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<document id="C426D35981D3CFBA701A3265A89A5B8B" ID-DOI="10.5852/ejt.2024.921.2413" ID-ISSN="2118-9773" ID-Zenodo-Dep="10590243" ID-ZooBank="80B3C408-DD2C-4373-96AE-15F067D723F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1706610496547" checkinUser="plazi" docAuthor="Bockwinkel, Jürgen &amp; Korn, Dieter" docDate="2024" docId="03D987F22922F649FD90272B76B8FC11" docLanguage="en" docName="EJT.2024.921.1-35.pdf" docOrigin="European Journal of Taxonomy 921 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2413/10641" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maenioceras beckeri Bockwinkel &amp; Korn 2024, sp. nov." docType="treatment" docUuid="48EAD7CA-D85E-40E0-B32C-7A6C5A078A7C" docUuidSource="ZooBank" docVersion="5" lastPageNumber="15" masterDocId="FFE0FF8A292EF647FFFB212F7550FF8C" masterDocTitle="Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco" masterLastPageNumber="35" masterPageNumber="1" pageNumber="13" updateTime="1706631334240" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3075D593EB9BB0C35B7803D22BCC194F">Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco</mods:title>
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<mods:namePart id="6D5452478DDFA63ABBC25CAECE5361CC">Bockwinkel, Jürgen</mods:namePart>
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<mods:affiliation id="028C4F77B0A394D3DC0B320CF27AF4BC">Dechant-Fein-Strasse 22, 51375 Leverkusen, Germany.</mods:affiliation>
<mods:nameIdentifier id="5A5646DB1B0DD8459CF4EB5C26DF1C5A" type="email">j.bockwinkel@t-online.de</mods:nameIdentifier>
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<mods:namePart id="5209477C892CF55A7622F32D1E62B9B2">Korn, Dieter</mods:namePart>
<mods:nameIdentifier id="3DB2B2F016B052BEAF3109D1D17EC3C8" type="LSID">286CA4F3-7EBC-4AEF-A66A-B2508D001367</mods:nameIdentifier>
<mods:affiliation id="A844B880F74E41A72A7A404B5FC1C5A7">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="3B6748E5F14F8C73B97FF8A033992EB7" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:title id="1E389920E62ADCBEE3E4549C55F01B90">European Journal of Taxonomy</mods:title>
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<mods:date id="CDBBB4BED7438640CF81C07649702F07">2024</mods:date>
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<mods:number id="A6BDD4768B3485D2002B917C5BA52AF5">2024-01-30</mods:number>
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<mods:number id="3AC4582D5C14FECD2A0DBE8A1241A37D">921</mods:number>
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<treatment id="03D987F22922F649FD90272B76B8FC11" LSID="urn:lsid:zoobank.org:act:48EAD7CA-D85E-40E0-B32C-7A6C5A078A7C" httpUri="http://treatment.plazi.org/id/03D987F22922F649FD90272B76B8FC11" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="C36A656F2922F64BFD90272B7699F992" box="[619,969,1539,1566]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BCF36E42922F64BFD90272B7699F992" blockId="12.[353,1234,1539,1637]" box="[619,969,1539,1566]" pageId="12" pageNumber="13">
<heading id="D08781882922F64BFD90272B7699F992" box="[619,969,1539,1566]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="4C704D672922F64BFD90272B7637F992" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[619,871,1539,1566]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" order="Agoniatitida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B904EAF62922F64BFD90272B7637F992" bold="true" box="[619,871,1539,1566]" italics="true" pageId="12" pageNumber="13">Maenioceras beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2922F64BFC95272B7699F992" box="[878,969,1540,1566]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2922F64BFE9A2708763FF9E8" pageId="12" pageNumber="13" type="description">
<paragraph id="8BCF36E42922F64BFE9A27087182F9CE" blockId="12.[353,1234,1539,1637]" box="[353,1234,1575,1602]" pageId="12" pageNumber="13">
<uri id="FFE13AE62922F64BFE9A27087182F9CE" box="[353,1234,1575,1602]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="FFD60C312922F64BFD7B27087182F9CE" box="[640,1234,1575,1602]" pageId="12" pageNumber="13">48EAD7CA-D85E-40E0-B32C-7A6C5A078A7C</uuid>
</uri>
</paragraph>
<paragraph id="8BCF36E42922F64BFD3E2765763FF9E8" blockId="12.[353,1234,1539,1637]" box="[709,879,1610,1637]" pageId="12" pageNumber="13">
<figureCitation id="134B2A612922F64BFD3E2765765BF9E9" box="[709,779,1610,1637]" captionStart="Fig" captionStartId="13.[189,232,1955,1981]" captionTargetBox="[243,1340,1305,1894]" captionTargetId="figure-233@13.[235,1353,1296,1911]" captionTargetPageId="13" captionText="Fig. 7. Maenioceras beckeri sp. nov. from Hassi Nebech.A. Holotype MB.C.31952.1. B. Paratype MB.C.31952.2. C. Holotype MB.C.31952.1; suture line at wh = 6.4 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590257" httpUri="https://zenodo.org/record/10590257/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
;
<tableCitation id="C6F2035F2922F64BFCE32765763FF9E8" box="[792,879,1610,1636]" captionStart="Table 4" captionStartId="14.[189,259,265,291]" captionTargetPageId="14" captionText="Table 4. Conch dimensions (in mm) and ratios of Maenioceras beckeri sp. nov." httpUri="http://table.plazi.org/id/DF0F666C2920F649FF4620267132FEA8" pageId="12" pageNumber="13" tableUuid="DF0F666C2920F649FF4620267132FEA8">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2922F64BFF4627A376C6F886" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8BCF36E42922F64BFF4627A37605F92A" blockId="12.[189,853,1675,1702]" box="[189,853,1675,1702]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB6011CA2922F64BFF4627A37605F92A" box="[189,853,1675,1702]" pageId="12" pageNumber="13">
<taxonomicName id="4C704D672922F64BFF4627A3748AF929" box="[189,474,1676,1701]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" order="Agoniatitida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="excavatum">
<emphasis id="B904EAF62922F64BFF4627A3748AF929" box="[189,474,1676,1701]" italics="true" pageId="12" pageNumber="13">Maenioceras excavatum</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AD110F52922F64BFE0327A377EDF92A" author="Petter G." box="[504,701,1675,1702]" page="124" pageId="12" pageNumber="13" year="1959">
<bibRefCitation id="EFE14B152922F64BFE0327A377EDF92A" author="Petter G." box="[504,701,1675,1702]" pageId="12" pageNumber="13" pagination="1 - 313" refId="ref17052" refString="Petter G. 1959. Goniatites devoniennes du Sahara. Publications du Service de la Carte geologique de l'Algerie (nouvelle serie), Paleontologie 2: 1 - 313." type="journal article" year="1959">Petter 1959: 124</bibRefCitation>
</treatmentCitation>
, pl. 6 fig. 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF36E42922F64BFF4627E271BCF964" blockId="12.[189,1260,1741,1803]" box="[189,1260,1741,1768]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB6011CA2922F64BFF4627E271BCF964" box="[189,1260,1741,1768]" pageId="12" pageNumber="13">
?
<taxonomicName id="4C704D672922F64BFF2A27E17744F96B" box="[209,532,1741,1767]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" isUncertain="true" kingdom="Animalia" order="Agoniatitida" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="excavatum">
<emphasis id="B904EAF62922F64BFF2A27E1743AF96B" box="[209,362,1742,1767]" italics="true" pageId="12" pageNumber="13">Maenioceras</emphasis>
cf.
<emphasis id="B904EAF62922F64BFE6C27E17744F96B" box="[407,532,1742,1767]" italics="true" pageId="12" pageNumber="13">excavatum</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AD110F52922F64BFDC927E177A6F964" author="Petter G." box="[562,758,1741,1768]" page="125" pageId="12" pageNumber="13" year="1959">
<bibRefCitation id="EFE14B152922F64BFDC927E177A6F964" author="Petter G." box="[562,758,1741,1768]" pageId="12" pageNumber="13" pagination="1 - 313" refId="ref17052" refString="Petter G. 1959. Goniatites devoniennes du Sahara. Publications du Service de la Carte geologique de l'Algerie (nouvelle serie), Paleontologie 2: 1 - 313." type="journal article" year="1959">Petter 1959: 125</bibRefCitation>
</treatmentCitation>
, pl. 6 fig. 13. —
<bibRefCitation id="EFE14B152922F64BFC4727E27131F96B" author="Bensaid M." box="[956,1121,1741,1767]" pageId="12" pageNumber="13" pagination="82 - 137" refId="ref16343" refString="Bensaid M. 1974. Etude sur des Goniatites a la limite du Devonien moyen et superieur, du Sud marocain. Notes du Service geologique du Maroc 36: 82 - 137." type="journal article" year="1974">Bensaïd 1974</bibRefCitation>
: pl. 2 fig. 5.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF36E42922F64BFF4627DF76C6F886" blockId="12.[189,1260,1741,1803]" box="[189,918,1776,1803]" pageId="12" pageNumber="13">
?
