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<document id="F7B40FA1260F3F2188BFC4A574B79A81" ID-DOI="10.5852/ejt.2024.921.2413" ID-ISSN="2118-9773" ID-Zenodo-Dep="10590243" ID-ZooBank="80B3C408-DD2C-4373-96AE-15F067D723F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1706610496547" checkinUser="plazi" docAuthor="Bockwinkel, Jürgen &amp; Korn, Dieter" docDate="2024" docId="03D987F22933F659FDD420267472FA9D" docLanguage="en" docName="EJT.2024.921.1-35.pdf" docOrigin="European Journal of Taxonomy 921 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2413/10641" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Trimaenioceras fuscina Bockwinkel &amp; Korn 2024, gen. et sp. nov." docType="treatment" docUuid="C5B0E733-6BD9-48EB-A89A-009451C83B6F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="31" masterDocId="FFE0FF8A292EF647FFFB212F7550FF8C" masterDocTitle="Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco" masterLastPageNumber="35" masterPageNumber="1" pageNumber="30" updateTime="1706631334240" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D610A36691880AC210A1516B68C6F021">Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco</mods:title>
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<mods:namePart id="58DCC6328A5955661BAEEF810FC7443C">Bockwinkel, Jürgen</mods:namePart>
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<mods:affiliation id="9C4F5FEEEA32736EF23A9F08E36C1CE1">Dechant-Fein-Strasse 22, 51375 Leverkusen, Germany.</mods:affiliation>
<mods:nameIdentifier id="6E976B241D90E97E56F80CFF8152B3E7" type="email">j.bockwinkel@t-online.de</mods:nameIdentifier>
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<mods:namePart id="AAAE193CE670B72D6E20DBE878C47726">Korn, Dieter</mods:namePart>
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<mods:affiliation id="2A068436D8288E737A56A5D0BF656435">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="38BBFF7A184E93F8B9DF4E77CA2F2A85" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:title id="58EC83DE331F227EEA0314ADB7CDAE64">European Journal of Taxonomy</mods:title>
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<mods:date id="9D48BAF0C734518A2E7399F59C750B5F">2024</mods:date>
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<mods:identifier id="440EEBEB9FF9297ACB5B0FBCEDE92D96" type="DOI">10.5852/ejt.2024.921.2413</mods:identifier>
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<treatment id="03D987F22933F659FDD420267472FA9D" LSID="urn:lsid:zoobank.org:act:C5B0E733-6BD9-48EB-A89A-009451C83B6F" httpUri="http://treatment.plazi.org/id/03D987F22933F659FDD420267472FA9D" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="C36A656F2933F65AFDD420267155FEAF" box="[559,1029,264,291]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BCF36E42933F65AFDD420267155FEAF" blockId="29.[364,1223,264,363]" box="[559,1029,264,291]" pageId="29" pageNumber="30">
<heading id="D08781882933F65AFDD420267155FEAF" box="[559,1029,264,291]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="4C704D672933F65AFDD42026761EFEAE" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[559,846,264,291]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" order="Agoniatitida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fuscina" status="gen. et sp. nov.">
<emphasis id="B904EAF62933F65AFDD42026761EFEAE" bold="true" box="[559,846,264,291]" italics="true" pageId="29" pageNumber="30">Trimaenioceras fuscina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2933F65AFCAE20267155FEAF" box="[853,1029,265,291]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2933F65AFE972003762DFEE7" pageId="29" pageNumber="30" type="description">
<paragraph id="8BCF36E42933F65AFE9720037197FECB" blockId="29.[364,1223,264,363]" box="[364,1223,300,327]" pageId="29" pageNumber="30">
<uri id="FFE13AE62933F65AFE9720037197FECB" box="[364,1223,300,327]" pageId="29" pageNumber="30">
urn:lsid:zoobank.org:act:
