1799,1825]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="5" pageNumber="38">25</figureCitation>
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) (body length of adult animals 478.8–685.0 m). Color white (colourless), large specimens often with granules of brown pigment. Most specimens with granular eye spots. Cuticle smooth, large specimens with well defined small areas of granulation on the ventral side. Typical distribution of these areas is the following: one area between legs I, one area between the 1st and the 2nd pairs of legs, two areas aligned in a transverse row, between legs II, three areas aligned transversally between the 2nd and the 3rd pairs of legs, one pair of areas, aligned as previously, between legs III, two rows of three areas between the 3rd and the 4th pairs of legs and a pair of areas laterally to anus. Legs with very fine dotlike sculpture (on dorsal surface and around the claw bases), better visible on legs IV.
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</paragraph>
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<caption id="DF0C667FD149FF8B8736FD78FB3AEB81" ID-Table-UUID="DF0C667FD149FF8B8736FD78FB3AEB81" box="[264,1194,652,678]" httpUri="http://table.plazi.org/id/DF0C667FD149FF8B8736FD78FB3AEB81" pageId="6" pageNumber="39" targetBox="[264,1323,714,1699]" targetIsTable="true" targetPageId="6">
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<paragraph id="8BCC36F7D149FF8B8736FD78FB3AEB81" blockId="6.[264,1194,652,678]" box="[264,1194,652,678]" pageId="6" pageNumber="39">
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<emphasis id="B907EAE5D149FF8B8736FD78FEEBEB81" bold="true" box="[264,379,654,678]" pageId="6" pageNumber="39">TABLE 2.</emphasis>
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Summary of morphometric data for
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<taxonomicName id="4C734D74D149FF8B853AFD78FB96EB82" box="[772,1030,654,677]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="39" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
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<emphasis id="B907EAE5D149FF8B853AFD78FB96EB82" box="[772,1030,654,677]" italics="true" pageId="6" pageNumber="39">Macrobiotus kirghizicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A234579ED149FF8B8232FD78FBF1EB81" box="[1036,1121,654,678]" pageId="6" pageNumber="39" rank="species">
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<emphasis id="B907EAE5D149FF8B8232FD78FBF1EB81" bold="true" box="[1036,1121,654,678]" pageId="6" pageNumber="39">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(n=9).
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</paragraph>
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</caption>
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<paragraph id="8BCC36F7D149FF8B871AFD3CFA97EF87" pageId="6" pageNumber="39">
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<table id="F973C457D14900728736FD3CFABBEF84" box="[264,1323,714,1699]" gridcols="4" gridrows="22" pageId="6" pageNumber="39">
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||
<tr id="354334B5D14900728736FD3CFABBEA23" box="[264,1323,714,772]" gridrow="0" pageId="6" pageNumber="39">
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||
<th id="76925DC9D14900728736FD3CFDC1EA23" box="[264,593,714,772]" gridcol="0" gridrow="0" pageId="6" pageNumber="39">Character</th>
|
||
<th id="76925DC9D149007284AFFD3CFD7AEA23" box="[657,746,714,772]" gridcol="1" gridrow="0" pageId="6" pageNumber="39">Holotype</th>
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||
<th id="76925DC9D14900728533FD3CFB37EA23" box="[781,1191,714,772]" gridcol="2" gridrow="0" pageId="6" pageNumber="39">Smallest Largest Mean (Range) specimen specimen</th>
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<th id="76925DC9D149007282F6FD3CFABBEA23" box="[1224,1323,714,772]" gridcol="3" gridrow="0" pageId="6" pageNumber="39">Standard error</th>
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||
</tr>
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||
<tr id="354334B5D14900728736FCEFFABBEA16" box="[264,1323,793,817]" gridrow="1" pageId="6" pageNumber="39">
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||
<th id="76925DC9D14900728736FCEFFDC1EA16" box="[264,593,793,817]" gridcol="0" gridrow="1" pageId="6" pageNumber="39">Body length (m)</th>
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<td id="76925DC9D149007284AFFCEFFD7AEA16" box="[657,746,793,817]" gridcol="1" gridrow="1" pageId="6" pageNumber="39">651.7</td>
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||
<td id="76925DC9D14900728533FCEFFB37EA16" box="[781,1191,793,817]" gridcol="2" gridrow="1" pageId="6" pageNumber="39">478.8 685.0 605.9 (478.8–685.0)</td>
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<td id="76925DC9D149007282F6FCEFFABBEA16" box="[1224,1323,793,817]" gridcol="3" gridrow="1" pageId="6" pageNumber="39">22.59</td>
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</tr>
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<tr id="354334B5D14900728736FCB1FABBEA79" box="[264,1323,839,862]" gridrow="2" pageId="6" pageNumber="39">
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<th id="76925DC9D14900728736FCB1FDC1EA79" box="[264,593,839,862]" gridcol="0" gridrow="2" pageId="6" pageNumber="39">Buccal tube length (m)</th>
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<td id="76925DC9D149007284AFFCB1FD7AEA79" box="[657,746,839,862]" gridcol="1" gridrow="2" pageId="6" pageNumber="39">54.0</td>
|
||
<td id="76925DC9D14900728533FCB1FB37EA79" box="[781,1191,839,862]" gridcol="2" gridrow="2" pageId="6" pageNumber="39">45.9 57.0 52.2 (45.9–57.0)</td>
|
||
<td id="76925DC9D149007282F6FCB1FABBEA79" box="[1224,1323,839,862]" gridcol="3" gridrow="2" pageId="6" pageNumber="39">1.26</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FC82FABBEAAB" box="[264,1323,884,908]" gridrow="3" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FC82FDC1EAAB" box="[264,593,884,908]" gridcol="0" gridrow="3" pageId="6" pageNumber="39">Ss (m)</th>
|
||
<td id="76925DC9D149007284AFFC82FD7AEAAB" box="[657,746,884,908]" gridcol="1" gridrow="3" pageId="6" pageNumber="39">40.3</td>
|
||
<td id="76925DC9D14900728533FC82FB37EAAB" box="[781,1191,884,908]" gridcol="2" gridrow="3" pageId="6" pageNumber="39">34.0 42.6 38.8 (34–42.6)</td>
|
||
<td id="76925DC9D149007282F6FC82FABBEAAB" box="[1224,1323,884,908]" gridcol="3" gridrow="3" pageId="6" pageNumber="39">0.94</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FC57FABBEA90" box="[264,1323,929,951]" gridrow="4" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FC57FDC1EA90" box="[264,593,929,951]" gridcol="0" gridrow="4" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FC54FE8DEA90" box="[267,285,930,951]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
Ss
|
||
</th>
|
||
<td id="76925DC9D149007284AFFC57FD7AEA90" box="[657,746,929,951]" gridcol="1" gridrow="4" pageId="6" pageNumber="39">74.7</td>
|
||
<td id="76925DC9D14900728533FC57FB37EA90" box="[781,1191,929,951]" gridcol="2" gridrow="4" pageId="6" pageNumber="39">74.2 74.7 74.3 (73.5–74.7)</td>
|
||
<td id="76925DC9D149007282F6FC57FABBEA90" box="[1224,1323,929,951]" gridcol="3" gridrow="4" pageId="6" pageNumber="39">0.15</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FC3AFABBEAC3" box="[264,1323,972,996]" gridrow="5" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FC3AFDC1EAC3" box="[264,593,972,996]" gridcol="0" gridrow="5" pageId="6" pageNumber="39">Outer buccal tube width (m)</th>
|
||
<td id="76925DC9D149007284AFFC3AFD7AEAC3" box="[657,746,972,996]" gridcol="1" gridrow="5" pageId="6" pageNumber="39">6.7</td>
|
||
<td id="76925DC9D14900728533FC3AFB37EAC3" box="[781,1191,972,996]" gridcol="2" gridrow="5" pageId="6" pageNumber="39">6.7 7.4 6.9 (6.4–7.4)</td>
|
||
<td id="76925DC9D149007282F6FC3AFABBEAC3" box="[1224,1323,972,996]" gridcol="3" gridrow="5" pageId="6" pageNumber="39">0.13</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FC0FFABBED28" box="[264,1323,1017,1039]" gridrow="6" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FC0FFDC1ED28" box="[264,593,1017,1039]" gridcol="0" gridrow="6" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FC0CFE8DED28" box="[267,285,1018,1039]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of outer buccal tube width
|
||
</th>
|
||
<td id="76925DC9D149007284AFFC0FFD7AED28" box="[657,746,1017,1039]" gridcol="1" gridrow="6" pageId="6" pageNumber="39">12.3</td>
|
||
<td id="76925DC9D14900728533FC0FFB37ED28" box="[781,1191,1017,1039]" gridcol="2" gridrow="6" pageId="6" pageNumber="39">14.5 13.0 13.3 (12.3–14.5)</td>
|
||
<td id="76925DC9D149007282F6FC0FFABBED28" box="[1224,1323,1017,1039]" gridcol="3" gridrow="6" pageId="6" pageNumber="39">0.31</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FBD2FABBED1B" box="[264,1323,1060,1084]" gridrow="7" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FBD2FDC1ED1B" box="[264,593,1060,1084]" gridcol="0" gridrow="7" pageId="6" pageNumber="39">Placoid row length (m)</th>
|
||
<td id="76925DC9D149007284AFFBD2FD7AED1B" box="[657,746,1060,1084]" gridcol="1" gridrow="7" pageId="6" pageNumber="39">29.8</td>
|
||
<td id="76925DC9D14900728533FBD2FB37ED1B" box="[781,1191,1060,1084]" gridcol="2" gridrow="7" pageId="6" pageNumber="39">25.2 33.3 28.7 (25.2–33.3)</td>
|
||
<td id="76925DC9D149007282F6FBD2FABBED1B" box="[1224,1323,1060,1084]" gridcol="3" gridrow="7" pageId="6" pageNumber="39">0.98</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FBA7FABBED40" box="[264,1323,1105,1127]" gridrow="8" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FBA7FDC1ED40" box="[264,593,1105,1127]" gridcol="0" gridrow="8" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FBA4FE8DED40" box="[267,285,1106,1127]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of placoid row length
|
||
</th>
|
||
<td id="76925DC9D149007284AFFBA7FD7AED40" box="[657,746,1105,1127]" gridcol="1" gridrow="8" pageId="6" pageNumber="39">55.2</td>
|
||
<td id="76925DC9D14900728533FBA7FB37ED40" box="[781,1191,1105,1127]" gridcol="2" gridrow="8" pageId="6" pageNumber="39">54.8 58.4 55.3 (50.0–58.4)</td>
|
||
<td id="76925DC9D149007282F6FBA7FABBED40" box="[1224,1323,1105,1127]" gridcol="3" gridrow="8" pageId="6" pageNumber="39">0.89</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FB8AFABBEDB3" box="[264,1323,1148,1172]" gridrow="9" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FB8AFDC1EDB3" box="[264,593,1148,1172]" gridcol="0" gridrow="9" pageId="6" pageNumber="39">Macroplacoid row length (m)</th>
|
||
<td id="76925DC9D149007284AFFB8AFD7AEDB3" box="[657,746,1148,1172]" gridcol="1" gridrow="9" pageId="6" pageNumber="39">24.4</td>
|
||
<td id="76925DC9D14900728533FB8AFB37EDB3" box="[781,1191,1148,1172]" gridcol="2" gridrow="9" pageId="6" pageNumber="39">20.4 26.6 24.1 (20.4–27.0)</td>
|
||
<td id="76925DC9D149007282F6FB8AFABBEDB3" box="[1224,1323,1148,1172]" gridcol="3" gridrow="9" pageId="6" pageNumber="39">0.80</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FB5FFABBED99" box="[264,1323,1193,1214]" gridrow="10" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FB5FFDC1ED99" box="[264,593,1193,1214]" gridcol="0" gridrow="10" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FB5FFE8DED99" box="[267,285,1193,1214]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of macroplacoid row length
|
||
</th>
|
||
<td id="76925DC9D149007284AFFB5FFD7AED99" box="[657,746,1193,1214]" gridcol="1" gridrow="10" pageId="6" pageNumber="39">45.2</td>
|
||
<td id="76925DC9D14900728533FB5FFB37ED99" box="[781,1191,1193,1214]" gridcol="2" gridrow="10" pageId="6" pageNumber="39">44.4 46.8 46.2 (41.7–49.7)</td>
|
||
<td id="76925DC9D149007282F6FB5FFABBED99" box="[1224,1323,1193,1214]" gridcol="3" gridrow="10" pageId="6" pageNumber="39">0.78</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FB22FABBEDCB" box="[264,1323,1236,1260]" gridrow="11" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FB22FDC1EDCB" box="[264,593,1236,1260]" gridcol="0" gridrow="11" pageId="6" pageNumber="39">1st macroplacoid length (m)</th>
|
||
<td id="76925DC9D149007284AFFB22FD7AEDCB" box="[657,746,1236,1260]" gridcol="1" gridrow="11" pageId="6" pageNumber="39">14.1</td>
|
||
<td id="76925DC9D14900728533FB22FB37EDCB" box="[781,1191,1236,1260]" gridcol="2" gridrow="11" pageId="6" pageNumber="39">11.8 14.8 13.6 (11.5–15.5)</td>
|
||
<td id="76925DC9D149007282F6FB22FABBEDCB" box="[1224,1323,1236,1260]" gridcol="3" gridrow="11" pageId="6" pageNumber="39">0.50</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FAF7FABBEC31" box="[264,1323,1281,1302]" gridrow="12" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FAF7FDC1EC31" box="[264,593,1281,1302]" gridcol="0" gridrow="12" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FAF7FE8DEC31" box="[267,285,1281,1302]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of 1st macroplacoid length
|
||
</th>
|
||
<td id="76925DC9D149007284AFFAF7FD7AEC31" box="[657,746,1281,1302]" gridcol="1" gridrow="12" pageId="6" pageNumber="39">26.0</td>
|
||
<td id="76925DC9D14900728533FAF7FB37EC31" box="[781,1191,1281,1302]" gridcol="2" gridrow="12" pageId="6" pageNumber="39">25.8 26.0 26.0 (21.5–28.6)</td>
|
||
<td id="76925DC9D149007282F6FAF7FABBEC31" box="[1224,1323,1281,1302]" gridcol="3" gridrow="12" pageId="6" pageNumber="39">0.63</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FADAFABBEC64" box="[264,1323,1324,1347]" gridrow="13" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FADAFDC1EC64" box="[264,593,1324,1347]" gridcol="0" gridrow="13" pageId="6" pageNumber="39">2nd macroplacoid length (m)</th>
|
||
<td id="76925DC9D149007284AFFADAFD7AEC64" box="[657,746,1324,1347]" gridcol="1" gridrow="13" pageId="6" pageNumber="39">8.1</td>
|
||
<td id="76925DC9D14900728533FADAFB37EC64" box="[781,1191,1324,1347]" gridcol="2" gridrow="13" pageId="6" pageNumber="39">7.4 9.6 8.4 (7.0–9.6)</td>
|
||
<td id="76925DC9D149007282F6FADAFABBEC64" box="[1224,1323,1324,1347]" gridcol="3" gridrow="13" pageId="6" pageNumber="39">0.32</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FAAFFABBEC49" box="[264,1323,1369,1390]" gridrow="14" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FAAFFDC1EC49" box="[264,593,1369,1390]" gridcol="0" gridrow="14" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FAAFFE8DEC49" box="[267,285,1369,1390]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of 2nd macroplacoid length
|
||
</th>
|
||
<td id="76925DC9D149007284AFFAAFFD7AEC49" box="[657,746,1369,1390]" gridcol="1" gridrow="14" pageId="6" pageNumber="39">15.1</td>
|
||
<td id="76925DC9D14900728533FAAFFB37EC49" box="[781,1191,1369,1390]" gridcol="2" gridrow="14" pageId="6" pageNumber="39">16.1 16.9 16.1 (14.6–17.6)</td>
|
||
