151 lines
20 KiB
XML
151 lines
20 KiB
XML
<document id="189FECB9197B7EFE3BDE2BD55526C428" ID-DOI="10.5281/zenodo.170519" ID-GBIF-Dataset="5c49c4b3-87e1-4e70-a3ab-af504169d977" ID-ISSN="11755326" ID-Zenodo-Dep="170519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460099730580" checkinUser="plazi" docAuthor="Arefina, Tatyana I." docDate="2005" docId="03DA87CCC30FB926FEF92FFABE2FF8D7" docLanguage="en" docName="zt01088p053.pdf" docOrigin="Zootaxa 1088" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Molanna byssa Arefina, 2005, new species" docType="treatment" docVersion="9" lastPageNumber="49" masterDocId="FFE3FFB4C30CB922FFF12B7CBB22FF86" masterDocTitle="Two new species and new records of caddisflies (Insecta: Trichoptera) from the Far East of Russia" masterLastPageNumber="53" masterPageNumber="45" pageNumber="48" updateTime="1698234811366" updateUser="plazi">
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<mods:titleInfo id="48056C31A0A6916AA057DABC98E2D564">
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<mods:title id="A23570AD3E9E28818B202FA9AC557768">Two new species and new records of caddisflies (Insecta: Trichoptera) from the Far East of Russia</mods:title>
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<mods:name id="84122C511C0D5BE4F2159E27B11C22FD" type="personal">
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<mods:roleTerm id="968A59CCBAB72DC402B804EFEC88FF47">Author</mods:roleTerm>
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<mods:namePart id="DA6D5848166C69C9AF0971446284549E">Arefina, Tatyana I.</mods:namePart>
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<mods:typeOfResource id="9FF62EFE8786BF558FFE0DADAA36A54C">text</mods:typeOfResource>
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<mods:title id="DB46F46317C14EAF1261B99B17B8F074">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F02438412A22D9298A2763292E8F67F3">
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<mods:date id="97C05EC707CA1BD34C085570735C0DA8">2005</mods:date>
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<mods:detail id="E0E4CBB19272ED60310FC3A000937CC0" type="volume">
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<mods:number id="6C302CEC5CC2D7E9FA3382F5C13A7E45">1088</mods:number>
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<mods:classification id="6C1F86E61978B65AD363A3E2C8F95884">journal article</mods:classification>
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<mods:identifier id="63631BCDBA8A7E1CFF5E452BD4C4D850" type="DOI">10.5281/zenodo.170519</mods:identifier>
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<mods:identifier id="0D2800DFBC4F5307E98714B79E41B9EE" type="ISSN">11755326</mods:identifier>
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<treatment id="03DA87CCC30FB926FEF92FFABE2FF8D7" ID-DOI="http://doi.org/10.5281/zenodo.6267483" ID-GBIF-Taxon="124857767" ID-Zenodo-Dep="6267483" LSID="urn:lsid:plazi:treatment:03DA87CCC30FB926FEF92FFABE2FF8D7" httpUri="http://treatment.plazi.org/id/03DA87CCC30FB926FEF92FFABE2FF8D7" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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<subSubSection id="C3696551C30FB921FEF92FFABA4CFB45" pageId="3" pageNumber="48" type="nomenclature">
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<paragraph id="8BCC36DAC30FB921FEF92FFAB97FFB26" blockId="3.[264,605,1158,1219]" box="[264,605,1158,1184]" pageId="3" pageNumber="48">
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<heading id="D08481B6C30FB921FEF92FFAB97FFB26" bold="true" box="[264,605,1158,1184]" fontSize="11" level="1" pageId="3" pageNumber="48" reason="1">
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<emphasis id="B907EAC8C30FB921FEF92FFAB97FFB26" bold="true" box="[264,605,1158,1184]" pageId="3" pageNumber="48">
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<taxonomicName id="4C734D59C30FB921FEF92FFABA9DFB26" box="[264,447,1158,1184]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="byssa" status="sp. nov.">
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<emphasis id="B907EAC8C30FB921FEF92FFABA9DFB26" bold="true" box="[264,447,1158,1184]" italics="true" pageId="3" pageNumber="48">Molanna byssa</emphasis>
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A23457B3C30FB921FE3F2FFAB97FFB26" box="[462,605,1158,1184]" pageId="3" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCC36DAC30FB921FEF92FD5BA4CFB45" blockId="3.[264,605,1158,1219]" box="[264,366,1193,1219]" pageId="3" pageNumber="48">
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<figureCitation id="13482A5FC30FB921FEF92FD5BA71FB45" box="[264,339,1193,1219]" captionStart="FIGURE 2" captionStartId="4.[264,368,1220,1244]" captionTargetBox="[396,1247,570,1219]" captionTargetId="figure@4.[319,1268,542,1231]" captionTargetPageId="4" captionText="FIGURE 2. Molanna byssa, new species, holotype (wing venation): A. Forewing; B. Hind wing." httpUri="https://zenodo.org/record/170521/files/figure.png" pageId="3" pageNumber="48">Figs 2</figureCitation>
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,
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<figureCitation id="13482A5FC30FB921FEAE2FD5BA4CFB45" box="[351,366,1193,1219]" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="3" pageNumber="48">3</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3696551C30FB926FEF92F85BE2FF8D7" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48" type="materials_examined">
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<paragraph id="8BCC36DAC30FB921FEF92F85B9EAFABD" blockId="3.[264,1323,1273,1899]" pageId="3" pageNumber="48">
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<emphasis id="B907EAC8C30FB921FEF92F85B870FA95" bold="true" box="[264,850,1273,1299]" pageId="3" pageNumber="48">
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<typeStatus id="54C88878C30FB921FEF92F85BA58FA95" box="[264,378,1273,1299]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
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male.