<uuid id="FFD60C312933F65AFD7620037197FECB" box="[653,1223,300,327]" pageId="29" pageNumber="30">C5B0E733-6BD9-48EB-A89A-009451C83B6F</uuid>
</uri>
</paragraph>
<paragraph id="8BCF36E42933F65AFD4D207F762DFEE7" blockId="29.[364,1223,264,363]" box="[694,893,336,363]" pageId="29" pageNumber="30">
<figureCitation id="134B2A612933F65AFD4D207F765CFEE7" box="[694,780,336,363]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[204,1381,1230,1876]" captionTargetId="figure-165@29.[195,1391,1228,1885]" captionTargetPageId="29" captionText="Fig. 15. Trimaenioceras fuscina gen. et sp. nov. from Jebel Ouaoufilal. A. Paratype MB.C.31970.2. B. Holotype MB.C.31970.1. C. Paratype MB.C.31970.2; suture line at wh = 9.5 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590277" httpUri="https://zenodo.org/record/10590277/files/figure.png" pageId="29" pageNumber="30">Fig. 15</figureCitation>
;
<tableCitation id="C6F2035F2933F65AFCE3207F762DFEE7" box="[792,893,336,363]" captionStart="Table 12" captionStartId="30.[189,259,265,291]" captionTargetPageId="30" captionText="Table 12. Conch dimensions (in mm) and ratios of Trimaenioceras fuscina gen. et sp. nov." httpUri="http://table.plazi.org/id/DF0F666C2930F659FF46202671B4FEA8" pageId="29" pageNumber="30" tableUuid="DF0F666C2930F659FF46202671B4FEA8">Table 12</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2933F65AFF4620BD71A7FD02" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BCF36E42933F65AFF4620BD7466FE20" blockId="29.[189,310,402,428]" box="[189,310,402,428]" pageId="29" pageNumber="30">
<heading id="D08781882933F65AFF4620BD7466FE20" bold="true" box="[189,310,402,428]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B904EAF62933F65AFF4620BD7466FE20" bold="true" box="[189,310,402,428]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42933F65AFF4620EE71A7FD02" blockId="29.[188,1398,449,661]" pageId="29" pageNumber="30">
Species of
<taxonomicName id="4C704D672933F65AFEB020ED775DFE57" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[331,525,450,475]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B904EAF62933F65AFEB020ED775DFE57" box="[331,525,450,475]" italics="true" pageId="29" pageNumber="30">Trimaenioceras</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2933F65AFDE320ED77DBFE50" box="[536,651,450,476]" pageId="29" pageNumber="30" rank="genus">gen. nov.</taxonomicNameLabel>
with thinly discoidal and subinvolute conch at
<quantity id="4C889B012933F65AFB1620ED7015FE50" box="[1261,1349,450,476]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="20.0">20 mm</quantity>
dm (ww/dm ~ 0.40; uw/dm ~ 0.15) with weakly compressed whorl profile (ww/wh ~ 0.85) and low coiling rate (WER ~ 1.70). Whorl profile at
<quantity id="4C889B012933F65AFD3423267671FDAF" box="[719,801,521,547]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="20.0">20 mm</quantity>
conch diameter horseshoe-shaped with flattened subparallel flanks, rounded ventrolateral shoulder and rounded venter. Ventrolateral shoulder without spiral groove. Growth lines very fine. Suture line with wide external lobe, moderately deep, cuneiform E
<subScript id="17F434A12933F65AFE5A23A874FAFD19" attach="left" box="[417,426,647,661]" fontSize="6" pageId="29" pageNumber="30">2</subScript>
lobe, rounded ventrolateral saddle and V-shaped, acute lateral lobe.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2933F65AFF46239971F2FD73" box="[189,1186,694,768]" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BCF36E42933F65AFF4623997414FD5C" blockId="29.[189,324,694,720]" box="[189,324,694,720]" pageId="29" pageNumber="30">
<heading id="D08781882933F65AFF4623997414FD5C" bold="true" box="[189,324,694,720]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B904EAF62933F65AFF4623997414FD5C" bold="true" box="[189,324,694,720]" pageId="29" pageNumber="30">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42933F65AFF4623CA71F2FD73" blockId="29.[189,1186,741,768]" box="[189,1186,741,768]" pageId="29" pageNumber="30">
After the Latin nomen
<taxonomicName id="4C704D672933F65AFE3123CA7779FD73" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[458,553,741,767]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fuscina">
<emphasis id="B904EAF62933F65AFE2F23CA7779FD73" box="[468,553,741,767]" italics="true" pageId="29" pageNumber="30">fuscina</emphasis>
</taxonomicName>
= trident, because of the shape of the external lobe.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2933F65AFF4622087006FB92" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BCF36E42933F65AFF462208743CFCCD" blockId="29.[189,364,807,833]" box="[189,364,807,833]" pageId="29" pageNumber="30">
<heading id="D08781882933F65AFF462208743CFCCD" bold="true" box="[189,364,807,833]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B904EAF62933F65AFF462208743CFCCD" bold="true" box="[189,364,807,833]" pageId="29" pageNumber="30">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B183CB92933F65AFF1122797165FC34" collectionCode="MB" collectorName="Bockwinkel &amp; Ebbighausen" country="Morocco" county="Ebbighausen" location="Anti-Atlas" municipality="Bockwinkel" pageId="29" pageNumber="30" specimenCode="MB.C.31970.1" specimenCount="1" typeStatus="holotype">
<paragraph id="8BCF36E42933F65AFF112279740CFCFC" blockId="29.[189,1399,854,952]" box="[234,348,854,880]" pageId="29" pageNumber="30">
<emphasis id="B904EAF62933F65AFF112279740CFCFC" bold="true" box="[234,348,854,880]" pageId="29" pageNumber="30">
<typeStatus id="54CB88462933F65AFF112279740CFCFC" box="[234,348,854,880]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF36E42933F65AFF4622557165FC34" blockId="29.[189,1399,854,952]" pageId="29" pageNumber="30">
<collectingCountry id="F36776742933F65AFF4622557401FC18" box="[189,337,890,916]" name="Morocco" pageId="29" pageNumber="30">MOROCCO</collectingCountry>
<location id="8EAF603F2933F65AFE93225574B7FC18" LSID="urn:lsid:plazi:treatment:03D987F22933F659FDD420267472FA9D:8EAF603F2933F65AFE93225574B7FC18" box="[360,487,890,916]" country="Morocco" county="Ebbighausen" municipality="Bockwinkel" name="Anti-Atlas" pageId="29" pageNumber="30">Anti-Atlas</location>
, south-eastern
<location id="8EAF603F2933F65AFD6F225577BAFC18" LSID="urn:lsid:plazi:treatment:03D987F22933F659FDD420267472FA9D:8EAF603F2933F65AFD6F225577BAFC18" box="[660,746,890,916]" country="Morocco" county="Ebbighausen" municipality="Bockwinkel" name="Tafilalt" pageId="29" pageNumber="30">Tafilalt</location>
,
<location id="8EAF603F2933F65AFD0E225576CAFC18" LSID="urn:lsid:plazi:treatment:03D987F22933F659FDD420267472FA9D:8EAF603F2933F65AFD0E225576CAFC18" box="[757,922,890,916]" country="Morocco" county="Ebbighausen" municipality="Bockwinkel" name="Hassi Nebech" pageId="29" pageNumber="30">Hassi Nebech</location>
, section 2; middle
<location id="8EAF603F2933F65AFB8E2255718EFC18" LSID="urn:lsid:plazi:treatment:03D987F22933F659FDD420267472FA9D:8EAF603F2933F65AFB8E2255718EFC18" box="[1141,1246,890,916]" country="Morocco" county="Ebbighausen" municipality="Bockwinkel" name="Givetian" pageId="29" pageNumber="30">Givetian</location>
;
<collectorName id="268553322933F65AFB1222557027FC18" box="[1257,1399,890,916]" pageId="29" pageNumber="30">Bockwinkel</collectorName>
and
<collectorName id="268553322933F65AFF1422B174D7FC34" box="[239,391,926,952]" pageId="29" pageNumber="30">Ebbighausen</collectorName>
20022004 Coll.;
<specimenCode id="DBD69E9F2933F65AFD9922B17642FC34" box="[610,786,926,952]" pageId="29" pageNumber="30">MB.C.31970.1</specimenCode>
(illustrated in
<figureCitation id="134B2A612933F65AFC4622B17178FC34" box="[957,1064,926,952]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[195,1397,477,1823]" captionTargetId="figure-38@27.[189,1398,464,1832]" captionTargetPageId="27" captionText="Fig. 14. Trimaenioceras eculeus gen. et sp. nov., from Hassi Nebech. A. Holotype MB.C.31969.1. B. Paratype MB.C.31969.2. C. Paratype MB.C.31969.3. D. Holotype MB.C.31969.1; suture line at wh = 13.5 mm. E. Paratype MB.C.31969.4; suture line at wh = 9.5 mm. F. Paratype JB HN 1; cross section. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590272" httpUri="https://zenodo.org/record/10590272/files/figure.png" pageId="29" pageNumber="30">Fig. 14B</figureCitation>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B183CB92933F65AFF1122CF7002FB92" collectionCode="MB" collectorName="Bockwinkel &amp; Ebbighausen" country="Morocco" county="Ebbighausen" location="MOROCCO" municipality="Bockwinkel" pageId="29" pageNumber="30" specimenCode="MB.C.31970.2" specimenCount="1" typeStatus="paratype">
<paragraph id="8BCF36E42933F65AFF1122CF740CFC76" blockId="29.[189,1366,992,1054]" box="[234,348,992,1018]" pageId="29" pageNumber="30">
<emphasis id="B904EAF62933F65AFF1122CF740CFC76" bold="true" box="[234,348,992,1018]" pageId="29" pageNumber="30">
<typeStatus id="54CB88462933F65AFF1122CF740CFC76" box="[234,348,992,1018]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF36E42933F65AFF46252B7006FB92" blockId="29.[189,1366,992,1054]" box="[189,1366,1027,1054]" pageId="29" pageNumber="30">
<collectingCountry id="F36776742933F65AFF46252B7401FB92" box="[189,337,1028,1054]" name="Morocco" pageId="29" pageNumber="30">MOROCCO</collectingCountry>
<specimenCount id="9D76FD6D2933F65AFE91252B74EFFB92" box="[362,447,1028,1054]" count="1" pageId="29" pageNumber="30" type="generic">1 spec.</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBD69E9F2933F65AFC7A252B7161FB92" box="[897,1073,1028,1054]" pageId="29" pageNumber="30">MB.C.31970.2</specimenCode>
(illustrated in
<figureCitation id="134B2A612933F65AFB26252C7019FB92" box="[1245,1353,1027,1054]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[195,1397,477,1823]" captionTargetId="figure-38@27.[189,1398,464,1832]" captionTargetPageId="27" captionText="Fig. 14. Trimaenioceras eculeus gen. et sp. nov., from Hassi Nebech. A. Holotype MB.C.31969.1. B. Paratype MB.C.31969.2. C. Paratype MB.C.31969.3. D. Holotype MB.C.31969.1; suture line at wh = 13.5 mm. E. Paratype MB.C.31969.4; suture line at wh = 9.5 mm. F. Paratype JB HN 1; cross section. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590272" httpUri="https://zenodo.org/record/10590272/files/figure.png" pageId="29" pageNumber="30">Fig. 14A</figureCitation>
).
</paragraph>
</materialsCitation>
</subSubSection>
<caption id="DF0F666C2933F65AFF4626A77459F865" ID-DOI="http://doi.org/10.5281/zenodo.10590277" ID-Zenodo-Dep="10590277" httpUri="https://zenodo.org/record/10590277/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1928,1954]" targetBox="[204,1381,1230,1876]" targetPageId="29">
<paragraph id="8BCF36E42933F65AFF4626A77459F865" blockId="29.[189,1399,1928,2025]" pageId="29" pageNumber="30">
<emphasis id="B904EAF62933F65AFF4626A7744EF82E" bold="true" box="[189,286,1928,1954]" pageId="29" pageNumber="30">Fig. 15.</emphasis>
<taxonomicName id="4C704D672933F65AFED326A67710F82E" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[296,576,1928,1954]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fuscina" status="gen. et sp. nov.">
<emphasis id="B904EAF62933F65AFED326A67710F82E" box="[296,576,1928,1954]" italics="true" pageId="29" pageNumber="30">Trimaenioceras fuscina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2933F65AFDB126A77655F82E" box="[586,773,1928,1954]" pageId="29" pageNumber="30" rank="species">gen. et sp. nov.</taxonomicNameLabel>
from Jebel Ouaoufilal.
<emphasis id="B904EAF62933F65AFBDC26A7716CF82E" bold="true" box="[1063,1084,1928,1954]" pageId="29" pageNumber="30">A</emphasis>
. Paratype MB.C.31970.2.
<emphasis id="B904EAF62933F65AFF4626847581F849" bold="true" box="[189,209,1963,1989]" pageId="29" pageNumber="30">B</emphasis>
. Holotype MB.C.31970.1.
<emphasis id="B904EAF62933F65AFDE826847778F849" bold="true" box="[531,552,1963,1989]" pageId="29" pageNumber="30">C</emphasis>
. Paratype MB.C.31970.2; suture line at wh = 9.5 mm. Scale bar units = 1 mm.
</paragraph>
</caption>
<caption id="DF0F666C2930F659FF46202671B4FEA8" ID-Table-UUID="DF0F666C2930F659FF46202671B4FEA8" box="[189,1252,265,292]" httpUri="http://table.plazi.org/id/DF0F666C2930F659FF46202671B4FEA8" pageId="30" pageNumber="31" startId="30.[189,259,265,291]" targetBox="[204,1371,336,451]" targetIsTable="true" targetPageId="30">
<paragraph id="8BCF36E42930F659FF46202671B4FEA8" blockId="30.[189,1252,265,292]" box="[189,1252,265,292]" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FF462026747FFEAF" bold="true" box="[189,303,265,291]" pageId="30" pageNumber="31">Table 12.</emphasis>
Conch dimensions (in mm) and ratios of
<taxonomicName id="4C704D672930F659FCE32025717DFEAF" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[792,1069,265,291]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fuscina" status="gen. et sp. nov.">
<emphasis id="B904EAF62930F659FCE32025717DFEAF" box="[792,1069,265,291]" italics="true" pageId="30" pageNumber="31">Trimaenioceras fuscina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2930F659FBCF202571B4FEA8" box="[1076,1252,266,292]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCF36E42930F659FF1C207F700BFE4F" pageId="30" pageNumber="31">
<table id="F970C444293009B8FF37207F700BFE4F" box="[204,1371,336,451]" gridcols="11" gridrows="3" pageId="30" pageNumber="31">
<tr id="354034A6293009B8FF37207F700BFEE4" box="[204,1371,336,360]" gridrow="0" pageId="30" pageNumber="31">
<th id="76915DDA293009B8FF37207F743BFEE4" box="[204,363,336,360]" gridcol="0" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FF1C207F7400FEE4" bold="true" box="[231,336,336,360]" pageId="30" pageNumber="31">specimen</emphasis>
</th>
<th id="76915DDA293009B8FE6B207F74EFFEE4" box="[400,447,336,360]" gridcol="1" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FE6F207F74E9FEE4" bold="true" box="[404,441,336,360]" pageId="30" pageNumber="31">dm</emphasis>
</th>
<th id="76915DDA293009B8FE15207F7745FEE4" box="[494,533,336,360]" gridcol="2" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FE15207F7745FEE4" bold="true" box="[494,533,336,360]" pageId="30" pageNumber="31">ww</emphasis>
</th>
<th id="76915DDA293009B8FDBE207F7724FEE4" box="[581,628,336,360]" gridcol="3" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FDB0207F773DFEE4" bold="true" box="[587,621,336,360]" pageId="30" pageNumber="31">wh</emphasis>
</th>
<th id="76915DDA293009B8FD5D207F7798FEE4" box="[678,712,336,360]" gridcol="4" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FD5D207F7798FEE4" bold="true" box="[678,712,336,360]" pageId="30" pageNumber="31">uw</emphasis>
</th>
<th id="76915DDA293009B8FCFA207F7672FEE4" box="[769,802,336,360]" gridcol="5" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FCFF207F7670FEE4" bold="true" box="[772,800,336,360]" pageId="30" pageNumber="31">ah</emphasis>
</th>
<th id="76915DDA293009B8FCA2207F76FCFEE4" box="[857,940,336,360]" gridcol="6" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FCA2207F76FCFEE4" bold="true" box="[857,940,336,360]" pageId="30" pageNumber="31">ww/dm</emphasis>
</th>
<th id="76915DDA293009B8FC2D207F7175FEE4" box="[982,1061,336,360]" gridcol="7" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FC2D207F7175FEE4" bold="true" box="[982,1061,336,360]" pageId="30" pageNumber="31">ww/wh</emphasis>
</th>
<th id="76915DDA293009B8FBBE207F71C3FEE4" box="[1093,1171,336,360]" gridcol="8" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FBBE207F71C3FEE4" bold="true" box="[1093,1171,336,360]" pageId="30" pageNumber="31">uw/dm</emphasis>
</th>
<th id="76915DDA293009B8FB40207F71ABFEE4" box="[1211,1275,336,360]" gridcol="9" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FB40207F71ABFEE4" bold="true" box="[1211,1275,336,360]" pageId="30" pageNumber="31">WER</emphasis>
</th>
<th id="76915DDA293009B8FAD7207F700BFEE4" box="[1324,1371,336,360]" gridcol="10" gridrow="0" pageId="30" pageNumber="31">
<emphasis id="B904EAF62930F659FAD7207F700BFEE4" bold="true" box="[1324,1371,336,360]" pageId="30" pageNumber="31">IZR</emphasis>
</th>
</tr>
<tr id="354034A6293009B8FF372054700BFE1A" box="[204,1371,379,406]" gridrow="1" pageId="30" pageNumber="31">
<th id="76915DDA293009B8FF372054743BFE1A" box="[204,363,379,406]" gridcol="0" gridrow="1" pageId="30" pageNumber="31">MB.C.31970.2</th>
<td id="76915DDA293009B8FE6B205474EFFE1A" box="[400,447,379,406]" gridcol="1" gridrow="1" pageId="30" pageNumber="31">21.3</td>
<td id="76915DDA293009B8FE1520547745FE1A" box="[494,533,379,406]" gridcol="2" gridrow="1" pageId="30" pageNumber="31">8.5</td>
<td id="76915DDA293009B8FDBE20547724FE1A" box="[581,628,379,406]" gridcol="3" gridrow="1" pageId="30" pageNumber="31">10.2</td>
<td id="76915DDA293009B8FD5D20547798FE1A" box="[678,712,379,406]" gridcol="4" gridrow="1" pageId="30" pageNumber="31">2.9</td>
<td id="76915DDA293009B8FCFA20547672FE1A" box="[769,802,379,406]" gridcol="5" gridrow="1" pageId="30" pageNumber="31">5.1</td>
<td id="76915DDA293009B8FCA2205476FCFE1A" box="[857,940,379,406]" gridcol="6" gridrow="1" pageId="30" pageNumber="31">0.40</td>
<td id="76915DDA293009B8FC2D20547175FE1A" box="[982,1061,379,406]" gridcol="7" gridrow="1" pageId="30" pageNumber="31">0.83</td>
<td id="76915DDA293009B8FBBE205471C3FE1A" box="[1093,1171,379,406]" gridcol="8" gridrow="1" pageId="30" pageNumber="31">0.14</td>
<td id="76915DDA293009B8FB40205471ABFE1A" box="[1211,1275,379,406]" gridcol="9" gridrow="1" pageId="30" pageNumber="31">1.73</td>
<td id="76915DDA293009B8FAD72054700BFE1A" box="[1324,1371,379,406]" gridcol="10" gridrow="1" pageId="30" pageNumber="31">0.50</td>
</tr>
<tr id="354034A6293009B8FF372084700BFE4F" box="[204,1371,427,451]" gridrow="2" pageId="30" pageNumber="31">
<th id="76915DDA293009B8FF372084743BFE4F" box="[204,363,427,451]" gridcol="0" gridrow="2" pageId="30" pageNumber="31">MB.C.31970.1</th>
<td id="76915DDA293009B8FE6B208474EFFE4F" box="[400,447,427,451]" gridcol="1" gridrow="2" pageId="30" pageNumber="31">20.8</td>
<td id="76915DDA293009B8FE1520847745FE4F" box="[494,533,427,451]" gridcol="2" gridrow="2" pageId="30" pageNumber="31">8.8</td>
<td id="76915DDA293009B8FDBE20847724FE4F" box="[581,628,427,451]" gridcol="3" gridrow="2" pageId="30" pageNumber="31">9.9</td>
<td id="76915DDA293009B8FD5D20847798FE4F" box="[678,712,427,451]" gridcol="4" gridrow="2" pageId="30" pageNumber="31">3.2</td>
<td id="76915DDA293009B8FCFA20847672FE4F" box="[769,802,427,451]" gridcol="5" gridrow="2" pageId="30" pageNumber="31">4.9</td>
<td id="76915DDA293009B8FCA2208476FCFE4F" box="[857,940,427,451]" gridcol="6" gridrow="2" pageId="30" pageNumber="31">0.42</td>
<td id="76915DDA293009B8FC2D20847175FE4F" box="[982,1061,427,451]" gridcol="7" gridrow="2" pageId="30" pageNumber="31">0.89</td>
<td id="76915DDA293009B8FBBE208471C3FE4F" box="[1093,1171,427,451]" gridcol="8" gridrow="2" pageId="30" pageNumber="31">0.15</td>
<td id="76915DDA293009B8FB40208471ABFE4F" box="[1211,1275,427,451]" gridcol="9" gridrow="2" pageId="30" pageNumber="31">1.71</td>
<td id="76915DDA293009B8FAD72084700BFE4F" box="[1324,1371,427,451]" gridcol="10" gridrow="2" pageId="30" pageNumber="31">0.51</td>
</tr>
</table>
</paragraph>
<subSubSection id="C36A656F2930F659FF46232B74F2FC62" pageId="30" pageNumber="31" type="description">
<paragraph id="8BCF36E42930F659FF46232B741EFD92" blockId="30.[189,334,516,542]" box="[189,334,516,542]" pageId="30" pageNumber="31">
<heading id="D08781882930F659FF46232B741EFD92" bold="true" box="[189,334,516,542]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B904EAF62930F659FF46232B741EFD92" bold="true" box="[189,334,516,542]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42930F659FF46231D771FFD55" blockId="30.[189,1401,562,729]" pageId="30" pageNumber="31">
<typeStatus id="54CB88462930F659FF46231D747AFDC0" box="[189,298,562,588]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
MB.C.31970.1 is a fully septate internal mould of a specimen with
<quantity id="4C889B012930F659FB99231D71E7FDC1" box="[1122,1207,562,589]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="30" pageNumber="31" unit="mm" value="21.0">21 mm</quantity>
conch diameter (
<figureCitation id="134B2A612930F659FF33237A7463FDFC" box="[200,307,597,624]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[204,1381,1230,1876]" captionTargetId="figure-165@29.[195,1391,1228,1885]" captionTargetPageId="29" captionText="Fig. 15. Trimaenioceras fuscina gen. et sp. nov. from Jebel Ouaoufilal. A. Paratype MB.C.31970.2. B. Holotype MB.C.31970.1. C. Paratype MB.C.31970.2; suture line at wh = 9.5 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590277" httpUri="https://zenodo.org/record/10590277/files/figure.png" pageId="30" pageNumber="31">Fig. 15B</figureCitation>
). The conch is thinly discoidal with a rather small umbilicus (ww/dm = 0.42; uw/dm = 0.15); a weakly compressed whorl profile (ww/wh = 0.89) and a low coiling rate (WER = 1.71). The specimen shows that the umbilicus is being narrowed, caused by increased overlap of the flanks upon the umbilicus, during the last preserved volution.
</paragraph>
<paragraph id="8BCF36E42930F659FF46222E74F2FC62" blockId="30.[188,1400,768,1006]" pageId="30" pageNumber="31">
<typeStatus id="54CB88462930F659FF46222E7474FC97" box="[189,292,769,795]" pageId="30" pageNumber="31" type="paratype">Paratype</typeStatus>
MB.C.31970.2 is a specimen with
<quantity id="4C889B012930F659FD2D222F7612FC97" box="[726,834,768,795]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" pageId="30" pageNumber="31" unit="mm" value="21.5">21.5 mm</quantity>
conch diameter (
<figureCitation id="134B2A612930F659FBEC222E71D7FC97" box="[1047,1159,768,795]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[204,1381,1230,1876]" captionTargetId="figure-165@29.[195,1391,1228,1885]" captionTargetPageId="29" captionText="Fig. 15. Trimaenioceras fuscina gen. et sp. nov. from Jebel Ouaoufilal. A. Paratype MB.C.31970.2. B. Holotype MB.C.31970.1. C. Paratype MB.C.31970.2; suture line at wh = 9.5 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590277" httpUri="https://zenodo.org/record/10590277/files/figure.png" pageId="30" pageNumber="31">Fig. 15A</figureCitation>
), which in size and proportions of the conch closely resembles the
<typeStatus id="54CB88462930F659FD1B220B7617FCB2" box="[736,839,804,830]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(ww/dm = 0.40; uw/dm = 0.14). The suture line shows a very wide external lobe that is subdivided into three almost equally wide and deep prongs, of which the E
<subScript id="17F434A12930F659FEB722527405FC07" attach="left" box="[332,341,893,907]" fontSize="6" pageId="30" pageNumber="31">2</subScript>
lobes are asymmetric and cuneiform. The median saddle reaches about two thirds of the depth of the external lobe. The ventrolateral saddle is nearly symmetric and inclined towards the venter; the lateral lobe is V-shaped with gently convex flanks (
<figureCitation id="134B2A612930F659FCB4229E76EDFC47" box="[847,957,944,971]" captionStart="Fig" captionStartId="29.[189,232,1928,1954]" captionTargetBox="[204,1381,1230,1876]" captionTargetId="figure-165@29.[195,1391,1228,1885]" captionTargetPageId="29" captionText="Fig. 15. Trimaenioceras fuscina gen. et sp. nov. from Jebel Ouaoufilal. A. Paratype MB.C.31970.2. B. Holotype MB.C.31970.1. C. Paratype MB.C.31970.2; suture line at wh = 9.5 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590277" httpUri="https://zenodo.org/record/10590277/files/figure.png" pageId="30" pageNumber="31">Fig. 15C</figureCitation>
). The last phragmocone volution has about 16 chambers.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2930F659FF4625397472FA9D" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BCF36E42930F659FF462539747FFBBC" blockId="30.[189,303,1046,1072]" box="[189,303,1046,1072]" pageId="30" pageNumber="31">
<heading id="D08781882930F659FF462539747FFBBC" bold="true" box="[189,303,1046,1072]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B904EAF62930F659FF462539747FFBBC" bold="true" box="[189,303,1046,1072]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42930F659FF4625697472FA9D" blockId="30.[189,1401,1093,1298]" pageId="30" pageNumber="31">
<taxonomicName id="4C704D672930F659FF4625697485FBD3" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[189,469,1093,1119]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fuscina" status="gen. et sp. nov.">
<emphasis id="B904EAF62930F659FF4625697485FBD3" box="[189,469,1093,1119]" italics="true" pageId="30" pageNumber="31">Trimaenioceras fuscina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2930F659FE24256A77CAFBD3" box="[479,666,1093,1119]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is the species of the genus with the most slender conch. At about
<quantity id="4C889B012930F659FEFD25467409FB0F" box="[262,345,1129,1155]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="mm" value="20.0">20 mm</quantity>
conch diameter, its ww/dm ratio is ~0.40, while it is~
<quantity id="4C889B012930F659FC202547717DFB0E" box="[987,1069,1128,1155]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="30" pageNumber="31" unit="in" value="0.5">0.50 in</quantity>
<taxonomicName id="4C704D672930F659FBCE254671DDFB0E" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[1077,1165,1128,1154]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="klugi" status="gen. et sp. nov.">
<emphasis id="B904EAF62930F659FBCE254671DDFB0E" box="[1077,1165,1128,1154]" italics="true" pageId="30" pageNumber="31">T. klugi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2930F659FB6F25467015FB0F" box="[1172,1349,1129,1155]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C704D672930F659FF4625A27462FB2A" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[189,306,1164,1190]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="eculeus" status="gen. et sp. nov.">
<emphasis id="B904EAF62930F659FF4625A27462FB2A" box="[189,306,1164,1190]" italics="true" pageId="30" pageNumber="31">T. eculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2930F659FEC225A274B8FB2B" box="[313,488,1165,1191]" pageId="30" pageNumber="31" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<taxonomicName id="4C704D672930F659FE1425A27654FB2A" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[495,772,1164,1190]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fuscina">
<emphasis id="B904EAF62930F659FE1425A27654FB2A" box="[495,772,1164,1190]" italics="true" pageId="30" pageNumber="31">Trimaenioceras fuscina</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C704D672930F659FC6525A276A5FB2A" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[926,1013,1164,1190]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="klugi">
<emphasis id="B904EAF62930F659FC6525A276A5FB2A" box="[926,1013,1164,1190]" italics="true" pageId="30" pageNumber="31">T. klugi</emphasis>
</taxonomicName>
in the less deep sutural elements of the external lobe. A suture line very similar to
<taxonomicName id="4C704D672930F659FCF6259E76D0FB46" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[781,896,1200,1226]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fuscina">
<emphasis id="B904EAF62930F659FCF6259E76D0FB46" box="[781,896,1200,1226]" italics="true" pageId="30" pageNumber="31">T. fuscina</emphasis>
</taxonomicName>
can be seen in
<taxonomicName id="4C704D672930F659FBC0259E71E0FB46" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[1083,1200,1200,1226]" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="eculeus">
<emphasis id="B904EAF62930F659FBC0259E71E0FB46" box="[1083,1200,1200,1226]" italics="true" pageId="30" pageNumber="31">T. eculeus</emphasis>
</taxonomicName>
, but this species differs in the wider umbilicus (uw/dm~ 0.20) at
<quantity id="4C889B012930F659FCF725FB7633FB62" box="[780,867,1236,1262]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="mm" value="20.0">20 mm</quantity>
conch diameter (uw/dm is only ~
<quantity id="4C889B012930F659FAFA25FB7006FB62" box="[1281,1366,1236,1263]" metricMagnitude="-3" metricUnit="m" metricValue="3.81" pageId="30" pageNumber="31" unit="in" value="0.15">0.15 in</quantity>
<taxonomicName id="4C704D672930F659FA9A25FA7442FA9E" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" class="Cephalopoda" family="Maenioceratidae" genus="Trimaenioceras" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fuscina">
<emphasis id="B904EAF62930F659FA9A25FA7442FA9E" italics="true" pageId="30" pageNumber="31">T. fuscina</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>