<td id="76925DC9D149007282F6FAAFFABBEC49" box="[1224,1323,1369,1390]" gridcol="3" gridrow="14" pageId="6" pageNumber="39">0.36</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FA75FABBECBC" box="[264,1323,1411,1435]" gridrow="15" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FA75FDC1ECBC" box="[264,593,1411,1435]" gridcol="0" gridrow="15" pageId="6" pageNumber="39">Microplacoid length (m)</th>
|
||
<td id="76925DC9D149007284AFFA75FD7AECBC" box="[657,746,1411,1435]" gridcol="1" gridrow="15" pageId="6" pageNumber="39">3.7</td>
|
||
<td id="76925DC9D14900728533FA75FB37ECBC" box="[781,1191,1411,1435]" gridcol="2" gridrow="15" pageId="6" pageNumber="39">3.0 4.1 3.6 (3.0–4.1)</td>
|
||
<td id="76925DC9D149007282F6FA75FABBECBC" box="[1224,1323,1411,1435]" gridcol="3" gridrow="15" pageId="6" pageNumber="39">0.16</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FA47FABBECE1" box="[264,1323,1457,1478]" gridrow="16" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FA47FDC1ECE1" box="[264,593,1457,1478]" gridcol="0" gridrow="16" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735FA47FE8DECE1" box="[267,285,1457,1478]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of microplacoid length
|
||
</th>
|
||
<td id="76925DC9D149007284AFFA47FD7AECE1" box="[657,746,1457,1478]" gridcol="1" gridrow="16" pageId="6" pageNumber="39">6.8</td>
|
||
<td id="76925DC9D14900728533FA47FB37ECE1" box="[781,1191,1457,1478]" gridcol="2" gridrow="16" pageId="6" pageNumber="39">6.5 7.1 6.9 (5.7–8.5)</td>
|
||
<td id="76925DC9D149007282F6FA47FABBECE1" box="[1224,1323,1457,1478]" gridcol="3" gridrow="16" pageId="6" pageNumber="39">0.31</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736FA2DFABBECD4" box="[264,1323,1499,1523]" gridrow="17" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736FA2DFDC1ECD4" box="[264,593,1499,1523]" gridcol="0" gridrow="17" pageId="6" pageNumber="39">External claw length, leg I (m)</th>
|
||
<td id="76925DC9D149007284AFFA2DFD7AECD4" box="[657,746,1499,1523]" gridcol="1" gridrow="17" pageId="6" pageNumber="39">24.1</td>
|
||
<td id="76925DC9D14900728533FA2DFB37ECD4" box="[781,1191,1499,1523]" gridcol="2" gridrow="17" pageId="6" pageNumber="39">19.2 25.9 24.3 (19.2–26.6)</td>
|
||
<td id="76925DC9D149007282F6FA2DFABBECD4" box="[1224,1323,1499,1523]" gridcol="3" gridrow="17" pageId="6" pageNumber="39">0.76</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736F9FFFABBEF39" box="[264,1323,1545,1566]" gridrow="18" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736F9FFFDC1EF39" box="[264,593,1545,1566]" gridcol="0" gridrow="18" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735F9FFFE8DEF39" box="[267,285,1545,1566]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of external claw length, leg I
|
||
</th>
|
||
<td id="76925DC9D149007284AFF9FFFD7AEF39" box="[657,746,1545,1566]" gridcol="1" gridrow="18" pageId="6" pageNumber="39">44.5</td>
|
||
<td id="76925DC9D14900728533F9FFFB37EF39" box="[781,1191,1545,1566]" gridcol="2" gridrow="18" pageId="6" pageNumber="39">41.9 45.5 46.4 (41.9–50.7)</td>
|
||
<td id="76925DC9D149007282F6F9FFFABBEF39" box="[1224,1323,1545,1566]" gridcol="3" gridrow="18" pageId="6" pageNumber="39">0.94</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736F9C5FABBEF6C" box="[264,1323,1587,1611]" gridrow="19" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736F9C5FDC1EF6C" box="[264,593,1587,1611]" gridcol="0" gridrow="19" pageId="6" pageNumber="39">External claw length, leg III (m)</th>
|
||
<td id="76925DC9D149007284AFF9C5FD7AEF6C" box="[657,746,1587,1611]" gridcol="1" gridrow="19" pageId="6" pageNumber="39">26.6</td>
|
||
<td id="76925DC9D14900728533F9C5FB37EF6C" box="[781,1191,1587,1611]" gridcol="2" gridrow="19" pageId="6" pageNumber="39">22.2 31.8 27.4 (22.2–31.8)</td>
|
||
<td id="76925DC9D149007282F6F9C5FABBEF6C" box="[1224,1323,1587,1611]" gridcol="3" gridrow="19" pageId="6" pageNumber="39">1.07</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736F997FABBEF51" box="[264,1323,1633,1654]" gridrow="20" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736F997FDC1EF51" box="[264,593,1633,1654]" gridcol="0" gridrow="20" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735F997FE8DEF51" box="[267,285,1633,1654]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of external claw length, leg III
|
||
</th>
|
||
<td id="76925DC9D149007284AFF997FD7AEF51" box="[657,746,1633,1654]" gridcol="1" gridrow="20" pageId="6" pageNumber="39">49.3</td>
|
||
<td id="76925DC9D14900728533F997FB37EF51" box="[781,1191,1633,1654]" gridcol="2" gridrow="20" pageId="6" pageNumber="39">48.4 55.8 52.8 (48.4–57.7)</td>
|
||
<td id="76925DC9D149007282F6F997FABBEF51" box="[1224,1323,1633,1654]" gridcol="3" gridrow="20" pageId="6" pageNumber="39">1.17</td>
|
||
</tr>
|
||
<tr id="354334B5D14900728736F97DFABBEF84" box="[264,1323,1675,1699]" gridrow="21" pageId="6" pageNumber="39">
|
||
<th id="76925DC9D14900728736F97DFDC1EF84" box="[264,593,1675,1699]" gridcol="0" gridrow="21" pageId="6" pageNumber="39">External claw length, leg IV (m)</th>
|
||
<td id="76925DC9D149007284AFF97DFD7AEF84" box="[657,746,1675,1699]" gridcol="1" gridrow="21" pageId="6" pageNumber="39">40.7</td>
|
||
<td id="76925DC9D14900728533F97DFB37EF84" box="[781,1191,1675,1699]" gridcol="2" gridrow="21" pageId="6" pageNumber="39">35.5 45.9 42.1 (35.5–45.9)</td>
|
||
<td id="76925DC9D149007282F6F97DFABBEF84" box="[1224,1323,1675,1699]" gridcol="3" gridrow="21" pageId="6" pageNumber="39">1.24</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D149FF8B8735F94FFA97EFEA" blockId="6.[264,1323,1720,1833]" box="[267,1287,1720,1742]" pageId="6" pageNumber="39">
|
||
<emphasis id="B907EAE5D149FF8B8735F94FFE8DEFE9" box="[267,285,1721,1742]" italics="true" pageId="6" pageNumber="39">pt</emphasis>
|
||
of external claw length, leg IV 75.3 77.4 80.5 80.6 (75.3–88) 1.77
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D149FF8B8736F916FC5CEFD4" blockId="6.[264,1323,1720,1833]" box="[264,972,1760,1779]" pageId="6" pageNumber="39">Ss = distance from cephalic end of buccal tube to insertion point of stylet supports.</paragraph>
|
||
<paragraph id="8BCC36F7D149FF8B8736F90DFDD2EE0E" blockId="6.[264,1323,1720,1833]" pageId="6" pageNumber="39">
|
||
Supplementary measurements for
|
||
<typeStatus id="54C88855D149FF8B8412F90DFDEAEE29" box="[556,634,1787,1806]" pageId="6" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
: length of strengthening bar on buccal tube: above
|
||
<quantity id="4C8B9B12D149FF8B820FF90DFBECEE29" box="[1073,1148,1787,1806]" metricMagnitude="1" metricUnit="m" metricValue="3.3299999999999996" pageId="6" pageNumber="39" unit="m" value="33.3">33.3 m</quantity>
|
||
, lunules on claws of the fourth pair of legs: 8.1 x
|
||
<quantity id="4C8B9B12D149FF8B87C2F8E0FDD2EE0E" box="[508,578,1814,1833]" metricMagnitude="1" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="39" unit="m" value="11.8">11.8 m</quantity>
|
||
</paragraph>
|
||
<caption id="DF0C667FD148FF8A8736F97AFB6DEE2B" httpUri="https://zenodo.org/record/169858/files/figure.png" pageId="7" pageNumber="40" targetBox="[271,1302,322,1594]" targetPageId="7">
|
||
<paragraph id="8BCC36F7D148FF8A8736F97AFB6DEE2B" blockId="7.[264,1323,1676,1804]" pageId="7" pageNumber="40">
|
||
<emphasis id="B907EAE5D148FF8A8736F97AFE5FEF83" bold="true" box="[264,463,1676,1700]" pageId="7" pageNumber="40">FIGURES 18–24</emphasis>
|
||
.
|
||
<taxonomicName id="4C734D74D148FF8A87DAF97BFD7EEF83" box="[484,750,1677,1700]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="40" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
|
||
<emphasis id="B907EAE5D148FF8A87DAF97BFD7EEF83" box="[484,750,1677,1700]" italics="true" pageId="7" pageNumber="40">Macrobiotus kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED148FF8A84C2F97AFCC8EF83" box="[764,856,1676,1700]" pageId="7" pageNumber="40" rank="species">
|
||
<emphasis id="B907EAE5D148FF8A84C2F97AFCC8EF83" bold="true" box="[764,856,1676,1700]" pageId="7" pageNumber="40">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
18 — habitus, 19 — buccopharyngeal apparatus, dorsal, 20 — part of the egg surface, 21 — claws of the II pair of legs, 22 — claw of the III pair of legs, lateral view, 23 — claws of the IV pair of legs (18, 19, 21, 23 — paratypes, 22 — specimen from the type series).
|
||
<taxonomicName id="4C734D74D148FF8A8462F903FC1AEE2B" box="[604,906,1781,1804]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="40" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D148FF8A8462F903FC1AEE2B" box="[604,906,1781,1804]" italics="true" pageId="7" pageNumber="40">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
24 — claws of the IV pair of legs.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0C667FD147FF858736F943FB0AEE49" httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="8" pageNumber="41" targetBox="[352,1265,283,1716]" targetPageId="8">
|
||
<paragraph id="8BCC36F7D147FF858736F943FB0AEE49" blockId="8.[264,1323,1717,1902]" pageId="8" pageNumber="41">
|
||
<emphasis id="B907EAE5D147FF858736F943FE5FEFEA" bold="true" box="[264,463,1717,1741]" pageId="8" pageNumber="41">FIGURES 25–33</emphasis>
|
||
.
|
||
<taxonomicName id="4C734D74D147FF8587DAF940FD7EEFEA" box="[484,750,1718,1741]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="41" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
|
||
<emphasis id="B907EAE5D147FF8587DAF940FD7EEFEA" box="[484,750,1718,1741]" italics="true" pageId="8" pageNumber="41">Macrobiotus kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED147FF8584C2F943FCC8EFEA" box="[764,856,1717,1741]" pageId="8" pageNumber="41" rank="species">
|
||
<emphasis id="B907EAE5D147FF8584C2F943FCC8EFEA" bold="true" box="[764,856,1717,1741]" pageId="8" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28–30 – paratypes).
|
||
<taxonomicName id="4C734D74D147FF8585C3F8E1FABAEE09" box="[1021,1322,1815,1838]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="41" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D147FF8585C3F8E1FABAEE09" box="[1021,1322,1815,1838]" italics="true" pageId="8" pageNumber="41">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36F7D146FF848706FEEAFD83EB69" blockId="9.[264,1324,284,1350]" pageId="9" pageNumber="42">
|
||
Buccopharyngeal apparatus of
|
||
<taxonomicName id="4C734D74D146FF848495FEEAFCD0E812" box="[683,832,284,309]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D146FF848495FEEAFCD0E812" box="[683,832,284,309]" italics="true" pageId="9" pageNumber="42">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54C88855D146FF848578FEEAFCE8E811" box="[838,888,284,310]" pageId="9" pageNumber="42">type</typeStatus>
|
||
(
|
||
<figureCitation id="13482A72D146FF8485B6FEEAFC79E811" box="[904,1001,284,310]" captionStart="FIGURES 18 – 24" captionStartId="7.[264,382,1676,1700]" captionTargetBox="[271,1302,322,1594]" captionTargetId="figure@7.[264,1323,304,1650]" captionTargetPageId="7" captionText="FIGURES 18 – 24. Macrobiotus kirghizicus sp. nov. 18 — habitus, 19 — buccopharyngeal apparatus, dorsal, 20 — part of the egg surface, 21 — claws of the II pair of legs, 22 — claw of the III pair of legs, lateral view, 23 — claws of the IV pair of legs (18, 19, 21, 23 — paratypes, 22 — specimen from the type series). Macrobiotus ariekammensis 24 — claws of the IV pair of legs." httpUri="https://zenodo.org/record/169858/files/figure.png" pageId="9" pageNumber="42">Figs. 19</figureCitation>
|
||
,
|
||
<figureCitation id="13482A72D146FF8485CBFEEAFB84E811" box="[1013,1044,284,310]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="9" pageNumber="42">26</figureCitation>
|
||
) (for all dimensions see
|
||
<tableCitation id="C6F1034CD146FF848736FEB2FEF8E879" box="[264,360,324,350]" captionStart="TABLE 2" captionStartId="6.[264,352,654,678]" captionTargetBox="[264,1323,714,1699]" captionTargetPageId="6" captionText="TABLE 2. Summary of morphometric data for Macrobiotus kirghizicus sp. nov. (n = 9)." httpUri="http://table.plazi.org/id/DF0C667FD149FF8B8736FD78FB3AEB81" pageId="9" pageNumber="42" tableUuid="DF0C667FD149FF8B8736FD78FB3AEB81">Table 2</tableCitation>
|
||
). Buccal cavity with ten scarcely visible peribuccal lamellae. Dorsal buccal armature with two granules in medial position without points and teeth (
|
||
<figureCitation id="13482A72D146FF848259FE9AFB50E8A1" box="[1127,1216,364,390]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="9" pageNumber="42">Fig. 27</figureCitation>
|
||
). Ventral buccal armature not visible. Ventral strengthening bar relatively long and exceeding half of the length of the buccal tube. Pharyngeal bulb suboval, with well developed apophyses, two macroplacoids (first with well developed constriction, longer than second) and elongated microplacoid. Between the apophyses and first macroplacoids there are thin slanting cuticular bars.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D146FF848706FDAAFC9CEA31" blockId="9.[264,1324,284,1350]" pageId="9" pageNumber="42">
|
||
Claws extremely similar to those described by
|
||
<bibRefCitation id="EFE24B06D146FF848553FDAAFBB5EB51" author="Dastych" box="[877,1061,604,630]" pageId="9" pageNumber="45" refString="Dastych, H. (1977) Macrobiotus adelges sp. nov., a new species of Tardigrada from Poland. Bulletin of the Polish Academy of Sciences, 25, 667 - 669." type="journal article" year="1977">Dastych (1977)</bibRefCitation>
|
||
in
|
||
<taxonomicName id="4C734D74D146FF848270FDAAFB42EB52" box="[1102,1234,604,629]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="adelges">
|
||
<emphasis id="B907EAE5D146FF848270FDAAFB42EB52" box="[1102,1234,604,629]" italics="true" pageId="9" pageNumber="42">M. adelges</emphasis>
|
||
</taxonomicName>
|
||
(junior synonym of
|
||
<taxonomicName id="4C734D74D146FF8487A3FD72FCD6EBB9" authority="Weglarska, 1965" authorityName="Weglarska" authorityYear="1965" box="[413,838,644,670]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF8487A3FD72FDE5EBBA" box="[413,629,644,669]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