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<collectingCountry id="F364764AC30FB921FE242F85B967FA95" box="[469,581,1273,1299]" name="Russia" pageId="3" pageNumber="48">
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<collectionCode id="ED62AE1FC30FB921FE242F85B967FA95" box="[469,581,1273,1299]" pageId="3" pageNumber="48">RUSSIA</collectionCode>
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</collectingCountry>
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: Amurskaya Oblast:
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</emphasis>
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<taxonomicName id="4C734D59C30FB921FCAC2F85B881FA95" box="[861,931,1273,1299]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="byssa" status="sp. nov.">Byssa</taxonomicName>
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River at Fevralsk Town (Amur River Basin),
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<date id="FFCD101AC30FB921FE5D2E5DB909FABD" box="[428,555,1313,1339]" pageId="3" pageNumber="48" value="2004-06-15">15.vi.2004</date>
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, T.I. Arefina.
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</paragraph>
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<paragraph id="8BCC36DAC30FB921FEC92E35B801FAE5" blockId="3.[264,1323,1273,1899]" box="[312,803,1353,1379]" pageId="3" pageNumber="48">
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<emphasis id="B907EAC8C30FB921FEC92E35BA93FAE5" bold="true" box="[312,433,1353,1379]" pageId="3" pageNumber="48">
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<typeStatus id="54C88878C30FB921FEC92E35BA8EFAE5" box="[312,428,1353,1379]" pageId="3" pageNumber="48" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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Same data as
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<typeStatus id="54C88878C30FB921FDA92E35B9E0FAE5" box="[600,706,1353,1379]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
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,
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<specimenCount id="9D75FD53C30FB921FD3C2E35B802FAE5" box="[717,800,1353,1379]" pageId="3" pageNumber="48" type="male">1 male</specimenCount>
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.
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</paragraph>
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<paragraph id="8BCC36DAC30FB921FEC92E0DBABAF9AD" blockId="3.[264,1323,1273,1899]" pageId="3" pageNumber="48">
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Adult. Length of fore wing: male 11.4 mm. Head and thorax dark brown, antennae, legs and abdomen lighter. Forewing brownish, hind wing paler, both covered with short dark hair all over membrane; venation as in
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<figureCitation id="13482A5FC30FB921FCF82EBDB872FA5D" box="[777,848,1473,1499]" captionStart="FIGURE 2" captionStartId="4.[264,368,1220,1244]" captionTargetBox="[396,1247,570,1219]" captionTargetId="figure@4.[319,1268,542,1231]" captionTargetPageId="4" captionText="FIGURE 2. Molanna byssa, new species, holotype (wing venation): A. Forewing; B. Hind wing." httpUri="https://zenodo.org/record/170521/files/figure.png" pageId="3" pageNumber="48">Fig. 2</figureCitation>
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.
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<collectingCountry id="F364764AC30FB921FCAA2EBDB8B4FA5D" box="[859,918,1473,1499]" name="India" pageId="3" pageNumber="48">Hind</collectingCountry>
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wing with row of small hamuli on front edge. Arrangement and form of setal warts on head and thorax typical for genus. Spurs 244.
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</paragraph>
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<paragraph id="8BCC36DAC30FB926FEC92D45B801FE50" blockId="3.[264,1323,1273,1899]" lastBlockId="4.[264,1323,284,470]" lastPageId="4" lastPageNumber="49" pageId="3" pageNumber="48">
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Male genitalia (
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<figureCitation id="13482A5FC30FB921FE022D45B919F9D5" box="[499,571,1593,1619]" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="3" pageNumber="48">Fig. 3</figureCitation>
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). Segment IX with anterior margin broadly triangular in lateral view; posterodorsally with mesal membranous lobe, which is slightly shorter in the
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<typeStatus id="54C88878C30FB921FEF92DF5BA4FF925" box="[264,365,1673,1699]" pageId="3" pageNumber="48" type="paratype">paratype</typeStatus>
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male. Preanal appendage about 1/2 length of inferior appendage, elongate, slightly wider at midlength than at base and apex; apex truncate. Intermediate appendage composed of flat thin plates; in lateral view plates curved and arched posteroventrally, gradually tapering to rounded apex in lateral view and bearing several short stout setae along lateral margin; flattened dorsomesal plate wide basally, tapering to triangular apex, bearing 2 stout distal spines; dorsomesal plate and intermediate appendage connected by lateral sclerite, having many long stout spines along posterior margin. Inferior appendage elongate, tapering in lateral view, with basoventral projection curved mesoventrally, apex of this projection rounded; inner knob of inferior appendage straight, fingerlike, with several short setae. Phallotheca rather robust, strongly curved, no spines in endotheca; phallotremal sclerite Ushaped in dorsal view.
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</paragraph>
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<caption id="DF0C6652C308B926FEF92FB8BE34FB5D" box="[264,1302,1220,1244]" httpUri="https://zenodo.org/record/170521/files/figure.png" pageId="4" pageNumber="49" targetBox="[396,1247,570,1219]" targetPageId="4">
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||
<paragraph id="8BCC36DAC308B926FEF92FB8BE34FB5D" blockId="4.[264,1302,1220,1244]" box="[264,1302,1220,1244]" pageId="4" pageNumber="49">
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<emphasis id="B907EAC8C308B926FEF92FB8BAA8FB5A" bold="true" box="[264,394,1220,1244]" pageId="4" pageNumber="49">FIGURE 2.</emphasis>
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<taxonomicName id="4C734D59C308B926FE602FB9B916FB5A" box="[401,564,1221,1244]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="byssa" status="sp. nov.">
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<emphasis id="B907EAC8C308B926FE602FB9B916FB5A" box="[401,564,1221,1244]" italics="true" pageId="4" pageNumber="49">Molanna byssa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A23457B3C308B926FDB02FB8B9E6FB5D" box="[577,708,1220,1243]" pageId="4" pageNumber="49" rank="species">new species</taxonomicNameLabel>
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, holotype (wing venation): A. Forewing; B. Hind wing.
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</paragraph>
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</caption>
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<paragraph id="8BCC36DAC308B926FEC92E53B820FACF" blockId="4.[264,1323,1327,1873]" box="[312,770,1327,1353]" pageId="4" pageNumber="49">Female and immature stages unknown.</paragraph>
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<paragraph id="8BCC36DAC308B926FEC92E2BB952F887" blockId="4.[264,1323,1327,1873]" pageId="4" pageNumber="49">
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Diagnosis. Placement of the new species among species of the genus
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<taxonomicName id="4C734D59C308B926FB512E24BE28FAF7" box="[1184,1290,1368,1393]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="genus">
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<emphasis id="B907EAC8C308B926FB512E24BE28FAF7" box="[1184,1290,1368,1393]" italics="true" pageId="4" pageNumber="49">Molanna</emphasis>
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</taxonomicName>
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is uncertain as it does not belong to any of the existing subgenera. The male genitalia of
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<taxonomicName id="4C734D59C308B926FEF92ED4BA98FA47" box="[264,442,1448,1473]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="byssa">
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<emphasis id="B907EAC8C308B926FEF92ED4BA98FA47" box="[264,442,1448,1473]" italics="true" pageId="4" pageNumber="49">Molanna byssa</emphasis>
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</taxonomicName>
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share some similarities with
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<taxonomicName id="4C734D59C308B926FCE32ED4BF71FA47" authority="Malicky 1994" authorityName="Malicky" authorityYear="1994" box="[786,1107,1447,1473]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="xiaguana">
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<emphasis id="B907EAC8C308B926FCE32ED4B884FA47" box="[786,934,1448,1473]" italics="true" pageId="4" pageNumber="49">M. xiaguana</emphasis>
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<bibRefCitation id="EFE24B2BC308B926FC5C2EDBBF71FA47" author="Malicky" box="[941,1107,1447,1473]" pageId="4" pageNumber="49" refString="Malicky, H. (1994) Neue Trichopteren aus Nepal, Vietnam, China, von den Philippinen und vom Bismarck-Archipel (Trichoptera). Entomologische Berichte Lunzen, 31, 163 - 172." type="journal article" year="1994">Malicky 1994</bibRefCitation>
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</taxonomicName>
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, whose placement in the genus is also uncertain (
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<bibRefCitation id="EFE24B2BC308B926FD9F2EB3B83AFA6F" author="Malicky" box="[622,792,1487,1513]" pageId="4" pageNumber="49" refString="Malicky, H. (1994) Neue Trichopteren aus Nepal, Vietnam, China, von den Philippinen und vom Bismarck-Archipel (Trichoptera). Entomologische Berichte Lunzen, 31, 163 - 172." type="journal article" year="1994">Malicky 1994</bibRefCitation>
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). Both species are similar in the shape of the preanal appendages in lateral view (
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<figureCitation id="13482A5FC308B926FD442E8BB9D8F997" box="[693,762,1527,1553]" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="4" pageNumber="49">Fig. 3</figureCitation>
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A) and the inferior appendages in ventral view (
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<figureCitation id="13482A5FC308B926FEE02D63BA75F9BF" box="[273,343,1567,1593]" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="4" pageNumber="49">Fig. 3</figureCitation>
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C); in the possession of the dorsomesal plates of the intermediate appendages (
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<figureCitation id="13482A5FC308B926FB092D63BA35F9E7" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="4" pageNumber="49">Fig. 3</figureCitation>
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B); and in the absence of endothecal spines (
|
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<figureCitation id="13482A5FC308B926FCD22D3BB844F9E7" box="[803,870,1607,1633]" captionStart="FIGURE 3" captionStartId="5.[264,367,1387,1411]" captionTargetBox="[350,1236,326,1341]" captionTargetId="figure@5.[319,1268,296,1363]" captionTargetPageId="5" captionText="FIGURE 3. Molanna byssa, new species, holotype (male genitalia): A. Lateral view; B. Dorsal view (right preanal appendage omitted); C. Ventral view; D. Caudal view; E. Phallus in lateral view; F. Phallus in dorsal view." httpUri="https://zenodo.org/record/170522/files/figure.png" pageId="4" pageNumber="49">Fig. 3</figureCitation>
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E & F). The new species differs in the forewing venation (
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<figureCitation id="13482A5FC308B926FE1E2D13B914F90F" box="[495,566,1647,1673]" captionStart="FIGURE 2" captionStartId="4.[264,368,1220,1244]" captionTargetBox="[396,1247,570,1219]" captionTargetId="figure@4.[319,1268,542,1231]" captionTargetPageId="4" captionText="FIGURE 2. Molanna byssa, new species, holotype (wing venation): A. Forewing; B. Hind wing." httpUri="https://zenodo.org/record/170521/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
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A) and in the absence of a long row of yellow hairs through the middle of the hind wing (
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<figureCitation id="13482A5FC308B926FD952DEBB997F937" box="[612,693,1687,1713]" captionStart="FIGURE 2" captionStartId="4.[264,368,1220,1244]" captionTargetBox="[396,1247,570,1219]" captionTargetId="figure@4.[319,1268,542,1231]" captionTargetPageId="4" captionText="FIGURE 2. Molanna byssa, new species, holotype (wing venation): A. Forewing; B. Hind wing." httpUri="https://zenodo.org/record/170521/files/figure.png" pageId="4" pageNumber="49">Fig. 2</figureCitation>
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B). In addition, the complicated structure of the intermediate appendages armed with many stout spines distinguishes
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<taxonomicName id="4C734D59C308B926FBB22DBCBF94F95F" box="[1091,1206,1728,1753]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="byssa">
|
||
<emphasis id="B907EAC8C308B926FBB22DBCBF94F95F" box="[1091,1206,1728,1753]" italics="true" pageId="4" pageNumber="49">M. byssa</emphasis>
|
||
</taxonomicName>
|
||
from this and other species of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BCC36DAC308B926FEC92C73B852F8AF" blockId="4.[264,1323,1327,1873]" box="[312,880,1807,1833]" pageId="4" pageNumber="49">
|
||
Distribution. Known only from the
|
||
<typeStatus id="54C88878C308B926FD282C73B82EF8AF" box="[729,780,1807,1833]" pageId="4" pageNumber="49">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph id="8BCC36DAC308B926FEC92C4BBE2FF8D7" blockId="4.[264,1323,1327,1873]" box="[312,1293,1847,1873]" pageId="4" pageNumber="49">
|
||
Etymology. The new species is named after the
|
||
<taxonomicName id="4C734D59C308B926FC982C4BB88DF8D7" box="[873,943,1847,1873]" class="Insecta" family="Molannidae" genus="Molanna" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="byssa" status="sp. nov.">Byssa</taxonomicName>
|
||
River where it was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |