treatments-xml/data/03/D9/87/03D987EFAE63935896F5FF39FD961DCD.xml

<document id="4719DC83213A6A02FDCB17BE3970D37F" ID-CLB-Dataset="5549" ID-DOI="10.11646/zootaxa.4990.2.1" ID-GBIF-Dataset="acda30e5-6f57-43bb-81e6-70d3f96762d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5025886" ID-ZooBank="2C8C9945-A40A-409F-AD90-DE6A792285B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624535705376" checkinUser="plazi" docAuthor="Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. &amp; Almeida, Lúcia M." docDate="2021" docId="03D987EFAE63935896F5FF39FD961DCD" docLanguage="en" docName="zootaxa.4990.2.1.pdf" docOrigin="Zootaxa 4990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys schneiderae Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida 2021, new species" docType="treatment" docVersion="7" lastPageNumber="224" masterDocId="FFE0FF97AE77934F9662FFAEFF9E1C38" masterDocTitle="Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species" masterLastPageNumber="226" masterPageNumber="201" pageNumber="221" updateTime="1699020235602" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9E655015706BBB8828EC5A7D3E7DDA54" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B431620A8AE4DFEA22A7FC68845536C0">
<mods:title id="908D496A6E4E320D75B549EA71A47614">Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species</mods:title>
</mods:titleInfo>
<mods:name id="93184CC01E743FD78A2D2274BDEE6B0D" type="personal">
<mods:role id="B057903F8D261123CFC833058BD07ED4">
<mods:roleTerm id="FF370956545D2F7C2F023290EB0CEA6F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12E6FF422111F6F877F53DFBEC645407">Cherman, Mariana A.</mods:namePart>
<mods:affiliation id="1C9D2E445E4AB0C0B926C380F67E87C7">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="F81FD9A0494EB81BC399C7A5F83F8838" type="personal">
<mods:role id="14FFD6831905730AA4F0B0236A70FE52">
<mods:roleTerm id="A5E6E06F4BEC37CB8B692B62CF8525B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E7855E4813050D25AF53D6385E3ADB2">Basílio, Daniel S.</mods:namePart>
<mods:affiliation id="CF91D23B7ACD310A35C2F5524E02AC31">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="E0EFF81207CE92755DF3D0FFDB38AB7C" type="personal">
<mods:role id="FE8C25B09EFCA97421605DFD231C1084">
<mods:roleTerm id="CF39BB3163A9CF50F39B3A40E4C8934A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B1381076275A06D7758C30510E5C474F">Mise, Kleber M.</mods:namePart>
<mods:affiliation id="48F13725BC838617ED64CD903C9CBE51">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="24F62B25979800949CC47B0F8D8823BA" type="personal">
<mods:role id="99F9227B4D20C57C7644D3F76432CB78">
<mods:roleTerm id="00684361F491E5C0EECDAE9789549330">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2CBF25FBD9257E0F0A600F774951A0FA">Frisch, Johannes</mods:namePart>
<mods:affiliation id="52EA2D1A2AEBF10F4F430819CDA67F1E">Museum für Naturkunde, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
</mods:name>
<mods:name id="8425EFD493C45669FDB5554EF4FF5082" type="personal">
<mods:role id="9E3DA3F2AD12A375C5966B2C5A6EDDB5">
<mods:roleTerm id="D22CB77754451A9736AD7F24117D86D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF628DC2C58C5EA1537B0AC3DA11BAB5">Smith, Andrew B. T.</mods:namePart>
<mods:nameIdentifier id="C047A30E355E1FCD01B2340D34606FFC" type="ORCID">0000-0002-8059-5133</mods:nameIdentifier>
<mods:affiliation id="CBB02F4A5A97B8238ED9828ECA694133">Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada. asmith @ nature. ca; https: // orcid. org / 0000 - 0002 - 8059 - 5133</mods:affiliation>
<mods:nameIdentifier id="3E6BBF9DFC5E355622A84A8019B838EB" type="email">asmith@nature.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="B71623FE201F3E6EC41F41B925590F6F" type="personal">
<mods:role id="6DCF7CC0E530FBD2896AC4253D5197BC">
<mods:roleTerm id="D89929FA750DA889A617681195F55DA4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC5FDD17B02EBBB04E2EA48C7F01035C">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="E202131B68E4A994D0277690873DC0DE">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="85E2CC490ECB13ECDAE223CDBF1383CB">text</mods:typeOfResource>
<mods:relatedItem id="1FBD6C3DA6B3E109A0097444996E92D9" type="host">
<mods:titleInfo id="CFDD4BB7CC111A186683FE019FBB43D6">
<mods:title id="95F4AB97425CA0BF2BF668A74AE547E0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F51D66CD4DECC4A4275DE0A9D8A84D59">
<mods:date id="CFD7F742A53081BB5A464C4FE1973BAE">2021</mods:date>
<mods:detail id="D5FF1FF84C59B866E14336F3C9A41906" type="pubDate">
<mods:number id="5618D4E216DC00B85F85D10AE21CBE5A">2021-06-21</mods:number>
</mods:detail>
<mods:detail id="6265C73EDB54D270B4C5D5E8394BDAAB" type="volume">
<mods:number id="01A2536C9FAD9B7F27F4A259A1526A3C">4990</mods:number>
</mods:detail>
<mods:detail id="92666B8024AC0723544649766CD4B306" type="issue">
<mods:number id="9F7B801F412927F9718DFF52BD8B0BE4">2</mods:number>
</mods:detail>
<mods:extent id="22AC57596EA16F840419C36905ED3011" unit="page">
<mods:start id="9AE13790C1F35E0D9B6CCEE78C5CA507">201</mods:start>
<mods:end id="6989CAA44C5A9DDF0A89ED3C53B0C4D7">226</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A29AA3E88A12CD3E6FB38F8AC9F3537C">journal article</mods:classification>
<mods:identifier id="D9BF61A8152C92A274D41DDA03A921D3" type="CLB-Dataset">5549</mods:identifier>
<mods:identifier id="7E6FF93C4C7EB70C6CE6C33F17092534" type="DOI">10.11646/zootaxa.4990.2.1</mods:identifier>
<mods:identifier id="07FDDAE6FFB9EA84561EB4395FEB96CA" type="GBIF-Dataset">acda30e5-6f57-43bb-81e6-70d3f96762d6</mods:identifier>
<mods:identifier id="89EC410F7B0A871A53AFB90B1265223A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="480F2C887DDF1EEDE55797DD8745E72C" type="Zenodo-Dep">5025886</mods:identifier>
<mods:identifier id="894245F5C19EE21CFE24D4CB1AF615BA" type="ZooBank">2C8C9945-A40A-409F-AD90-DE6A792285B3</mods:identifier>
</mods:mods>
<treatment id="03D987EFAE63935896F5FF39FD961DCD" ID-DOI="http://doi.org/10.5281/zenodo.5025914" ID-GBIF-Taxon="183501235" ID-Zenodo-Dep="5025914" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD" httpUri="http://treatment.plazi.org/id/03D987EFAE63935896F5FF39FD961DCD" lastPageId="23" lastPageNumber="224" pageId="20" pageNumber="221">
<subSubSection id="C36A6572AE63935B96F5FF39FD271C89" box="[151,697,151,178]" pageId="20" pageNumber="221" type="nomenclature">
<paragraph id="8BCF36F9AE63935B96F5FF39FD271C89" blockId="20.[151,697,151,213]" box="[151,697,151,178]" pageId="20" pageNumber="221">
<heading id="D0878195AE63935B96F5FF39FD271C89" bold="true" box="[151,697,151,178]" fontSize="11" level="1" pageId="20" pageNumber="221" reason="1">
<emphasis id="B904EAEBAE63935B96F5FF39FD271C89" bold="true" box="[151,697,151,178]" pageId="20" pageNumber="221">
<taxonomicName id="4C704D7AAE63935B96F5FF39FD831C89" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2021" box="[151,541,151,178]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="schneiderae" status="sp. nov.">
<emphasis id="B904EAEBAE63935B96F5FF39FE031C8A" bold="true" box="[151,413,151,178]" italics="true" pageId="20" pageNumber="221">Liogenys schneiderae</emphasis>
Cherman
</taxonomicName>
, 
<taxonomicNameLabel id="A2375790AE63935B9448FF39FD271C89" box="[554,697,151,177]" pageId="20" pageNumber="221" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE63935B96F5FF15FE981CED" box="[151,262,187,213]" pageId="20" pageNumber="221" type="description">
<paragraph id="8BCF36F9AE63935B96F5FF15FE981CED" blockId="20.[151,697,151,213]" box="[151,262,187,213]" pageId="20" pageNumber="221">
<figureCitation id="134B2A7CAE63935B96F5FF15FF7A1CED" box="[151,228,187,213]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="20" pageNumber="221">Figs. 8</figureCitation>
, 
<figureCitation id="134B2A7CAE63935B9693FF15FE9C1CED" box="[241,258,187,213]" captionStart="FIGURE 9" captionStartId="21.[151,250,1756,1780]" captionTargetBox="[184,1431,215,1732]" captionTargetId="figure-17@21.[151,1435,181,1732]" captionTargetPageId="21" captionText="FIGURE 9. Geographical distribution of the Liogenys of A) the northern South America and Central America, overlapping records in B) Panama and in C) Venezuela." figureDoi="http://doi.org/10.5281/zenodo.5025920" httpUri="https://zenodo.org/record/5025920/files/figure.png" pageId="20" pageNumber="221">9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE63935A96F5FEADFBCC1BC9" lastPageId="21" lastPageNumber="222" pageId="20" pageNumber="221" type="materials_examined">
<paragraph id="8BCF36F9AE63935B96F5FEADFBCD1B81" blockId="20.[151,1437,258,1978]" pageId="20" pageNumber="221">
<emphasis id="B904EAEBAE63935B96F5FEADFEDD1D25" bold="true" box="[151,323,259,285]" pageId="20" pageNumber="221">Type material.</emphasis>
<materialsCitation id="3B183CA4AE63935B972AFEADFC631D5D" ID-GBIF-Occurrence="3320554379" collectionCode="MLUH" country="Colombia" location="Colombia" pageId="20" pageNumber="221" specimenCount="2" typeStatus="holotype">
<typeStatus id="54CB885BAE63935B972AFEADFE2E1D25" box="[328,432,259,285]" pageId="20" pageNumber="221" type="holotype">Holotype</typeStatus>
male, labeled: [green, handwritten] “4-dens Fbr / 4-dentata / kl. / Pg.[?] / Col. [
<collectingCountry id="F3677669AE63935B936CFEACFA191D25" box="[1294,1415,258,285]" name="Colombia" pageId="20" pageNumber="221">Colombia</collectingCountry>
]”, [white, typeset and handwritten] “MLU Halle / WB Zoologie / S. –Nr. 8/3/11”, [red typeset and handwritten] “ 
<taxonomicName id="4C704D7AAE63935B96C3FEE5FEAE1D5D" box="[161,304,331,357]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ SCHNEIDERAE / 
<typeStatus id="54CB885BAE63935B9440FEE5FD281D5D" box="[546,694,331,357]" pageId="20" pageNumber="221" type="holotype">HOLOTYPE</typeStatus>
/ Cherman M. A.” (
<collectionCode id="ED61AE3CAE63935B95FEFEE5FC6B1D5D" box="[924,1013,331,357]" country="Germany" name="Martin Luther Universitaet" pageId="20" pageNumber="221">MLUH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B926AFEE2FE411DE9" ID-GBIF-Occurrence="3320554390" collectionCode="ZMHB" collectorName="One" country="Venezuela" determinerName="G. Frey" location="Venezuela" pageId="20" pageNumber="221" specimenCount="2" typeStatus="paratype">
<typeStatus id="54CB885BAE63935B926AFEE2FBEB1D5E" box="[1032,1141,332,358]" pageId="20" pageNumber="221" type="paratype">Paratypes</typeStatus>
(81), all bearing the label [yellow, typeset and handwritten] “ 
<taxonomicName id="4C704D7AAE63935B9446FEC1FD2D1DB1" box="[548,691,367,393]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ SCHNEIDERAE / 
<typeStatus id="54CB885BAE63935B95CAFEC1FBAD1DB1" box="[936,1075,367,393]" pageId="20" pageNumber="221" type="paratype">PARATYPE</typeStatus>
/ Cherman M. A.”. 
<collectorName id="2685532FAE63935B937BFEC1FAD61DB1" box="[1305,1352,367,393]" pageId="20" pageNumber="221">One</collectorName>
female with the labels: [white handwritten] “ 
<collectingCountry id="F3677669AE63935B9425FE3DFD5C1D95" box="[583,706,403,429]" name="Venezuela" pageId="20" pageNumber="221">Venezuela</collectingCountry>
”, [white typeset and handwritten] “ 
<taxonomicName id="4C704D7AAE63935B9239FE3DFB581D95" box="[1115,1222,403,429]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
quadri- / dens / F det. 
<determinerName id="7E13DA3DAE63935B96AAFE19FE811DEA" box="[200,287,439,466]" pageId="20" pageNumber="221">G. Frey</determinerName>
, 1968” (
<collectionCode id="ED61AE3CAE63935B97E2FE19FE491DE9" box="[384,471,439,465]" pageId="20" pageNumber="221">ZMHB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9788FE19FA071DE9" ID-GBIF-Occurrence="3320554380" box="[490,1433,438,466]" collectionCode="ZMHB" collectorName="F. Kummerow" country="Venezuela" location="One" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B9788FE19FD871DE9" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9788FE19FD871DE9" box="[490,537,439,465]" country="Venezuela" name="One" pageId="20" pageNumber="221">One</location>
female with the label: [green typeset] “ 
<collectingCountry id="F3677669AE63935B95B5FE19FBD31DE9" box="[983,1101,439,465]" name="Venezuela" pageId="20" pageNumber="221">Venezuela</collectingCountry>
/ 
<collectorName id="2685532FAE63935B923CFE19FB611DEA" box="[1118,1279,439,466]" pageId="20" pageNumber="221">F. Kummerow</collectorName>
S.” (
<collectionCode id="ED61AE3CAE63935B9358FE19FA0F1DE9" box="[1338,1425,439,465]" pageId="20" pageNumber="221">ZMHB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B96F5FE72FB3A1DCD" ID-GBIF-Occurrence="3320554302" box="[151,1188,474,502]" collectionCode="ZMHB" country="Guyana" location="Two" pageId="20" pageNumber="221" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8EAF6022AE63935B96F5FE72FF561DCE" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B96F5FE72FF561DCE" box="[151,200,476,502]" country="Guyana" name="Two" pageId="20" pageNumber="221">Two</location>
females and 
<specimenCount id="9D76FD70AE63935B973EFE72FE5F1DCD" box="[348,449,475,502]" pageId="20" pageNumber="221" type="male">one male</specimenCount>
with the following label: [white handwritten] “ 
<collectingCountry id="F3677669AE63935B95B2FE75FBB21DCD" box="[976,1068,475,501]" name="Guyana" pageId="20" pageNumber="221">Guyana</collectingCountry>
” (
<collectionCode id="ED61AE3CAE63935B9227FE75FB021DCD" box="[1093,1180,475,501]" pageId="20" pageNumber="221">ZMHB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B92CFFE72FD561E05" ID-GBIF-Occurrence="3320554333" collectionCode="ZMHB" country="Guyana" location="Two" pageId="20" pageNumber="221" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8EAF6022AE63935B92CFFE72FB401DCE" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B92CFFE72FB401DCE" box="[1197,1246,476,502]" country="Guyana" name="Two" pageId="20" pageNumber="221">Two</location>
females and 
<specimenCount id="9D76FD70AE63935B9310FE72FF461E21" pageId="20" pageNumber="221" type="male">two males</specimenCount>
with the label: [green, outlined, typeset] “Hist. Coll. (
<taxonomicName id="4C704D7AAE63935B955FFE51FC221E22" box="[829,956,511,538]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="20" pageNumber="221" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 24131 / 
<taxonomicName id="4C704D7AAE63935B9201FE51FA031E22" authority="Burm." authorityName=": Burmeister" authorityYear="1855" box="[1123,1437,511,538]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="quadridens">Liogenys quadridens Burm.</taxonomicName>
/ Columb., Moritz. / Zool. Mus. Berlin” (
<collectionCode id="ED61AE3CAE63935B940BFD8DFD5E1E05" box="[617,704,547,573]" pageId="20" pageNumber="221">ZMHB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B94B1FD8DFEE11EBD" ID-GBIF-Occurrence="3320554355" collectionCode="ZMHB" country="Guyana" location="One" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B94B1FD8DFC9C1E05" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B94B1FD8DFC9C1E05" box="[723,770,547,573]" country="Guyana" name="One" pageId="20" pageNumber="221">One</location>
male with the labels: [white typeset] “24131”, [dark green handwritten] “ 
<taxonomicName id="4C704D7AAE63935B975AFDE9FE531E59" box="[312,461,583,609]" class="Insecta" family="Scarabaeidae" genus="Amphicrania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="218" phylum="Arthropoda" rank="genus">Amphicrania</taxonomicName>
/ 4 dentata Nr / Columb. Moritz”, [white handwritten] “quadridens / Burm / Melol. 4 dens Fab.” (
<collectionCode id="ED61AE3CAE63935B9742FDC5FEE91EBD" box="[288,375,619,645]" pageId="20" pageNumber="221">ZMHB</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B97E8FDC2FDFA1E91" ID-GBIF-Occurrence="3320554348" collectionCode="ZMUC" country="Colombia" location="Two" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B97E8FDC2FE251EBE" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B97E8FDC2FE251EBE" box="[394,443,620,646]" country="Colombia" name="Two" pageId="20" pageNumber="221">Two</location>
males bearing the labels: [white handwritten] “ 
<collectingCountry id="F3677669AE63935B95B7FDC5FBD81EBD" box="[981,1094,619,645]" name="Colombia" pageId="20" pageNumber="221">Colombia</collectingCountry>
/ Mus. Ber. / 4-dentata / Kl.”, [white typeset] “Mus. Western” (
<collectionCode id="ED61AE3CAE63935B9466FD21FDC51E91" box="[516,603,655,681]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="20" pageNumber="221">ZMUC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B940FFD21FCFA1F01" ID-GBIF-Occurrence="3320554316" collectionCode="SDEI" collectorName="Coll. Kraatz" country="Guyana" determinerName="Moser &amp; G. Frey" location="One" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B940FFD21FD021E91" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B940FFD21FD021E91" box="[621,668,655,681]" country="Guyana" name="One" pageId="20" pageNumber="221">One</location>
male bearing: [white typeset] “ 
<collectorName id="2685532FAE63935B959AFD21FB191E91" box="[1016,1159,655,681]" pageId="20" pageNumber="221">Coll. Kraatz</collectorName>
”, [white typeset] “ 
<determinerName id="7E13DA3DAE63935B9330FD21FA021E91" box="[1362,1436,655,681]" pageId="20" pageNumber="221">Moser</determinerName>
det”, [white typeset] “Dtsch. Ent. Inst. / Eberswalde”, [white outlined green, handwritten] “quadridens. / 
<collectingCountry id="F3677669AE63935B9327FD1DFA021EF5" box="[1349,1436,691,717]" name="Guyana" pageId="20" pageNumber="221">Guyana</collectingCountry>
/F.”, [white handwritten] “ 
<taxonomicName id="4C704D7AAE63935B97A0FD79FDAE1EC9" box="[450,560,727,753]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
. /Guerin.”, [white handwritten] “ 
<taxonomicName id="4C704D7AAE63935B95C5FD79FB8C1EC9" box="[935,1042,727,753]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ 
<taxonomicName id="4C704D7AAE63935B924EFD79FB401EC9" authority="Fab" authorityName="Fab" box="[1068,1246,727,753]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="quadridens">quadridens Fab</taxonomicName>
”, [white, typeset and handwritten] “ 
<taxonomicName id="4C704D7AAE63935B9704FD55FE4F1F2D" box="[358,465,763,789]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
quadri- / dens / F det. 
<determinerName id="7E13DA3DAE63935B94ADFD55FCBB1F2E" box="[719,805,763,790]" pageId="20" pageNumber="221">G. Frey</determinerName>
, 1968”, [white handwritten] “ 
<taxonomicName id="4C704D7AAE63935B9213FD55FB471F2D" box="[1137,1241,763,789]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/quadridens / F.”, [light green typeset] “DEI Münchenberg / Col – 03566” (
<collectionCode id="ED61AE3CAE63935B957AFCB1FCC21F01" box="[792,860,799,825]" pageId="20" pageNumber="221">SDEI</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B950DFCB1FA901FB9" ID-GBIF-Occurrence="3320554349" collectionCode="CMNC" collectorName="Bordon &amp; A. Martinez" country="Guyana" location="One" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Sucre" typeStatus="paratype">
<location id="8EAF6022AE63935B950DFCB1FC001F01" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B950DFCB1FC001F01" box="[879,926,799,825]" country="Guyana" name="One" pageId="20" pageNumber="221" stateProvince="Sucre">One</location>
male with the following data: [white typeset] “CARIACO / (Edo. 
<collectingRegion id="49B4F81BAE63935B97EEFCEDFE731F65" box="[396,493,835,861]" country="Venezuela" name="Sucre" pageId="20" pageNumber="221">SUCRE</collectingRegion>
)”, [white typeset] “ 
<collectorName id="2685532FAE63935B94B3FCEDFCB61F65" box="[721,808,835,861]" pageId="20" pageNumber="221">Bordon</collectorName>
leg.” / [handwritten] “10-IV-59” [white handwritten] “4”, [white, outlined, typeset] “H. &amp; A. HOWDEN/COLLECTION / 
<emphasis id="B904EAEBAE63935B95CBFCC6FC571FB9" box="[937,969,872,897]" italics="true" pageId="20" pageNumber="221">ex.</emphasis>
<collectorName id="2685532FAE63935B95B2FCC9FBC61FB9" box="[976,1112,871,897]" pageId="20" pageNumber="221">A. Martinez</collectorName>
coll.” (
<collectionCode id="ED61AE3CAE63935B92CFFCC9FA981FB9" box="[1197,1286,871,897]" pageId="20" pageNumber="221">CMNC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9378FCC9FAC01FF1" ID-GBIF-Occurrence="3320554377" collectionCode="CMNC" collectorName="A. Martinez" country="Guyana" location="One" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Sucre" typeStatus="paratype">
<location id="8EAF6022AE63935B9378FCC9FAD71FB9" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9378FCC9FAD71FB9" box="[1306,1353,871,897]" country="Guyana" name="One" pageId="20" pageNumber="221" stateProvince="Sucre">One</location>
female with the data: [white typeset] “QUIAMARE[Libertad] / (BARCELONA)”, [white typeset] “5-4-58/BORDON LEG.”, [white, outlined, typeset] “H. &amp; A. HOWDEN / COLLECTION / 
<emphasis id="B904EAEBAE63935B9593FC1EFB8F1FF1" box="[1009,1041,944,969]" italics="true" pageId="20" pageNumber="221">ex.</emphasis>
<collectorName id="2685532FAE63935B9278FC01FB3A1FF1" box="[1050,1188,943,969]" pageId="20" pageNumber="221">A. Martinez</collectorName>
coll.” (
<collectionCode id="ED61AE3CAE63935B929FFC01FAC81FF1" box="[1277,1366,943,969]" pageId="20" pageNumber="221">CMNC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9309FC1EFBBB1FD5" ID-GBIF-Occurrence="3320554344" collectingDate="1978-05" collectionCode="USNM" country="Venezuela" location="Two" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Aragua" typeStatus="paratype">
<location id="8EAF6022AE63935B9309FC1EFA021FF2" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9309FC1EFA021FF2" box="[1387,1436,944,970]" country="Venezuela" name="Two" pageId="20" pageNumber="221" stateProvince="Aragua">Two</location>
females with the data: [white, typeset] “ 
<collectingRegion id="49B4F81BAE63935B943DFC7DFD2A1FD5" box="[607,692,979,1005]" country="Venezuela" name="Aragua" pageId="20" pageNumber="221">Aragua</collectingRegion>
/ 
<collectingCountry id="F3677669AE63935B94AFFC7DFCDE1FD5" box="[717,832,979,1005]" name="Venezuela" pageId="20" pageNumber="221">Venezuela</collectingCountry>
/ 
<date id="FFCE1039AE63935B953BFC7DFC351FD5" box="[857,939,979,1005]" pageId="20" pageNumber="221" value="1978-05">
<collectingDate id="EF8AE9D1AE63935B953BFC7DFC351FD5" box="[857,939,979,1005]" pageId="20" pageNumber="221" value="1978-05">V.1978</collectingDate>
</date>
” (
<collectionCode id="ED61AE3CAE63935B95A4FC7DFB831FD5" box="[966,1053,979,1005]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="221" type="Museum">USNM</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9250FC7DFBBF1861" ID-GBIF-Occurrence="3320554325" collectionCode="CMNC" collectorName="A. Martinez" country="Venezuela" location="One" pageId="20" pageNumber="221" specimenCount="1" specimenCount-male="1" stateProvince="Guarico" typeStatus="paratype">
<location id="8EAF6022AE63935B9250FC7DFBFF1FD5" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9250FC7DFBFF1FD5" box="[1074,1121,979,1005]" country="Venezuela" name="One" pageId="20" pageNumber="221" stateProvince="Guarico">One</location>
male with the data: [white, typeset] “Chaguarramas / (Edo. 
<collectingRegion id="49B4F81BAE63935B9464FC59FDF9182A" box="[518,615,1015,1042]" country="Venezuela" name="Guarico" pageId="20" pageNumber="221">Guarico</collectingRegion>
)”, [white typeset] “13-3-60 / BORDON LEG.”, [white, handwritten and typeset] “ 
<taxonomicName id="4C704D7AAE63935B9762FBB5FE73180D" box="[256,493,1051,1077]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="quadridens">Liogenys quadridens</taxonomicName>
<specimenCount id="9D76FD70AE63935B9791FBB2FD96180E" box="[499,520,1052,1078]" pageId="20" pageNumber="221" type="male">♂</specimenCount>
/ (Fab.) / A MARTINEZ-DET. 1974”, [white with black border, typeset] “H. &amp; A. HOWDEN / COLLECTION / 
<emphasis id="B904EAEBAE63935B978EFBEEFD9B1861" box="[492,517,1088,1113]" italics="true" pageId="20" pageNumber="221">ex</emphasis>
. 
<collectorName id="2685532FAE63935B9473FB91FD061861" box="[529,664,1087,1113]" pageId="20" pageNumber="221">A. Martinez</collectorName>
coll.”, genitalia mounted (
<collectionCode id="ED61AE3CAE63935B95A2FB91FB871861" box="[960,1049,1087,1113]" pageId="20" pageNumber="221">CMNC</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9248FBEEFE9F18D1" ID-GBIF-Occurrence="3320554341" collectingDate="1914-06-16" collectingDateMax="1914-06-17" collectingDateMin="1914-06-16" collectionCode="BMNH, CMNC" collectorName="W. E. Broadway." country="Trinidad and Tobago" location="Two" pageId="20" pageNumber="221" specimenCount="7" specimenCount-female="5" specimenCount-male="2" typeStatus="paratype">
<location id="8EAF6022AE63935B9248FBEEFBC51862" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9248FBEEFBC51862" box="[1066,1115,1088,1114]" country="Trinidad and Tobago" name="Two" pageId="20" pageNumber="221">Two</location>
males and 
<specimenCount id="9D76FD70AE63935B92BAFB91FAFA1861" box="[1240,1380,1087,1113]" pageId="20" pageNumber="221" type="female">four females</specimenCount>
with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B9403FBCDFD271845" box="[609,697,1123,1149]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / Scarborough. / 
<date id="FFCE1039AE63935B95E2FBCDFB971845" box="[896,1033,1123,1149]" pageId="20" pageNumber="221" value="1914-06-16">
<collectingDate id="EF8AE9D1AE63935B95E2FBCDFB971845" box="[896,1033,1123,1149]" pageId="20" pageNumber="221" value="1914-06-16">16. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B9241FBCAFB431845" box="[1059,1245,1123,1150]" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<specimenCount id="9D76FD70AE63935B975FFB29FE161899" box="[317,392,1159,1185]" pageId="20" pageNumber="221" type="male">1 male</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B97A2FB29FDB31899" box="[448,557,1159,1185]" pageId="20" pageNumber="221" type="female">2 females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B9430FB26FD34189A" box="[594,682,1160,1186]" pageId="20" pageNumber="221">BMNH</collectionCode>
; 
<specimenCount id="9D76FD70AE63935B94D7FB29FC9F1899" box="[693,769,1159,1185]" pageId="20" pageNumber="221" type="male">1 male</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B9558FB29FC391899" box="[826,935,1159,1185]" pageId="20" pageNumber="221" type="female">2 females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B95AEFB29FBBA1899" box="[972,1060,1159,1185]" pageId="20" pageNumber="221">CMNC</collectionCode>
). One male with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B97CEFB05FD9A18FD" box="[428,516,1195,1221]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / Scarborough. / 
<date id="FFCE1039AE63935B94B0FB05FCFE18FD" box="[722,864,1195,1221]" pageId="20" pageNumber="221" value="1914-06-17">
<collectingDate id="EF8AE9D1AE63935B94B0FB05FCFE18FD" box="[722,864,1195,1221]" pageId="20" pageNumber="221" value="1914-06-17">17. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B951FFB02FBA718FD" box="[893,1081,1195,1222]" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<collectionCode id="ED61AE3CAE63935B96FDFB61FF6718D1" box="[159,249,1231,1257]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B976FFB61FD711935" ID-GBIF-Occurrence="3320554320" collectingDate="1914-06-10" collectionCode="BMNH" collectorName="W. E. Broadway." country="Trinidad and Tobago" location="One" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B976FFB61FEA218D1" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B976FFB61FEA218D1" box="[269,316,1231,1257]" country="Trinidad and Tobago" name="One" pageId="20" pageNumber="221">One</location>
female with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B95FDFB61FC6918D1" box="[927,1015,1231,1257]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / Scarborough. / 
<date id="FFCE1039AE63935B92A2FB61FAD418D1" box="[1216,1354,1231,1257]" pageId="20" pageNumber="221" value="1914-06-10">
<collectingDate id="EF8AE9D1AE63935B92A2FB61FAD418D1" box="[1216,1354,1231,1257]" pageId="20" pageNumber="221" value="1914-06-10">10. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B9307FB7EFE8D1935" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<collectionCode id="ED61AE3CAE63935B94EEFB5DFD781935" box="[652,742,1267,1293]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B949EFB5DFB521909" ID-GBIF-Occurrence="3320554353" collectingDate="1914-06-12" collectionCode="BMNH" collectorName="W. E. Broadway." country="Trinidad and Tobago" location="One" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B949EFB5DFCB51935" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B949EFB5DFCB51935" box="[764,811,1267,1293]" country="Trinidad and Tobago" name="One" pageId="20" pageNumber="221">One</location>
female with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B96F5FAB9FF711909" box="[151,239,1303,1329]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / Scarborough. / 
<date id="FFCE1039AE63935B97D3FAB9FDA61909" box="[433,568,1303,1329]" pageId="20" pageNumber="221" value="1914-06-12">
<collectingDate id="EF8AE9D1AE63935B97D3FAB9FDA61909" box="[433,568,1303,1329]" pageId="20" pageNumber="221" value="1914-06-12">12. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B9432FAB6FC971909" box="[592,777,1303,1330]" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<collectionCode id="ED61AE3CAE63935B9208FAB9FB5A1909" box="[1130,1220,1303,1329]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B92B4FAB6FE001941" ID-GBIF-Occurrence="3320554339" collectingDate="1914-06-15" collectionCode="BMNH" collectorName="W. E. Broadway." country="Trinidad and Tobago" location="Two" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B92B4FAB6FA99190A" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B92B4FAB6FA99190A" box="[1238,1287,1304,1330]" country="Trinidad and Tobago" name="Two" pageId="20" pageNumber="221">Two</location>
females with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B9403FA95FD27196D" box="[609,697,1339,1365]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / Scarborough. / 
<date id="FFCE1039AE63935B95E2FA95FB97196D" box="[896,1033,1339,1366]" pageId="20" pageNumber="221" value="1914-06-15">
<collectingDate id="EF8AE9D1AE63935B95E2FA95FB97196D" box="[896,1033,1339,1366]" pageId="20" pageNumber="221" value="1914-06-15">15. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B9241FA92FB43196D" box="[1059,1245,1339,1366]" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<collectionCode id="ED61AE3CAE63935B975EFAF1FE081941" box="[316,406,1375,1401]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B97C8FACEFD4E19A5" ID-GBIF-Occurrence="3320554305" collectingDate="1914-06-09" collectionCode="BMNH" collectorName="W. E. Broadway." country="Trinidad and Tobago" location="Two" pageId="20" pageNumber="221" specimenCount="1" typeStatus="paratype">
<location id="8EAF6022AE63935B97C8FACEFE451942" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B97C8FACEFE451942" box="[426,475,1376,1402]" country="Trinidad and Tobago" name="Two" pageId="20" pageNumber="221">Two</location>
females with the data: [white, typeset] “West Indies: / 
<collectingCountry id="F3677669AE63935B9237FAF1FB331941" box="[1109,1197,1375,1401]" name="Trinidad and Tobago" pageId="20" pageNumber="221">Tobago</collectingCountry>
: / 
<date id="FFCE1039AE63935B92A8FAF1FAD61941" box="[1226,1352,1375,1401]" pageId="20" pageNumber="221" value="1914-06-09">
<collectingDate id="EF8AE9D1AE63935B92A8FAF1FAD61941" box="[1226,1352,1375,1401]" pageId="20" pageNumber="221" value="1914-06-09">9. vi. 1914</collectingDate>
</date>
. / 
<collectorName id="2685532FAE63935B9307FACEFE8D19A5" pageId="20" pageNumber="221">W.E. Broadway.</collectorName>
” and “Brit. Mus. / 1920–487.” (
<collectionCode id="ED61AE3CAE63935B940CFA2DFD5619A5" box="[622,712,1411,1437]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B94BBFA2DFAA319DD" ID-GBIF-Occurrence="3320554387" collectionCode="BMNH" country="Trinidad and Tobago" location="One" pageId="20" pageNumber="221" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8EAF6022AE63935B94BBFA2DFC9619A5" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B94BBFA2DFC9619A5" box="[729,776,1411,1437]" country="Trinidad and Tobago" name="One" pageId="20" pageNumber="221">One</location>
male with the data: [white, handwritten] “quadridens / Fabr. / 4-dentata, / Klug. / Columbia”, [white, typeset] “So named / in Reiches / Collection. / C.W.”, [green, handwritten] “ 
<specimenCount id="9D76FD70AE63935B96C1FA65FF2419DD" box="[163,186,1483,1509]" pageId="20" pageNumber="221" type="male">♂</specimenCount>
”, [white, handwritten] “nov. sp.”, [light blue, handwritten] “2258.”, [white, typeset] “67.45” (
<collectionCode id="ED61AE3CAE63935B92B9FA65FAAB19DD" box="[1243,1333,1483,1509]" pageId="20" pageNumber="221">BMNH</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B932AFA65FE941A4E" ID-GBIF-Occurrence="3320554365" collectingDate="1988-06-03" collectingDateMax="1988-06-05" collectingDateMin="1988-06-03" collectionCode="USNM, CMNC, DZUP, CEMT" collectorName="M. Epstein" country="Venezuela" latitude="8.57" location="Sixteen" longLatPrecision="781" longitude="-67.58" pageId="20" pageNumber="221" specimenCount="23" specimenCount-female="19" specimenCount-male="4" stateProvince="Guarico" typeStatus="paratype">
<location id="8EAF6022AE63935B932AFA65FA0219DD" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B932AFA65FA0219DD" box="[1352,1436,1483,1509]" country="Venezuela" latitude="8.57" longLatPrecision="781" longitude="-67.58" name="Sixteen" pageId="20" pageNumber="221" stateProvince="Guarico">Sixteen</location>
males and 
<specimenCount id="9D76FD70AE63935B976FFA41FE181A31" box="[269,390,1519,1546]" pageId="20" pageNumber="221" type="female">15 females</specimenCount>
with the data: [white, typeset] “ 
<collectingCountry id="F3677669AE63935B9487FA41FC0D1A31" box="[741,915,1519,1545]" name="Venezuela" pageId="20" pageNumber="221">VENEZUELA</collectingCountry>
: 
<collectingRegion id="49B4F81BAE63935B95FEFA41FC671A31" box="[924,1017,1519,1545]" country="Venezuela" name="Guarico" pageId="20" pageNumber="221">Guarico</collectingRegion>
, Hato / Masaguaral, 45kmS Calabozo / 
<geoCoordinate id="EE44503EAE63935B96C5F9BDFF731A16" box="[167,237,1555,1582]" degrees="8.57" direction="north" orientation="latitude" pageId="20" pageNumber="221" precision="555" value="8.57">8.57N</geoCoordinate>
, 
<geoCoordinate id="EE44503EAE63935B9693F9BDFED01A16" box="[241,334,1555,1582]" degrees="67.58" direction="west" orientation="longitude" pageId="20" pageNumber="221" precision="555" value="-67.58">67.58W</geoCoordinate>
, GalryForest / #28, 75m, 
<date id="FFCE1039AE63935B9410F9BDFC8A1A15" box="[626,788,1555,1582]" pageId="20" pageNumber="221" value="1988-06-03" valueMax="1988-06-05" valueMin="1988-06-03">
<collectingDate id="EF8AE9D1AE63935B9410F9BDFC8A1A15" box="[626,788,1555,1582]" pageId="20" pageNumber="221" value="1988-06-03" valueMax="1988-06-05" valueMin="1988-06-03">3-5 June 1988</collectingDate>
</date>
/ uv light, 
<collectorName id="2685532FAE63935B95F1F9BAFB8C1A15" box="[915,1042,1555,1582]" pageId="20" pageNumber="221">M. Epstein</collectorName>
” (
<specimenCount id="9D76FD70AE63935B924EF9BDFB311A15" box="[1068,1199,1555,1581]" pageId="20" pageNumber="221" type="male">eight males</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B928AF9BDFA1E1A15" box="[1256,1408,1555,1581]" pageId="20" pageNumber="221" type="female">eight females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B96F5F999FF751A69" box="[151,235,1591,1617]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="221" type="Museum">USNM</collectionCode>
, 
<specimenCount id="9D76FD70AE63935B9695F999FEF01A69" box="[247,366,1591,1617]" pageId="20" pageNumber="221" type="male">four males</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B97C7F999FDA21A69" box="[421,572,1591,1617]" pageId="20" pageNumber="221" type="female">three females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B943DF999FD2A1A69" box="[607,692,1591,1617]" pageId="20" pageNumber="221">CMNC</collectionCode>
, 
<specimenCount id="9D76FD70AE63935B94A2F996FCAC1A69" box="[704,818,1591,1618]" pageId="20" pageNumber="221" type="male">two males</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B9508F996FC6C1A69" box="[874,1010,1591,1618]" pageId="20" pageNumber="221" type="female">two females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B9277F996FBC01A6A" box="[1045,1118,1592,1618]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="221" type="University or college">DZUP</collectionCode>
, 
<specimenCount id="9D76FD70AE63935B9208F996FB421A69" box="[1130,1244,1591,1618]" pageId="20" pageNumber="221" type="male">two males</specimenCount>
and 
<specimenCount id="9D76FD70AE63935B9376F996FA021A69" box="[1300,1436,1591,1618]" pageId="20" pageNumber="221" type="female">two females</specimenCount>
at 
<collectionCode id="ED61AE3CAE63935B96D0F9F5FE9E1A4D" box="[178,256,1627,1653]" pageId="20" pageNumber="221">CEMT</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B183CA4AE63935B9774F9F2FE711A4E" ID-GBIF-Occurrence="3320554361" box="[278,495,1627,1654]" collectionCode="DZUP" country="Venezuela" pageId="20" pageNumber="221" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
<typeStatus id="54CB885BAE63935B9774F9F2FE1D1A4E" box="[278,387,1628,1654]" pageId="20" pageNumber="221" type="paratype">Paratypes</typeStatus>
of 
<collectionCode id="ED61AE3CAE63935B97C5F9F2FE711A4E" box="[423,495,1628,1654]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="221" type="University or college">DZUP</collectionCode>
</materialsCitation>
<materialsCitation id="3B183CA4AE63935B9797F9F5FBD11B81" ID-GBIF-Occurrence="3320554389" collectingDate="1964-05-01" collectingDateMax="1996-06-16" collectingDateMin="1964-05-01" collectionCode="USNM, CMNC" country="Venezuela" county="Hato" latitude="8.57" location="Calabozo" longLatPrecision="781" longitude="-67.58" municipality="Flores Morades" pageId="20" pageNumber="221" specimenCode="DZUP / 402753, DZUP / 402756" specimenCount="3" specimenCount-female="3" stateProvince="Guarico" typeStatus="paratype">
with the sequence of voucher numbers: [white typeset] “ 
<specimenCode id="DBD69E82AE63935B9207F9F5FA821A4E" box="[1125,1308,1627,1654]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="221" type="University or college">DZUP / 402753</specimenCode>
” to “ 
<specimenCode id="DBD69E82AE63935B9333F9F5FE9D1AA2" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="20" pageNumber="221" type="University or college">DZUP / 402756</specimenCode>
”. One male and 
<specimenCount id="9D76FD70AE63935B97AAF9D1FDFC1AA1" box="[456,610,1663,1689]" pageId="20" pageNumber="221" type="female">eight females</specimenCount>
with the data: [white, typeset] “ 
<collectingCountry id="F3677669AE63935B95BEF9D1FB141AA1" box="[988,1162,1663,1689]" name="Venezuela" pageId="20" pageNumber="221">VENEZUELA</collectingCountry>
: 
<collectingRegion id="49B4F81BAE63935B92F5F9D1FB6A1AA1" box="[1175,1268,1663,1689]" country="Venezuela" name="Guarico" pageId="20" pageNumber="221">Guarico</collectingRegion>
, 
<collectingCounty id="62AE4E75AE63935B9361F92EFAA71AA2" box="[1283,1337,1664,1690]" pageId="20" pageNumber="221">Hato</collectingCounty>
/ 
<collectingMunicipality id="6BABAC83AE63935B9334F9D1FF621A85" pageId="20" pageNumber="221">Flores Morades</collectingMunicipality>
, 
<quantity id="4C889B1CAE63935B9764F90DFED31A85" box="[262,333,1699,1726]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="221" unit="km" value="45.0">45 km</quantity>
S / 
<location id="8EAF6022AE63935B9715F90DFE7A1A85" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B9715F90DFE7A1A85" box="[375,484,1699,1725]" country="Venezuela" county="Hato" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Flores Morades" name="Calabozo" pageId="20" pageNumber="221" stateProvince="Guarico">Calabozo</location>
, 
<geoCoordinate id="EE44503EAE63935B978CF90DFDAB1A86" box="[494,565,1699,1726]" degrees="8.57" direction="north" orientation="latitude" pageId="20" pageNumber="221" precision="555" value="8.57">8.57N</geoCoordinate>
, 
<geoCoordinate id="EE44503EAE63935B9422F90DFD071A86" box="[576,665,1699,1726]" degrees="67.58" direction="west" orientation="longitude" pageId="20" pageNumber="221" precision="555" value="-67.58">67.58W</geoCoordinate>
/ 
<location id="8EAF6022AE63935B94CCF90DFCA41A86" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B94CCF90DFCA41A86" box="[686,826,1699,1726]" country="Venezuela" county="Hato" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Flores Morades" name="Galry Forest" pageId="20" pageNumber="221" stateProvince="Guarico">Galry Forest</location>
#2, 75 m / 
<date id="FFCE1039AE63935B95D4F90DFBF21A85" box="[950,1132,1699,1725]" pageId="20" pageNumber="221" value="1989-06-24" valueMax="1989-06-26" valueMin="1989-06-24">
<collectingDate id="EF8AE9D1AE63935B95D4F90DFBF21A85" box="[950,1132,1699,1725]" pageId="20" pageNumber="221" value="1989-06-24" valueMax="1989-06-26" valueMin="1989-06-24">24-26June 1989</collectingDate>
</date>
, uv light / 
<collectorName id="2685532FAE63935B9282F90AFAC41A85" box="[1248,1370,1699,1726]" pageId="20" pageNumber="221">M. Epstein</collectorName>
&amp; 
<collectorName id="2685532FAE63935B931EF90AFE9B1AD9" pageId="20" pageNumber="221">M. Rodiguez</collectorName>
” (
<collectionCode id="ED61AE3CAE63935B977CF969FEEB1AD9" box="[286,373,1735,1761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="221" type="Museum">USNM</collectionCode>
). One male and 
<specimenCount id="9D76FD70AE63935B944EF966FD2A1AD9" box="[556,692,1735,1762]" pageId="20" pageNumber="221" type="female">two females</specimenCount>
with the data: [white, typeset and handwritten] “ex? / intercepted / at Miami Fla / 
<date id="FFCE1039AE63935B972CF945FE471B3D" box="[334,473,1771,1798]" pageId="20" pageNumber="221" value="1964-05-01">
<collectingDate id="EF8AE9D1AE63935B972CF945FE471B3D" box="[334,473,1771,1798]" pageId="20" pageNumber="221" value="1964-05-01">1 May 1964</collectingDate>
</date>
” (
<collectionCode id="ED61AE3CAE63935B9790F945FDD71B3D" box="[498,585,1771,1797]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="221" type="Museum">USNM</collectionCode>
). One male and 
<specimenCount id="9D76FD70AE63935B9562F942FC161B3D" box="[768,904,1771,1798]" pageId="20" pageNumber="221" type="female">two females</specimenCount>
with the data: [white, typeset and handwritten] “ 
<collectingCountry id="F3677669AE63935B96C3F8A1FED11B11" box="[161,335,1807,1833]" name="Venezuela" pageId="20" pageNumber="221">VENEZUELA</collectingCountry>
: 
<collectingRegion id="49B4F81BAE63935B9738F8A1FE4D1B11" box="[346,467,1807,1833]" country="Venezuela" name="Bolivar" pageId="20" pageNumber="221">BOLIVAR</collectingRegion>
/ Guri, 
<date id="FFCE1039AE63935B944DF8A1FD311B11" box="[559,687,1807,1833]" pageId="20" pageNumber="221" value="1996-06-16">
<collectingDate id="EF8AE9D1AE63935B944DF8A1FD311B11" box="[559,687,1807,1833]" pageId="20" pageNumber="221" value="1996-06-16">16.VI.1996</collectingDate>
</date>
/ 
<collectorName id="2685532FAE63935B94AAF8A1FC031B12" box="[712,925,1807,1834]" pageId="20" pageNumber="221">H.&amp;A. Howden Lt</collectorName>
/ 
<location id="8EAF6022AE63935B95D7F8BEFBB31B11" LSID="urn:lsid:plazi:treatment:03D987EFAE63935896F5FF39FD961DCD:8EAF6022AE63935B95D7F8BEFBB31B11" box="[949,1069,1807,1834]" country="Venezuela" county="Hato" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Flores Morades" name="Wet forest" pageId="20" pageNumber="221" stateProvince="Guarico">Wet forest</location>
” (
<collectionCode id="ED61AE3CAE63935B9225F8A1FB3E1B11" box="[1095,1184,1807,1833]" pageId="20" pageNumber="221">CMNC</collectionCode>
). One female with the data: [white, typeset and handwritten] “ 
<collectingCountry id="F3677669AE63935B942CF89DFD621B75" box="[590,764,1843,1869]" name="Venezuela" pageId="20" pageNumber="221">VENEZUELA</collectingCountry>
: 
<collectingRegion id="49B4F81BAE63935B9567F89DFC1B1B75" box="[773,901,1843,1869]" country="Venezuela" name="Guarico" pageId="20" pageNumber="221">GUARICO</collectingRegion>
/ 
<locationDeviation id="9A7D2FDBAE63935B95F8F89DFBA41B75" box="[922,1082,1843,1870]" location="Valle" pageId="20" pageNumber="221">
<quantity id="4C889B1CAE63935B95F8F89DFC441B75" box="[922,986,1843,1869]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="221" unit="km" value="12.0">12km</quantity>
W Valle
</locationDeviation>
de la / Pascua, 21-22.VI.96 Lt / 
<collectorName id="2685532FAE63935B96F5F8F9FED31B49" box="[151,333,1879,1905]" pageId="20" pageNumber="221">H.&amp;A. Howden</collectorName>
” (
<collectionCode id="ED61AE3CAE63935B9704F8F9FE211B49" box="[358,447,1879,1905]" pageId="20" pageNumber="221">CMNC</collectionCode>
). One female with the data: [white, typeset] “VEN: 
<collectingRegion id="49B4F81BAE63935B9265F8F9FBC31B49" box="[1031,1117,1879,1905]" country="Venezuela" name="Bolivar" pageId="20" pageNumber="221">Bolivar</collectingRegion>
, 
<quantity id="4C889B1CAE63935B920AF8F9FB371B49" box="[1128,1193,1879,1905]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="221" unit="km" value="20.0">20km</quantity>
/ SW Ciudad 
<collectingRegion id="49B4F81BAE63935B932AF8F9FA021B49" box="[1352,1436,1879,1905]" country="Venezuela" name="Bolivar" pageId="20" pageNumber="221">Bolivar</collectingRegion>
/ 
<date id="FFCE1039AE63935B96C4F8D5FEA91BAD" box="[166,311,1915,1942]" pageId="20" pageNumber="221" value="1987-06-19">
<collectingDate id="EF8AE9D1AE63935B96C4F8D5FEA91BAD" box="[166,311,1915,1942]" pageId="20" pageNumber="221" value="1987-06-19">19. VI. 1987</collectingDate>
</date>
, S&amp;JPeck / woodland on sand, UV” (
<collectionCode id="ED61AE3CAE63935B9482F8D5FCA71BAD" box="[736,825,1915,1941]" pageId="20" pageNumber="221">CMNC</collectionCode>
). One female with the data: [white, typeset] “VENEZ: 
<collectingRegion id="49B4F81BAE63935B96F5F831FE861B81" box="[151,280,1951,1977]" country="Venezuela" name="Anzoategui" pageId="20" pageNumber="221">Anzoategui</collectingRegion>
/ 
<locationDeviation id="9A7D2FDBAE63935B974CF831FD9A1B81" box="[302,516,1951,1978]" location="Pariaguan" pageId="20" pageNumber="221">
<quantity id="4C889B1CAE63935B974CF831FEEB1B81" box="[302,373,1951,1977]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="20" pageNumber="221" unit="km" value="18.0">18 km</quantity>
E Pariaguan
</locationDeviation>
/ 
<date id="FFCE1039AE63935B9478F831FD3D1B81" box="[538,675,1951,1977]" pageId="20" pageNumber="221" value="1987-06-24">
<collectingDate id="EF8AE9D1AE63935B9478F831FD3D1B81" box="[538,675,1951,1977]" pageId="20" pageNumber="221" value="1987-06-24">
24JUN 
<quantity id="4C889B1CAE63935B940EF831FD3D1B81" box="[620,675,1951,1977]" metricMagnitude="6" metricUnit="m" metricValue="1.98724" pageId="20" pageNumber="221" unit="m" value="1987240.0">1987</quantity>
</collectingDate>
</date>
, 240m / 
<collectorName id="2685532FAE63935B9567F831FCEB1B81" box="[773,885,1951,1977]" pageId="20" pageNumber="221">M.A. Ivie</collectorName>
, beating” (
<collectionCode id="ED61AE3CAE63935B958CF831FBD91B81" box="[1006,1095,1951,1977]" pageId="20" pageNumber="221">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF0F6671AE62935A96F5F972FDD21B20" ID-DOI="http://doi.org/10.5281/zenodo.5025920" ID-Zenodo-Dep="5025920" httpUri="https://zenodo.org/record/5025920/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,250,1756,1780]" targetBox="[184,1431,215,1732]" targetPageId="21">
<paragraph id="8BCF36F9AE62935A96F5F972FDD21B20" blockId="21.[151,1436,1756,1816]" pageId="21" pageNumber="222">
<emphasis id="B904EAEBAE62935A96F5F972FE8B1ACC" bold="true" box="[151,277,1756,1780]" pageId="21" pageNumber="222">FIGURE 9.</emphasis>
Geographical distribution of the 
<taxonomicName id="4C704D7AAE62935A9410F973FD531ACC" box="[626,717,1757,1780]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAEBAE62935A9410F973FD531ACC" box="[626,717,1757,1780]" italics="true" pageId="21" pageNumber="222">Liogenys</emphasis>
</taxonomicName>
of A) the northern South America and Central America, overlapping records in B) Panama and in C) Venezuela.
</paragraph>
</caption>
<paragraph id="8BCF36F9AE62935A96A5F8E8FC281B91" blockId="21.[151,1437,1862,2033]" pageId="21" pageNumber="222">
<materialsCitation id="3B183CA4AE62935A96A5F8E8FC281B91" ID-GBIF-Occurrence="3320554303" collectionCode="BMNH, CMNC, DZUP, CEMT, USNM, ZMHB, ZMUC" pageId="21" pageNumber="222" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CB885BAE62935A96A5F8E8FEB11B58" box="[199,303,1862,1888]" pageId="21" pageNumber="222" type="holotype">Holotype</typeStatus>
deposited at MLUH. Eleven 
<typeStatus id="54CB885BAE62935A94EEF8E9FD691B59" box="[652,759,1863,1889]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
at 
<collectionCode id="ED61AE3CAE62935A9540F8E9FCE41B59" box="[802,890,1863,1889]" pageId="21" pageNumber="222">BMNH</collectionCode>
. Nineteen 
<typeStatus id="54CB885BAE62935A959AF8E9FBFD1B59" box="[1016,1123,1863,1889]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
deposited at 
<collectionCode id="ED61AE3CAE62935A9361F8E9FAC71B59" box="[1283,1369,1863,1889]" pageId="21" pageNumber="222">CMNC</collectionCode>
. Four 
<typeStatus id="54CB885BAE62935A96F5F8C5FE9C1BBD" box="[151,258,1899,1925]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
at 
<collectionCode id="ED61AE3CAE62935A9740F8C5FEF51BBD" box="[290,363,1899,1925]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="21" pageNumber="222" type="University or college">DZUP</collectionCode>
, 
<specimenCount id="9D76FD70AE62935A9717F8C4FD8A1BBD" box="[373,532,1898,1925]" pageId="21" pageNumber="222" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
at 
<collectionCode id="ED61AE3CAE62935A9456F8C5FD1D1BBD" box="[564,643,1899,1925]" pageId="21" pageNumber="222">CEMT</collectionCode>
. One 
<typeStatus id="54CB885BAE62935A94A3F8C5FCBF1BBD" box="[705,801,1899,1925]" pageId="21" pageNumber="222" type="paratype">paratype</typeStatus>
deposited at SDEI. Thirty 
<typeStatus id="54CB885BAE62935A922AF8C5FB2D1BBD" box="[1096,1203,1899,1925]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
deposited at 
<collectionCode id="ED61AE3CAE62935A9326F8C5FA071BBD" box="[1348,1433,1899,1925]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="222" type="Museum">USNM</collectionCode>
. Ten 
<typeStatus id="54CB885BAE62935A96AAF821FEAD1B91" box="[200,307,1935,1961]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
deposited at 
<collectionCode id="ED61AE3CAE62935A97AAF821FD831B91" box="[456,541,1935,1961]" pageId="21" pageNumber="222">ZMHB</collectionCode>
. Two 
<typeStatus id="54CB885BAE62935A943CF821FD571B91" box="[606,713,1935,1961]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
deposited at 
<collectionCode id="ED61AE3CAE62935A953CF821FC311B91" box="[862,943,1935,1961]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="21" pageNumber="222">ZMUC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCF36F9AE62935A96A5F81CFBCC1BC9" blockId="21.[151,1437,1862,2033]" pageId="21" pageNumber="222">
<emphasis id="B904EAEBAE62935A96A5F81CFE2B1BF4" bold="true" box="[199,437,1970,1996]" pageId="21" pageNumber="222">Additional material</emphasis>
. One male (
<collectionCode id="ED61AE3CAE62935A9426F81DFD051BF5" box="[580,667,1971,1997]" pageId="21" pageNumber="222">BMNH</collectionCode>
) missing the head and pronotum with the data: [white, typeset and handwritten] “Scarborough, / 
<collectingCountry id="F3677669AE62935A9787F878FDA21BC8" box="[485,572,2006,2032]" name="Trinidad and Tobago" pageId="21" pageNumber="222">Tobago</collectingCountry>
. / 
<date id="FFCE1039AE62935A9437F878FD701BC9" box="[597,750,2006,2033]" pageId="21" pageNumber="222" value="1914-05-15">15 May 1914</date>
. / W.E. Broadway. / 1914-403”.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE61935996A5FF38FEE21DCC" pageId="22" pageNumber="223" type="diagnosis">
<paragraph id="8BCF36F9AE61935996A5FF38FEE21DCC" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="223">
<emphasis id="B904EAEBAE61935996A5FF38FEDC1C88" bold="true" box="[199,322,150,176]" pageId="22" pageNumber="223">Diagnosis.</emphasis>
Body brown, elongate, sides almost parallel in males (
<figureCitation id="134B2A7CAE61935995DEFF38FB861C88" box="[956,1048,150,176]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8A</figureCitation>
), slightly wider on posterior third in females; elytra brown; pronotum reddish to purplish brown; clypeus quadridentate, clypeal emargination deep, rounded, and narrow; distance between lateral and anterior clypeal teeth slightly longer than basal width of one anterior tooth; right angle between lateral and anterior clypeal teeth; pronotal corners rounded (
<figureCitation id="134B2A7CAE61935992A3FEACFA871D24" box="[1217,1305,258,284]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8B</figureCitation>
), appearing obsolete especially in females; mesotibia quadrate in cross section (
<figureCitation id="134B2A7CAE61935995FDFE88FC671D78" box="[927,1017,294,320]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8C</figureCitation>
); pygidium weakly convex; pygidial disc glabrous, with scattered bristles on apex (
<figureCitation id="134B2A7CAE61935994CEFEE4FC961D5C" box="[684,776,330,356]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8D</figureCitation>
); ventrites II, III, and sometimes IV undulate on midline, especially ventrite II (
<figureCitation id="134B2A7CAE61935997F0FEC0FE751DB0" box="[402,491,366,392]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8E</figureCitation>
); in males protarsomere II up to twice the length of protarsomere I in dorsal view; metatibia subbasally produced towards apex; basal region of parameres strongly narrowed, parameres more than five times the length of their apex; apex harpoon-shaped, lateral spine short, projected almost perpendicular to parameres (
<figureCitation id="134B2A7CAE6193599778FE74FEF11DCC" box="[282,367,474,500]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE61935996A5FE50FC201A4C" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="8BCF36F9AE61935996A5FE50FC201A4C" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="223">
<emphasis id="B904EAEBAE61935996A5FE50FEE11E20" bold="true" box="[199,383,510,536]" pageId="22" pageNumber="223">
<typeStatus id="54CB885BAE61935996A5FE50FEA81E20" box="[199,310,510,536]" pageId="22" pageNumber="223" type="holotype">Holotype</typeStatus>
. Male
</emphasis>
. Length: 
<quantity id="4C889B1CAE6193599788FE50FDDC1E21" box="[490,578,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="22" pageNumber="223" unit="mm" value="9.4">9.4 mm</quantity>
, width: 
<quantity id="4C889B1CAE61935994F9FE50FD6F1E21" box="[667,753,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="22" pageNumber="223" unit="mm" value="4.4">4.4 mm</quantity>
. Brown. 
<emphasis id="B904EAEBAE619359953AFE50FC001E20" box="[856,926,510,536]" italics="true" pageId="22" pageNumber="223">Head:</emphasis>
distance between eyes nearly twice the width of one eye; frons longer than clypeus; clypeus rugose, clypeal emargination deep, rounded, and narrow; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; distance between lateral and anterior teeth slightly longer than basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye longer than one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere maximum width slightly wider than at apex; fovea shallow, extending past the transverse midline of the palpomere; antenna with 10 antennomeres, club lighter in color and as long as the funicle. 
<emphasis id="B904EAEBAE6193599697FCB0FED11F00" box="[245,335,798,824]" italics="true" pageId="22" pageNumber="223">Thorax:</emphasis>
pronotal anterior margin slightly produced medially, undulate and flanged (
<figureCitation id="134B2A7CAE61935992DBFCB0FA8D1F00" box="[1209,1299,798,824]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8B</figureCitation>
); maximum length of pronotum exceeding the length of tarsomeres I–III combined; disc glabrous, punctures very sparse and coarse, denser towards anterior margin; pronotal posterior corners rounded, appearing obsolete; hypomere with long bristles and thin scales; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular, few, scattered, fine punctures at the sides. 
<emphasis id="B904EAEBAE61935997F1FC7CFE7C1FD4" box="[403,482,978,1004]" italics="true" pageId="22" pageNumber="223">Elytra:</emphasis>
shiny, uniform light brown, lighter in color than the pronotum; elytra more than three times longer than the pronotum; elytral suture darker than the elytron and slightly elevated; all four elytral ridges weakly defined. 
<emphasis id="B904EAEBAE619359972DFBB5FE10180C" box="[335,398,1051,1076]" italics="true" pageId="22" pageNumber="223">Legs:</emphasis>
procoxa scaly, thin scales on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia subquadrate in cross section, surface finely sculptured, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout, basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia carinate and abruptly subbasally produced towards apex, apical inner surface glabrous, metatibial surface coarsely sculptured on outer margin; metatibial transverse carina present posteriorly; protarsomere II very elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres equal in width to the mesotarsomeres, slightly wider than the metatarsomeres; metatarsomere I shorter than metatarsomere II; claw bifid, outer tooth of a claw longer and narrower than the inner tooth; distance between teeth shorter than the inner tooth. 
<emphasis id="B904EAEBAE6193599296FAF0FAF91940" box="[1268,1383,1374,1400]" italics="true" pageId="22" pageNumber="223">Abdomen:</emphasis>
disc of ventrites with sparse bristles; ventrites II–III and sometimes IV with tubercles on disc (
<figureCitation id="134B2A7CAE61935992FFFA2CFB6B19A4" box="[1181,1269,1410,1436]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8E</figureCitation>
); propygidium with bristles, pygidium slightly convex, subtrapezoidal, wide; pygidial width exceeding distance between spiracles of propygidium; pygidial disc glabrous, few bristles on apex, pygidial disc finely punctate; pygidial apex quadrate. 
<emphasis id="B904EAEBAE61935996F5FA41FE8F1A30" box="[151,273,1519,1544]" italics="true" pageId="22" pageNumber="223">Parameres</emphasis>
: basal region dorsally narrowed, narrower than both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; parameres more than five times the length of their apex; inner margins opened, convergent; apex harpoon-shaped, lateral spine thin, short, projected almost perpendicular to parameres (approximately 80º angle). Parameres concave in lateral view (
<figureCitation id="134B2A7CAE6193599535F9F4FC2F1A4C" box="[855,945,1626,1652]" captionStart="FIGURE 8" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@19.[203,1385,181,1920]" captionTargetPageId="19" captionText="FIGURE 8. Liogenys schneiderae Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) lateral, D) pygidium, E) ventral of abdomen, black arrows indicate the tubercles at ventrites II–III, and parameres F) dorsal and G) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025918" httpUri="https://zenodo.org/record/5025918/files/figure.png" pageId="22" pageNumber="223">Fig. 8G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE61935996A5F9D0FB381B10" pageId="22" pageNumber="223" type="description">
<paragraph id="8BCF36F9AE61935996A5F9D0FB381B10" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="223">
<emphasis id="B904EAEBAE61935996A5F9D0FEA61AA0" bold="true" box="[199,312,1662,1688]" pageId="22" pageNumber="223">Variation</emphasis>
. Female length: 
<quantity id="4C889B1CAE6193599793F9D0FD161AA1" box="[497,648,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" metricValueMax="10.2" metricValueMin="9.2" pageId="22" pageNumber="223" unit="mm" value="9.7" valueMax="10.2" valueMin="9.2">9.2–10.2 mm</quantity>
, width 
<quantity id="4C889B1CAE61935994BAF9D0FCFF1AA1" box="[728,865,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.3" metricValueMin="4.5" pageId="22" pageNumber="223" unit="mm" value="4.9" valueMax="5.3" valueMin="4.5">4.5–5.3 mm</quantity>
. females as the 
<typeStatus id="54CB885BAE619359926CF9D0FBEE1AA0" box="[1038,1136,1662,1688]" pageId="22" pageNumber="223" type="holotype">holotype</typeStatus>
except the clypeal anterior teeth wider; canthus exceeding the outer margin of the eye; pronotal corners more marked; metatibia coarsely sculptured, metatibial apical carina complete; ventrite tubercles more noticeable and pygidium wider, more trapezoidal. Male length: 9.0– 
<quantity id="4C889B1CAE6193599791F944FDD01B3D" box="[499,590,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="22" pageNumber="223" unit="mm" value="9.5">9.5 mm</quantity>
, width 
<quantity id="4C889B1CAE61935994C5F944FCAB1B3D" box="[679,821,1770,1797]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="22" pageNumber="223" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4–4.8 mm</quantity>
. Males as the 
<typeStatus id="54CB885BAE6193599582F944FBDC1B3C" box="[992,1090,1770,1796]" pageId="22" pageNumber="223" type="holotype">holotype</typeStatus>
except the head and pronotal punctures coarser; pronotal corners more marked, and pygidium flatter and bent downwards.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE61935996A5F89CFB211B81" pageId="22" pageNumber="223" type="etymology">
<paragraph id="8BCF36F9AE61935996A5F89CFB211B81" blockId="22.[151,1437,150,2013]" pageId="22" pageNumber="223">
<emphasis id="B904EAEBAE61935996A5F89CFED01B74" bold="true" box="[199,334,1842,1868]" pageId="22" pageNumber="223">Etymology.</emphasis>
The species is dedicated to Dr. Karla Schneider, biologist and curator of the Zoological Collection of the Martin-Luther-Universität, Halle-Wittenberg, Halle, 
<collectingCountry id="F3677669AE619359954FF8F8FC091B48" box="[813,919,1878,1904]" name="Germany" pageId="22" pageNumber="223">Germany</collectingCountry>
. Karla made the discovery of this new species possible by loaning the 
<typeStatus id="54CB885BAE61935997C0F8D4FD9A1BAC" box="[418,516,1914,1940]" pageId="22" pageNumber="223" type="holotype">holotype</typeStatus>
from the MLUH to the ZHMB. This allowed its thorough examination during the visit of M.A.C. to these collections. This name should be treated as a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE61935896A5F86CFD6B1CEC" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="materials_examined">
<paragraph id="8BCF36F9AE61935996A5F86CFE7B1BE4" blockId="22.[151,1437,150,2013]" box="[199,485,1986,2013]" pageId="22" pageNumber="223">
<emphasis id="B904EAEBAE61935996A5F86CFEFE1BE4" bold="true" box="[199,352,1986,2012]" pageId="22" pageNumber="223">Type locality</emphasis>
: 
<collectingCountry id="F3677669AE619359970DF86CFE7F1BE4" box="[367,481,1986,2012]" name="Colombia" pageId="22" pageNumber="223">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8BCF36F9AE60935896A5FF38FD6B1CEC" blockId="23.[151,1437,150,501]" pageId="23" pageNumber="224">
<emphasis id="B904EAEBAE60935896A5FF38FD881C88" bold="true" box="[199,534,150,176]" pageId="23" pageNumber="224">Geographical distribution.</emphasis>
<collectingCountry id="F3677669AE609358944FFF38FC0D1C88" box="[557,915,150,176]" name="Trinidad and Tobago" pageId="23" pageNumber="224">TRINIDAD AND TOBAGO</collectingCountry>
(Scarborough), 
<collectingCountry id="F3677669AE6093589209FF38FA9D1C88" box="[1131,1283,150,176]" name="Colombia" pageId="23" pageNumber="224">COLOMBIA</collectingCountry>
, 
<collectingCountry id="F3677669AE609358937DFF38FA061C88" box="[1311,1432,150,176]" name="Guyana" pageId="23" pageNumber="224">GUYANA</collectingCountry>
, 
<collectingCountry id="F3677669AE60935896F5FF14FEA21CEC" box="[151,316,186,212]" name="Venezuela" pageId="23" pageNumber="224">VENEZUELA</collectingCountry>
(
<collectingRegion id="49B4F81BAE6093589729FF14FE4F1CEC" box="[331,465,186,212]" country="Venezuela" name="Anzoategui" pageId="23" pageNumber="224">Anzoátegui</collectingRegion>
, 
<collectingRegion id="49B4F81BAE60935897B9FF14FDAF1CEC" box="[475,561,186,212]" country="Venezuela" name="Bolivar" pageId="23" pageNumber="224">Bolívar</collectingRegion>
, 
<collectingRegion id="49B4F81BAE609358945EFF14FD071CEC" box="[572,665,186,212]" country="Venezuela" name="Guarico" pageId="23" pageNumber="224">Guaricó</collectingRegion>
, 
<collectingRegion id="49B4F81BAE60935894C6FF14FD771CEC" box="[676,745,186,212]" country="Venezuela" name="Sucre" pageId="23" pageNumber="224">Sucre</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE60935896A5FF70FD961DCD" pageId="23" pageNumber="224" type="discussion">
<paragraph id="8BCF36F9AE60935896A5FF70FD961DCD" blockId="23.[151,1437,150,501]" pageId="23" pageNumber="224">
<emphasis id="B904EAEBAE60935896A5FF70FEAA1CC0" bold="true" box="[199,308,222,248]" pageId="23" pageNumber="224">Remarks</emphasis>
. 
<taxonomicName id="4C704D7AAE609358972BFF71FD241CC0" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2021" box="[329,698,222,248]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="schneiderae" status="sp. nov.">
<emphasis id="B904EAEBAE609358972BFF71FDDC1CC0" box="[329,578,222,248]" italics="true" pageId="23" pageNumber="224">Liogenys schneiderae</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="B904EAEBAE60935894A9FF70FCC51CC0" bold="true" box="[715,859,222,248]" pageId="23" pageNumber="224">
<taxonomicNameLabel id="A2375790AE60935894A9FF70FCC51CC0" box="[715,859,222,248]" pageId="23" pageNumber="224" rank="species">new species</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C704D7AAE6093589584FF71FB371CC0" authorityName="Blanchard" authorityYear="1851" box="[998,1193,222,248]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B904EAEBAE6093589584FF71FB371CC0" box="[998,1193,222,248]" italics="true" pageId="23" pageNumber="224">L. quadridentata</emphasis>
</taxonomicName>
mainly in the body shape and size, color of the body and elytra, and in the quadridentate clypeus. However, 
<taxonomicName id="4C704D7AAE60935892C0FEADFAD71D24" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2021" box="[1186,1353,258,284]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="schneiderae">
<emphasis id="B904EAEBAE60935892C0FEADFAD71D24" box="[1186,1353,258,284]" italics="true" pageId="23" pageNumber="224">L. schneiderae</emphasis>
</taxonomicName>
differs (
<taxonomicName id="4C704D7AAE60935896C3FE89FEC11D78" authorityName="Blanchard" authorityYear="1851" box="[161,351,294,320]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="species" species="quadridentata">
<emphasis id="B904EAEBAE60935896C3FE89FEC11D78" box="[161,351,294,320]" italics="true" pageId="23" pageNumber="224">L. quadridentata</emphasis>
</taxonomicName>
in parenthesis) in the clypeus being more rugose, with the impressions next to the frons more marked (not rugose); antenna with club shorter; anterior margin of pronotum slightly depressed throughout (not as so); elytral suture medially darker than the elytron (medially unicolored with elytron); metatibia flatter with inner margin abruptly subbasally produced towards apex (produced straight towards apex); ventrites I–IV with tubercles (without tubercles); and pygidium never inflated, with apex quadrate (pygidium convex in both sexes, inflated in females, pygidial apex rounded).
</paragraph>
</subSubSection>
</treatment>
</document>