<bibRefCitation id="EFE24B06D146FF848443FD72FCD6EBB9" author="Weglarska" box="[637,838,644,670]" pageId="9" pageNumber="46" refString="Weglarska, B. (1965) Die Tardigraden (Tardigrada) Spitsbergens. Acta Zoologica Cracoviensia, 11, 43 - 51." type="journal article" year="1965">Weglarska, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
), large, with poorly developed accessory points (
|
||
<figureCitation id="13482A72D146FF848761FD5AFE7DEBE1" box="[351,493,684,710]" captionStart="FIGURES 18 – 24" captionStartId="7.[264,382,1676,1700]" captionTargetBox="[271,1302,322,1594]" captionTargetId="figure@7.[264,1323,304,1650]" captionTargetPageId="7" captionText="FIGURES 18 – 24. Macrobiotus kirghizicus sp. nov. 18 — habitus, 19 — buccopharyngeal apparatus, dorsal, 20 — part of the egg surface, 21 — claws of the II pair of legs, 22 — claw of the III pair of legs, lateral view, 23 — claws of the IV pair of legs (18, 19, 21, 23 — paratypes, 22 — specimen from the type series). Macrobiotus ariekammensis 24 — claws of the IV pair of legs." httpUri="https://zenodo.org/record/169858/files/figure.png" pageId="9" pageNumber="42">Figs. 21–23</figureCitation>
|
||
,
|
||
<figureCitation id="13482A72D146FF8487C7FD5AFDD8EBE1" box="[505,584,684,710]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="9" pageNumber="42">28–30</figureCitation>
|
||
). All claws with large lunules, especially in legs IV. Lunules of legs IV with distinct teeth, lunules of legs I–III with slightly waved edges. Legs I–III with band of thickened cuticle below claw bases.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D146FF848706FCD2FC02EDD1" blockId="9.[264,1324,284,1350]" pageId="9" pageNumber="42">
|
||
Spherical eggs ornamented and laid free (
|
||
<figureCitation id="13482A72D146FF848523FCD2FC3DEA19" box="[797,941,804,830]" captionStart="FIGURES 34 – 39" captionStartId="10.[264,383,1713,1737]" captionTargetBox="[285,1300,310,1687]" captionTargetId="figure@10.[264,1323,304,1709]" captionTargetPageId="10" captionText="FIGURES 34 – 39. Macrobiotus kirghizicus sp. nov. 34 — general appearance of the egg, 35 — part of the egg surface, 36 — egg processes. Macrobiotus ariekammensis 37 — general appearance of the egg, 38 — part of the egg surface, 39 — egg processes (phase contrast)." httpUri="https://zenodo.org/record/169860/files/figure.png" pageId="9" pageNumber="42">Figs. 34–36</figureCitation>
|
||
). Egg processes consists of short domeshaped basal part and rigid spinelike apical part, clearly separated from the base with single internal septum. Without septation of apical part of processes. Basal part lacks reticulation, lateral surfaces of apical part with irregularly distributed minute spines. Bases of processes with wreath of little dark granules and two rows of light pores. The first row is located on the process base above the granules, the second row is located below the granules on the shell surface. Pores of both rows are ordered opposite each other. Shell of eggs between processes smooth, with rare, scarcely visible pores, similar to the pores at the process bases. Usually egg surface and basal parts of processes poorly visible, covered with debris particles. Diameter of eggs (without processes):
|
||
<quantity id="4C8B9B12D146FF8485D8FB7AFB3EED81" box="[998,1198,1164,1190]" metricMagnitude="2" metricUnit="m" metricValue="1.1405" metricValueMax="1.267" metricValueMin="1.014" pageId="9" pageNumber="42" unit="m" value="114.05" valueMax="126.7" valueMin="101.4">101.4–126.7 m</quantity>
|
||
; height of processes (about 25 on the circumference of egg):
|
||
<quantity id="4C8B9B12D146FF848546FB42FBB4EDE9" box="[888,1060,1204,1230]" metricMagnitude="1" metricUnit="m" metricValue="2.195" metricValueMax="2.87" metricValueMin="1.52" pageId="9" pageNumber="42" unit="m" value="21.95" valueMax="28.7" valueMin="15.2">15.2–28.7 m</quantity>
|
||
, their basal diameter:
|
||
<quantity id="4C8B9B12D146FF848736FB2AFE31EDD1" box="[264,417,1244,1270]" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="1.35" metricValueMin="0.85" pageId="9" pageNumber="42" unit="m" value="11.0" valueMax="13.5" valueMin="8.5">8.5–13.5 m</quantity>
|
||
. Height of process basal part:
|
||
<quantity id="4C8B9B12D146FF84853DFB2AFC1EEDD1" box="[771,910,1244,1270]" metricMagnitude="0" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="9" pageNumber="42" unit="m" value="4.65" valueMax="5.1" valueMin="4.2">4.2–5.1 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD146FF818706FAF2FD17EBE1" lastPageId="12" lastPageNumber="45" pageId="9" pageNumber="42" type="discussion">
|
||
<paragraph id="8BCC36F7D146FF848706FAF2FABBEC61" blockId="9.[264,1324,284,1350]" pageId="9" pageNumber="42">
|
||
<emphasis id="B907EAE5D146FF848706FAF2FE21EC39" bold="true" box="[312,433,1284,1310]" pageId="9" pageNumber="42">Remarks.</emphasis>
|
||
The new species is most similar to
|
||
<taxonomicName id="4C734D74D146FF8485F7FAF2FAB3EC3A" box="[969,1315,1284,1309]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF8485F7FAF2FAB3EC3A" box="[969,1315,1284,1309]" italics="true" pageId="9" pageNumber="42">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C734D74D146FF848736FADAFDBAEC62" box="[264,554,1324,1349]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
|
||
<emphasis id="B907EAE5D146FF848736FADAFDBAEC62" box="[264,554,1324,1349]" italics="true" pageId="9" pageNumber="42">Macrobiotus kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED146FF84840FFADAFD1DEC61" box="[561,653,1324,1350]" pageId="9" pageNumber="42" rank="species">
|
||
<emphasis id="B907EAE5D146FF84840FFADAFD1DEC61" bold="true" box="[561,653,1324,1350]" pageId="9" pageNumber="42">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C734D74D146FF848519FADAFC6EEC62" box="[807,1022,1324,1349]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF848519FADAFC6EEC62" box="[807,1022,1324,1349]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
in the following features:
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D146FF848736FA8AFDE0EF29" blockId="9.[264,1323,1404,1830]" pageId="9" pageNumber="42">
|
||
1. Two granules in the buccal armature, while
|
||
<taxonomicName id="4C734D74D146FF84855AFA8AFBD1ECB2" box="[868,1089,1404,1429]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF84855AFA8AFBD1ECB2" box="[868,1089,1404,1429]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
has a single large triangular median tooth (
|
||
<figureCitation id="13482A72D146FF848460FA52FD26EC99" box="[606,694,1444,1470]" captionStart="FIGURES 25 – 33" captionStartId="8.[264,382,1717,1741]" captionTargetBox="[352,1265,283,1716]" captionTargetId="figure@8.[333,1285,280,1716]" captionTargetPageId="8" captionText="FIGURES 25 – 33. Macrobiotus kirghizicus sp. nov. 25 — habitus, 26 — buccopharyngeal apparatus (arrow points to the slanting cuticular bar), 27 – buccal armature, dorsal, 28 — claws of the II pair of legs, 29 — claws of the I pair of legs (arrow points to the cuticular thickening), 30 — claws of the IV pair of legs (26, 27 – holotype, 25, 28 – 30 – paratypes). Macrobiotus ariekammensis 31 — buccal armature (arrow points to the dorsal tooth), 32 — claws of the I pair of legs (arrow points to the cuticular thickening), 33 — claws of the IV pair of legs (phase contrast)." httpUri="https://zenodo.org/record/169859/files/figure.png" pageId="9" pageNumber="42">Fig. 31</figureCitation>
|
||
). This character was not pointed out in the published descriptions of
|
||
<taxonomicName id="4C734D74D146FF8487CDFA3AFD5EECC2" box="[499,718,1484,1509]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF8487CDFA3AFD5EECC2" box="[499,718,1484,1509]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D74D146FF848532FA3AFC01ECC2" box="[780,913,1484,1509]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="adelges">
|
||
<emphasis id="B907EAE5D146FF848532FA3AFC01ECC2" box="[780,913,1484,1509]" italics="true" pageId="9" pageNumber="42">M. adelges</emphasis>
|
||
</taxonomicName>
|
||
, but it is shown on a drawing by
|
||
<bibRefCitation id="EFE24B06D146FF848706FA02FDFCEF29" author="Dastych" box="[312,620,1524,1550]" pageId="9" pageNumber="45" refString="Dastych, H. (1977) Macrobiotus adelges sp. nov., a new species of Tardigrada from Poland. Bulletin of the Polish Academy of Sciences, 25, 667 - 669." type="journal article" year="1977">
|
||
Dastych (1977,
|
||
<figureCitation id="13482A72D146FF8487CCFA02FDF1EF29" box="[498,609,1524,1550]" captionStart="FIGURES 1 – 8" captionStartId="3.[264,383,1720,1744]" captionTargetBox="[276,1286,323,1652]" captionTargetId="figure@3.[264,1323,302,1694]" captionTargetPageId="3" captionText="FIGURES 1 – 8. Macrobiotus barabanovi sp. nov. 1 – habitus, 2 – buccopharyngeal apparatus, ventral, 3 – buccal armature, dorsal, 4 – buccal armature, ventral, 5 – claws of the II pair of legs, 6 – claws of the IV pair of legs, 7 – general appearance of the egg, 8 – part of the egg surface (3, 4, 6 – holotype; 1, 2 – paratypes, 5 – specimen from the type series)." httpUri="https://zenodo.org/record/169856/files/figure.png" pageId="9" pageNumber="42">Figs. 2, 5</figureCitation>
|
||
)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D146FF848736F9EAFD26EF11" blockId="9.[264,1323,1404,1830]" box="[264,694,1564,1590]" pageId="9" pageNumber="42">2. Slightly longer claws on legs IV.</paragraph>
|
||
<paragraph id="8BCC36F7D146FF848736F9B2FDE5EE01" blockId="9.[264,1323,1404,1830]" pageId="9" pageNumber="42">
|
||
3. Differences in egg morphology. Processes of
|
||
<taxonomicName id="4C734D74D146FF8485B5F9B2FBACEF7A" box="[907,1084,1604,1629]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="kirghizicus">
|
||
<emphasis id="B907EAE5D146FF8485B5F9B2FBACEF7A" box="[907,1084,1604,1629]" italics="true" pageId="9" pageNumber="42">M. kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
eggs with clearly separated basal and apical parts, while in
|
||
<taxonomicName id="4C734D74D146FF848572F99AFBBAEFA2" box="[844,1066,1644,1669]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF848572F99AFBBAEFA2" box="[844,1066,1644,1669]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
processes are bottle shaped or conic (
|
||
<figureCitation id="13482A72D146FF84843BF962FD07EF89" box="[517,663,1684,1710]" captionStart="FIGURES 34 – 39" captionStartId="10.[264,383,1713,1737]" captionTargetBox="[285,1300,310,1687]" captionTargetId="figure@10.[264,1323,304,1709]" captionTargetPageId="10" captionText="FIGURES 34 – 39. Macrobiotus kirghizicus sp. nov. 34 — general appearance of the egg, 35 — part of the egg surface, 36 — egg processes. Macrobiotus ariekammensis 37 — general appearance of the egg, 38 — part of the egg surface, 39 — egg processes (phase contrast)." httpUri="https://zenodo.org/record/169860/files/figure.png" pageId="9" pageNumber="42">Figs. 37–39</figureCitation>
|
||
). In
|
||
<taxonomicName id="4C734D74D146FF8484F3F962FC36EF8A" box="[717,934,1684,1709]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D146FF8484F3F962FC36EF8A" box="[717,934,1684,1709]" italics="true" pageId="9" pageNumber="42">M. ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
the apical parts of the processes have internal septae and dull tips with tufts or rosettes of spines, while in
|
||
<taxonomicName id="4C734D74D146FF848335F94AFE27EFDA" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="42" phylum="Tardigrada" rank="species" species="kirghizicus">
|
||
<emphasis id="B907EAE5D146FF848335F94AFE27EFDA" italics="true" pageId="9" pageNumber="42">M. kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
spinelike apical parts of the processes lack internal septae and have only laterally distributed spines.
|
||
</paragraph>
|
||
<caption id="DF0C667FD145FF878736F947FBA0EE29" httpUri="https://zenodo.org/record/169860/files/figure.png" pageId="10" pageNumber="43" targetBox="[285,1300,310,1687]" targetPageId="10">
|
||
<paragraph id="8BCC36F7D145FF878736F947FBA0EE29" blockId="10.[264,1323,1713,1806]" pageId="10" pageNumber="43">
|
||
<emphasis id="B907EAE5D145FF878736F947FE57EFEE" bold="true" box="[264,455,1713,1737]" pageId="10" pageNumber="43">FIGURES 34–39</emphasis>
|
||
.
|
||
<taxonomicName id="4C734D74D145FF8787EAF944FD45EFEE" box="[468,725,1714,1737]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="43" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
|
||
<emphasis id="B907EAE5D145FF8787EAF944FD45EFEE" box="[468,725,1714,1737]" italics="true" pageId="10" pageNumber="43">Macrobiotus kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED145FF8784E5F947FCA0EFEE" box="[731,816,1713,1737]" pageId="10" pageNumber="43" rank="species">
|
||
<emphasis id="B907EAE5D145FF8784E5F947FCA0EFEE" bold="true" box="[731,816,1713,1737]" pageId="10" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
34 — general appearance of the egg, 35 — part of the egg surface, 36 — egg processes.
|
||
<taxonomicName id="4C734D74D145FF8784FCF922FC61EFCC" box="[706,1009,1748,1771]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="43" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D145FF8784FCF922FC61EFCC" box="[706,1009,1748,1771]" italics="true" pageId="10" pageNumber="43">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
37 — general appearance of the egg, 38 — part of the egg surface, 39 — egg processes (phase contrast).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36F7D144FF868706FEEAFBD1E8A1" blockId="11.[264,1323,284,510]" pageId="11" pageNumber="44">
|
||
<emphasis id="B907EAE5D144FF868706FEEAFD28E811" box="[312,696,284,310]" italics="true" pageId="11" pageNumber="44">
|
||
<taxonomicName id="4C734D74D144FF868706FEEAFDC3E812" box="[312,595,284,309]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">Macrobiotus kirghizicus</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED144FF868462FEEAFD28E811" box="[604,696,284,310]" pageId="11" pageNumber="44" rank="species">
|
||
<emphasis id="B907EAE5D144FF868462FEEAFD28E811" bold="true" box="[604,696,284,310]" italics="true" pageId="11" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</emphasis>
|
||
also has some similarity in claw appearance with
|
||
<taxonomicName id="4C734D74D144FF868335FEEAFE1FE87A" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="barabanovi" status="sp. nov.">
|
||
<emphasis id="B907EAE5D144FF868335FEEAFE1FE87A" italics="true" pageId="11" pageNumber="44">M. barabanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED144FF8687A6FEB5FE68E87A" box="[408,504,323,349]" pageId="11" pageNumber="44" rank="species">
|
||
<emphasis id="B907EAE5D144FF8687A6FEB5FE68E87A" bold="true" box="[408,504,323,349]" pageId="11" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, but it could be easily distinguished from this species in having two macroplacoids, very different buccal armature and egg shell sculpture.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868706FE65FC75E8D9" blockId="11.[264,1323,284,510]" pageId="11" pageNumber="44">
|
||
<emphasis id="B907EAE5D144FF868706FE65FE53E88A" bold="true" box="[312,451,403,429]" pageId="11" pageNumber="44">Discussion.</emphasis>
|
||
<taxonomicName id="4C734D74D144FF8687F2FE62FC89E88A" box="[460,793,404,429]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D144FF8687F2FE62FC89E88A" box="[460,793,404,429]" italics="true" pageId="11" pageNumber="44">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
was included by Maucci in the
|
||
<taxonomicName id="4C734D74D144FF8682A9FE62FB4DE88A" box="[1175,1245,404,429]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="tenuis">
|
||
<emphasis id="B907EAE5D144FF8682A9FE62FB4DE88A" box="[1175,1245,404,429]" italics="true" pageId="11" pageNumber="44">tenuis</emphasis>
|
||
</taxonomicName>
|
||
group of the genus
|
||
<taxonomicName id="4C734D74D144FF868788FE4AFDDBE8F2" box="[438,587,444,469]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D144FF868788FE4AFDDBE8F2" box="[438,587,444,469]" italics="true" pageId="11" pageNumber="44">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
. This group of species was separated on the base of the combination of the following features (
|
||
<bibRefCitation id="EFE24B06D144FF8684ECFE12FCEDE8D9" author="Maucci" box="[722,893,484,510]" pageId="11" pageNumber="46" refString="Maucci, W. (1987 a) A contribution to the knowledge of the North American Tardigrada with emphasis on the fauna of Yellowstone National Park (Wyoming). U. Z. I. Selected Symposia and Monographs, 1, 187 - 210." type="journal article" year="1987" yearSuffix="a">Maucci 1987a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE24B06D144FF8685B7FE12FC48E8D9" author="Maucci" box="[905,984,484,510]" pageId="11" pageNumber="46" refString="Maucci, W. (1987 b) Tardigradi della Mongolia Esterna, con descrizione di Macrobiotus mongolicus sp. nov. Bolletino del Museo Civico di Storia Naturale Verona, 14, 339 - 349." type="journal article" year="1987" yearSuffix="b">1987b</bibRefCitation>
|
||
):
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FDC2FD6FEB69" blockId="11.[264,1323,564,990]" box="[264,767,564,590]" pageId="11" pageNumber="44">1. Large dimensions of the adult animals.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FDAAFD36EB51" blockId="11.[264,1323,564,990]" box="[264,678,604,630]" pageId="11" pageNumber="44">2. Yellowish or light brown color.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FD72FB94EBE1" blockId="11.[264,1323,564,990]" pageId="11" pageNumber="44">3. Buccal tube with short strengthening bar, and with insertion point of the stylet supports placed far from the posterior end of the buccal tube.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FD22FD44EA19" blockId="11.[264,1323,564,990]" pageId="11" pageNumber="44">4. Two macroplacoids, first with constriction, which could divide it in two, forming as a result three macroplacoids. Pharyngeal bulb with thin slanting cuticular bars between apophyses and first macroplacoids.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FCBAFE69EA91" blockId="11.[264,1323,564,990]" pageId="11" pageNumber="44">5. Long and thin claws, with well differentiated primary and secondary branches, secondary branch attached to the claw at right angle, claws IV are distinctly longer than claws I–III.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF868736FC32FCADEAF9" blockId="11.[264,1323,564,990]" box="[264,829,964,990]" pageId="11" pageNumber="44">6. Large dentate lunules, especially on legs IV.</paragraph>
|
||
<paragraph id="8BCC36F7D144FF818706FBE2FC77E8DA" blockId="11.[264,1323,1044,1870]" lastBlockId="12.[264,1323,284,710]" lastPageId="12" lastPageNumber="45" pageId="11" pageNumber="44">
|
||
Most of the above listed characters are of small phylogenetic significance. Large dimensions, coloured body, large dentate lunules and pharyngeal bulb with two macroplacoids are widely distributed within the genus
|
||
<taxonomicName id="4C734D74D144FF8685A3FB92FBA1ED5A" box="[925,1073,1124,1149]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D144FF8685A3FB92FBA1ED5A" box="[925,1073,1124,1149]" italics="true" pageId="11" pageNumber="44">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
. The significance of the presence of slanting cuticular bars in the pharyngeal bulb is unclear, because this structure is also present in other species of the genus
|
||
<taxonomicName id="4C734D74D144FF868582FB42FBC1EDEA" box="[956,1105,1204,1229]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D144FF868582FB42FBC1EDEA" box="[956,1105,1204,1229]" italics="true" pageId="11" pageNumber="44">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
(for example
|
||
<taxonomicName id="4C734D74D144FF868335FB42FD4BEDD1" authority="Bertolani & Rebecchi, 1993" authorityName="Bertolani & Rebecchi" authorityYear="1993" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="macrocalix">
|
||
<emphasis id="B907EAE5D144FF868335FB42FE1AEDD2" italics="true" pageId="11" pageNumber="44">M. macrocalix</emphasis>
|
||
<bibRefCitation id="EFE24B06D144FF8687ACFB2AFD4BEDD1" author="Bertolani" box="[402,731,1244,1270]" pageId="11" pageNumber="45" refString="Bertolani, R. & Rebecchi, L. (1993) A revision of the Macrobiotus hufelandi group (Tardigrada, Macrobiotidae), with some observations on the taxonomic characters of eutardigrades. Zoologica Scripta, 22, 127 - 152." type="journal article" year="1993">Bertolani & Rebecchi, 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C734D74D144FF86851CFB2AFC01EDD2" box="[802,913,1244,1269]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="hufelandi">
|
||
<emphasis id="B907EAE5D144FF86851CFB2AFC01EDD2" box="[802,913,1244,1269]" italics="true" pageId="11" pageNumber="44">hufelandi</emphasis>
|
||
</taxonomicName>
|
||
group). Moreover, these structures are present in some species, belonging to the other genera of Eutardigrada (e.g.
|
||
<taxonomicName id="4C734D74D144FF868292FAF2FABBEC3A" box="[1196,1323,1284,1309]" class="Eutardigrada" family="Macrobiotidae" genus="Minibiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D144FF868292FAF2FABBEC3A" box="[1196,1323,1284,1309]" italics="true" pageId="11" pageNumber="44">Minibiotus</emphasis>
|
||
</taxonomicName>
|
||
sp.; Tunisian population, author’s collection). Well developed baculiform structures visible on SEM photographs of buccopharyngeal apparatus of
|
||
<taxonomicName id="4C734D74D144FF86821FFAA2FD66ECB1" authority="Schuster Toftner & Grigarick 1977" authorityName="Schuster Toftner & Grigarick" authorityYear="1977" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="saltursus">
|
||
<emphasis id="B907EAE5D144FF86821FFAA2FABBEC4A" box="[1057,1323,1364,1389]" italics="true" pageId="11" pageNumber="44">Isohypsibius saltursus</emphasis>
|
||
<bibRefCitation id="EFE24B06D144FF868736FA8AFD39ECB1" author="Schuster" box="[264,681,1404,1430]" pageId="11" pageNumber="46" refString="Schuster, R. O., Toftner, E. C. & Grigarick, A. A. (1977 (1978 )) Tardigrada of Pope Beach, Lake Tahoe, California. Wasmann Journal of Biology, 35 (1), 115 - 136." type="journal article" year="1977">Schuster Toftner & Grigarick 1977</bibRefCitation>
|
||
(1978)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D74D144FF86850CFA8AFBC4ECB2" box="[818,1108,1404,1429]" class="Eutardigrada" family="Macrobiotidae" genus="Dactylobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="grandipes">
|
||
<emphasis id="B907EAE5D144FF86850CFA8AFBC4ECB2" box="[818,1108,1404,1429]" italics="true" pageId="11" pageNumber="44">Dactylobiotus grandipes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE24B06D144FF86825BFA8AFE4FEC99" author="Schuster" pageId="11" pageNumber="46" refString="Schuster, R. O., Toftner, E. C. & Grigarick, A. A. (1977 (1978 )) Tardigrada of Pope Beach, Lake Tahoe, California. Wasmann Journal of Biology, 35 (1), 115 - 136." type="journal article" year="1977">Schuster Toftner & Grigarick 1977</bibRefCitation>
|
||
(1978)) (
|
||
<bibRefCitation id="EFE24B06D144FF868479FA52FCAAEC99" author="Schuster" box="[583,826,1444,1470]" pageId="11" pageNumber="46" refString="Schuster, R. O., Nelson, D. R., Grigarick, A. A., Christenberry, D. (1980) Systematic criteria of the Eutardigrada. Transactions of the American Microscopical Society, 99, 284 - 303." type="journal article" year="1980">
|
||
Schuster
|
||
<emphasis id="B907EAE5D144FF868486FA52FD7EEC9A" box="[696,750,1444,1469]" italics="true" pageId="11" pageNumber="44">et al</emphasis>
|
||
. 1980
|
||
</bibRefCitation>
|
||
) seems to be homologous to the slanting cuticular bars, visible in optical microscope. The claw length cannot be considered as a base for the grouping of species without taking into account the exact structure of the claw. Recently it was stated that details of the claw structure should be taken into consideration when discussing the taxonomic position of
|
||
<taxonomicName id="4C734D74D144FF86852BF9B2FC50EF79" box="[789,960,1604,1630]" class="Eutardigrada" family="Macrobiotidae" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="family">Macrobiotidae</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EFE24B06D144FF86820EF9B2FED7EFA1" author="Guidetti" pageId="11" pageNumber="45" refString="Guidetti, R. & Bertolani, R. (2001) An evolutionary line of the Macrobiotinae (Tardigrada, Macrobiotidae): Calcarobiotus and related species. Italian Journal of Zoology, 68, 229 - 233." type="journal article" year="2001">Guidetti & Bertolani 2001</bibRefCitation>
|
||
). In my opinion the structure of the claws of
|
||
<taxonomicName id="4C734D74D144FF868545F99AFB5BEFA2" box="[891,1227,1644,1669]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D144FF868545F99AFB5BEFA2" box="[891,1227,1644,1669]" italics="true" pageId="11" pageNumber="44">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D74D144FF868335F99AFE17EF8A" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="kirghizicus">
|
||
<emphasis id="B907EAE5D144FF868335F99AFE17EF8A" italics="true" pageId="11" pageNumber="44">M. kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
is very different from the structure of the claws of other species of the
|
||
<taxonomicName id="4C734D74D144FF8682DAF962FABAEF8A" box="[1252,1322,1684,1709]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="tenuis">
|
||
<emphasis id="B907EAE5D144FF8682DAF962FABAEF8A" box="[1252,1322,1684,1709]" italics="true" pageId="11" pageNumber="44">tenuis</emphasis>
|
||
</taxonomicName>
|
||
group. In both species poorly differentiated main and secondary branches diverge from the very short and wide common tract. Distal part of the basal portion is very poorly marked, flattened in the frontal plane of the claw, and in the claws of the legs IV it is nearly totally fused with the stalk. On the other hand, other species of the
|
||
<taxonomicName id="4C734D74D144FF8685D9F8C2FBBDEE6A" box="[999,1069,1844,1869]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="11" pageNumber="44" phylum="Tardigrada" rank="species" species="tenuis">
|
||
<emphasis id="B907EAE5D144FF8685D9F8C2FBBDEE6A" box="[999,1069,1844,1869]" italics="true" pageId="11" pageNumber="44">tenuis</emphasis>
|
||
</taxonomicName>
|
||
group have different claw morphology, with well differentiated main and secondary branches, long thin common tract (secondary branch diverge at about half of the claw length) and typically developed distal part of the basal portion. Furthermore, all species of the
|
||
<taxonomicName id="4C734D74D143FF8182AAFE9AFB4AE8A2" box="[1172,1242,364,389]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="tenuis">
|
||
<emphasis id="B907EAE5D143FF8182AAFE9AFB4AE8A2" box="[1172,1242,364,389]" italics="true" pageId="12" pageNumber="45">tenuis</emphasis>
|
||
</taxonomicName>
|
||
group possesses a common feature in their claw structure – the stalk, connecting the claw base with the lunule, form a prominent frontal appendage on the claw base. This structure is absent in both
|
||
<taxonomicName id="4C734D74D143FF81878AFE12FC90E8DA" box="[436,768,484,509]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D143FF81878AFE12FC90E8DA" box="[436,768,484,509]" italics="true" pageId="12" pageNumber="45">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D74D143FF818507FE12FC73E8DA" box="[825,995,484,510]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="kirghizicus">
|
||
<emphasis id="B907EAE5D143FF818507FE12FCC1E8DA" box="[825,849,484,509]" italics="true" pageId="12" pageNumber="45">M</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D143FF81855EFE12FC73E8DA" box="[864,995,484,509]" italics="true" pageId="12" pageNumber="45">kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D143FF818706FDFAFD17EBE1" blockId="12.[264,1323,284,710]" pageId="12" pageNumber="45">
|
||
Taking into account those differences in claw morphology,
|
||
<taxonomicName id="4C734D74D143FF8182A8FDFAFE28EB6A" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D143FF8182A8FDFAFE28EB6A" italics="true" pageId="12" pageNumber="45">Macrobiotus ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C734D74D143FF8187CBFDC2FD0EEB6A" box="[501,670,564,589]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="kirghizicus" status="sp. nov.">
|
||
<emphasis id="B907EAE5D143FF8187CBFDC2FD0EEB6A" box="[501,670,564,589]" italics="true" pageId="12" pageNumber="45">M. kirghizicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED143FF818498FDC5FC96EB6A" box="[678,774,563,589]" pageId="12" pageNumber="45" rank="species">
|
||
<emphasis id="B907EAE5D143FF818498FDC5FC96EB6A" bold="true" box="[678,774,563,589]" pageId="12" pageNumber="45">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
should not be included in the
|
||
<taxonomicName id="4C734D74D143FF81824BFDC2FB2BEB6A" box="[1141,1211,564,589]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="tenuis">
|
||
<emphasis id="B907EAE5D143FF81824BFDC2FB2BEB6A" box="[1141,1211,564,589]" italics="true" pageId="12" pageNumber="45">tenuis</emphasis>
|
||
</taxonomicName>
|
||
group of the genus
|
||
<taxonomicName id="4C734D74D143FF8187B8FDAAFD8AEB52" box="[390,538,604,629]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D143FF8187B8FDAAFD8AEB52" box="[390,538,604,629]" italics="true" pageId="12" pageNumber="45">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
. Both species are undoubtedly closely related to each other and well detached from other species of
|
||
<taxonomicName id="4C734D74D143FF818493FD72FCD1EBBA" box="[685,833,644,669]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B907EAE5D143FF818493FD72FCD1EBBA" box="[685,833,644,669]" italics="true" pageId="12" pageNumber="45">Macrobiotus</emphasis>
|
||
</taxonomicName>
|
||
, so they should be combined into the new
|
||
<taxonomicName id="4C734D74D143FF818736FD5AFE28EBE2" box="[264,440,684,709]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="12" pageNumber="45" phylum="Tardigrada" rank="species" species="ariekammensis">
|
||
<emphasis id="B907EAE5D143FF818736FD5AFE28EBE2" box="[264,440,684,709]" italics="true" pageId="12" pageNumber="45">ariekammensis</emphasis>
|
||
</taxonomicName>
|
||
group of species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |