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<document id="8851772C4DEBDFAEC58B9F00C2A88D44" ID-CLB-Dataset="99517" ID-DOI="10.11646/zootaxa.5165.4.1" ID-GBIF-Dataset="cd84b880-d77f-401a-bf4c-a98021a5e166" ID-ISSN="1175-5326" ID-Zenodo-Dep="6853942" ID-ZooBank="DBE812FF-7B25-4522-97FE-45A86ECAEFD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127043518" checkinUser="plazi" docAuthor="Yang, Qiao-Zhi &amp; Pathomwattananurak, Wuttipon" docDate="2022" docId="03D9879BE853177D7EFDF9B269C0A337" docLanguage="en" docName="zootaxa.5165.4.1.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eophileurus malyi Endrodi 1978" docType="treatment" docVersion="6" lastPageNumber="468" masterDocId="FFE0FFE3E85D176C7E6AFFEE6B79A22A" masterDocTitle="Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species" masterLastPageNumber="485" masterPageNumber="451" pageNumber="465" updateTime="1699360166553" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8C222F7E2CA0A37FA686BD9B11B6024E">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<mods:namePart id="A79900D498C9C9C45E9B90D1687AFF83">Yang, Qiao-Zhi</mods:namePart>
<mods:affiliation id="774D8C830E49D09456FAE2D999E62366">3 B, Pingguoshequ, Baiziwan Road, Chaoyang Dist., Beijing, 100022, P. R. China.</mods:affiliation>
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<mods:namePart id="C07CBC5A3C8B2A6B964D7847F56B61D9">Pathomwattananurak, Wuttipon</mods:namePart>
<mods:affiliation id="0508F545F018847CC4112010DBAE459F">1051 / 38 Wiang, Mueang Dist., Chiang Rai Province, 57000, Thailand.</mods:affiliation>
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<mods:date id="97DB57FBAB1309982EC2F6675A7E4B6A">2022</mods:date>
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<mods:number id="3E15859F36249205022FD81F6F1CA9D0">2022-07-18</mods:number>
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<mods:identifier id="BE1010F1FE8338A2B308AFC0CF58D527" type="DOI">10.11646/zootaxa.5165.4.1</mods:identifier>
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<treatment id="03D9879BE853177D7EFDF9B269C0A337" ID-DOI="http://doi.org/10.5281/zenodo.6853929" ID-GBIF-Taxon="197819084" ID-Zenodo-Dep="6853929" LSID="urn:lsid:plazi:treatment:03D9879BE853177D7EFDF9B269C0A337" httpUri="http://treatment.plazi.org/id/03D9879BE853177D7EFDF9B269C0A337" lastPageId="17" lastPageNumber="468" pageId="14" pageNumber="465">
<subSubSection id="C36A6506E85317627EFDF9B26948A45C" box="[151,561,1627,1655]" pageId="14" pageNumber="465" type="nomenclature">
<paragraph id="8BCF368DE85317627EFDF9B26948A45C" blockId="14.[151,561,1627,1690]" box="[151,561,1627,1655]" pageId="14" pageNumber="465">
<heading id="D08781E1E85317627EFDF9B26948A45C" bold="true" box="[151,561,1627,1655]" fontSize="11" level="1" pageId="14" pageNumber="465" reason="1">
<taxonomicName id="4C704D0EE85317627EFDF9B26948A45C" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" box="[151,561,1627,1655]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85317627EFDF9B26948A45C" bold="true" box="[151,561,1627,1655]" pageId="14" pageNumber="465">
<emphasis id="B904EA9FE85317627EFDF9B26A01A45D" bold="true" box="[151,376,1628,1655]" italics="true" pageId="14" pageNumber="465">Eophileurus malyi</emphasis>
<bibRefCitation id="EFE14B7CE85317627F15F9B56948A45C" author="Endrodi, S." box="[383,561,1627,1654]" pageId="14" pageNumber="465" pagination="183 - 185" refId="ref16975" refString="Endrodi, S. (1978) Zwei neue Eophileurus Arten aus Thailand. Annales historico-naturales Musei nationalis hungarici, 70, 183 - 185." type="journal article" year="1978">Endrődi, 1978</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85317627EFDF96E6A57A4B0" box="[151,302,1664,1690]" pageId="14" pageNumber="465" type="description">
<paragraph id="8BCF368DE85317627EFDF96E6A57A4B0" blockId="14.[151,561,1627,1690]" box="[151,302,1664,1690]" pageId="14" pageNumber="465">
(
<figureCitation id="134B2A08E85317627EF5F96E6A5FA4B0" box="[159,294,1664,1690]" captionStart="FIGURES 71–78" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1375,186,1893]" captionTargetId="figure-18@15.[202,1386,181,1920]" captionTargetPageId="15" captionText="FIGURES 71–78. Type specimens of Eophileurus malyi Endrődi, 1978 (images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels." figureDoi="http://doi.org/10.5281/zenodo.6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="14" pageNumber="465">Figs. 71–78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85317627EFDF92B68CEA4D1" pageId="14" pageNumber="465" type="reference_group">
<paragraph id="8BCF368DE85317627EFDF92B68CEA4D1" blockId="14.[151,1437,1733,1787]" pageId="14" pageNumber="465">
<treatmentCitationGroup id="AB6011A3E85317627EFDF92B68CEA4D1" pageId="14" pageNumber="465">
<treatmentCitation id="0AD1109CE85317627EFDF92B6953A4F7" author="Endrodi, S." box="[151,554,1733,1757]" page="184" pageId="14" pageNumber="465" year="1978">
<taxonomicName id="4C704D0EE85317627EFDF92B6953A4F7" authority="Endrodi, 1978: 184" authorityName="Endrodi" authorityPageNumber="184" authorityYear="1978" box="[151,554,1733,1757]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85317627EFDF92B6A2CA4F7" box="[151,341,1733,1757]" italics="true" pageId="14" pageNumber="465">Eophileurus malyi</emphasis>
<bibRefCitation id="EFE14B7CE85317627F35F92B6953A4F7" author="Endrodi, S." box="[351,554,1733,1757]" pageId="14" pageNumber="465" pagination="183 - 185" refId="ref16975" refString="Endrodi, S. (1978) Zwei neue Eophileurus Arten aus Thailand. Annales historico-naturales Musei nationalis hungarici, 70, 183 - 185." type="journal article" year="1978">Endrődi, 1978: 184</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54CB882FE85317627C50F92B691EA4F7" box="[570,615,1733,1757]" pageId="14" pageNumber="465">type</typeStatus>
locality: “
<collectingRegion id="49B4F86FE85317627CBDF92B6862A4F7" box="[727,795,1733,1757]" country="Thailand" name="Pattani" pageId="14" pageNumber="465">Patani</collectingRegion>
, Sudsiam”) fig. 2 (parameres);
<treatmentCitation id="0AD1109CE85317627A03F92B6E56A4F7" author="Endrodi, S." box="[1129,1327,1733,1757]" page="667" pageId="14" pageNumber="465" year="1985">
<bibRefCitation id="EFE14B7CE85317627A03F92B6E56A4F7" author="Endrodi, S." box="[1129,1327,1733,1757]" pageId="14" pageNumber="465" refId="ref17001" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 667</bibRefCitation>
</treatmentCitation>
, fig. 2017 (parameres);
<treatmentCitation id="0AD1109CE85317627F24F90D6A85A4D1" author="Krajcik, M." box="[334,508,1763,1787]" page="64" pageId="14" pageNumber="465" year="2005">
<bibRefCitation id="EFE14B7CE85317627F24F90D6A85A4D1" author="Krajcik, M." box="[334,508,1763,1787]" pageId="14" pageNumber="465" pagination="1 - 122" refId="ref17162" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčik 2005: 64</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85317627C6DF90D69BBA4D1" author="Krajcik, M." box="[519,706,1763,1787]" page="102" pageId="14" pageNumber="465" year="2012">
<bibRefCitation id="EFE14B7CE85317627C6DF90D69BBA4D1" author="Krajcik, M." box="[519,706,1763,1787]" pageId="14" pageNumber="465" pagination="1 - 278" refId="ref17193" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčik 2012: 102</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85317627CA6F90D68CAA4D1" author="Jakl, S. &amp; Zidek, J." box="[716,947,1763,1787]" page="19" pageId="14" pageNumber="465" year="2015">
<bibRefCitation id="EFE14B7CE85317627CA6F90D68CAA4D1" author="Jakl, S. &amp; Zidek, J." box="[716,947,1763,1787]" pageId="14" pageNumber="465" pagination="14 - 26" refId="ref17117" refString="Jakl, S. &amp; Zidek, J. (2015) Contribution to the knowledge of Oriental and Austro-Papuan Eophileurus Arrow, with descriptions of two new species. Folia Heyrovskyana, Series A, 23 (2), 14 - 26." type="journal article" year="2015">Jákl &amp; Zídek 2015: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85317627EFDF8C46906A5FE" pageId="14" pageNumber="465" type="materials_examined">
<paragraph id="8BCF368DE85317627EFDF8C4698BA585" blockId="14.[151,1437,1833,2039]" pageId="14" pageNumber="465">
<emphasis id="B904EA9FE85317627EFDF8C46ABEA569" bold="true" box="[151,455,1833,1860]" pageId="14" pageNumber="465">Type material examined.</emphasis>
<materialsCitation id="3B183CD0E85317627FBBF8C46F95A54D" ID-GBIF-Occurrence="3863916344" collectionCode="HNHM" location="Otto Merkl" pageId="14" pageNumber="465" specimenCount="1" specimenCount-male="1" stateProvince="Pattani" typeStatus="holotype">
<emphasis id="B904EA9FE85317627FBBF8C46947A56E" bold="true" box="[465,574,1834,1860]" pageId="14" pageNumber="465">
<typeStatus id="54CB882FE85317627FBBF8C46947A56E" box="[465,574,1834,1860]" pageId="14" pageNumber="465" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D76FD04E85317627C38F8C46910A56E" box="[594,617,1834,1860]" pageId="14" pageNumber="465" type="male">♂</specimenCount>
,
<collectionCode id="ED61AE48E85317627C13F8C469A8A56E" box="[633,721,1834,1860]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="465">HNHM</collectionCode>
,
<figureCitation id="134B2A08E85317627C8AF8C76812A56E" box="[736,875,1833,1860]" captionStart="FIGURES 71–78" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1375,186,1893]" captionTargetId="figure-18@15.[202,1386,181,1920]" captionTargetPageId="15" captionText="FIGURES 71–78. Type specimens of Eophileurus malyi Endrődi, 1978 (images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels." figureDoi="http://doi.org/10.5281/zenodo.6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="14" pageNumber="465">Figs. 71–75</figureCitation>
), labeled: “
<collectingRegion id="49B4F86FE85317627D9AF8C76F40A569" box="[1008,1081,1833,1859]" country="Thailand" name="Pattani" pageId="14" pageNumber="465">Patani</collectingRegion>
Südsiam Friederichs // I. q. //
<typeStatus id="54CB882FE85317627EFDF8A36A75A54D" box="[151,268,1869,1895]" pageId="14" pageNumber="465" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C704D0EE85317627F73F8A36942A54D" authority="Endr." authorityName="Endrodi" authorityYear="1978" box="[281,571,1869,1895]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85317627F73F8A36A94A54D" box="[281,493,1869,1895]" italics="true" pageId="14" pageNumber="465">Eophileurus malyi</emphasis>
Endr.
</taxonomicName>
”, examined through photographs provided by
<location id="8EAF6056E85317627A0CF8A06F95A54D" LSID="urn:lsid:plazi:treatment:03D9879BE853177D7EFDF9B269C0A337:8EAF6056E85317627A0CF8A06F95A54D" box="[1126,1260,1869,1896]" name="Otto Merkl" pageId="14" pageNumber="465" stateProvince="Pattani">Ottó Merkl</location>
</materialsCitation>
.
<materialsCitation id="3B183CD0E85317627A97F8A06997A585" ID-GBIF-Occurrence="3863916349" collectionCode="HNHM" location="Otto Merkl" pageId="14" pageNumber="465" specimenCount="1" specimenCount-female="1" stateProvince="Pattani" typeStatus="paratype">
<emphasis id="B904EA9FE85317627A97F8A06E13A542" bold="true" box="[1277,1386,1870,1896]" pageId="14" pageNumber="465">
<typeStatus id="54CB882FE85317627A97F8A06E13A542" box="[1277,1386,1870,1896]" pageId="14" pageNumber="465" type="paratype">Paratype</typeStatus>
</emphasis>
(
<specimenCount id="9D76FD04E85317627BE8F8A06EECA542" box="[1410,1429,1870,1896]" pageId="14" pageNumber="465" type="female">♀</specimenCount>
,
<collectionCode id="ED61AE48E85317627EFDF89C6B96A5A6" box="[151,239,1906,1932]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="14" pageNumber="465">HNHM</collectionCode>
,
<figureCitation id="134B2A08E85317627F6AF89F6AF4A5A1" box="[256,397,1905,1931]" captionStart="FIGURES 71–78" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1375,186,1893]" captionTargetId="figure-18@15.[202,1386,181,1920]" captionTargetPageId="15" captionText="FIGURES 71–78. Type specimens of Eophileurus malyi Endrődi, 1978 (images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels." figureDoi="http://doi.org/10.5281/zenodo.6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="14" pageNumber="465">Figs. 76–78</figureCitation>
), labeled: “
<collectingRegion id="49B4F86FE85317627C7EF89F6924A5A1" box="[532,605,1905,1931]" country="Thailand" name="Pattani" pageId="14" pageNumber="465">Patani</collectingRegion>
Südsiam Friederichs //
<typeStatus id="54CB882FE85317627D10F89C6891A5A6" box="[890,1000,1906,1932]" pageId="14" pageNumber="465" type="paratype">Paratypus</typeStatus>
<taxonomicName id="4C704D0EE85317627D9FF89F6E6CA5A1" authority="Endr." authorityName="Endrodi" authorityYear="1978" box="[1013,1301,1905,1931]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="465" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85317627D9FF89F6FB1A5A1" box="[1013,1224,1905,1931]" italics="true" pageId="14" pageNumber="465">Eophileurus malyi</emphasis>
Endr.
</taxonomicName>
”, examined through photographs provided by the late
<location id="8EAF6056E85317627C07F8786997A585" LSID="urn:lsid:plazi:treatment:03D9879BE853177D7EFDF9B269C0A337:8EAF6056E85317627C07F8786997A585" box="[621,750,1941,1968]" name="Otto Merkl" pageId="14" pageNumber="465" stateProvince="Pattani">Ottó Merkl</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF368DE85317627EADF8576906A5FE" blockId="14.[151,1437,1833,2039]" box="[199,639,1977,2004]" pageId="14" pageNumber="465">
<emphasis id="B904EA9FE85317627EADF8576954A5F9" bold="true" box="[199,557,1977,2003]" pageId="14" pageNumber="465">Additional material examined</emphasis>
. None.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85317627EADF8336AABA5DD" box="[199,466,2013,2039]" pageId="14" pageNumber="465" type="distribution">
<paragraph id="8BCF368DE85317627EADF8336AABA5DD" blockId="14.[151,1437,1833,2039]" box="[199,466,2013,2039]" pageId="14" pageNumber="465">
<emphasis id="B904EA9FE85317627EADF8336A18A5DD" bold="true" box="[199,353,2013,2039]" pageId="14" pageNumber="465">Distribution.</emphasis>
<collectingCountry id="F367761DE85317627F0DF8336AB7A5DD" box="[359,462,2013,2039]" name="Thailand" pageId="14" pageNumber="465">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF0F6605E85217637EFDF8796910A5D2" ID-DOI="http://doi.org/10.5281/zenodo.6853966" ID-Zenodo-Dep="6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="15" pageNumber="466" startId="15.[151,264,1943,1967]" targetBox="[209,1375,186,1893]" targetPageId="15">
<paragraph id="8BCF368DE85217637EFDF8796910A5D2" blockId="15.[151,1436,1943,2040]" pageId="15" pageNumber="466">
<emphasis id="B904EA9FE85217637EFDF8796A2DA585" bold="true" box="[151,340,1943,1967]" pageId="15" pageNumber="466">FIGURES 71–78.</emphasis>
Type specimens of
<taxonomicName id="4C704D0EE85217637C71F8766815A59A" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" box="[539,876,1944,1968]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85217637C71F87669ACA59A" box="[539,725,1944,1968]" italics="true" pageId="15" pageNumber="466">Eophileurus malyi</emphasis>
<bibRefCitation id="EFE14B7CE85217637CB1F8766815A59A" author="Endrodi, S." box="[731,876,1944,1968]" pageId="15" pageNumber="466" pagination="183 - 185" refId="ref16975" refString="Endrodi, S. (1978) Zwei neue Eophileurus Arten aus Thailand. Annales historico-naturales Musei nationalis hungarici, 70, 183 - 185." type="journal article" year="1978">Endrődi, 1978</bibRefCitation>
</taxonomicName>
(images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels.
</paragraph>
</caption>
<caption id="DF0F6605E84D177C7EFDF87969F6A5D2" ID-DOI="http://doi.org/10.5281/zenodo.6853968" ID-Zenodo-Dep="6853968" httpUri="https://zenodo.org/record/6853968/files/figure.png" pageId="16" pageNumber="467" startId="16.[151,264,1943,1967]" targetBox="[180,1404,199,1908]" targetPageId="16">
<paragraph id="8BCF368DE84D177C7EFDF87969F6A5D2" blockId="16.[151,1436,1943,2040]" pageId="16" pageNumber="467">
<emphasis id="B904EA9FE84D177C7EFDF8796A2AA585" bold="true" box="[151,339,1943,1967]" pageId="16" pageNumber="467">FIGURES 79–86.</emphasis>
Type specimens of
<taxonomicName id="4C704D0EE84D177C7C7DF87668D4A59A" authority="Endrodi, 1977" authorityName="Endrodi" authorityYear="1977" box="[535,941,1944,1968]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="nicobarensis">
<emphasis id="B904EA9FE84D177C7C7DF8766860A59A" box="[535,793,1944,1968]" italics="true" pageId="16" pageNumber="467">Eophileurus nicobarensis</emphasis>
<bibRefCitation id="EFE14B7CE84D177C7D77F87668D4A59A" author="Endrodi, S." box="[797,941,1944,1968]" pageId="16" pageNumber="467" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi, 1977</bibRefCitation>
</taxonomicName>
(images ©HNHM/Ottó Merkl). 69–83, holotype, male, 79, habitus, dorsal; 80, habitus, lateral; 81, parameres, apical; 82, parameres, lateral; 83, labels; 84–86, allotype, female, 84, habitus, dorsal; 85, habitus, lateral; 86, labels.
</paragraph>
</caption>
<subSubSection id="C36A6506E84C177D7EADFF7969C0A337" pageId="17" pageNumber="468" type="discussion">
<paragraph id="8BCF368DE84C177D7EADFF7969C0A337" blockId="17.[151,1437,151,285]" pageId="17" pageNumber="468">
<emphasis id="B904EA9FE84C177D7EADFF796A42A29B" bold="true" box="[199,315,151,177]" pageId="17" pageNumber="468">Remarks.</emphasis>
<taxonomicName id="4C704D0EE84C177D7F28FF7969CFA29B" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" box="[322,694,151,177]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE84C177D7F28FF796976A29B" box="[322,527,151,177]" italics="true" pageId="17" pageNumber="468">Eophileurus malyi</emphasis>
<bibRefCitation id="EFE14B7CE84C177D7C7CFF7969CFA29B" author="Endrodi, S." box="[534,694,151,177]" pageId="17" pageNumber="468" pagination="183 - 185" refId="ref16975" refString="Endrodi, S. (1978) Zwei neue Eophileurus Arten aus Thailand. Annales historico-naturales Musei nationalis hungarici, 70, 183 - 185." type="journal article" year="1978">Endrődi, 1978</bibRefCitation>
</taxonomicName>
was described from southern
<collectingCountry id="F367761DE84C177D7A6FFF796F10A29B" box="[1029,1129,151,177]" name="Thailand" pageId="17" pageNumber="468">Thailand</collectingCountry>
with no additional material known to literature since its discovery. The
<typeStatus id="54CB882FE84C177D7CFEFF5569BDA2FF" box="[660,708,187,213]" pageId="17" pageNumber="468">type</typeStatus>
series (
<figureCitation id="134B2A08E84C177D7D4AFF5568D3A2FF" box="[800,938,187,213]" captionStart="FIGURES 71–78" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1375,186,1893]" captionTargetId="figure-18@15.[202,1386,181,1920]" captionTargetPageId="15" captionText="FIGURES 71–78. Type specimens of Eophileurus malyi Endrődi, 1978 (images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels." figureDoi="http://doi.org/10.5281/zenodo.6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="17" pageNumber="468">Figs. 71–78</figureCitation>
) clearly suggest its close relationship to
<taxonomicName id="4C704D0EE84C177D7BEEFF556A0DA2D3" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE84C177D7BEEFF556B89A2D3" italics="true" pageId="17" pageNumber="468">E. confinis</emphasis>
<bibRefCitation id="EFE14B7CE84C177D7E92FF316A0DA2D3" author="Prell, H." box="[248,372,223,249]" pageId="17" pageNumber="468" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
but the parameres are structurally different in having an extra pair of backward process at the center of the parameres in frontal view (
<figureCitation id="134B2A08E84C177D7C32FEED69D4A337" box="[600,685,259,285]" captionStart="FIGURES 71–78" captionStartId="15.[151,264,1943,1967]" captionTargetBox="[209,1375,186,1893]" captionTargetId="figure-18@15.[202,1386,181,1920]" captionTargetPageId="15" captionText="FIGURES 71–78. Type specimens of Eophileurus malyi Endrődi, 1978 (images ©HNHM/Ottó Merkl). 71–75, holotype, male, 71, habitus, dorsal; 72, habitus, lateral; 73, parameres, apical; 74, parameres, lateral; 75, labels; 76–77, paratype, female, 76, habitus, dorsal; 77, habitus, lateral; 78, labels." figureDoi="http://doi.org/10.5281/zenodo.6853966" httpUri="https://zenodo.org/record/6853966/files/figure.png" pageId="17" pageNumber="468">Fig. 73</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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