394 lines
51 KiB
XML
394 lines
51 KiB
XML
<document id="D2B350DFA8CBCECAD296661977C7A242" ID-CLB-Dataset="45928" ID-DOI="10.11646/zootaxa.3793.2.3" ID-GBIF-Dataset="cec96d21-19fa-4e66-8305-9e70166f04f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="230581" ID-ZooBank="5E6DB00C-E98F-4D9D-958B-95B634E6D9E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460412815762" checkinUser="plazi" docAuthor="Baca, Stephen M., Gustafson, Grey T., Toledo, Mario & Miller, Kelly B." docDate="2014" docId="03D9AD22FFBEFFFC75A8FBA0FA5C9222" docLanguage="en" docName="zt03793p246.pdf" docOrigin="Zootaxa 3793 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocanthydrus nanops Baca, Gustafson, Toledo & Miller, 2014, sp. n." docType="treatment" docVersion="9" lastPageNumber="244" masterDocId="FFE0D55AFFB2FFF1753FFF9FFF65972F" masterDocTitle="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species" masterLastPageNumber="246" masterPageNumber="231" pageNumber="243" updateTime="1698301202102" updateUser="ExternalLinkService">
|
||
<mods:mods id="695D076030E158EC19F719C0B7C82071" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CE336E4488E026BEBCC29D4C88E2C8DE">
|
||
<mods:title id="2E2292B6A921A28C7552F6C6A2A1FEAD">Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1A28814114D9F608EA82B7C58839C818" type="personal">
|
||
<mods:role id="7E52758DC34FDC47964DFEC1136E75DD">
|
||
<mods:roleTerm id="96C7EB695993B54C85634B3609E7A42C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="41EE6B33BA600092E6534DC1332D6E94">Baca, Stephen M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="18055BE93CFFEE35BC5F11AE24EE9EB6" type="personal">
|
||
<mods:role id="A1D25D7A4F75488FCF1D59A0624113C1">
|
||
<mods:roleTerm id="E13CAE29E34D927D1D6ACD517A029EF4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1FE7E612C7A9D8A3AF68A293878017AA">Gustafson, Grey T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A130E424FE985E52A084C00BD81977B4" type="personal">
|
||
<mods:role id="662920272EC500F59FB5F44A7F7B6703">
|
||
<mods:roleTerm id="AB5EB5918DF6E727F174C05053A5BBDE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="298C634E03FBD2CC342A608920EC1B32">Toledo, Mario</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6BA36CC3D10637E2C50533D8E58312AB" type="personal">
|
||
<mods:role id="BDA3C0A3B5840BD0A8795C4A8B66BCB1">
|
||
<mods:roleTerm id="51390A5B74F01B7346DE7967E69F3C75">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="362D5B3E903F983222E8A865B5332699">Miller, Kelly B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="AE04E8AE3FD876CCECF28A90FD4F3937">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3369F4DF083A2CB8FB60EEB86D649C48" type="host">
|
||
<mods:titleInfo id="47365801BFC3533F3D7CDE81515D41A6">
|
||
<mods:title id="B64347E57F35B9AD339F3CD44718E237">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9444EC76C1F57A3E5493D6D71BC983F8">
|
||
<mods:date id="99590785EE19B7AF8D6DC52F22F86AED">2014</mods:date>
|
||
<mods:detail id="E93276CF97C0C38136543F6CE73BD544" type="volume">
|
||
<mods:number id="8BCB9F45165C8F6CEBF58FDB4BE3E914">3793</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="0933E0E66DC7768E3A9CAA32162FC893" type="issue">
|
||
<mods:number id="AAE50AC120669D3CED2703BD5BD2D7B7">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="DE9966F33AB360A61F58B00E83CCDC88" unit="page">
|
||
<mods:start id="179F5EADCBDC919BE833714D5DECC9A4">231</mods:start>
|
||
<mods:end id="551FE6940A63801A8E6B5FB65479B0B7">246</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="24E7DC1C96E82F94BE1707D6E2D690ED">journal article</mods:classification>
|
||
<mods:identifier id="93823C93AA8765F9242363974B217397" type="CLB-Dataset">45928</mods:identifier>
|
||
<mods:identifier id="FBD0A7108172A9FF2940DE3BFC288F91" type="DOI">10.11646/zootaxa.3793.2.3</mods:identifier>
|
||
<mods:identifier id="116DEA78FDB5AF2F4344A7BB3827FA1E" type="GBIF-Dataset">cec96d21-19fa-4e66-8305-9e70166f04f7</mods:identifier>
|
||
<mods:identifier id="D27F7947514C20B6793FF843542AC249" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="46C37CDA79F2360043AD88B457173200" type="Zenodo-Dep">230581</mods:identifier>
|
||
<mods:identifier id="1301EA557C57FE9BE7967164D6B05677" type="ZooBank">5E6DB00C-E98F-4D9D-958B-95B634E6D9E4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D9AD22FFBEFFFC75A8FBA0FA5C9222" ID-DOI="http://doi.org/10.5281/zenodo.6131937" ID-GBIF-Taxon="119423980" ID-Zenodo-Dep="6131937" LSID="urn:lsid:plazi:treatment:03D9AD22FFBEFFFC75A8FBA0FA5C9222" httpUri="http://treatment.plazi.org/id/03D9AD22FFBEFFFC75A8FBA0FA5C9222" lastPageId="13" lastPageNumber="244" pageId="12" pageNumber="243">
|
||
<subSubSection id="C36A4FBFFFBEFFFD75A8FBA0FE8A9375" box="[151,495,1087,1114]" pageId="12" pageNumber="243" type="nomenclature">
|
||
<paragraph id="8BCF1C34FFBEFFFD75A8FBA0FE8A9375" blockId="12.[151,495,1087,1114]" box="[151,495,1087,1114]" pageId="12" pageNumber="243">
|
||
<heading id="D087AB58FFBEFFFD75A8FBA0FE8A9375" bold="true" box="[151,495,1087,1114]" fontSize="11" level="1" pageId="12" pageNumber="243" reason="1">
|
||
<emphasis id="B904C026FFBEFFFD75A8FBA0FE8A9375" bold="true" box="[151,495,1087,1114]" pageId="12" pageNumber="243">
|
||
<taxonomicName id="4C7067B7FFBEFFFD75A8FBA0FEC39376" box="[151,422,1087,1113]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops" status="sp. nov.">
|
||
<emphasis id="B904C026FFBEFFFD75A8FBA0FEC39376" bold="true" box="[151,422,1087,1113]" italics="true" pageId="12" pageNumber="243">Liocanthydrus nanops</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377D5DFFBEFFFD7492FBDFFE8A9375" box="[429,495,1088,1114]" pageId="12" pageNumber="243" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBEFFFD75A8FB19FBEB9128" pageId="12" pageNumber="243" type="materials_examined">
|
||
<paragraph id="8BCF1C34FFBEFFFD75A8FB19FC0F93B0" blockId="12.[151,1437,1158,2011]" box="[151,874,1158,1183]" pageId="12" pageNumber="243">
|
||
<emphasis id="B904C026FFBEFFFD75A8FB19FE5093B0" bold="true" box="[151,309,1158,1183]" pageId="12" pageNumber="243">
|
||
<typeStatus id="54CBA296FFBEFFFD75A8FB19FFB793B0" box="[151,210,1158,1183]" pageId="12" pageNumber="243">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F3675CA4FFBEFFFD7403FB18FEC093B0" box="[316,421,1159,1183]" name="Paraguay" pageId="12" pageNumber="243">Paraguay</collectingCountry>
|
||
, Paraguari Dept., Ybycui National Park.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBEFFFD75F8FB35FBEB9128" blockId="12.[151,1437,1158,2011]" pageId="12" pageNumber="243">
|
||
<emphasis id="B904C026FFBEFFFD75F8FB35FE1C93EC" bold="true" box="[199,377,1194,1219]" pageId="12" pageNumber="243">
|
||
<typeStatus id="54CBA296FFBEFFFD75F8FB35FE6793EC" box="[199,258,1194,1219]" pageId="12" pageNumber="243">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54CBA296FFBEFFFD74BAFB35FE8893EC" box="[389,493,1194,1219]" pageId="12" pageNumber="243" type="holotype">Holotype</typeStatus>
|
||
(male): “
|
||
<collectingCountry id="F3675CA4FFBEFFFD7760FB34FD9F93EC" box="[607,762,1195,1219]" name="Paraguay" pageId="12" pageNumber="243">PARAGUAY</collectingCountry>
|
||
: Paraguari Dept./Ybycui (
|
||
<quantity id="4C88B1D1FFBEFFFD7114FB35FB1D93EC" box="[1067,1144,1194,1219]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="243" unit="km" value="25.0">25 km</quantity>
|
||
SE), in/Ybycui National Park/
|
||
<date id="FFCE3AF4FFBEFFFD75E9FB51FEC793C8" box="[214,418,1230,1255]" pageId="12" pageNumber="243" value="1980-04-12" valueMax="1980-04-24" valueMin="1980-04-12">12-24 April 1980</date>
|
||
/P. J. Spangler et al.” [printed], “
|
||
<typeStatus id="54CBA296FFBEFFFD7609FB50FCA993C8" box="[822,972,1231,1255]" pageId="12" pageNumber="243" type="holotype">HOLOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD76EBFB51FB1393C8" box="[980,1142,1230,1255]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBEFFFD76EBFB51FB1393C8" box="[980,1142,1230,1255]" italics="true" pageId="12" pageNumber="243">Liocanthydrus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD7141FB50FBAA93C9" box="[1150,1231,1231,1254]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD7141FB50FBAA93C9" box="[1150,1231,1231,1254]" italics="true" pageId="12" pageNumber="243">nanops</emphasis>
|
||
</taxonomicName>
|
||
/Baca, Gustafson, Toledo/ & Miller, 2014” [red with black line border; printed] (
|
||
<collectionCode id="ED6184F1FFBEFFFD7648FB6CFCAB9224" LSID="urn:lsid:biocol.org:col:34871" box="[887,974,1267,1291]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="243">USNM</collectionCode>
|
||
).