<taxonomicName id="4C704D672922F64BFF2A27DE743AF886" ID-CoL="5K26" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[209,362,1777,1802]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" order="Agoniatitida" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus" status="n. sp.">
<emphasis id="B904EAF62922F64BFF2A27DE743AF886" box="[209,362,1777,1802]" italics="true" pageId="12" pageNumber="13">Maenioceras</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2922F64BFE8927DE74E1F887" box="[370,433,1777,1803]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
III –
<bibRefCitation id="EFE14B152922F64BFE0827DF779EF886" author="Becker R. T. &amp; Aboussalam Z. S. &amp; Bockwinkel J. &amp; Ebbighausen V. &amp; El Hassani A. &amp; Nubel H." box="[499,718,1776,1803]" pageId="12" pageNumber="13" pagination="29 - 43" refId="ref16173" refString="Becker R. T., Aboussalam Z. S., Bockwinkel J., Ebbighausen V., El Hassani A. &amp; Nubel H. 2004. The Givetian and Frasnian at Oued Mzerreb (Tata region, eastern Dra Valley). Documents de l'Institut scientifique Rabat 19: 29 - 43." type="journal article" year="2004">
Becker
<emphasis id="B904EAF62922F64BFDB627DE77D0F886" box="[589,640,1776,1802]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
. 2004
</bibRefCitation>
: pl. 1 figs 11–12.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2922F64AFF46261E7719FEE5" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="8BCF36E42922F64BFF46261E7466F8C7" blockId="12.[189,310,1841,1867]" box="[189,310,1841,1867]" pageId="12" pageNumber="13">
<heading id="D08781882922F64BFF46261E7466F8C7" bold="true" box="[189,310,1841,1867]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B904EAF62922F64BFF46261E7466F8C7" bold="true" box="[189,310,1841,1867]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42922F64AFF46264F7719FEE5" blockId="12.[189,1399,1888,2020]" lastBlockId="13.[188,1399,265,361]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
Species of
<taxonomicName id="4C704D672922F64BFEBF264E748FF8F6" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[324,479,1889,1914]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">
<emphasis id="B904EAF62922F64BFEBF264E748FF8F6" box="[324,479,1889,1914]" italics="true" pageId="12" pageNumber="13">Maenioceras</emphasis>
</taxonomicName>
with thickly discoidal and subinvolute conch at
<quantity id="4C889B012922F64BFBCA264F71D7F8F7" box="[1073,1159,1888,1915]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="mm" value="17.0">17 mm</quantity>
dm (ww/dm ~0.50; uw/dm ~ 0.30) with weakly depressed whorl profile (ww/wh ~ 1.20) and low coiling rate (WER~ 1.55). Whorl profile at
<quantity id="4C889B012922F64BFE68268874BBF84D" box="[403,491,1959,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="13" unit="mm" value="17.0">17 mm</quantity>
conch diameter horseshoe-shaped with flattened, nearly parallel, weakly convergent flanks, subangular ventrolateral shoulder and broad, nearly flat venter. Ventrolateral shoulder with a faint groove in adult specimens. Growth lines coarse, strongly biconvex. Suture line with extremely wide external lobe, deep, U-shaped E
<subScript id="17F434A12923F64AFCD620107666FEC1" attach="left" box="[813,822,319,333]" fontSize="6" pageId="13" pageNumber="14">2</subScript>
lobe, inverted U-shaped ventrolateral saddle and blunt, tongue-shaped lateral lobe.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2923F64AFF4620BE71F9FE56" box="[189,1193,401,474]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BCF36E42923F64AFF4620BE7414FE27" blockId="13.[189,324,401,427]" box="[189,324,401,427]" pageId="13" pageNumber="14">
<heading id="D08781882923F64AFF4620BE7414FE27" bold="true" box="[189,324,401,427]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B904EAF62923F64AFF4620BE7414FE27" bold="true" box="[189,324,401,427]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42923F64AFF46209071F9FE56" blockId="13.[189,1193,447,474]" box="[189,1193,447,474]" pageId="13" pageNumber="14">Named after R. Thomas Becker, who possibly already recognised this species as new.</paragraph>
</subSubSection>
<subSubSection id="C36A656F2923F64AFF46232E71AAFD79" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BCF36E42923F64AFF46232E743CFD96" blockId="13.[189,364,512,539]" box="[189,364,512,539]" pageId="13" pageNumber="14">
<heading id="D08781882923F64AFF46232E743CFD96" bold="true" box="[189,364,512,539]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B904EAF62923F64AFF46232E743CFD96" bold="true" box="[189,364,512,539]" pageId="13" pageNumber="14">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B183CB92923F64AFF112300779CFD1C" collectionCode="MB" collectorName="Bockwinkel &amp; Ebbighausen" country="Morocco" county="Bockwinkel" location="Anti-Atlas" municipality="Givetian" pageId="13" pageNumber="14" specimenCode="MB.C.31952.1" specimenCount="1" typeStatus="holotype">
<paragraph id="8BCF36E42923F64AFF112300740CFDC5" blockId="13.[189,1399,559,656]" box="[234,348,559,585]" pageId="13" pageNumber="14">
<emphasis id="B904EAF62923F64AFF112300740CFDC5" bold="true" box="[234,348,559,585]" pageId="13" pageNumber="14">
<typeStatus id="54CB88462923F64AFF112300740CFDC5" box="[234,348,559,585]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF36E42923F64AFF46237C779CFD1C" blockId="13.[189,1399,559,656]" pageId="13" pageNumber="14">
<collectingCountry id="F36776742923F64AFF46237C7401FDE1" box="[189,337,595,621]" name="Morocco" pageId="13" pageNumber="14">MOROCCO</collectingCountry>
•
<location id="8EAF603F2923F64AFE8A237D74A0FDE0" LSID="urn:lsid:plazi:treatment:03D987F22922F649FD90272B76B8FC11:8EAF603F2923F64AFE8A237D74A0FDE0" box="[369,496,594,620]" country="Morocco" county="Bockwinkel" municipality="Givetian" name="Anti-Atlas" pageId="13" pageNumber="14">Anti-Atlas</location>
,
<location id="8EAF603F2923F64AFDFB237D77FAFDE0" LSID="urn:lsid:plazi:treatment:03D987F22922F649FD90272B76B8FC11:8EAF603F2923F64AFDFB237D77FAFDE0" box="[512,682,594,620]" country="Morocco" county="Bockwinkel" municipality="Givetian" name="Hassi Nebech" pageId="13" pageNumber="14">Hassi Nebech</location>
, section 2; middle
<collectingMunicipality id="6BABAC9E2923F64AFC6C237D7150FDE0" box="[919,1024,594,620]" pageId="13" pageNumber="14">Givetian</collectingMunicipality>
;
<collectorName id="268553322923F64AFBEB237D71CEFDE0" box="[1040,1182,594,620]" pageId="13" pageNumber="14">Bockwinkel</collectorName>
and
<collectorName id="268553322923F64AFB24237D7027FDE0" box="[1247,1399,594,620]" pageId="13" pageNumber="14">Ebbighausen</collectorName>
Coll.;
<specimenCode id="DBD69E9F2923F64AFEFD235A74E6FD1C" box="[262,438,629,656]" pageId="13" pageNumber="14">MB.C.31952.1</specimenCode>
(illustrated in
<figureCitation id="134B2A612923F64AFD9A23597790FD1C" box="[609,704,630,656]" captionStart="Fig" captionStartId="13.[189,232,1955,1981]" captionTargetBox="[243,1340,1305,1894]" captionTargetId="figure-233@13.[235,1353,1296,1911]" captionTargetPageId="13" captionText="Fig. 7. Maenioceras beckeri sp. nov. from Hassi Nebech.A. Holotype MB.C.31952.1. B. Paratype MB.C.31952.2. C. Holotype MB.C.31952.1; suture line at wh = 6.4 mm. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10590257" httpUri="https://zenodo.org/record/10590257/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
).
</paragraph>
</materialsCitation>
<materialsCitation id="3B183CB92923F64AFF11239871A7FD79" collectionCode="MB" collectorName="Bockwinkel &amp; Ebbighausen" country="Morocco" county="Bockwinkel" location="MOROCCO" municipality="Givetian" pageId="13" pageNumber="14" specimenCode="MB.C.31952.2, MB.C.31952.3" specimenCount="2" typeStatus="paratype">
<paragraph id="8BCF36E42923F64AFF1123987437FD5D" blockId="13.[189,1274,695,757]" box="[234,359,695,721]" pageId="13" pageNumber="14">
<emphasis id="B904EAF62923F64AFF1123987437FD5D" bold="true" box="[234,359,695,721]" pageId="13" pageNumber="14">
<typeStatus id="54CB88462923F64AFF1123987437FD5D" box="[234,359,695,721]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF36E42923F64AFF4623F571AAFD79" blockId="13.[189,1274,695,757]" box="[189,1274,730,757]" pageId="13" pageNumber="14">
<collectingCountry id="F36776742923F64AFF4623F57401FD78" box="[189,337,730,756]" name="Morocco" pageId="13" pageNumber="14">MOROCCO</collectingCountry>
•
<specimenCount id="9D76FD6D2923F64AFE9123F57429FD78" box="[362,377,730,756]" pageId="13" pageNumber="14">2</specimenCount>
specs; same collection data as for holotype;
<specimenCode id="DBD69E9F2923F64AFC7D23F57168FD79" box="[902,1080,730,757]" pageId="13" pageNumber="14">MB.C.31952.2</specimenCode>
,
<specimenCode id="DBD69E9F2923F64AFBB823F571A7FD79" box="[1091,1271,730,757]" pageId="13" pageNumber="14">MB.C.31952.3</specimenCode>
.
</paragraph>
</materialsCitation>
</subSubSection>
<subSubSection id="C36A656F2923F649FF46223471CDFC85" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="8BCF36E42923F64AFF462234741EFCB9" blockId="13.[189,334,795,821]" box="[189,334,795,821]" pageId="13" pageNumber="14">
<heading id="D08781882923F64AFF462234741EFCB9" bold="true" box="[189,334,795,821]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B904EAF62923F64AFF462234741EFCB9" bold="true" box="[189,334,795,821]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42923F64AFF46226574EAFBF2" blockId="13.[188,1399,842,1150]" pageId="13" pageNumber="14">
<typeStatus id="54CB88462923F64AFF462265747AFCE8" box="[189,298,842,868]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
MB.C.31952.1 is a limonitic internal mould of a fully chambered specimen (Fig, 7A) with
<quantity id="4C889B012923F64AFAA2226575BBFC04" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="mm" value="17.0">17 mm</quantity>
conch diameter; it is rather well-preserved only on the right side. The conch is thickly discoidal and subinvolute (ww/dm = 0.49; uw/dm = 0.24) with low coiling rate (WER = 1.57). The whorl profile is weakly depressed (ww/wh = 1.18) with slightly convex flanks. The ventrolateral shoulder forms a rather distinct margin separating the flanks and the almost flat, but slightly convex venter. The growth lines are bundled with a strongly biconvex course. The suture line is somewhat unusual for
<taxonomicName id="4C704D672923F64AFB5922D4706BFB98" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[1186,1339,1019,1044]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="genus">
<emphasis id="B904EAF62923F64AFB5922D4706BFB98" box="[1186,1339,1019,1044]" italics="true" pageId="13" pageNumber="14">Maenioceras</emphasis>
</taxonomicName>
with an extremely wide external lobe and a symmetrically rounded E
<subScript id="17F434A12923F64AFC50251F76E4FBB2" attach="left" box="[939,948,1072,1086]" fontSize="6" pageId="13" pageNumber="14">2</subScript>
lobe that is half as deep as the E
<subScript id="17F434A12923F64AFACB251F7069FBB2" attach="left" box="[1328,1337,1072,1086]" fontSize="6" pageId="13" pageNumber="14">1</subScript>
lobe. On the flank follow a narrowly rounded asymmetrical ventrolateral saddle and a deep, tongue-shaped lateral lobe (Fig, 7C).
</paragraph>
<caption id="DF0F666C2923F64AFF46268C7063F86D" ID-DOI="http://doi.org/10.5281/zenodo.10590257" ID-Zenodo-Dep="10590257" httpUri="https://zenodo.org/record/10590257/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1955,1981]" targetBox="[243,1340,1305,1894]" targetPageId="13">
<paragraph id="8BCF36E42923F64AFF46268C7063F86D" blockId="13.[189,1399,1955,2017]" pageId="13" pageNumber="14">
<emphasis id="B904EAF62923F64AFF46268C7440F831" bold="true" box="[189,272,1955,1981]" pageId="13" pageNumber="14">Fig. 7.</emphasis>
<taxonomicName id="4C704D672923F64AFEE6268B7748F831" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[285,536,1955,1981]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B904EAF62923F64AFEE6268B7748F831" box="[285,536,1955,1981]" italics="true" pageId="13" pageNumber="14">Maenioceras beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2923F64AFDDE268C77D4F831" box="[549,644,1955,1981]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
from Hassi Nebech.
<emphasis id="B904EAF62923F64AFC7F268C76C9F831" bold="true" box="[900,921,1955,1981]" pageId="13" pageNumber="14">A</emphasis>
. Holotype MB.C.31952.1.
<emphasis id="B904EAF62923F64AFB12268C71ADF831" bold="true" box="[1257,1277,1955,1981]" pageId="13" pageNumber="14">B</emphasis>
. Paratype MB.C.31952.2.
<emphasis id="B904EAF62923F64AFE8026E974C0F86C" bold="true" box="[379,400,1990,2016]" pageId="13" pageNumber="14">C</emphasis>
. Holotype MB.C.31952.1; suture line at wh = 6.4 mm. Scale bar units = 1 mm.
</paragraph>
</caption>
<caption id="DF0F666C2920F649FF4620267132FEA8" ID-Table-UUID="DF0F666C2920F649FF4620267132FEA8" box="[189,1122,265,292]" httpUri="http://table.plazi.org/id/DF0F666C2920F649FF4620267132FEA8" pageId="14" pageNumber="15" startId="14.[189,259,265,291]" targetBox="[205,1378,341,502]" targetIsTable="true" targetPageId="14">
<paragraph id="8BCF36E42920F649FF4620267132FEA8" blockId="14.[189,1122,265,292]" box="[189,1122,265,292]" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FF4620267470FEAF" bold="true" box="[189,288,265,291]" pageId="14" pageNumber="15">Table 4.</emphasis>
Conch dimensions (in mm) and ratios of
<taxonomicName id="4C704D672920F649FCF220257150FEAF" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[777,1024,265,291]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B904EAF62920F649FCF220257150FEAF" box="[777,1024,265,291]" italics="true" pageId="14" pageNumber="15">Maenioceras beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2920F649FBFC20257132FEA8" box="[1031,1122,266,292]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BCF36E42920F649FF13207A7032FE7A" pageId="14" pageNumber="15">
<table id="F970C444292009B8FF36207A7032FE7A" box="[205,1378,341,502]" gridcols="11" gridrows="4" pageId="14" pageNumber="15">
<tr id="354034A6292009B8FF36207A7032FEE2" box="[205,1378,341,366]" gridrow="0" pageId="14" pageNumber="15">
<th id="76915DDA292009B8FF36207A743DFEE2" box="[205,365,341,366]" gridcol="0" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FF13207A7401FEE1" bold="true" box="[232,337,341,365]" pageId="14" pageNumber="15">specimen</emphasis>
</th>
<th id="76915DDA292009B8FE6E207A7494FEE2" box="[405,452,341,366]" gridcol="1" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FE61207974EFFEE2" bold="true" box="[410,447,342,366]" pageId="14" pageNumber="15">dm</emphasis>
</th>
<th id="76915DDA292009B8FE0D207A774DFEE2" box="[502,541,341,366]" gridcol="2" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FE0D2079774DFEE2" bold="true" box="[502,541,342,366]" pageId="14" pageNumber="15">ww</emphasis>
</th>
<th id="76915DDA292009B8FDA8207A7725FEE2" box="[595,629,341,366]" gridcol="3" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FDA820797725FEE2" bold="true" box="[595,629,342,366]" pageId="14" pageNumber="15">wh</emphasis>
</th>
<th id="76915DDA292009B8FD56207A779FFEE2" box="[685,719,341,366]" gridcol="4" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FD562079779FFEE2" bold="true" box="[685,719,342,366]" pageId="14" pageNumber="15">uw</emphasis>
</th>
<th id="76915DDA292009B8FCF3207A7679FEE2" box="[776,809,341,366]" gridcol="5" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FCF020797677FEE2" bold="true" box="[779,807,342,366]" pageId="14" pageNumber="15">ah</emphasis>
</th>
<th id="76915DDA292009B8FCA2207A76FCFEE2" box="[857,940,341,366]" gridcol="6" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FCA2207A76FCFEE1" bold="true" box="[857,940,341,365]" pageId="14" pageNumber="15">ww/dm</emphasis>
</th>
<th id="76915DDA292009B8FC2E207A7174FEE2" box="[981,1060,341,366]" gridcol="7" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FC2E207A7174FEE1" bold="true" box="[981,1060,341,365]" pageId="14" pageNumber="15">ww/wh</emphasis>
</th>
<th id="76915DDA292009B8FBAB207A71CEFEE2" box="[1104,1182,341,366]" gridcol="8" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FBAB207A71CEFEE1" bold="true" box="[1104,1182,341,365]" pageId="14" pageNumber="15">uw/dm</emphasis>
</th>
<th id="76915DDA292009B8FB31207A705AFEE2" box="[1226,1290,341,366]" gridcol="9" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FB312079705AFEE2" bold="true" box="[1226,1290,342,366]" pageId="14" pageNumber="15">WER</emphasis>
</th>
<th id="76915DDA292009B8FAC9207A7032FEE2" box="[1330,1378,341,366]" gridcol="10" gridrow="0" pageId="14" pageNumber="15">
<emphasis id="B904EAF62920F649FAC920797031FEE2" bold="true" box="[1330,1377,342,366]" pageId="14" pageNumber="15">IZR</emphasis>
</th>
</tr>
<tr id="354034A6292009B8FF3620AF7032FE17" box="[205,1378,384,411]" gridrow="1" pageId="14" pageNumber="15">
<th id="76915DDA292009B8FF3620AF743DFE17" box="[205,365,384,411]" gridcol="0" gridrow="1" pageId="14" pageNumber="15">MB.C.31952.1</th>
<td id="76915DDA292009B8FE6E20AF7494FE17" box="[405,452,384,411]" gridcol="1" gridrow="1" pageId="14" pageNumber="15">17.3</td>
<td id="76915DDA292009B8FE0D20AF774DFE17" box="[502,541,384,411]" gridcol="2" gridrow="1" pageId="14" pageNumber="15">8.5</td>
<td id="76915DDA292009B8FDA820AF7725FE17" box="[595,629,384,411]" gridcol="3" gridrow="1" pageId="14" pageNumber="15">7.2</td>
<td id="76915DDA292009B8FD5620AF779FFE17" box="[685,719,384,411]" gridcol="4" gridrow="1" pageId="14" pageNumber="15">4.2</td>
<td id="76915DDA292009B8FCF320AF7679FE17" box="[776,809,384,411]" gridcol="5" gridrow="1" pageId="14" pageNumber="15">3.5</td>
<td id="76915DDA292009B8FCA220AF76FCFE17" box="[857,940,384,411]" gridcol="6" gridrow="1" pageId="14" pageNumber="15">0.49</td>
<td id="76915DDA292009B8FC2E20AF7174FE17" box="[981,1060,384,411]" gridcol="7" gridrow="1" pageId="14" pageNumber="15">1.18</td>
<td id="76915DDA292009B8FBAB20AF71CEFE17" box="[1104,1182,384,411]" gridcol="8" gridrow="1" pageId="14" pageNumber="15">0.24</td>
<td id="76915DDA292009B8FB3120AF705AFE17" box="[1226,1290,384,411]" gridcol="9" gridrow="1" pageId="14" pageNumber="15">1.57</td>
<td id="76915DDA292009B8FAC920AF7032FE17" box="[1330,1378,384,411]" gridcol="10" gridrow="1" pageId="14" pageNumber="15">0.51</td>
</tr>
<tr id="354034A6292009B8FF3620827032FE44" box="[205,1378,429,456]" gridrow="2" pageId="14" pageNumber="15">
<th id="76915DDA292009B8FF362082743DFE44" box="[205,365,429,456]" gridcol="0" gridrow="2" pageId="14" pageNumber="15">MB.C.31952.2</th>
<td id="76915DDA292009B8FE6E20827494FE44" box="[405,452,429,456]" gridcol="1" gridrow="2" pageId="14" pageNumber="15">15.2</td>
<td id="76915DDA292009B8FE0D2082774DFE44" box="[502,541,429,456]" gridcol="2" gridrow="2" pageId="14" pageNumber="15">7.5</td>
<td id="76915DDA292009B8FDA820827725FE44" box="[595,629,429,456]" gridcol="3" gridrow="2" pageId="14" pageNumber="15">7.4</td>
<td id="76915DDA292009B8FD562082779FFE44" box="[685,719,429,456]" gridcol="4" gridrow="2" pageId="14" pageNumber="15">–</td>
<td id="76915DDA292009B8FCF320827679FE44" box="[776,809,429,456]" gridcol="5" gridrow="2" pageId="14" pageNumber="15">–</td>
<td id="76915DDA292009B8FCA2208276FCFE44" box="[857,940,429,456]" gridcol="6" gridrow="2" pageId="14" pageNumber="15">0.49</td>
<td id="76915DDA292009B8FC2E20827174FE44" box="[981,1060,429,456]" gridcol="7" gridrow="2" pageId="14" pageNumber="15">1.01</td>
<td id="76915DDA292009B8FBAB208271CEFE44" box="[1104,1182,429,456]" gridcol="8" gridrow="2" pageId="14" pageNumber="15">–</td>
<td id="76915DDA292009B8FB312082705AFE44" box="[1226,1290,429,456]" gridcol="9" gridrow="2" pageId="14" pageNumber="15">–</td>
<td id="76915DDA292009B8FAC920827032FE44" box="[1330,1378,429,456]" gridcol="10" gridrow="2" pageId="14" pageNumber="15">–</td>
</tr>
<tr id="354034A6292009B8FF3620F47032FE7A" box="[205,1378,475,502]" gridrow="3" pageId="14" pageNumber="15">
<th id="76915DDA292009B8FF3620F4743DFE7A" box="[205,365,475,502]" gridcol="0" gridrow="3" pageId="14" pageNumber="15">MB.C.31952.3</th>
<td id="76915DDA292009B8FE6E20F47494FE7A" box="[405,452,475,502]" gridcol="1" gridrow="3" pageId="14" pageNumber="15">14.6</td>
<td id="76915DDA292009B8FE0D20F4774DFE7A" box="[502,541,475,502]" gridcol="2" gridrow="3" pageId="14" pageNumber="15">6.5</td>
<td id="76915DDA292009B8FDA820F47725FE7A" box="[595,629,475,502]" gridcol="3" gridrow="3" pageId="14" pageNumber="15">4.7</td>
<td id="76915DDA292009B8FD5620F4779FFE7A" box="[685,719,475,502]" gridcol="4" gridrow="3" pageId="14" pageNumber="15">5.3</td>
<td id="76915DDA292009B8FCF320F47679FE7A" box="[776,809,475,502]" gridcol="5" gridrow="3" pageId="14" pageNumber="15">3.3</td>
<td id="76915DDA292009B8FCA220F476FCFE7A" box="[857,940,475,502]" gridcol="6" gridrow="3" pageId="14" pageNumber="15">0.45</td>
<td id="76915DDA292009B8FC2E20F47174FE7A" box="[981,1060,475,502]" gridcol="7" gridrow="3" pageId="14" pageNumber="15">1.38</td>
<td id="76915DDA292009B8FBAB20F471CEFE7A" box="[1104,1182,475,502]" gridcol="8" gridrow="3" pageId="14" pageNumber="15">0.36</td>
<td id="76915DDA292009B8FB3120F4705AFE7A" box="[1226,1290,475,502]" gridcol="9" gridrow="3" pageId="14" pageNumber="15">1.67</td>
<td id="76915DDA292009B8FAC920F47032FE7A" box="[1330,1378,475,502]" gridcol="10" gridrow="3" pageId="14" pageNumber="15">0.30</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCF36E42920F649FF46236A74F8FD29" blockId="14.[189,1398,580,677]" pageId="14" pageNumber="15">
<typeStatus id="54CB88462920F649FF46236A7474FDD3" box="[189,292,581,607]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
MB.C.31952.2 shows the same conch proportions as the
<typeStatus id="54CB88462920F649FC32236B7160FDD2" box="[969,1072,580,606]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
with a ww/dm ratio of 0.49 and an estimated uw/dm ratio of 0.30 at
<quantity id="4C889B012920F649FD5C234777ABFD0E" box="[679,763,616,643]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="mm" value="15.0">15 mm</quantity>
dm. Numerous dense growth lines are visible in this specimen (Fig, 7B).
</paragraph>
<paragraph id="8BCF36E42920F649FF4623E271CDFC85" blockId="14.[189,1398,716,778]" pageId="14" pageNumber="15">
<typeStatus id="54CB88462920F649FF4623E27474FD6B" box="[189,292,717,743]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
MB.C.31952.3 is, with a ww/dm ratio of 0.45 and a uw/dm ratio of 0.36 at
<quantity id="4C889B012920F649FB4623E37077FD6B" box="[1213,1319,716,743]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="14" pageNumber="15" unit="mm" value="14.5">14.5 mm</quantity>
conch diameter, slightly more slender and more evolute than the other
<specimenCount id="9D76FD6D2920F649FC5523DF7144FC86" box="[942,1044,752,778]" count="2" pageId="14" pageNumber="15" type="generic">two type</specimenCount>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F2920F649FF46221E76B8FC11" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BCF36E42920F649FF46221E747FFCC7" blockId="14.[189,303,817,843]" box="[189,303,817,843]" pageId="14" pageNumber="15">
<heading id="D08781882920F649FF46221E747FFCC7" bold="true" box="[189,303,817,843]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B904EAF62920F649FF46221E747FFCC7" bold="true" box="[189,303,817,843]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E42920F649FF46227076B8FC11" blockId="14.[189,1399,863,925]" pageId="14" pageNumber="15">
The deep E
<subScript id="17F434A12920F649FEBF225D741DFC0C" attach="left" box="[324,333,882,896]" fontSize="6" pageId="14" pageNumber="15">2</subScript>
lobe in
<taxonomicName id="4C704D672920F649FE4B224F77F7FCF5" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[432,679,863,889]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B904EAF62920F649FE4B224F77F7FCF5" box="[432,679,863,889]" italics="true" pageId="14" pageNumber="15">Maenioceras beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D2920F649FD54224F765BFCF6" box="[687,779,864,890]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is the cardinal character to separate the new species from the other species of the genus, which possess a shallow E 2 lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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