|
||
<typeStatus id="54CBA296FFBEFFFD76DAFB6CFB339224" box="[997,1110,1267,1291]" pageId="12" pageNumber="243" type="paratype">Paratypes</typeStatus>
|
||
: Same data as
|
||
<typeStatus id="54CBA296FFBEFFFD7036FB6DFA0E9224" box="[1289,1387,1266,1291]" pageId="12" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="9D76D7BDFFBEFFFD7041FB6CFFB99200" pageId="12" pageNumber="243" type="male">27 males</specimenCount>
|
||
;
|
||
<specimenCount id="9D76D7BDFFBEFFFD75D8FA88FE079200" box="[231,354,1302,1327]" pageId="12" pageNumber="243" type="female">30 females</specimenCount>
|
||
exs.
|
||
<collectionCode id="ED6184F1FFBEFFFD74A2FA88FE969200" LSID="urn:lsid:biocol.org:col:34871" box="[413,499,1303,1327]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="243">USNM</collectionCode>
|
||
); “
|
||
<date id="FFCE3AF4FFBEFFFD772EFA88FD1F9200" box="[529,634,1303,1327]" pageId="12" pageNumber="243" value="33-12-29">29.12.33</date>
|
||
”, “2643”, “
|
||
<collectingCountry id="F3675CA4FFBEFFFD77C6FA89FCFC9200" box="[761,921,1302,1327]" name="Paraguay" pageId="12" pageNumber="243">PARAGUAY</collectingCountry>
|
||
:/Hohenau./Alto-Parana./H. Jacob./B.M. 1954- 814.”, “
|
||
<taxonomicName id="4C7067B7FFBEFFFD75D0FAA5FE17927C" box="[239,370,1338,1363]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBEFFFD75D0FAA5FE17927C" box="[239,370,1338,1363]" italics="true" pageId="12" pageNumber="243">Canthydrus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD7445FAA3FD61927C" box="[378,516,1340,1363]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBEFFFD7445FAA3FD61927C" box="[378,516,1340,1363]" italics="true" pageId="12" pageNumber="243">octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
/Zimmermann/Luis E. Grosso det./86” [label description not available] (
|
||
<specimenCount id="9D76D7BDFFBEFFFD700BFAA4FAFC927C" box="[1332,1433,1338,1363]" pageId="12" pageNumber="243" type="female">1 female</specimenCount>
|
||
:
|
||
<collectionCode id="ED6184F1FFBEFFFD75A8FAC0FFB89258" box="[151,221,1375,1399]" pageId="12" pageNumber="243">NHM</collectionCode>
|
||
); “
|
||
<collectingCountry id="F3675CA4FFBEFFFD743EFAC1FE039258" box="[257,358,1374,1399]" name="Brazil" pageId="12" pageNumber="243">BRASIL</collectingCountry>
|
||
/Guarapuava/Guarauna/10-
|
||
<date id="FFCE3AF4FFBEFFFD77A6FAC1FDD19258" box="[665,692,1374,1399]" bridgedPair="/" pageId="12" pageNumber="243" value="1957-05">57</date>
|
||
/
|
||
<collectingCountry id="F3675CA4FFBEFFFD7782FAC1FD9B9259" box="[701,766,1374,1399]" name="Vietnam" pageId="12" pageNumber="243">V. N.</collectingCountry>
|
||
Alin” [printed except “10” (handwritten)] (
|
||
<specimenCount id="9D76D7BDFFBEFFFD702AFAC0FA009258" box="[1301,1381,1374,1399]" pageId="12" pageNumber="243" type="male">1 male</specimenCount>
|
||
ex.,
|
||
<collectionCode id="ED6184F1FFBEFFFD75A8FA1CFF8892B4" LSID="urn:lsid:biocol.org:col:34871" box="[151,237,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="243">USNM</collectionCode>
|
||
); same data except with “
|
||
<taxonomicName id="4C7067B7FFBEFFFD7721FA1DFD0F92B4" box="[542,618,1410,1435]" class="Insecta" family="Noteridae" genus="Siolius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBEFFFD7721FA1DFD0F92B4" box="[542,618,1410,1435]" italics="true" pageId="12" pageNumber="243">Siolius</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD774DFA1DFDDE92B4" box="[626,699,1410,1435]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD774DFA1DFDDE92B4" box="[626,699,1410,1435]" italics="true" pageId="12" pageNumber="243">clayae</emphasis>
|
||
</taxonomicName>
|
||
/J.B.-B/Det. FN Young” [handwritten] (
|
||
<specimenCount id="9D76D7BDFFBEFFFD71B5FA1CFB8992B4" box="[1162,1260,1410,1435]" pageId="12" pageNumber="243" type="female">1 female</specimenCount>
|
||
ex.,
|
||
<collectionCode id="ED6184F1FFBEFFFD7014FA1CFAE592B4" LSID="urn:lsid:biocol.org:col:34871" box="[1323,1408,1411,1435]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="243">USNM</collectionCode>
|
||
;
|
||
<specimenCount id="9D76D7BDFFBEFFFD70B1FA1CFF869290" pageId="12" pageNumber="243" type="female">1 female</specimenCount>
|
||
ex.,
|
||
<collectionCode id="ED6184F1FFBEFFFD7426FA38FE019290" box="[281,356,1447,1471]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="12" pageNumber="243">FSCA</collectionCode>
|
||
); same data except no det. label, instead “FNY/Kept2/adbt2” [handwritten in pencil] (
|
||
<specimenCount id="9D76D7BDFFBEFFFD701BFA38FA089290" box="[1316,1389,1446,1471]" pageId="12" pageNumber="243" type="male">1 male</specimenCount>
|
||
ex.,
|
||
<collectionCode id="ED6184F1FFBEFFFD75A8FA54FF8F92CC" LSID="urn:lsid:biocol.org:col:34871" box="[151,234,1483,1507]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="243">USNM</collectionCode>
|
||
) All
|
||
<typeStatus id="54CBA296FFBEFFFD741DFA54FEEB92CC" box="[290,398,1483,1507]" pageId="12" pageNumber="243" type="paratype">paratypes</typeStatus>
|
||
labeled with: “
|
||
<typeStatus id="54CBA296FFBEFFFD7709FA54FDA192CC" box="[566,708,1483,1507]" pageId="12" pageNumber="243" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD77F3FA55FC0B92CC" box="[716,878,1482,1507]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904C026FFBEFFFD77F3FA55FC0B92CC" box="[716,878,1482,1507]" italics="true" pageId="12" pageNumber="243">Liocanthydrus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBEFFFD764AFA53FCA392CC" box="[885,966,1484,1507]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD764AFA53FCA392CC" box="[885,966,1484,1507]" italics="true" pageId="12" pageNumber="243">nanops</emphasis>
|
||
</taxonomicName>
|
||
/Baca, Gustafson, Toledo/& Miller, 2014” [blue with black line border; printed]
|
||
<emphasis id="B904C026FFBEFFFD7703FA71FD189128" bold="true" box="[572,637,1518,1543]" pageId="12" pageNumber="243">Note:</emphasis>
|
||
All label descriptions are included in brackets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBEFFFD75F8F98DFB4E90F4" pageId="12" pageNumber="243" type="diagnosis">
|
||
<paragraph id="8BCF1C34FFBEFFFD75F8F98DFE699194" blockId="12.[151,1437,1158,2011]" pageId="12" pageNumber="243">
|
||
<emphasis id="B904C026FFBEFFFD75F8F98DFE5F9104" bold="true" box="[199,314,1554,1579]" pageId="12" pageNumber="243">Diagnosis</emphasis>
|
||
. Elytra maculate, distinguishable from other species by the following combination of characters: a) eyes small relative to head capsule, HW/EW = 1.47–1.52 (
|
||
<figureCitation id="134B00B1FFBEFFFD760CF9A9FC189160" box="[819,893,1590,1615]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="12" pageNumber="243">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="134B00B1FFBEFFFD76B5F9A9FCCC9160" box="[906,937,1590,1615]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="12" pageNumber="243">15</figureCitation>
|
||
); b) posterior elytral bands strongly oblique and extending anterolaterally to basal halves of elytra (
|
||
<figureCitation id="134B00B1FFBEFFFD77C4F9C5FC5A915C" box="[763,831,1626,1651]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="12" pageNumber="243">Fig. 8</figureCitation>
|
||
); c) aedeagus as in
|
||
<figureCitation id="134B00B1FFBEFFFD7126F9C5FB0E915C" box="[1049,1131,1626,1651]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="12" pageNumber="243">Figs 21</figureCitation>
|
||
a–e; median lobe sinuate in dorsal aspect (
|
||
<figureCitation id="134B00B1FFBEFFFD7408F9E1FEEF91B8" box="[311,394,1662,1687]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="12" pageNumber="243">Fig. 21</figureCitation>
|
||
b); left lateral lobe broad (
|
||
<figureCitation id="134B00B1FFBEFFFD7792F9E1FD9B91B8" box="[685,766,1662,1687]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="12" pageNumber="243">Fig. 21</figureCitation>
|
||
d), right lateral lobe with distal angle rounded and projected (
|
||
<figureCitation id="134B00B1FFBEFFFD75A0F93DFF979194" box="[159,242,1698,1723]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="12" pageNumber="243">Fig. 21</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBEFFFD75F8F959FB4E90F4" blockId="12.[151,1437,1158,2011]" pageId="12" pageNumber="243">
|
||
<taxonomicName id="4C7067B7FFBEFFFD75F8F959FEA091F1" box="[199,453,1734,1759]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD75F8F959FEA091F1" box="[199,453,1734,1759]" italics="true" pageId="12" pageNumber="243">Liocanthydrus nanops</emphasis>
|
||
</taxonomicName>
|
||
is externally very similar to
|
||
<taxonomicName id="4C7067B7FFBEFFFD761DF958FCAA91F1" box="[802,975,1735,1758]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBEFFFD761DF958FCAA91F1" box="[802,975,1735,1758]" italics="true" pageId="12" pageNumber="243">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
and also, though less so, to
|
||
<taxonomicName id="4C7067B7FFBEFFFD7015F958FAF091F0" box="[1322,1429,1734,1759]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD7015F958FAF091F0" box="[1322,1429,1734,1759]" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
. Externally
|
||
<taxonomicName id="4C7067B7FFBEFFFD7429F973FEED902C" box="[278,392,1772,1795]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD7429F973FEED902C" box="[278,392,1772,1795]" italics="true" pageId="12" pageNumber="243">L. nanops</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C7067B7FFBEFFFD77AEF973FD9E902C" box="[657,763,1770,1795]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD77AEF973FD9E902C" box="[657,763,1770,1795]" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
by the smaller relative eye size and the patterning of the posterior elytral maculae, which are more obliquely oriented.
|
||
<taxonomicName id="4C7067B7FFBEFFFD766CF891FB2B9009" box="[851,1102,1806,1831]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD766CF891FB2B9009" box="[851,1102,1806,1831]" italics="true" pageId="12" pageNumber="243">Liocanthydrus nanops</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from both
|
||
<taxonomicName id="4C7067B7FFBEFFFD70BAF890FF859064" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD70BAF890FF859064" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7067B7FFBEFFFD7429F8ABFEDA9064" box="[278,447,1844,1867]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBEFFFD7429F8ABFEDA9064" box="[278,447,1844,1867]" italics="true" pageId="12" pageNumber="243">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
by its distinct aedeagus, with a median lobe that is sinuate in dorsal aspect, similar to
|
||
<taxonomicName id="4C7067B7FFBEFFFD70BAF8ABFE479041" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBEFFFD70BAF8ABFE479041" italics="true" pageId="12" pageNumber="243">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
but not
|
||
<taxonomicName id="4C7067B7FFBEFFFD74BFF8C8FE829040" box="[384,487,1878,1903]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD74BFF8C8FE829040" box="[384,487,1878,1903]" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
, and is also attenuated distally to the apex in lateral aspect, similar to
|
||
<emphasis id="B904C026FFBEFFFD703DF8C8FA159040" box="[1282,1392,1878,1903]" italics="true" pageId="12" pageNumber="243">
|
||
<taxonomicName id="4C7067B7FFBEFFFD703DF8C8FA099040" box="[1282,1388,1878,1903]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">L. clayae</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but not very elongate and subparallel like that of
|
||
<emphasis id="B904C026FFBEFFFD77A7F8E3FC2C90BC" box="[664,841,1916,1939]" italics="true" pageId="12" pageNumber="243">
|
||
<taxonomicName id="4C7067B7FFBEFFFD77A7F8E3FC2090BC" box="[664,837,1916,1939]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">L. octoguttatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The aedeagus of
|
||
<taxonomicName id="4C7067B7FFBEFFFD712EF8E3FBE590BC" box="[1041,1152,1916,1939]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBEFFFD712EF8E3FBE590BC" box="[1041,1152,1916,1939]" italics="true" pageId="12" pageNumber="243">L. nanops</emphasis>
|
||
</taxonomicName>
|
||
also differs from both
|
||
<taxonomicName id="4C7067B7FFBEFFFD70BAF8E3FF859098" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD70BAF8E3FF859098" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7067B7FFBEFFFD7421F800FEAF9099" box="[286,458,1951,1974]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">
|
||
<emphasis id="B904C026FFBEFFFD7421F800FEAF9099" box="[286,458,1951,1974]" italics="true" pageId="12" pageNumber="243">L. octoguttatus</emphasis>
|
||
</taxonomicName>
|
||
by its subtriangular right lateral lobe with a distinctly projected and rounded distal angle, not triangular with an acute distal angle as in
|
||
<taxonomicName id="4C7067B7FFBEFFFD77E0F85BFC2090F4" box="[735,837,1986,2011]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="clayae">
|
||
<emphasis id="B904C026FFBEFFFD77E0F85BFC2090F4" box="[735,837,1986,2011]" italics="true" pageId="12" pageNumber="243">L. clayae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B904C026FFBEFFFD7643F85BFB4E90F4" box="[892,1067,1988,2011]" italics="true" pageId="12" pageNumber="243">
|
||
<taxonomicName id="4C7067B7FFBEFFFD7643F85BFB4290F4" box="[892,1063,1988,2011]" class="Insecta" family="Noteridae" genus="Canthydrus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="243" phylum="Arthropoda" rank="species" species="octoguttatus">L. octoguttatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFC75F8FF08FC2A9587" pageId="13" pageNumber="244" type="description">
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FF08FAF897D7" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FF08FE34979F" bold="true" box="[199,337,151,176]" pageId="13" pageNumber="244">Description</emphasis>
|
||
. Measurements. TL =
|
||
<quantity id="4C88B1D1FFBFFFFC776DFF08FD8C9780" box="[594,745,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.025" metricValueMax="3.35" metricValueMin="2.7" pageId="13" pageNumber="244" unit="mm" value="3.025" valueMax="3.35" valueMin="2.7">2.7–3.35 mm</quantity>
|
||
, male max. =
|
||
<quantity id="4C88B1D1FFBFFFFC76B7FF08FC8B9780" box="[904,1006,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="13" pageNumber="244" unit="mm" value="3.05">3.05 mm</quantity>
|
||
, female max. =
|
||
<quantity id="4C88B1D1FFBFFFFC719CFF08FA6C9780" box="[1187,1289,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="13" pageNumber="244" unit="mm" value="3.35">3.35 mm</quantity>
|
||
, male min. =
|
||
<quantity id="4C88B1D1FFBFFFFC75A8FF23FF9597FB" box="[151,240,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="13" pageNumber="244" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, female min. =
|
||
<quantity id="4C88B1D1FFBFFFFC749AFF23FD6997FB" box="[421,524,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="13" pageNumber="244" unit="mm" value="2.95">2.95 mm</quantity>
|
||
, GW =
|
||
<quantity id="4C88B1D1FFBFFFFC7756FF23FD9697FB" box="[617,755,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="13" pageNumber="244" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
|
||
, TL/GW = 2.03–2.15, HW = 0.8–1.0 mm, EW =
|
||
<quantity id="4C88B1D1FFBFFFFC7013FF23FFA097D8" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="6.5" metricValueMin="5.5" pageId="13" pageNumber="244" unit="mm" value="0.6" valueMax="0.65" valueMin="0.55">0.55–0.65 mm</quantity>
|
||
, HW/EW = 1.47–1.52. PNWpost. =
|
||
<quantity id="4C88B1D1FFBFFFFC7760FF40FD9397D8" box="[607,758,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" pageId="13" pageNumber="244" unit="mm" value="1.325" valueMax="1.4" valueMin="1.25">1.25–1.4 mm</quantity>
|
||
, PNWant. =
|
||
<quantity id="4C88B1D1FFBFFFFC76BBFF40FB7E97D8" box="[900,1051,223,248]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="13" pageNumber="244" unit="mm" value="0.875" valueMax="0.95" valueMin="0.8">0.8–0.95 mm</quantity>
|
||
, PNWpost./PNWant. = 1.53–1.62.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FE9BFC8C96A7" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FE9BFE279632" box="[199,322,260,285]" italics="true" pageId="13" pageNumber="244">Coloration</emphasis>
|
||
. Elytra maculate, with transverse bands; color of elytra brown; color of head pronotum and elytral maculae yellow; band pattern as in
|
||
<figureCitation id="134B00B1FFBFFFFC7704FEB8FDE7966F" box="[571,642,295,320]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Fig. 8</figureCitation>
|
||
, discontinuous across elytral suture, posterior elytral bands strongly oblique and extending anterolaterally onto basal half of elytra; anteromedial region of pronotal disc usually with dark maculation (
|
||
<figureCitation id="134B00B1FFBFFFFC7464FEF0FEFA96A7" box="[347,415,367,392]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Fig. 8</figureCitation>
|
||
); color of venter yellowish brown to reddish brown.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FE0BFE2896FF" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FE0BFF9A9682" box="[199,255,404,429]" italics="true" pageId="13" pageNumber="244">Body</emphasis>
|
||
. Body shape as in
|
||
<figureCitation id="134B00B1FFBFFFFC74E0FE0BFD439683" box="[479,550,404,429]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Fig. 8</figureCitation>
|
||
, oval, moderately narrow, posteriorly attenuate; moderately convex in lateral aspect (
|
||
<figureCitation id="134B00B1FFBFFFFC75D3FE28FE5A96FF" box="[236,319,439,464]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="13" pageNumber="244">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FE43FA2C96DA" blockId="13.[151,1437,151,1293]" box="[199,1353,476,501]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FE43FE6196DA" box="[199,260,476,501]" italics="true" pageId="13" pageNumber="244">Head</emphasis>
|
||
. Eye size small relative to head capsule, EW =
|
||
<quantity id="4C88B1D1FFBFFFFC7626FE43FCDB96DB" box="[793,958,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="6" metricValueMax="6.5" metricValueMin="5.5" pageId="13" pageNumber="244" unit="mm" value="0.6" valueMax="0.65" valueMin="0.55">0.55–0.65 mm</quantity>
|
||
, HW/EW = 1.47–1.52 (
|
||
<figureCitation id="134B00B1FFBFFFFC71F7FE43FA7596DB" box="[1224,1296,476,501]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="134B00B1FFBFFFFC7022FE43FA5996DA" box="[1309,1340,476,501]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="13" pageNumber="244">15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FE60FBD79537" blockId="13.[151,1437,151,1293]" box="[199,1202,511,536]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FE60FE459537" box="[199,288,511,536]" italics="true" pageId="13" pageNumber="244">Thorax.</emphasis>
|
||
Elytra with lateral margins strongly ascendant medially in lateral aspect (
|
||
<figureCitation id="134B00B1FFBFFFFC7167FE60FBCD9537" box="[1112,1192,511,536]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="13" pageNumber="244">Fig. 15</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FDBBFC2A9587" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FDBBFE549512" box="[199,305,548,573]" italics="true" pageId="13" pageNumber="244">Abdomen</emphasis>
|
||
. Aedeagus as in
|
||
<figureCitation id="134B00B1FFBFFFFC74C5FDBBFD359513" box="[506,592,548,573]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="13" pageNumber="244">Figs 21</figureCitation>
|
||
a–e; median lobe sinuate in dorsal aspect (
|
||
<figureCitation id="134B00B1FFBFFFFC7101FDBBFBF79512" box="[1086,1170,548,573]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="13" pageNumber="244">Fig. 21</figureCitation>
|
||
b), attenuate to apex in lateral aspect (
|
||
<figureCitation id="134B00B1FFBFFFFC747EFDD8FEFE954F" box="[321,411,583,608]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="13" pageNumber="244">Figs 21</figureCitation>
|
||
a, c); left lateral lobe broad medially; setal fringe extending nearly to median of dorsal margin, apically expanded to small field (
|
||
<figureCitation id="134B00B1FFBFFFFC7755FDF3FDD895AA" box="[618,701,620,645]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="13" pageNumber="244">Fig. 21</figureCitation>
|
||
d); right lateral lobe subtriangular, distal angle strongly projected, dorsal margin broadly emarginated along its length (
|
||
<figureCitation id="134B00B1FFBFFFFC77DEFD10FC519587" box="[737,820,655,680]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="13" pageNumber="244">Fig. 21</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFC75F8FD2BFCC5943A" pageId="13" pageNumber="244" type="etymology">
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FD2BFCC5943A" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FD2BFE2B95E2" bold="true" box="[199,334,692,717]" pageId="13" pageNumber="244">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C7067B7FFBFFFFC7704FD2AFDE995E3" box="[571,652,693,716]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="nanops">
|
||
<emphasis id="B904C026FFBFFFFC7704FD2AFDE995E3" box="[571,652,693,716]" italics="true" pageId="13" pageNumber="244">nanops</emphasis>
|
||
</taxonomicName>
|
||
is a combination of the Greek words
|
||
<emphasis id="B904C026FFBFFFFC7109FD2AFB1C95E3" box="[1078,1145,693,716]" italics="true" pageId="13" pageNumber="244">nanos</emphasis>
|
||
, meaning small or dwarf, and
|
||
<emphasis id="B904C026FFBFFFFC75F8FD46FF8B95DF" box="[199,238,729,752]" italics="true" pageId="13" pageNumber="244">ops</emphasis>
|
||
, meaning eye. It is named in reference to the significantly smaller eyes bore by members of the species. It is treated as a noun in the nominative singular standing in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFC75F8FC80FDFC9472" pageId="13" pageNumber="244" type="biology_ecology">
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FC80FDFC9472" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FC80FE429417" bold="true" box="[199,295,799,824]" pageId="13" pageNumber="244">Biology.</emphasis>
|
||
While there are no collection data indicating the habitat of this species, it is very likely that it is found in similar habitats to the other known species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFC75F8FCF8FBB094AF" box="[199,1237,871,896]" pageId="13" pageNumber="244" type="distribution">
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FCF8FBB094AF" blockId="13.[151,1437,151,1293]" box="[199,1237,871,896]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FCF8FE0594AF" bold="true" box="[199,352,871,896]" pageId="13" pageNumber="244">Distribution.</emphasis>
|
||
Known from
|
||
<collectingCountry id="F3675CA4FFBFFFFC74C1FCF7FD0394AF" box="[510,614,872,896]" name="Paraguay" pageId="13" pageNumber="244">Paraguay</collectingCountry>
|
||
and the Guarapuava region of southern
|
||
<collectingCountry id="F3675CA4FFBFFFFC7117FCF8FB0994AF" box="[1064,1132,871,896]" name="Brazil" pageId="13" pageNumber="244">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="134B00B1FFBFFFFC7143FCF8FBA894AF" box="[1148,1229,871,896]" captionStart="FIGURE 23" captionStartId="7.[151,250,1491,1513]" captionText="FIGURE 23. Distribution map of Liocanthydrus species." pageId="13" pageNumber="244">Fig. 23</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBFFFFC75F8FC13FA5C9222" pageId="13" pageNumber="244" type="description">
|
||
<paragraph id="8BCF1C34FFBFFFFC75F8FC13FA5C9222" blockId="13.[151,1437,151,1293]" pageId="13" pageNumber="244">
|
||
<emphasis id="B904C026FFBFFFFC75F8FC13FE5E948A" bold="true" box="[199,315,908,933]" pageId="13" pageNumber="244">Variation.</emphasis>
|
||
Of the specimens examined, those from
|
||
<collectingCountry id="F3675CA4FFBFFFFC762CFC13FC32948A" box="[787,855,908,933]" name="Brazil" pageId="13" pageNumber="244">Brazil</collectingCountry>
|
||
were somewhat smaller than those from
|
||
<collectingCountry id="F3675CA4FFBFFFFC7010FC12FAFC948A" box="[1327,1433,909,933]" name="Paraguay" pageId="13" pageNumber="244">Paraguay</collectingCountry>
|
||
. The specimens were still observed to have a nearly identical patterning, body outline, small eyes relative to the head capsule with a HW/EW ratio that fell within the range of those from
|
||
<collectingCountry id="F3675CA4FFBFFFFC76DFFC4AFB2F94C2" box="[992,1098,981,1005]" name="Paraguay" pageId="13" pageNumber="244">Paraguay</collectingCountry>
|
||
. Examination of the aedeagus showed that the same diagnostic characters are also present, such as the median and left lateral lobes’ shape and the projecting distal angle of the right lateral lobe. Some very minor variation was seen in the shape of the distal portion of the median lobe, and overall the aedeagus was slightly smaller than those of the
|
||
<collectingCountry id="F3675CA4FFBFFFFC7190FBDFFA729377" box="[1199,1303,1088,1112]" name="Paraguay" pageId="13" pageNumber="244">Paraguay</collectingCountry>
|
||
specimens. Due to the limited amount of material from
|
||
<collectingCountry id="F3675CA4FFBFFFFC77FBFBFBFC6E9352" box="[708,779,1124,1149]" name="Brazil" pageId="13" pageNumber="244">Brazil</collectingCountry>
|
||
, geographic proximity of the two series, and lack of differentiating characters these differences were considered a part of normal variation within
|
||
<emphasis id="B904C026FFBFFFFC7194FB16FA44938F" box="[1195,1313,1161,1184]" italics="true" pageId="13" pageNumber="244">
|
||
<taxonomicName id="4C7067B7FFBFFFFC7194FB16FA79938F" box="[1195,1308,1161,1184]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="244" phylum="Arthropoda" rank="species" species="nanops">L. nanops</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Very little variation was observed otherwise. Individuals varied slightly in overall color some individuals slightly lighter or darker than others. Maculae generally appeared as in
|
||
<figureCitation id="134B00B1FFBFFFFC764CFB50FCDD93C7" box="[883,952,1231,1256]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="13" pageNumber="244">Fig. 8</figureCitation>
|
||
, though reduced in some individuals. The respective orientation of the bands/spots remained consistent within all observed members of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |