treatments-xml/data/03/D9/2E/03D92E39FFB0FF805354FB65FE6615E1.xml
2024-06-21 12:22:17 +02:00

1454 lines
178 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="1513CDC1C1F187CF691162AC1FF94C5F" ID-CLB-Dataset="297539" ID-DOI="10.5281/zenodo.11390643" ID-GBIF-Dataset="e4d4534e-bedb-43bf-ae38-ceb9fc6fc0cc" ID-ISSN="1525-9153" ID-Zenodo-Dep="11390643" ID-ZooBank="737F8861-364F-48EC-8DCD-BBDCAD3573C4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716909751475" checkinUser="felipe" docAuthor="Arias, Erick, Chaves, Gerardo &amp; Parra-Olea, Gabriela" docDate="2023" docId="03D92E39FFB0FF805354FB65FE6615E1" docLanguage="en" docName="AmphibReptileCons.17.1-2.e327.pdf" docOrigin="Amphibian &amp; Reptile Conservation (e 327) 17 (1)" docStyle="DocumentStyle:D97478CDCB68366E5D4AECD282BFFB6F.2:AmphibReptileConse.2015-.journal_article.0cover" docStyleId="D97478CDCB68366E5D4AECD282BFFB6F" docStyleName="AmphibReptileConse.2015-.journal_article.0cover" docStyleVersion="2" docTitle="Bolitoglossa bolanosi Arias &amp; Chaves &amp; Parra-Olea 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="154" masterDocId="FFE05641FFB3FF8B5043FFF6FFF71347" masterDocTitle="A new species of salamander (Caudata: Plethodontidae: Bolitoglossa) from the subalpine rain páramo of the Cordillera de Talamanca, Costa Rica" masterLastPageNumber="160" masterPageNumber="143" pageNumber="146" updateTime="1717006729590" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="3847E5AA3279D38ED655CF87D546BD81" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C1FB659953CE5C2E01EE601D75453DF9">
<mods:title id="F4EC431E8A720D78224AA1C19CCA5EF8">A new species of salamander (Caudata: Plethodontidae: Bolitoglossa) from the subalpine rain páramo of the Cordillera de Talamanca, Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="4ED30D82E12AA093D6719CA71B4E1151" type="personal">
<mods:role id="12E5D777B19E78DC9FE25955D72C6188">
<mods:roleTerm id="9C600DE762EB2807BB83F8FE5F209102">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36E4397DA1155F6DE741841B4FECED0D">Arias, Erick</mods:namePart>
<mods:affiliation id="3219562D51AF8CB0B1E82CBAC020B469">Centro de Investigaciones en Biodiversidad y Ecología Tropical (Museo de Zoología), Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, COSTA RICA &amp; Escuela de Biología, Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, COSTA RICA &amp; Departamento de Zoología, Instituto de Biología, UNAM, AP 70 - 153 Ciudad Universitaria, CP 04510, México, D. F., MEXICO</mods:affiliation>
</mods:name>
<mods:name id="469D59D57974C7EE3A86F414E82FCB8A" type="personal">
<mods:role id="A892A83C343771AE741548BA6DA84C2C">
<mods:roleTerm id="57CB3308D6A81F0B734AA2525EFF9C62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="101336DB4F4BCC14A5331BEDD6E07932">Chaves, Gerardo</mods:namePart>
<mods:affiliation id="0EC9291123C451B9551C7A7068FCF019">Centro de Investigaciones en Biodiversidad y Ecología Tropical (Museo de Zoología), Universidad de Costa Rica, San Pedro, 11501 - 2060 San José, COSTA RICA</mods:affiliation>
</mods:name>
<mods:name id="F920973ECEEC8B11DB1709ED3BD14294" type="personal">
<mods:role id="D88A36AE9B517104F24A614EF4BC9FE1">
<mods:roleTerm id="FC157C61E6DAA49CA95554DDC72E7E50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0146A456FD62D1A4124F406ACC3221BE">Parra-Olea, Gabriela</mods:namePart>
<mods:affiliation id="BF126B684106156519FC239862843A57">Departamento de Zoología, Instituto de Biología, UNAM, AP 70 - 153 Ciudad Universitaria, CP 04510, México, D. F., MEXICO</mods:affiliation>
</mods:name>
<mods:typeOfResource id="228F27659E26C9BFE1B27EA2CA4C32D4">text</mods:typeOfResource>
<mods:relatedItem id="1EB4FACCC5879C7013D0D126B9244E36" type="host">
<mods:titleInfo id="34280FB8B81911A5451823E6ED5D0FE4">
<mods:title id="C348B0DC4A42DAF8A507B573AD64F3A8">Amphibian &amp; Reptile Conservation</mods:title>
</mods:titleInfo>
<mods:part id="D3C61D44FCB67369A91FC0996E77F0CF">
<mods:date id="03977AE11122367EA3BBFC4E718FEFFC">2023</mods:date>
<mods:detail id="20A56497DC6A7CDAF20086979C0FAAC9" type="series">
<mods:title id="E8ECA1519B126A603F3A16C0E3CE9989">e 327</mods:title>
</mods:detail>
<mods:detail id="52BC6D8E44EE69FCB4594C0E65BB7C7D" type="pubDate">
<mods:number id="E6ADC942EC6DB7D1F2B8F6BBA9BC3BBC">2023-12-24</mods:number>
</mods:detail>
<mods:detail id="3CD550E7A0407143FD2CC8FEC297ED20" type="volume">
<mods:number id="B8BEAECFE31B7C6FBA8D0F2B5A36E679">17</mods:number>
</mods:detail>
<mods:detail id="E1AE6EC57B9AA4F1A0C8D05DBEED0691" type="issue">
<mods:number id="F92901260BF91AC2B6046EEC7DAC284C">1</mods:number>
</mods:detail>
<mods:extent id="8F0BA99C1826D135F3CEE4E4DD31EAE3" unit="page">
<mods:start id="662112F11F8B9E040B6B0735C118DD63">143</mods:start>
<mods:end id="2F0DD24DC91E64FDF77EE4E1C303A7A4">160</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="95B05929609F9AD8FF94D96CB577647F">journal article</mods:classification>
<mods:identifier id="888B82BB57A9F06974F5024EA2F2EDED" type="CLB-Dataset">297539</mods:identifier>
<mods:identifier id="0AE667A375C31130B48C4C38ADE9AE58" type="DOI">10.5281/zenodo.11390643</mods:identifier>
<mods:identifier id="0D9B17C08F394175C398012054820E17" type="GBIF-Dataset">e4d4534e-bedb-43bf-ae38-ceb9fc6fc0cc</mods:identifier>
<mods:identifier id="BE501AF12695838A4F39E32226CEB3CD" type="ISSN">1525-9153</mods:identifier>
<mods:identifier id="F0404B19341F41570E6ED0BD159E677E" type="Zenodo-Dep">11390643</mods:identifier>
<mods:identifier id="288873050AB9D8CAE6578BAD7F55E938" type="ZooBank">737F8861-364F-48EC-8DCD-BBDCAD3573C4</mods:identifier>
</mods:mods>
<treatment id="03D92E39FFB0FF805354FB65FE6615E1" ID-DOI="http://doi.org/10.5281/zenodo.11390651" ID-Zenodo-Dep="11390651" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1" httpUri="http://treatment.plazi.org/id/03D92E39FFB0FF805354FB65FE6615E1" lastPageId="11" lastPageNumber="154" pageId="3" pageNumber="146">
<subSubSection id="C36ACCA4FFB0FF885354FB65FBA917EC" box="[791,1118,1171,1195]" pageId="3" pageNumber="146" type="nomenclature">
<paragraph id="8BCF9F2FFFB0FF885354FB65FBA917EC" blockId="3.[791,1149,1171,1254]" box="[791,1118,1171,1195]" pageId="3" pageNumber="146">
<heading id="D0872843FFB0FF885354FB65FBA917EC" box="[791,1118,1171,1195]" fontSize="10" level="2" pageId="3" pageNumber="146" reason="4">
<emphasis id="B904433DFFB0FF885354FB65FBA917EC" bold="true" box="[791,1118,1171,1195]" pageId="3" pageNumber="146">
<taxonomicName id="4C70E4ACFFB0FF885354FB65FBF617EC" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[791,1025,1171,1195]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="3" pageNumber="146" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB0FF885354FB65FBF617EC" bold="true" box="[791,1025,1171,1195]" italics="true" pageId="3" pageNumber="146">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FE46FFB0FF88544BFB65FBA917EC" box="[1032,1118,1171,1195]" pageId="3" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFB0FF885354FB47FB8A178E" box="[791,1149,1201,1225]" pageId="3" pageNumber="146" type="vernacular_names">
<paragraph id="8BCF9F2FFFB0FF885354FB47FB8A178E" blockId="3.[791,1149,1171,1254]" box="[791,1149,1201,1225]" pageId="3" pageNumber="146">Bolaños Web-footed Salamander</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFB0FF885354FB38FA9A1658" pageId="3" pageNumber="146" type="description">
<paragraph id="8BCF9F2FFFB0FF885354FB38FC7B17A1" blockId="3.[791,1149,1171,1254]" box="[791,908,1230,1254]" pageId="3" pageNumber="146">
(
<figureCitation id="134B83AAFFB0FF88535CFB38FC7317A1" box="[799,900,1230,1254]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="10.[141,176,688,710]" captionStartId-1="10.[141,176,1873,1895]" captionStartId-2="11.[141,176,1002,1024]" captionTargetBox-0="[159,1364,165,650]" captionTargetBox-1="[141,1394,786,1860]" captionTargetBox-2="[189,1347,161,975]" captionTargetId-0="figure-21@10.[138,1397,138,675]" captionTargetId-1="figure-43@10.[141,1394,786,1860]" captionTargetId-2="figure-495@11.[186,1350,158,978]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Fig. 4. Holotype of Bolitoglossa bolanosi sp. nov. (UCR 22965) in life on a white background. Photograph by Erick Arias." captionText-1="Fig. 5. Dorsal views of the hand and foot of the holotypes of (AB) Bolitoglossa bolanosi sp. nov. (UCR 22965) and (CD) B. pesrubra in preservation. Photograph by Erick Arias." captionText-2="Fig. 6. In-life photographs of variations within Bolitoglossa bolanosi sp. nov. (A) Paratopotype male subadult UCR 22966, (B) paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and (D) paratype adult female UCR 24245. Photographs A and C by Erick Arias, B and D by Omar Zúñiga." figureDoi-0="http://doi.org/10.5281/zenodo.11390653" figureDoi-1="http://doi.org/10.5281/zenodo.11390655" figureDoi-2="http://doi.org/10.5281/zenodo.11390657" httpUri-0="https://zenodo.org/record/11390653/files/figure.png" httpUri-1="https://zenodo.org/record/11390655/files/figure.png" httpUri-2="https://zenodo.org/record/11390657/files/figure.png" pageId="3" pageNumber="146">Figs. 46</figureCitation>
)
</paragraph>
<paragraph id="8BCF9F2FFFB0FF885354FAF8FA9A1658" blockId="3.[791,1389,1294,1311]" box="[791,1389,1294,1311]" pageId="3" pageNumber="146">
<emphasis id="B904433DFFB0FF885354FAF8FA9A1658" bold="true" box="[791,1389,1294,1311]" pageId="3" pageNumber="146">
<uri id="FFE1932DFFB0FF885354FAF8FA9A1658" box="[791,1389,1294,1311]" pageId="3" pageNumber="146">
urn:lsid:zoobank.org:act:
<uuid id="FFD6A5FAFFB0FF8853A6FAF8FA9A1658" box="[997,1389,1294,1311]" pageId="3" pageNumber="146">3F7E0D7E-906F-4FCD-B4CE-0A48F134DEA1</uuid>
</uri>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFB0FF8F5354FAB5FB7B1420" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146" type="materials_examined">
<paragraph id="8BCF9F2FFFB0FF885354FAB5FC0416A9" blockId="3.[791,1395,1347,1518]" pageId="3" pageNumber="146">
<materialsCitation id="3B189572FFB0FF885354FAB5FC0416A9" collectingDate="2016-10-19" collectorName="Erick Arias &amp; Omar Zuniga" country="Costa Rica" county="Canton de Buenos Aires" elevation="2600" location="Cerro Arbolado" municipality="Distrito de Buenos Aires" pageId="3" pageNumber="146" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="B904433DFFB0FF885354FAB5FC72161C" bold="true" box="[791,901,1347,1371]" pageId="3" pageNumber="146">
<typeStatus id="54CB218DFFB0FF885354FAB5FC76161C" box="[791,897,1347,1371]" pageId="3" pageNumber="146" type="holotype">Holotype</typeStatus>
.
</emphasis>
UCR 22965, an adult male from
<collectingCountry id="F367DFBFFFB0FF8854B6FAB5FA98161C" box="[1269,1391,1347,1371]" name="Costa Rica" pageId="3" pageNumber="146">Costa Rica</collectingCountry>
:
<collectingRegion id="49B451CDFFB0FF885354FA97FBC5163F" box="[791,1074,1377,1401]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="146">Provincia de Puntarenas</collectingRegion>
:
<collectingCounty id="62AEE7A3FFB0FF88540BFA97FA98163E" box="[1096,1391,1377,1401]" pageId="3" pageNumber="146">Cantón de Buenos Aires</collectingCounty>
:
<collectingMunicipality id="6BAB0555FFB0FF885354FA88FBD616D1" box="[791,1057,1406,1430]" pageId="3" pageNumber="146">Distrito de Buenos Aires</collectingMunicipality>
: the summit of
<location id="8EAFC9F4FFB0FF885485FA88FA9916D1" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB0FF885485FA88FA9916D1" box="[1222,1390,1406,1430]" country="Costa Rica" county="Canton de Buenos Aires" municipality="Distrito de Buenos Aires" name="Cerro Arbolado" pageId="3" pageNumber="146" stateProvince="Puntarenas">Cerro Arbolado</location>
,
<location id="8EAFC9F4FFB0FF885354FA6AFB6016F4" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB0FF885354FA6AFB6016F4" box="[791,1175,1435,1459]" country="Costa Rica" county="Canton de Buenos Aires" municipality="Distrito de Buenos Aires" name="Parque Internacional La Amistad" pageId="3" pageNumber="146" stateProvince="Puntarenas">Parque Internacional La Amistad</location>
, (9.320º, -83.216º;
<elevation id="005D781CFFB0FF885354FA4FFC601696" box="[791,919,1465,1489]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="146" unit="m" value="2600.0">
<quantity id="4C8832CAFFB0FF885354FA4FFC981697" box="[791,879,1465,1489]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="146" unit="m" value="2600.0">2,600 m</quantity>
asl
</elevation>
), collected by
<collectorName id="2685FAF9FFB0FF885477FA4FFB591696" box="[1076,1198,1465,1489]" pageId="3" pageNumber="146">Erick Arias</collectorName>
and
<collectorName id="2685FAF9FFB0FF8854A0FA4FFA841696" box="[1251,1395,1465,1489]" pageId="3" pageNumber="146">Omar Zúñiga</collectorName>
on
<date id="FFCEB9EFFFB0FF88537BFA20FC1816A9" box="[824,1007,1494,1518]" pageId="3" pageNumber="146" value="2016-10-19">
<collectingDate id="EF8A4007FFB0FF88537BFA20FC1816A9" box="[824,1007,1494,1518]" pageId="3" pageNumber="146" value="2016-10-19">19 October 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCF9F2FFFB0FF885354F9E7FC6F1501" blockId="3.[791,1394,1553,1606]" pageId="3" pageNumber="146">
<emphasis id="B904433DFFB0FF885354F9E7FC4F156E" bold="true" box="[791,952,1553,1577]" pageId="3" pageNumber="146">
<typeStatus id="54CB218DFFB0FF885354F9E7FC44156E" box="[791,947,1553,1577]" pageId="3" pageNumber="146" type="paratopotype">Paratopotype</typeStatus>
.
</emphasis>
UCR 22964, a subadult male, same data as holotype.
</paragraph>
<paragraph id="8BCF9F2FFFB0FF8F5354F99FFB7B1420" blockId="3.[791,1396,1641,1958]" lastBlockId="4.[791,1381,1695,1896]" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146">
<materialsCitation id="3B189572FFB0FF885354F99FFC1E142C" collectingDate="2015-03-30" collectorName="Erick Arias &amp; Gerardo Chaves &amp; Olmer Cordero &amp; Omar Zuniga" country="Costa Rica" elevation="2870" location="Distrito de Buenos Aires" municipality="Canton de Buenos Aires" pageId="3" pageNumber="146" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B904433DFFB0FF885354F99FFC6715C6" bold="true" box="[791,912,1641,1665]" pageId="3" pageNumber="146">
<typeStatus id="54CB218DFFB0FF885354F99FFC7C15C6" box="[791,907,1641,1665]" pageId="3" pageNumber="146" type="paratype">Paratypes</typeStatus>
.
</emphasis>
UCR 22424, an adult male; UCR 22423, an adult female; UCR 22425, a subadult female; UCR 22422 and UCR 22426, subadult males; and UCR 22427, a juvenile from
<collectingCountry id="F367DFBFFFB0FF885392F937FBB8159E" box="[977,1103,1729,1753]" name="Costa Rica" pageId="3" pageNumber="146">Costa Rica</collectingCountry>
:
<collectingRegion id="49B451CDFFB0FF88541DF937FA98159F" box="[1118,1391,1729,1753]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="146">Provincia de Puntarenas</collectingRegion>
:
<collectingMunicipality id="6BAB0555FFB0FF885354F928FBD115B1" box="[791,1062,1758,1782]" pageId="3" pageNumber="146">Cantón de Buenos Aires</collectingMunicipality>
:
<location id="8EAFC9F4FFB0FF885471F928FAB115B1" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB0FF885471F928FAB115B1" box="[1074,1350,1758,1782]" country="Costa Rica" municipality="Canton de Buenos Aires" name="Distrito de Buenos Aires" pageId="3" pageNumber="146" stateProvince="Puntarenas">Distrito de Buenos Aires</location>
: the summit of
<collectorName id="2685FAF9FFB0FF8853C0F90DFBE71453" box="[899,1040,1787,1812]" pageId="3" pageNumber="146">Cerro Utyum</collectorName>
,
<collectorName id="2685FAF9FFB0FF885454F90AFA981454" box="[1047,1391,1787,1811]" pageId="3" pageNumber="146">Parque Internacional LaAmistad</collectorName>
, (9.323º, -83.187º;
<elevation id="005D781CFFB0FF885399F8EFFBAF1476" box="[986,1112,1817,1841]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="3" pageNumber="146" unit="m" value="2870.0">
<quantity id="4C8832CAFFB0FF885399F8EFFBC61477" box="[986,1073,1817,1841]" metricMagnitude="3" metricUnit="m" metricValue="2.87" pageId="3" pageNumber="146" unit="m" value="2870.0">2,870 m</quantity>
asl
</elevation>
), collected by
<collectorName id="2685FAF9FFB0FF8854B0F8EFFA991476" box="[1267,1390,1817,1841]" pageId="3" pageNumber="146">Erick Arias</collectorName>
,
<collectorName id="2685FAF9FFB0FF885354F8C0FC381409" box="[791,975,1846,1870]" pageId="3" pageNumber="146">Gerardo Chaves</collectorName>
,
<collectorName id="2685FAF9FFB0FF8853A0F8C0FB661409" box="[995,1169,1846,1870]" pageId="3" pageNumber="146">Olmer Cordero</collectorName>
, and
<collectorName id="2685FAF9FFB0FF885498F8C0FA841409" box="[1243,1395,1846,1870]" pageId="3" pageNumber="146">Omar Zúñiga</collectorName>
on
<date id="FFCEB9EFFFB0FF885378F8A5FC1E142C" box="[827,1001,1875,1899]" pageId="3" pageNumber="146" value="2015-03-30">
<collectingDate id="EF8A4007FFB0FF885378F8A5FC1E142C" box="[827,1001,1875,1899]" pageId="3" pageNumber="146" value="2015-03-30">30 March 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B189572FFB0FF8F53B5F8A5FD5515B5" collectingDate="2015-03-30" collectorName="Erick Arias &amp; Gerardo Chaves &amp; Olmer Cordero &amp; Omar Zuniga" country="Costa Rica" county="Canton de Talamanca" elevation="2913" lastPageId="4" lastPageNumber="147" location="Cerro Utyum" municipality="Distrito de Telire" pageId="3" pageNumber="146" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
UCR 22421, an adult female from
<collectingCountry id="F367DFBFFFB0FF885354F887FC6614CE" box="[791,913,1905,1929]" name="Costa Rica" pageId="3" pageNumber="146">Costa Rica</collectingCountry>
:
<collectingRegion id="49B451CDFFB0FF8853DFF887FB8014CE" box="[924,1143,1905,1929]" country="Costa Rica" name="Limon" pageId="3" pageNumber="146">Provincia de Limón</collectingRegion>
:
<collectingCounty id="62AEE7A3FFB0FF8854C1F887FA9814CE" box="[1154,1391,1905,1929]" pageId="3" pageNumber="146">Cantón de Talamanca</collectingCounty>
:
<collectingMunicipality id="6BAB0555FFB0FF885354F878FC2114E1" box="[791,982,1934,1958]" pageId="3" pageNumber="146">Distrito de Telire</collectingMunicipality>
: the summit of
<location id="8EAFC9F4FFB0FF8854CBF878FAEC14E1" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB0FF8854CBF878FAEC14E1" box="[1160,1307,1934,1958]" country="Costa Rica" county="Canton de Talamanca" municipality="Distrito de Telire" name="Cerro Utyum" pageId="3" pageNumber="146" stateProvince="Limon">Cerro Utyum</location>
,
<location id="8EAFC9F4FFB0FF8F556AF878FE5D15F0" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB0FF8F556AF878FE5D15F0" country="Costa Rica" county="Canton de Talamanca" lastPageId="4" lastPageNumber="147" municipality="Distrito de Telire" name="Parque Internacional La Amistad" pageId="3" pageNumber="146" stateProvince="Limon">Parque Internacional La Amistad</location>
, (9.333º, -83.180º;
<elevation id="005D781CFFB7FF8F52CBF956FF591592" metricMagnitude="3" metricUnit="m" metricValue="2.913" pageId="4" pageNumber="147" unit="m" value="2913.0">
<quantity id="4C8832CAFFB7FF8F52CBF956FD1115F0" box="[648,742,1696,1720]" metricMagnitude="3" metricUnit="m" metricValue="2.913" pageId="4" pageNumber="147" unit="m" value="2913.0">2,913 m</quantity>
asl
</elevation>
), collected by
<collectorName id="2685FAF9FFB7FF8F5117F94BFE251592" box="[340,466,1725,1749]" pageId="4" pageNumber="147">Erick Arias</collectorName>
,
<collectorName id="2685FAF9FFB7FF8F51A2F94BFD641592" box="[481,659,1725,1749]" pageId="4" pageNumber="147">Gerardo Chaves</collectorName>
,
<collectorName id="2685FAF9FFB7FF8F52E2F94BFF1015B5" pageId="4" pageNumber="147">Olmer Cordero</collectorName>
, and
<collectorName id="2685FAF9FFB7FF8F5169F92CFE4815B5" box="[298,447,1754,1778]" pageId="4" pageNumber="147">Omar Zúñiga</collectorName>
on
<date id="FFCEB9EFFFB7FF8F51B2F92CFD5515B5" box="[497,674,1754,1778]" pageId="4" pageNumber="147" value="2015-03-30">
<collectingDate id="EF8A4007FFB7FF8F51B2F92CFD5515B5" box="[497,674,1754,1778]" pageId="4" pageNumber="147" value="2015-03-30">30 March 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B189572FFB7FF8F52F2F92CFD1214E5" collectingDate="2015-12-28" collectorName="Erick Arias &amp; Omar Zuniga" country="Costa Rica" county="Canton de Buenos Aires" elevation="2660" location="Parque Internacional La Amistad" municipality="Distrito de Buenos Aires" pageId="4" pageNumber="147" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
UCR 22745, an adult male; UCR 227414, adult females; and UCR 22746, a juvenile from
<collectingCountry id="F367DFBFFFB7FF8F5196F8E3FDA6146A" box="[469,593,1813,1837]" name="Costa Rica" pageId="4" pageNumber="147">Costa Rica</collectingCountry>
:
<collectingRegion id="49B451CDFFB7FF8F521CF8E3FEF1140D" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="147">Provincia de Puntarenas</collectingRegion>
:
<collectingCounty id="62AEE7A3FFB7FF8F5153F8C4FDE0140D" box="[272,535,1842,1866]" pageId="4" pageNumber="147">Cantón de Buenos Aires</collectingCounty>
:
<collectingMunicipality id="6BAB0555FFB7FF8F5263F8C4FF3C1420" pageId="4" pageNumber="147">Distrito de Buenos Aires</collectingMunicipality>
: the summit of Cerro Hakú,
<location id="8EAFC9F4FFB7FF8F524BF8A6FEE714C2" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB7FF8F524BF8A6FEE714C2" country="Costa Rica" county="Canton de Buenos Aires" municipality="Distrito de Buenos Aires" name="Parque Internacional La Amistad" pageId="4" pageNumber="147" stateProvince="Puntarenas">Parque Internacional La Amistad</location>
, (9.322º, -83.203º;
<elevation id="005D781CFFB7FF8F51A9F89BFD8614C2" box="[490,625,1901,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.66" pageId="4" pageNumber="147" unit="m" value="2660.0">
<quantity id="4C8832CAFFB7FF8F51A9F89BFDB214C3" box="[490,581,1901,1925]" metricMagnitude="3" metricUnit="m" metricValue="2.66" pageId="4" pageNumber="147" unit="m" value="2660.0">2,660 m</quantity>
asl
</elevation>
), collected by
<collectorName id="2685FAF9FFB7FF8F50ECF87CFEDE14E5" box="[175,297,1930,1954]" pageId="4" pageNumber="147">Erick Arias</collectorName>
and
<collectorName id="2685FAF9FFB7FF8F511EF87CFE1A14E5" box="[349,493,1930,1954]" pageId="4" pageNumber="147">Omar Zúñiga</collectorName>
on
<date id="FFCEB9EFFFB7FF8F5255F87CFD1214E5" box="[534,741,1930,1954]" pageId="4" pageNumber="147" value="2015-12-28">
<collectingDate id="EF8A4007FFB7FF8F5255F87CFD1214E5" box="[534,741,1930,1954]" pageId="4" pageNumber="147" value="2015-12-28">28 December 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B189572FFB7FF8F5354F956FB7F1420" collectingDate="2016-01-13" collectorName="Cerro Durika &amp; Omar Zuniga" country="Costa Rica" county="Canton de Buenos Aires" elevation="3240" location="Parque Internacional La Amistad" municipality="Distrito de Buenos Aires" pageId="4" pageNumber="147" specimenCount="1" stateProvince="Puntarenas" typeStatus="paratype">
UCR 24245, an adult female; UCR 24246, an adult male; UCR 24247, a subadult male; and UCR 24248, a juvenile from
<collectingCountry id="F367DFBFFFB7FF8F5388F92CFBB015B5" box="[971,1095,1754,1778]" name="Costa Rica" pageId="4" pageNumber="147">Costa Rica</collectingCountry>
:
<collectingRegion id="49B451CDFFB7FF8F5417F92CFA9615B5" box="[1108,1377,1754,1778]" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="147">Provincia de Puntarenas</collectingRegion>
:
<collectingCounty id="62AEE7A3FFB7FF8F5354F90EFBC61448" box="[791,1073,1783,1808]" pageId="4" pageNumber="147">Cantón de Buenos Aires</collectingCounty>
:
<collectingMunicipality id="6BAB0555FFB7FF8F5401F901FA961448" box="[1090,1377,1783,1807]" pageId="4" pageNumber="147">Distrito de Buenos Aires</collectingMunicipality>
: the summit of
<collectorName id="2685FAF9FFB7FF8F53FFF8E3FBB8146A" box="[956,1103,1813,1837]" pageId="4" pageNumber="147">Cerro Dúrika</collectorName>
,
<location id="8EAFC9F4FFB7FF8F541EF8E3FC83140D" LSID="urn:lsid:plazi:treatment:03D92E39FFB0FF805354FB65FE6615E1:8EAFC9F4FFB7FF8F541EF8E3FC83140D" country="Costa Rica" county="Canton de Buenos Aires" municipality="Distrito de Buenos Aires" name="Parque Internacional La Amistad" pageId="4" pageNumber="147" stateProvince="Puntarenas">Parque Internacional La Amistad</location>
, (9.374º, -83.303º;
<elevation id="005D781CFFB7FF8F540BF8C4FB3C140D" box="[1096,1227,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="4" pageNumber="147" unit="m" value="3240.0">
<quantity id="4C8832CAFFB7FF8F540BF8C4FB55140D" box="[1096,1186,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="4" pageNumber="147" unit="m" value="3240.0">3,240 m</quantity>
asl
</elevation>
), collected by
<collectorName id="2685FAF9FFB7FF8F5354F8A6FC511420" box="[791,934,1871,1896]" pageId="4" pageNumber="147">Omar Zúñiga</collectorName>
on
<date id="FFCEB9EFFFB7FF8F538DF8A6FB7F1420" box="[974,1160,1871,1896]" pageId="4" pageNumber="147" value="2016-01-13">
<collectingDate id="EF8A4007FFB7FF8F538DF8A6FB7F1420" box="[974,1160,1871,1896]" pageId="4" pageNumber="147" value="2016-01-13">13 January 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0FCFA7FFB7FF8F50CCF9F0FE9915C8" ID-DOI="http://doi.org/10.5281/zenodo.11390647" ID-Zenodo-Dep="11390647" httpUri="https://zenodo.org/record/11390647/files/figure.png" pageId="4" pageNumber="147" startId="4.[143,178,1542,1564]" targetBox="[157,1367,154,1523]" targetPageId="4" targetType="figure">
<paragraph id="8BCF9F2FFFB7FF8F50CCF9F0FE9915C8" blockId="4.[143,1393,1542,1679]" pageId="4" pageNumber="147">
<emphasis id="B904433DFFB7FF8F50CCF9F0FF27155C" bold="true" box="[143,208,1542,1564]" pageId="4" pageNumber="147">Fig. 2.</emphasis>
Bayesian phylogenetic inference shows the relationships of the
<taxonomicName id="4C70E4ACFFB7FF8F5305F9F0FBC2155C" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[838,1077,1542,1563]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="147" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B904433DFFB7FF8F5305F9F0FBC2155C" box="[838,1077,1542,1563]" italics="true" pageId="4" pageNumber="147">Bolitoglossa subpalmata</emphasis>
</taxonomicName>
species group based on the 16S and cyt
<emphasis id="B904433DFFB7FF8F5099F9D5FF11157F" box="[218,230,1571,1592]" italics="true" pageId="4" pageNumber="147">b</emphasis>
mitochondrial DNA gene fragments. Bootstrap proportions from maximum likelihood are shown above the branches, and numbers below the branches are posterior probabilities (multiplied by 100) from the MrBayes analysis. The scale bar refers to the estimated substitutions per site. The support values of any node within the species are not shown. The asterisks represent support&gt;99. NV = no voucher.
</paragraph>
</caption>
<subSubSection id="C36ACCA4FFB7FF8E5354F87CFBC413A3" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147" type="discussion">
<paragraph id="8BCF9F2FFFB7FF8E5354F87CFBC413A3" blockId="4.[791,1380,1930,1955]" lastBlockId="5.[791,1395,146,228]" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147">
<emphasis id="B904433DFFB7FF8F5354F87CFC0414E5" bold="true" box="[791,1011,1930,1954]" pageId="4" pageNumber="147">Generic Placement.</emphasis>
Assigned to the genus
<taxonomicName id="4C70E4ACFFB7FF8F549CF87CFA9314E5" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[1247,1380,1930,1954]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="4" pageNumber="147" phylum="Chordata" rank="genus">
<emphasis id="B904433DFFB7FF8F549CF87CFA9314E5" box="[1247,1380,1930,1954]" italics="true" pageId="4" pageNumber="147">Bolitoglossa</emphasis>
</taxonomicName>
due to having 14 costal grooves and lacking a sublingual fold, and to the subgenus
<taxonomicName id="4C70E4ACFFB6FF8E545DFF59FB891380" box="[1054,1150,175,199]" class="Amphibia" family="Plethodontidae" genus="Eladinea" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="genus">
<emphasis id="B904433DFFB6FF8E545DFF59FB891380" box="[1054,1150,175,199]" italics="true" pageId="5" pageNumber="148">Eladinea</emphasis>
</taxonomicName>
based on the molecular evidence presented herein.
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFB6FF835354FEF1FDFE128C" lastPageId="8" lastPageNumber="151" pageId="5" pageNumber="148" type="diagnosis">
<paragraph id="8BCF9F2FFFB6FF8E5354FEF1FACC114D" blockId="5.[791,1395,263,522]" pageId="5" pageNumber="148">
<emphasis id="B904433DFFB6FF8E5354FEF1FC7C1258" bold="true" box="[791,907,263,287]" pageId="5" pageNumber="148">Diagnosis.</emphasis>
The combination of the following characteristics can be used to distinguish
<taxonomicName id="4C70E4ACFFB6FF8E54AEFED2FC84121D" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E54AEFED2FC84121D" italics="true" pageId="5" pageNumber="148">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
from the other described species of the genus
<taxonomicName id="4C70E4ACFFB6FF8E5354FEA9FC6B1230" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[791,924,351,375]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="genus">
<emphasis id="B904433DFFB6FF8E5354FEA9FC6B1230" box="[791,924,351,375]" italics="true" pageId="5" pageNumber="148">Bolitoglossa</emphasis>
</taxonomicName>
: (1) having broad hands and feet, with the distal phalanges on the fingers and toes free of palmar and plantar tissue; (2) dorsal coloration highly variable, rarely black brownish uniform and usually mottled with yellow spots, but never with red on hind limbs or forelimbs; and (3) 16S and cyt
<emphasis id="B904433DFFB6FF8E5421FE04FB98114D" box="[1122,1135,498,522]" italics="true" pageId="5" pageNumber="148">b</emphasis>
mtDNA distances.
</paragraph>
<paragraph id="8BCF9F2FFFB6FF8E5354FDDAFBC6102E" blockId="5.[791,1395,556,1959]" pageId="5" pageNumber="148">
<emphasis id="B904433DFFB6FF8E5354FDDAFC421103" bold="true" box="[791,949,556,580]" pageId="5" pageNumber="148">Comparisons.</emphasis>
<taxonomicName id="4C70E4ACFFB6FF8E5380FDDAFB431103" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[963,1204,556,580]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E5380FDDAFB431103" box="[963,1204,556,580]" italics="true" pageId="5" pageNumber="148">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
is differentiated from members of the subgenus
<taxonomicName id="4C70E4ACFFB6FF8E542DFDBCFB391125" box="[1134,1230,586,610]" class="Amphibia" family="Plethodontidae" genus="Eladinea" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="genus">
<emphasis id="B904433DFFB6FF8E542DFDBCFB391125" box="[1134,1230,586,610]" italics="true" pageId="5" pageNumber="148">Eladinea</emphasis>
</taxonomicName>
by its 16S and cyt
<emphasis id="B904433DFFB6FF8E537DFD91FCBC1138" box="[830,843,615,639]" italics="true" pageId="5" pageNumber="148">b</emphasis>
mtDNA distances. Since
<taxonomicName id="4C70E4ACFFB6FF8E541EFD9EFB211138" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[1117,1238,615,639]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E541EFD9EFB211138" box="[1117,1238,615,639]" italics="true" pageId="5" pageNumber="148">B. bolanosi</emphasis>
</taxonomicName>
is only known to occur in
<collectingCountry id="F367DFBFFFB6FF8E53D7FD72FBFD11DB" box="[916,1034,644,668]" name="Costa Rica" pageId="5" pageNumber="148">Costa Rica</collectingCountry>
and molecular evidence strongly supports it forming part of the
<taxonomicName id="4C70E4ACFFB6FF8E542BFD54FA8511FD" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[1128,1394,674,698]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B904433DFFB6FF8E542BFD54FA8511FD" box="[1128,1394,674,698]" italics="true" pageId="5" pageNumber="148">Bolitoglossa subpalmata</emphasis>
</taxonomicName>
species group within the subgenus
<taxonomicName id="4C70E4ACFFB6FF8E54CDFD49FB191190" box="[1166,1262,703,727]" class="Amphibia" family="Plethodontidae" genus="Eladinea" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="genus">
<emphasis id="B904433DFFB6FF8E54CDFD49FB191190" box="[1166,1262,703,727]" italics="true" pageId="5" pageNumber="148">Eladinea</emphasis>
</taxonomicName>
, phenotypic comparisons are presented here only concerning the members of that clade (
<emphasis id="B904433DFFB6FF8E5452FD0CFAB11068" italics="true" pageId="5" pageNumber="148">
<taxonomicName id="4C70E4ACFFB6FF8E5452FD0CFB8A1055" box="[1041,1149,762,786]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bramei">B. bramei</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E54C4FD0CFB041056" box="[1159,1267,762,785]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gomezi">B. gomezi</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E54BEFD0CFA981055" box="[1277,1391,762,786]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">B. gracilis</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E5354FCEEFC8B1068" box="[791,892,791,815]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="kamuk">B. kamuk</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E53C4FCEEFBFF1068" baseAuthorityName="Taylor" baseAuthorityYear="1952" box="[903,1032,791,815]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="pesrubra">B. pesrubra</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E5451FCEEFB6D1068" box="[1042,1178,791,815]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="splendida">B. splendida</taxonomicName>
,
<taxonomicName id="4C70E4ACFFB6FF8E54E6FCEEFAB61068" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[1189,1345,791,815]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="subpalmata">B. subpalmata</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C70E4ACFFB6FF8E5354FCC3FCAA100B" box="[791,861,821,844]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="tica">
<emphasis id="B904433DFFB6FF8E5354FCC3FCAA100B" box="[791,861,821,844]" italics="true" pageId="5" pageNumber="148">B. tica</emphasis>
</taxonomicName>
), which are endemic to mountain ranges of
<collectingCountry id="F367DFBFFFB6FF8E5575FCC2FCBF102D" name="Costa Rica" pageId="5" pageNumber="148">Costa Rica</collectingCountry>
and western
<collectingCountry id="F367DFBFFFB6FF8E5396FCA4FBDA102E" box="[981,1069,850,873]" name="Panama" pageId="5" pageNumber="148">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8BCF9F2FFFB6FF8E5374FC99FB50159D" blockId="5.[791,1395,556,1959]" pageId="5" pageNumber="148">
Contrasting characteristics for
<taxonomicName id="4C70E4ACFFB6FF8E54CBFC99FA8410C0" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[1160,1395,879,903]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E54CBFC99FA8410C0" box="[1160,1395,879,903]" italics="true" pageId="5" pageNumber="148">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
are presented in parentheses.
<taxonomicName id="4C70E4ACFFB6FF8E5415FC7AFAD810E3" box="[1110,1327,908,932]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bramei">
<emphasis id="B904433DFFB6FF8E5415FC7AFAD810E3" box="[1110,1327,908,932]" italics="true" pageId="5" pageNumber="148">Bolitoglossa bramei</emphasis>
</taxonomicName>
<bibRefCitation id="EFE1E2DEFFB6FF8E5574FC7AFC681085" author="Wake DB &amp; Savage JM &amp; Hanken J." pageId="5" pageNumber="148" pagination="556 - 565" refId="ref12264" refString="Wake DB, Savage JM, Hanken J. 2007. Montane salamanders from the Costa Rica-Panama border region, with descriptions of two new species of Bolitoglossa. Copeia 2007: 556 - 565." type="journal article" year="2007">
<collectingCountry id="F367DFBFFFB6FF8E5574FC7AFA8410E3" box="[1335,1395,908,932]" name="Wake Island" pageId="5" pageNumber="148">Wake</collectingCountry>
et al., 2007
</bibRefCitation>
can be distinguished from
<taxonomicName id="4C70E4ACFFB6FF8E54B3FC5CFA841085" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[1264,1395,938,962]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E54B3FC5CFA841085" box="[1264,1395,938,962]" italics="true" pageId="5" pageNumber="148">B. bolanosi</emphasis>
</taxonomicName>
by having a rounded snout in males (snout strongly truncated in males); dorsal ground color dark brown to brownish red, usually with darker mottling or frosting of silvery-gray (dorsal ground color black to dark brown, usually with mottling or blotches of yellow to red).
<taxonomicName id="4C70E4ACFFB6FF8E5354FBACFC031736" box="[791,1012,1114,1138]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gomezi">
<emphasis id="B904433DFFB6FF8E5354FBACFC031736" box="[791,1012,1114,1138]" italics="true" pageId="5" pageNumber="148">Bolitoglossa gomezi</emphasis>
</taxonomicName>
<bibRefCitation id="EFE1E2DEFFB6FF8E53BCFBACFB311735" author="Wake DB &amp; Savage JM &amp; Hanken J." box="[1023,1222,1114,1138]" pageId="5" pageNumber="148" pagination="556 - 565" refId="ref12264" refString="Wake DB, Savage JM, Hanken J. 2007. Montane salamanders from the Costa Rica-Panama border region, with descriptions of two new species of Bolitoglossa. Copeia 2007: 556 - 565." type="journal article" year="2007">
<collectingCountry id="F367DFBFFFB6FF8E53BCFBACFBCC1735" box="[1023,1083,1114,1138]" name="Wake Island" pageId="5" pageNumber="148">Wake</collectingCountry>
et al., 2007
</bibRefCitation>
has fore limbs relatively shorter, FLL/SL 20% (FLL/SL 22.326.1%), snout rounded (snout truncate in males).
<taxonomicName id="4C70E4ACFFB6FF8E54AEFB62FB9F178D" authority="Bolanos et al., 1987" authorityName="Bolanos" authorityYear="1987" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B904433DFFB6FF8E54AEFB62FC9F178D" italics="true" pageId="5" pageNumber="148">Bolitoglossa gracilis</emphasis>
Bolaños et al., 1987
</taxonomicName>
has dorsum yellowish ground color and a distinct dark midventral stripe (dorsum variable but never with yellowish ground color and never with a distinct midventral stripe).
<taxonomicName id="4C70E4ACFFB6FF8E54AEFAFCFB551678" authority="Boza-Oviedo et al., 2012" authorityName="Boza-Oviedo" authorityYear="2012" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="splendida">
<emphasis id="B904433DFFB6FF8E54AEFAFCFC881678" italics="true" pageId="5" pageNumber="148">Bolitoglossa splendida</emphasis>
<bibRefCitation id="EFE1E2DEFFB6FF8E53C9FAD1FB551678" author="Boza-Oviedo E &amp; Rovito SM &amp; Chaves G &amp; Garcia-Rodriguez A &amp; Artavia LG &amp; Bolanos F &amp; Wake DB" box="[906,1186,1319,1343]" pageId="5" pageNumber="148" pagination="36 - 61" refId="ref11034" refString="Boza-Oviedo E, Rovito SM, Chaves G, Garcia-Rodriguez A, Artavia LG, Bolanos F, Wake DB. 2012. Salamanders from the eastern Cordillera de Talamanca, Costa Rica, with descriptions of five new species (Plethodontidae: Bolitoglossa, Nototriton, and Oedipina) and natural history notes from recent expeditions. Zootaxa 3309: 36 - 61." type="journal article" year="2012">Boza-Oviedo et al., 2012</bibRefCitation>
</taxonomicName>
has a shiny black dorsum with a bright reddish-orange broad dorsal band extending from the back of the head to the base of the tail, and with bright enamel-yellow spots scattered along the lateral and ventrolateral surfaces (dorsum variable but none with a reddish dorsal band extending from the head to the tail).
<taxonomicName id="4C70E4ACFFB6FF8E539EFA21FCA2154B" authority="(Boulenger, 1896)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B904433DFFB6FF8E539EFA21FB1E16A8" box="[989,1257,1495,1519]" italics="true" pageId="5" pageNumber="148">Bolitoglossa subpalmata</emphasis>
(Boulenger, 1896)
</taxonomicName>
has forelimbs that are relatively shorter, FLL/SL 18.723.1% (FLL/SL 22.326.1%).
<taxonomicName id="4C70E4ACFFB6FF8E54F6F9E4FBED1500" authority="Garcia-Paris et al., 2008" authorityName="Garcia-Paris" authorityYear="2008" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="tica">
<emphasis id="B904433DFFB6FF8E54F6F9E4FA84156E" box="[1205,1395,1554,1578]" italics="true" pageId="5" pageNumber="148">Bolitoglossa tica</emphasis>
<bibRefCitation id="EFE1E2DEFFB6FF8E5354F9D9FBED1500" author="Garcia-Paris M &amp; Parra-Olea G &amp; Wake DB" box="[791,1050,1583,1607]" pageId="5" pageNumber="148" pagination="23 - 31" refId="ref11269" refString="Garcia-Paris M, Parra-Olea G, Wake DB. 2008. Description of a new species of the Bolitoglossa subpalmata group (Caudata: Plethodontidae) from Costa Rica. The Herpetological Journal 18: 23 - 31." type="journal article" year="2008">García-París et al., 2008</bibRefCitation>
</taxonomicName>
has dorsal ground color usually a uniform reddish brown with a darker tail, rarely with mottling or blotches contrasting (dorsal ground color black to dark brown, usually with yellow to red mottling or blotchy contrasting); prominent whitish spots on the venter (venter without whitish spots).
</paragraph>
<paragraph id="8BCF9F2FFFB6FF8E50A8F97AFD45126F" pageId="5" pageNumber="148">
<table id="F9706D8FFFB6007450A8FF59FD4014E3" box="[235,695,175,1956]" gridcols="10" gridrows="10" pageId="5" pageNumber="148" textDirection="bu">
<tr id="35409D6DFFB6007450A8FF59FEF514E3" box="[235,258,175,1956]" gridrow="0" pageId="5" pageNumber="148" rowspan-0="1" textDirection="bu">
<th id="7691F411FFB6007450A8F9D3FEF515DA" box="[235,258,1573,1693]" gridcol="1" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E50A8F97AFEF515DA" box="[235,258,1676,1693]" italics="true" pageId="5" pageNumber="148">B</emphasis>
.
<taxonomicName id="4C70E4ACFFB6FF8E50A8F9D3FEF515C5" box="[235,258,1573,1666]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="splendida">splendida</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FA8BFEF5169C" box="[235,258,1405,1499]" gridcol="2" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
<taxonomicName id="4C70E4ACFFB6FF8E50A8FA3CFEF51686" box="[235,258,1405,1499]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bramei">B. bramei</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FB1BFEF5160B" box="[235,258,1261,1356]" gridcol="3" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FB1BFEF51676" box="[235,258,1261,1329]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gomezi">gomezi</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FB92FEF5178F" box="[235,258,1124,1224]" gridcol="4" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E50A8FB41FEF5178F" box="[235,258,1207,1224]" italics="true" pageId="5" pageNumber="148">B</emphasis>
.
<emphasis id="B904433DFFB6FF8E50A8FB92FEF517EA" box="[235,258,1124,1197]" italics="true" pageId="5" pageNumber="148">gracilis</emphasis>
</th>
<th id="7691F411FFB6007450A8FC69FEF5176E" box="[235,258,927,1065]" gridcol="5" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FC69FEF51748" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[235,258,927,1039]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="subpalmata">
<emphasis id="B904433DFFB6FF8E50A8FC69FEF51748" box="[235,258,927,1039]" italics="true" pageId="5" pageNumber="148">subpalmata</emphasis>
</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FCE7FEF51008" box="[235,258,785,847]" gridcol="6" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FCE7FEF51072" box="[235,258,785,821]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="tica">tica</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FD9CFEF51184" box="[235,258,618,707]" gridcol="7" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E50A8FD44FEF51184" box="[235,258,690,707]" italics="true" pageId="5" pageNumber="148">B</emphasis>
.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FD9CFEF511EE" box="[235,258,618,681]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="kamuk">kamuk</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FE58FEF51158" box="[235,258,430,543]" gridcol="8" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FE58FEF51142" baseAuthorityName="Taylor" baseAuthorityYear="1952" box="[235,258,430,517]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="pesrubra">pesrubra</taxonomicName>
</th>
<th id="7691F411FFB6007450A8FF59FEF51228" box="[235,258,175,367]" gridcol="9" gridrow="0" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E50A8FEA8FEF51228" box="[235,258,350,367]" italics="true" pageId="5" pageNumber="148">B</emphasis>
.
<taxonomicName id="4C70E4ACFFB6FF8E50A8FEF4FEF51212" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[235,258,258,341]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E50A8FEF4FEF51212" box="[235,258,258,341]" italics="true" pageId="5" pageNumber="148">bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB6FF8E50A8FF12FEF513BB" bold="true" box="[235,258,228,252]" pageId="5" pageNumber="148">sp</emphasis>
<emphasis id="B904433DFFB6FF8E50A8FF29FEF513A3" bold="true" box="[235,258,223,228]" pageId="5" pageNumber="148">.</emphasis>
<emphasis id="B904433DFFB6FF8E50A8FF42FEF5139D" bold="true" box="[235,258,180,218]" pageId="5" pageNumber="148">nov</emphasis>
.
</th>
</tr>
<tr id="35409D6DFFB600745158FF59FEC014E3" box="[283,311,175,1956]" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745158F912FEC014E3" box="[283,311,1764,1956]" gridcol="0" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5158F8DAFEC514CE" box="[283,306,1836,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="splendida">splendida</taxonomicName>
</th>
<td id="7691F411FFB600745158F9D3FEC015DA" box="[283,311,1573,1693]" gridcol="1" gridrow="1" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745158FA8BFEC0169C" box="[283,311,1405,1499]" gridcol="2" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">3.88</td>
<td id="7691F411FFB600745158FB1BFEC0160B" box="[283,311,1261,1356]" gridcol="3" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">3.49</td>
<td id="7691F411FFB600745158FB92FEC0178F" box="[283,311,1124,1224]" gridcol="4" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">4.42</td>
<td id="7691F411FFB600745158FC69FEC0176E" box="[283,311,927,1065]" gridcol="5" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">2.63</td>
<td id="7691F411FFB600745158FCE7FEC01008" box="[283,311,785,847]" gridcol="6" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">4.19</td>
<td id="7691F411FFB600745158FD9CFEC01184" box="[283,311,618,707]" gridcol="7" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">2.53</td>
<td id="7691F411FFB600745158FE58FEC01158" box="[283,311,430,543]" gridcol="8" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">2.74</td>
<td id="7691F411FFB600745158FF59FEC01228" box="[283,311,175,367]" gridcol="9" gridrow="1" pageId="5" pageNumber="148" textDirection="bu">3.14</td>
</tr>
<tr id="35409D6DFFB600745108FF59FE9014E3" box="[331,359,175,1956]" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745108F912FE9014E3" box="[331,359,1764,1956]" gridcol="0" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5108F8B3FE9514CE" box="[331,354,1861,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bramei">bramei</taxonomicName>
</th>
<td id="7691F411FFB600745108F9D3FE9015DA" box="[331,359,1573,1693]" gridcol="1" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">10.01</td>
<td id="7691F411FFB600745108FA8BFE90169C" box="[331,359,1405,1499]" gridcol="2" gridrow="2" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745108FB1BFE90160B" box="[331,359,1261,1356]" gridcol="3" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">3.36</td>
<td id="7691F411FFB600745108FB92FE90178F" box="[331,359,1124,1224]" gridcol="4" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">3.20</td>
<td id="7691F411FFB600745108FC69FE90176E" box="[331,359,927,1065]" gridcol="5" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">2.66</td>
<td id="7691F411FFB600745108FCE7FE901008" box="[331,359,785,847]" gridcol="6" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">3.76</td>
<td id="7691F411FFB600745108FD9CFE901184" box="[331,359,618,707]" gridcol="7" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">2.32</td>
<td id="7691F411FFB600745108FE58FE901158" box="[331,359,430,543]" gridcol="8" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">3.36</td>
<td id="7691F411FFB600745108FF59FE901228" box="[331,359,175,367]" gridcol="9" gridrow="2" pageId="5" pageNumber="148" textDirection="bu">3.44</td>
</tr>
<tr id="35409D6DFFB600745138FF59FE6014E3" box="[379,407,175,1956]" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745138F912FE6014E3" box="[379,407,1764,1956]" gridcol="0" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5138F8B3FE6514CE" box="[379,402,1861,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gomezi">gomezi</taxonomicName>
</th>
<td id="7691F411FFB600745138F9D3FE6015DA" box="[379,407,1573,1693]" gridcol="1" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">10.10</td>
<td id="7691F411FFB600745138FA8BFE60169C" box="[379,407,1405,1499]" gridcol="2" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">5.95</td>
<td id="7691F411FFB600745138FB1BFE60160B" box="[379,407,1261,1356]" gridcol="3" gridrow="3" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745138FB92FE60178F" box="[379,407,1124,1224]" gridcol="4" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">4.58</td>
<td id="7691F411FFB600745138FC69FE60176E" box="[379,407,927,1065]" gridcol="5" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">3.09</td>
<td id="7691F411FFB600745138FCE7FE601008" box="[379,407,785,847]" gridcol="6" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">4.16</td>
<td id="7691F411FFB600745138FD9CFE601184" box="[379,407,618,707]" gridcol="7" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">2.56</td>
<td id="7691F411FFB600745138FE58FE601158" box="[379,407,430,543]" gridcol="8" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">3.60</td>
<td id="7691F411FFB600745138FF59FE601228" box="[379,407,175,367]" gridcol="9" gridrow="3" pageId="5" pageNumber="148" textDirection="bu">3.99</td>
</tr>
<tr id="35409D6DFFB6007451E8FF59FE3014E3" box="[427,455,175,1956]" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB6007451E8F912FE3014E3" box="[427,455,1764,1956]" gridcol="0" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E51E8F8B6FE3514CE" box="[427,450,1856,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="gracilis">gracilis</taxonomicName>
</th>
<td id="7691F411FFB6007451E8F9D3FE3015DA" box="[427,455,1573,1693]" gridcol="1" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">9.30</td>
<td id="7691F411FFB6007451E8FA8BFE30169C" box="[427,455,1405,1499]" gridcol="2" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">7.84</td>
<td id="7691F411FFB6007451E8FB1BFE30160B" box="[427,455,1261,1356]" gridcol="3" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">7.06</td>
<td id="7691F411FFB6007451E8FB92FE30178F" box="[427,455,1124,1224]" gridcol="4" gridrow="4" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB6007451E8FC69FE30176E" box="[427,455,927,1065]" gridcol="5" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">2.75</td>
<td id="7691F411FFB6007451E8FCE7FE301008" box="[427,455,785,847]" gridcol="6" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">3.73</td>
<td id="7691F411FFB6007451E8FD9CFE301184" box="[427,455,618,707]" gridcol="7" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">3.63</td>
<td id="7691F411FFB6007451E8FE58FE301158" box="[427,455,430,543]" gridcol="8" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">4.61</td>
<td id="7691F411FFB6007451E8FF59FE301228" box="[427,455,175,367]" gridcol="9" gridrow="4" pageId="5" pageNumber="148" textDirection="bu">4.71</td>
</tr>
<tr id="35409D6DFFB600745198FF59FE0014E3" box="[475,503,175,1956]" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745198F912FE0014E3" box="[475,503,1764,1956]" gridcol="0" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5198F8EFFE0514CE" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[475,498,1817,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="subpalmata">subpalmata</taxonomicName>
</th>
<td id="7691F411FFB600745198F9D3FE0015DA" box="[475,503,1573,1693]" gridcol="1" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">9.53</td>
<td id="7691F411FFB600745198FA8BFE00169C" box="[475,503,1405,1499]" gridcol="2" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">7.71</td>
<td id="7691F411FFB600745198FB1BFE00160B" box="[475,503,1261,1356]" gridcol="3" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">6.96</td>
<td id="7691F411FFB600745198FB92FE00178F" box="[475,503,1124,1224]" gridcol="4" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">8.12</td>
<td id="7691F411FFB600745198FC69FE00176E" box="[475,503,927,1065]" gridcol="5" gridrow="5" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745198FCE7FE001008" box="[475,503,785,847]" gridcol="6" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">1.94</td>
<td id="7691F411FFB600745198FD9CFE001184" box="[475,503,618,707]" gridcol="7" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">1.75</td>
<td id="7691F411FFB600745198FE58FE001158" box="[475,503,430,543]" gridcol="8" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">2.71</td>
<td id="7691F411FFB600745198FF59FE001228" box="[475,503,175,367]" gridcol="9" gridrow="5" pageId="5" pageNumber="148" textDirection="bu">2.82</td>
</tr>
<tr id="35409D6DFFB600745248FF59FDD014E3" box="[523,551,175,1956]" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745248F912FDD014E3" box="[523,551,1764,1956]" gridcol="0" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5248F893FDD514CE" box="[523,546,1893,1929]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="tica">tica</taxonomicName>
</th>
<td id="7691F411FFB600745248F9D3FDD015DA" box="[523,551,1573,1693]" gridcol="1" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">9.47</td>
<td id="7691F411FFB600745248FA8BFDD0169C" box="[523,551,1405,1499]" gridcol="2" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">6.60</td>
<td id="7691F411FFB600745248FB1BFDD0160B" box="[523,551,1261,1356]" gridcol="3" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">6.63</td>
<td id="7691F411FFB600745248FB92FDD0178F" box="[523,551,1124,1224]" gridcol="4" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">7.29</td>
<td id="7691F411FFB600745248FC69FDD0176E" box="[523,551,927,1065]" gridcol="5" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">6.48</td>
<td id="7691F411FFB600745248FCE7FDD01008" box="[523,551,785,847]" gridcol="6" gridrow="6" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745248FD9CFDD01184" box="[523,551,618,707]" gridcol="7" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">3.21</td>
<td id="7691F411FFB600745248FE58FDD01158" box="[523,551,430,543]" gridcol="8" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">3.88</td>
<td id="7691F411FFB600745248FF59FDD01228" box="[523,551,175,367]" gridcol="9" gridrow="6" pageId="5" pageNumber="148" textDirection="bu">3.39</td>
</tr>
<tr id="35409D6DFFB600745278FF59FDA014E3" box="[571,599,175,1956]" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745278F912FDA014E3" box="[571,599,1764,1956]" gridcol="0" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5278F8BDFDA514CD" box="[571,594,1867,1930]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="kamuk">kamuk</taxonomicName>
</th>
<td id="7691F411FFB600745278F9D3FDA015DA" box="[571,599,1573,1693]" gridcol="1" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">10.29</td>
<td id="7691F411FFB600745278FA8BFDA0169C" box="[571,599,1405,1499]" gridcol="2" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">6.56</td>
<td id="7691F411FFB600745278FB1BFDA0160B" box="[571,599,1261,1356]" gridcol="3" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">6.71</td>
<td id="7691F411FFB600745278FB92FDA0178F" box="[571,599,1124,1224]" gridcol="4" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">7.75</td>
<td id="7691F411FFB600745278FC69FDA0176E" box="[571,599,927,1065]" gridcol="5" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">8.46</td>
<td id="7691F411FFB600745278FCE7FDA01008" box="[571,599,785,847]" gridcol="6" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">7.43</td>
<td id="7691F411FFB600745278FD9CFDA01184" box="[571,599,618,707]" gridcol="7" gridrow="7" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745278FE58FDA01158" box="[571,599,430,543]" gridcol="8" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">1.25</td>
<td id="7691F411FFB600745278FF59FDA01228" box="[571,599,175,367]" gridcol="9" gridrow="7" pageId="5" pageNumber="148" textDirection="bu">1.45</td>
</tr>
<tr id="35409D6DFFB600745228FF59FD7014E3" box="[619,647,175,1956]" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB600745228F912FD7014E3" box="[619,647,1764,1956]" gridcol="0" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E5228F865FD7514CD" box="[619,642,1843,1956]" italics="true" pageId="5" pageNumber="148">
B.
<taxonomicName id="4C70E4ACFFB6FF8E5228F8C5FD7514CD" baseAuthorityName="Taylor" baseAuthorityYear="1952" box="[619,642,1843,1930]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="pesrubra">pesrubra</taxonomicName>
</emphasis>
</th>
<td id="7691F411FFB600745228F9D3FD7015DA" box="[619,647,1573,1693]" gridcol="1" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">9.11</td>
<td id="7691F411FFB600745228FA8BFD70169C" box="[619,647,1405,1499]" gridcol="2" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">6.51</td>
<td id="7691F411FFB600745228FB1BFD70160B" box="[619,647,1261,1356]" gridcol="3" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">6.72</td>
<td id="7691F411FFB600745228FB92FD70178F" box="[619,647,1124,1224]" gridcol="4" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">7.35</td>
<td id="7691F411FFB600745228FC69FD70176E" box="[619,647,927,1065]" gridcol="5" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">7.78</td>
<td id="7691F411FFB600745228FCE7FD701008" box="[619,647,785,847]" gridcol="6" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">7.29</td>
<td id="7691F411FFB600745228FD9CFD701184" box="[619,647,618,707]" gridcol="7" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">7.06</td>
<td id="7691F411FFB600745228FE58FD701158" box="[619,647,430,543]" gridcol="8" gridrow="8" pageId="5" pageNumber="148" textDirection="bu"></td>
<td id="7691F411FFB600745228FF59FD701228" box="[619,647,175,367]" gridcol="9" gridrow="8" pageId="5" pageNumber="148" textDirection="bu">1.69</td>
</tr>
<tr id="35409D6DFFB6007452D8FF59FD4014E3" box="[667,695,175,1956]" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">
<th id="7691F411FFB6007452D8F912FD4014E3" box="[667,695,1764,1956]" gridcol="0" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">
<emphasis id="B904433DFFB6FF8E52D8F865FD4514CD" box="[667,690,1847,1956]" italics="true" pageId="5" pageNumber="148">
B.
<taxonomicName id="4C70E4ACFFB6FF8E52D8F8C1FD4514CD" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[667,690,1847,1930]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">bolanosi</taxonomicName>
</emphasis>
<emphasis id="B904433DFFB6FF8E52D8F8EFFD451476" bold="true" box="[667,690,1817,1841]" pageId="5" pageNumber="148">sp</emphasis>
. nov.
</th>
<td id="7691F411FFB6007452D8F9D3FD4015DA" box="[667,695,1573,1693]" gridcol="1" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">9.09</td>
<td id="7691F411FFB6007452D8FA8BFD40169C" box="[667,695,1405,1499]" gridcol="2" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">6.50</td>
<td id="7691F411FFB6007452D8FB1BFD40160B" box="[667,695,1261,1356]" gridcol="3" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">6.34</td>
<td id="7691F411FFB6007452D8FB92FD40178F" box="[667,695,1124,1224]" gridcol="4" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">7.04</td>
<td id="7691F411FFB6007452D8FC69FD40176E" box="[667,695,927,1065]" gridcol="5" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">7.57</td>
<td id="7691F411FFB6007452D8FCE7FD401008" box="[667,695,785,847]" gridcol="6" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">6.51</td>
<td id="7691F411FFB6007452D8FD9CFD401184" box="[667,695,618,707]" gridcol="7" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">5.87</td>
<td id="7691F411FFB6007452D8FE58FD401158" box="[667,695,430,543]" gridcol="8" gridrow="9" pageId="5" pageNumber="148" textDirection="bu">5.03</td>
<td id="7691F411FFB6007452D8FF59FD401228" box="[667,695,175,367]" gridcol="9" gridrow="9" pageId="5" pageNumber="148" textDirection="bu"></td>
</tr>
</table>
</paragraph>
<paragraph id="8BCF9F2FFFB6FF835374F929FDFE128C" blockId="5.[791,1395,556,1959]" lastBlockId="8.[144,750,142,459]" lastPageId="8" lastPageNumber="151" pageId="5" pageNumber="148">
<taxonomicName id="4C70E4ACFFB6FF8E5374F929FBEA15B0" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[823,1053,1759,1783]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB6FF8E5374F929FBEA15B0" box="[823,1053,1759,1783]" italics="true" pageId="5" pageNumber="148">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
differs from its closest relatives as follows:
<taxonomicName id="4C70E4ACFFB6FF8E53D2F90AFA841453" authority="Boza-Oviedo et al., 2012" authorityName="Boza-Oviedo" authorityYear="2012" box="[913,1395,1788,1812]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B904433DFFB6FF8E53D2F90AFB961453" box="[913,1121,1788,1812]" italics="true" pageId="5" pageNumber="148">Bolitoglossa kamuk</emphasis>
<bibRefCitation id="EFE1E2DEFFB6FF8E542BF90AFA841453" author="Boza-Oviedo E &amp; Rovito SM &amp; Chaves G &amp; Garcia-Rodriguez A &amp; Artavia LG &amp; Bolanos F &amp; Wake DB" box="[1128,1395,1788,1812]" pageId="5" pageNumber="148" pagination="36 - 61" refId="ref11034" refString="Boza-Oviedo E, Rovito SM, Chaves G, Garcia-Rodriguez A, Artavia LG, Bolanos F, Wake DB. 2012. Salamanders from the eastern Cordillera de Talamanca, Costa Rica, with descriptions of five new species (Plethodontidae: Bolitoglossa, Nototriton, and Oedipina) and natural history notes from recent expeditions. Zootaxa 3309: 36 - 61." type="journal article" year="2012">Boza-Oviedo et al., 2012</bibRefCitation>
</taxonomicName>
is smaller and more slender with mean SL
<quantity id="4C8832CAFFB6FF8E549AF8ECFA851476" box="[1241,1394,1818,1842]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.84" metricValueMin="3.46" pageId="5" pageNumber="148" unit="mm" value="36.5" valueMax="38.4" valueMin="34.6">34.638.4 mm</quantity>
(larger and robust, SL =
<quantity id="4C8832CAFFB6FF8E5471F8C1FB071408" box="[1074,1264,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="4.4719999999999995" metricValueMax="5.0009999999999994" metricValueMin="3.943" pageId="5" pageNumber="148" unit="mm" value="44.72" valueMax="50.01" valueMin="39.43">39.4350.01 mm</quantity>
); internarial distance relatively shorter, IND/HeW = 0.26±0.05 (internarial distance relatively longer, IND/HeW = 0.31±0.05); shorter tail TL/SL 9899% (tail long, TL/SL 102.3117.2%); dorsal ground color relatively uniform, orange to black (dorsal ground usually mottled or blotchy); males with rounded snout (snout strongly truncated in males).
<taxonomicName id="4C70E4ACFFBBFF8350A4FF10FD8F13BA" authority="(Taylor, 1952)" baseAuthorityName="Taylor" baseAuthorityYear="1952" box="[231,632,229,254]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="151" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B904433DFFBBFF8350A4FF10FE2313B9" box="[231,468,230,254]" italics="true" pageId="8" pageNumber="151">Bolitoglossa pesrubra</emphasis>
(Taylor, 1952)
</taxonomicName>
has hands and feet more webbed (
<figureCitation id="134B83AAFFBBFF8351CEFEF5FDF6125C" box="[397,513,259,283]" captionStart="Fig" captionStartId="10.[141,176,1873,1895]" captionTargetBox="[141,1394,786,1860]" captionTargetId="figure-43@10.[141,1394,786,1860]" captionTargetPageId="10" captionText="Fig. 5. Dorsal views of the hand and foot of the holotypes of (AB) Bolitoglossa bolanosi sp. nov. (UCR 22965) and (CD) B. pesrubra in preservation. Photograph by Erick Arias." figureDoi="http://doi.org/10.5281/zenodo.11390655" httpUri="https://zenodo.org/record/11390655/files/figure.png" pageId="8" pageNumber="151">Fig. 5CD</figureCitation>
), usually less than the ultimate phalange free (with at least the ultimate phalange free beyond the interdigital tissue margin,
<figureCitation id="134B83AAFFBBFF835204FEC8FD4E1212" box="[583,697,317,342]" captionStart="Fig" captionStartId="10.[141,176,1873,1895]" captionTargetBox="[141,1394,786,1860]" captionTargetId="figure-43@10.[141,1394,786,1860]" captionTargetPageId="10" captionText="Fig. 5. Dorsal views of the hand and foot of the holotypes of (AB) Bolitoglossa bolanosi sp. nov. (UCR 22965) and (CD) B. pesrubra in preservation. Photograph by Erick Arias." figureDoi="http://doi.org/10.5281/zenodo.11390655" httpUri="https://zenodo.org/record/11390655/files/figure.png" pageId="8" pageNumber="151">Fig. 5AB</figureCitation>
); has red color on forelimbs and usually on hind limbs (dorsum variable but never with red on hind limbs or forelimbs). Hands relatively wider, HaW/HeW = 0.61±0.07 (hands narrower, HaW/HeW = 0.56±0.06).
</paragraph>
</subSubSection>
<caption id="DF0FCFA7FFB6FF8E50FBF88FFF3813A0" ID-Table-UUID="DF0FCFA7FFB6FF8E50FBF88FFF3813A0" box="[184,207,228,1969]" httpUri="http://table.plazi.org/id/DF0FCFA7FFB6FF8E50FBF88FFF3813A0" pageId="5" pageNumber="148" startId="5.[184,207,1913,1969]" targetBox="[235,695,175,1956]" targetIsTable="true" targetPageId="5">
<paragraph id="8BCF9F2FFFB6FF8E50FBF88FFF3813A0" blockId="5.[184,207,228,1969]" box="[184,207,228,1969]" pageId="5" pageNumber="148">
<emphasis id="B904433DFFB6FF8E50FBF88FFF381434" bold="true" box="[184,207,1895,1969]" pageId="5" pageNumber="148">Table 1</emphasis>
. Mean uncorrected genetic distances, as percentages, among species of the
<taxonomicName id="4C70E4ACFFB6FF8E50FBFBEFFF3817D6" authorityName="Dumeril, Bibron &amp; Dumeril" authorityYear="1854" box="[184,207,1049,1169]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="148" phylum="Chordata" rank="genus">Bolitoglossa</taxonomicName>
subpalmata species group, using the 16 S (right) and cyt b (left) mitochondrial genes.
</paragraph>
</caption>
<caption id="DF0FCFA7FFB5FF8D50CEFF78FE731387" ID-Table-UUID="DF0FCFA7FFB5FF8D50CEFF78FE731387" httpUri="http://table.plazi.org/id/DF0FCFA7FFB5FF8D50CEFF78FE731387" pageId="6" pageNumber="149" startId="6.[141,197,142,163]" targetBox="[151,1360,220,1934]" targetIsTable="true" targetPageId="6" targetType="table">
<paragraph id="8BCF9F2FFFB5FF8D50CEFF78FE731387" blockId="6.[141,1395,141,192]" pageId="6" pageNumber="149">
<emphasis id="B904433DFFB5FF8D50CEFF78FF1613E5" bold="true" box="[141,225,141,163]" pageId="6" pageNumber="149">Table 2.</emphasis>
Morphometric data for
<taxonomicName id="4C70E4ACFFB5FF8D5192FF78FD5213E4" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[465,677,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB5FF8D5192FF78FD5213E4" box="[465,677,142,163]" italics="true" pageId="6" pageNumber="149">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB5FF8D52EEFF78FD0B13E4" bold="true" box="[685,764,142,163]" pageId="6" pageNumber="149">
<taxonomicNameLabel id="A237FE46FFB5FF8D52EEFF78FD0B13E4" box="[685,764,142,163]" pageId="6" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B904433DFFB5FF8D5348FF78FC9913E4" box="[779,878,142,163]" italics="true" pageId="6" pageNumber="149">
<taxonomicName id="4C70E4ACFFB5FF8D5348FF78FC9D13E4" box="[779,874,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="kamuk">B. kamuk</taxonomicName>
,
</emphasis>
and
<emphasis id="B904433DFFB5FF8D53E2FF78FBEB13E4" box="[929,1052,142,163]" italics="true" pageId="6" pageNumber="149">
<taxonomicName id="4C70E4ACFFB5FF8D53E2FF78FBEF13E4" baseAuthorityName="Taylor" baseAuthorityYear="1952" box="[929,1048,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="pesrubra">B. pesrubra</taxonomicName>
.
</emphasis>
These data were obtained prior to morphometric correction.
</paragraph>
</caption>
<paragraph id="8BCF9F2FFFB5FF8D50DBFF2AFAB314C9" pageId="6" pageNumber="149">
<table id="F9706D8FFFB5007450D4FF2AFAA714C9" box="[151,1360,220,1934]" colsContinueIn="7.[153,1355,215,1862]" gridcols="7" gridrows="27" pageId="6" pageNumber="149">
<tr id="35409D6DFFB5007450D4FF2AFAA713B5" box="[151,1360,220,242]" gridrow="0" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FF2AFF1F13B5" box="[151,232,220,242]" gridcol="0" gridrow="0" pageId="6" pageNumber="149">Variable</th>
<th id="7691F411FFB50074510EFF2AFD6113B5" box="[333,662,220,242]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="6" pageNumber="149">
<taxonomicName id="4C70E4ACFFB5FF8D51CCFF2BFE0A13B6" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[399,509,220,242]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB5FF8D51CCFF2BFE0A13B6" box="[399,509,220,242]" italics="true" pageId="6" pageNumber="149">B. bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB5FF8D5241FF2AFDB813B6" bold="true" box="[514,591,220,241]" pageId="6" pageNumber="149">
<taxonomicNameLabel id="A237FE46FFB5FF8D5241FF2AFDB813B6" box="[514,591,220,241]" pageId="6" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="7691F411FFB50074528EFF2AFC2213B5" box="[717,981,220,242]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="6" pageNumber="149">
<taxonomicName id="4C70E4ACFFB5FF8D5366FF2BFC8913B6" box="[805,894,220,242]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B904433DFFB5FF8D5366FF2BFC8913B6" box="[805,894,220,242]" italics="true" pageId="6" pageNumber="149">B. kamuk</emphasis>
</taxonomicName>
</th>
<th id="7691F411FFB50074545FFF2AFAA713B5" box="[1052,1360,220,242]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="6" pageNumber="149">
<taxonomicName id="4C70E4ACFFB5FF8D5435FF2BFB1F13B6" authority="Mean" authorityName="Mean" box="[1142,1256,220,242]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="149" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B904433DFFB5FF8D5435FF2BFB1F13B6" box="[1142,1256,220,242]" italics="true" pageId="6" pageNumber="149">B. pesrubra</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35409D6DFFB5007450D4FEF5FAA7125E" box="[151,1360,259,281]" gridrow="1" pageId="6" pageNumber="149" rowspan-0="1">
<td id="7691F411FFB50074510EFEF5FD61125E" box="[333,662,259,281]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="6" pageNumber="149">Mean±S.D. (Range)</td>
<td id="7691F411FFB50074528EFEF5FC22125E" box="[717,981,259,281]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="6" pageNumber="149">Mean±S.D. (Range)</td>
<td id="7691F411FFB50074545FFEF5FAA7125E" box="[1052,1360,259,281]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="6" pageNumber="149">Mean±S.D. (Range)</td>
</tr>
<tr id="35409D6DFFB5007450D4FEDFFAA71278" box="[151,1360,297,319]" gridrow="2" pageId="6" pageNumber="149" rowspan-0="1">
<td id="7691F411FFB50074510EFEDFFE221278" box="[333,469,297,319]" gridcol="1" gridrow="2" pageId="6" pageNumber="149">♀ (N = 8)</td>
<td id="7691F411FFB50074524DFEDFFD611278" box="[526,662,297,319]" gridcol="2" gridrow="2" pageId="6" pageNumber="149">♂ (N = 5)</td>
<td id="7691F411FFB50074528EFEDFFCDC1278" box="[717,811,297,319]" gridcol="3" gridrow="2" pageId="6" pageNumber="149">♀ (N = 1)</td>
<td id="7691F411FFB500745326FEDFFC221278" box="[869,981,297,319]" gridcol="4" gridrow="2" pageId="6" pageNumber="149">♂ (N = 2)</td>
<td id="7691F411FFB50074545FFEDFFB7A1278" box="[1052,1165,297,319]" gridcol="5" gridrow="2" pageId="6" pageNumber="149">♀ (N = 7)</td>
<td id="7691F411FFB500745490FEDFFAA71278" box="[1235,1360,297,319]" gridcol="6" gridrow="2" pageId="6" pageNumber="149">♂ (N = 8)</td>
</tr>
<tr id="35409D6DFFB5007450D4FEB9FAA712C5" box="[151,1360,335,386]" gridrow="3" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FEB9FF1F12C5" box="[151,232,335,386]" gridcol="0" gridrow="3" pageId="6" pageNumber="149">SL</th>
<td id="7691F411FFB50074510EFEB9FE2212C5" box="[333,469,335,386]" gridcol="1" gridrow="3" pageId="6" pageNumber="149">43.05±5.54 (32.7150.01)</td>
<td id="7691F411FFB50074524DFEB9FD6112C5" box="[526,662,335,386]" gridcol="2" gridrow="3" pageId="6" pageNumber="149">41.83±6.55 (34.0149.04)</td>
<td id="7691F411FFB50074528EFEB9FCDC12C5" box="[717,811,335,386]" gridcol="3" gridrow="3" pageId="6" pageNumber="149">38.4</td>
<td id="7691F411FFB500745326FEB9FC2212C5" box="[869,981,335,386]" gridcol="4" gridrow="3" pageId="6" pageNumber="149">35.2±0.85 (34.635.8)</td>
<td id="7691F411FFB50074545FFEB9FB7A12C5" box="[1052,1165,335,386]" gridcol="5" gridrow="3" pageId="6" pageNumber="149">50.6±3 (47.654.3)</td>
<td id="7691F411FFB500745490FEB9FAA712C5" box="[1235,1360,335,386]" gridcol="6" gridrow="3" pageId="6" pageNumber="149">48.34±3.61 (47.654.78)</td>
</tr>
<tr id="35409D6DFFB5007450D4FE64FAA71282" box="[151,1360,402,453]" gridrow="4" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FE64FF1F1282" box="[151,232,402,453]" gridcol="0" gridrow="4" pageId="6" pageNumber="149">ShW</th>
<td id="7691F411FFB50074510EFE64FE221282" box="[333,469,402,453]" gridcol="1" gridrow="4" pageId="6" pageNumber="149">5.64±0.98 (4.257.49)</td>
<td id="7691F411FFB50074524DFE64FD611282" box="[526,662,402,453]" gridcol="2" gridrow="4" pageId="6" pageNumber="149">5.34±1.02 (4.47.09)</td>
<td id="7691F411FFB50074528EFE64FCDC1282" box="[717,811,402,453]" gridcol="3" gridrow="4" pageId="6" pageNumber="149">4.9</td>
<td id="7691F411FFB500745326FE64FC221282" box="[869,981,402,453]" gridcol="4" gridrow="4" pageId="6" pageNumber="149">3.9±0.14 (3.84)</td>
<td id="7691F411FFB50074545FFE64FB7A1282" box="[1052,1165,402,453]" gridcol="5" gridrow="4" pageId="6" pageNumber="149">6.3±0.69 (5.27.3)</td>
<td id="7691F411FFB500745490FE64FAA71282" box="[1235,1360,402,453]" gridcol="6" gridrow="4" pageId="6" pageNumber="149">5.46±0.52 (5.26.43)</td>
</tr>
<tr id="35409D6DFFB5007450D4FE23FAA7114E" box="[151,1360,469,521]" gridrow="5" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FE23FF1F114E" box="[151,232,469,521]" gridcol="0" gridrow="5" pageId="6" pageNumber="149">HeW</th>
<td id="7691F411FFB50074510EFE23FE22114E" box="[333,469,469,521]" gridcol="1" gridrow="5" pageId="6" pageNumber="149">6.74±0.7 (5.457.7)</td>
<td id="7691F411FFB50074524DFE23FD61114E" box="[526,662,469,521]" gridcol="2" gridrow="5" pageId="6" pageNumber="149">6.43±0.72 (5.467.3)</td>
<td id="7691F411FFB50074528EFE23FCDC114E" box="[717,811,469,521]" gridcol="3" gridrow="5" pageId="6" pageNumber="149">6</td>
<td id="7691F411FFB500745326FE23FC22114E" box="[869,981,469,521]" gridcol="4" gridrow="5" pageId="6" pageNumber="149">5.15±0.21 (55.3)</td>
<td id="7691F411FFB50074545FFE23FB7A114E" box="[1052,1165,469,521]" gridcol="5" gridrow="5" pageId="6" pageNumber="149">7.17±0.75 (6.098.3)</td>
<td id="7691F411FFB500745490FE23FAA7114E" box="[1235,1360,469,521]" gridcol="6" gridrow="5" pageId="6" pageNumber="149">7.02±0.41 (6.097.76)</td>
</tr>
<tr id="35409D6DFFB5007450D4FDEEFAA7110B" box="[151,1360,536,588]" gridrow="6" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FDEEFF1F110B" box="[151,232,536,588]" gridcol="0" gridrow="6" pageId="6" pageNumber="149">NeW</th>
<td id="7691F411FFB50074510EFDEEFE22110B" box="[333,469,536,588]" gridcol="1" gridrow="6" pageId="6" pageNumber="149">5.62±0.84 (3.996.72)</td>
<td id="7691F411FFB50074524DFDEEFD61110B" box="[526,662,536,588]" gridcol="2" gridrow="6" pageId="6" pageNumber="149">5.17±0.74 (4.266.17)</td>
<td id="7691F411FFB50074528EFDEEFCDC110B" box="[717,811,536,588]" gridcol="3" gridrow="6" pageId="6" pageNumber="149">5</td>
<td id="7691F411FFB500745326FDEEFC22110B" box="[869,981,536,588]" gridcol="4" gridrow="6" pageId="6" pageNumber="149">4.05±0.07 (44.1)</td>
<td id="7691F411FFB50074545FFDEEFB7A110B" box="[1052,1165,536,588]" gridcol="5" gridrow="6" pageId="6" pageNumber="149">6.2±0.77 (5.37.4)</td>
<td id="7691F411FFB500745490FDEEFAA7110B" box="[1235,1360,536,588]" gridcol="6" gridrow="6" pageId="6" pageNumber="149">5.38±0.52 (5.36.53)</td>
</tr>
<tr id="35409D6DFFB5007450D4FDAAFAA711C8" box="[151,1360,604,655]" gridrow="7" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FDAAFF1F11C8" box="[151,232,604,655]" gridcol="0" gridrow="7" pageId="6" pageNumber="149">EW</th>
<td id="7691F411FFB50074510EFDAAFE2211C8" box="[333,469,604,655]" gridcol="1" gridrow="7" pageId="6" pageNumber="149">2.32±0.28 (1.782.59)</td>
<td id="7691F411FFB50074524DFDAAFD6111C8" box="[526,662,604,655]" gridcol="2" gridrow="7" pageId="6" pageNumber="149">2.5±0.48 (2.013.18)</td>
<td id="7691F411FFB50074528EFDAAFCDC11C8" box="[717,811,604,655]" gridcol="3" gridrow="7" pageId="6" pageNumber="149">2.5</td>
<td id="7691F411FFB500745326FDAAFC2211C8" box="[869,981,604,655]" gridcol="4" gridrow="7" pageId="6" pageNumber="149">2.15±0.21 (22.3)</td>
<td id="7691F411FFB50074545FFDAAFB7A11C8" box="[1052,1165,604,655]" gridcol="5" gridrow="7" pageId="6" pageNumber="149">2.56±0.22 (2.22.8)</td>
<td id="7691F411FFB500745490FDAAFAA711C8" box="[1235,1360,604,655]" gridcol="6" gridrow="7" pageId="6" pageNumber="149">2.51±0.22 (2.22.8)</td>
</tr>
<tr id="35409D6DFFB5007450D4FD69FAA71194" box="[151,1360,671,723]" gridrow="8" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FD69FF1F1194" box="[151,232,671,723]" gridcol="0" gridrow="8" pageId="6" pageNumber="149">SnL</th>
<td id="7691F411FFB50074510EFD69FE221194" box="[333,469,671,723]" gridcol="1" gridrow="8" pageId="6" pageNumber="149">2.75±0.32 (2.263.09)</td>
<td id="7691F411FFB50074524DFD69FD611194" box="[526,662,671,723]" gridcol="2" gridrow="8" pageId="6" pageNumber="149">2.88±0.43 (2.313.43)</td>
<td id="7691F411FFB50074528EFD69FCDC1194" box="[717,811,671,723]" gridcol="3" gridrow="8" pageId="6" pageNumber="149">2.3</td>
<td id="7691F411FFB500745326FD69FC221194" box="[869,981,671,723]" gridcol="4" gridrow="8" pageId="6" pageNumber="149">1.95±0.07 (1.92)</td>
<td id="7691F411FFB50074545FFD69FB7A1194" box="[1052,1165,671,723]" gridcol="5" gridrow="8" pageId="6" pageNumber="149">3.03±0.18 (2.83.3)</td>
<td id="7691F411FFB500745490FD69FAA71194" box="[1235,1360,671,723]" gridcol="6" gridrow="8" pageId="6" pageNumber="149">3±0.76 (2.83.78)</td>
</tr>
<tr id="35409D6DFFB5007450D4FD14FAA71051" box="[151,1360,738,790]" gridrow="9" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FD14FF1F1051" box="[151,232,738,790]" gridcol="0" gridrow="9" pageId="6" pageNumber="149">JSL</th>
<td id="7691F411FFB50074510EFD14FE221051" box="[333,469,738,790]" gridcol="1" gridrow="9" pageId="6" pageNumber="149">7±0.64 (5.998.15)</td>
<td id="7691F411FFB50074524DFD14FD611051" box="[526,662,738,790]" gridcol="2" gridrow="9" pageId="6" pageNumber="149">7±0.88 (6.018.21)</td>
<td id="7691F411FFB50074528EFD14FCDC1051" box="[717,811,738,790]" gridcol="3" gridrow="9" pageId="6" pageNumber="149">6.8</td>
<td id="7691F411FFB500745326FD14FC221051" box="[869,981,738,790]" gridcol="4" gridrow="9" pageId="6" pageNumber="149">5.95±0.49 (5.66.3)</td>
<td id="7691F411FFB50074545FFD14FB7A1051" box="[1052,1165,738,790]" gridcol="5" gridrow="9" pageId="6" pageNumber="149">8.09±0.5 (7.69)</td>
<td id="7691F411FFB500745490FD14FAA71051" box="[1235,1360,738,790]" gridcol="6" gridrow="9" pageId="6" pageNumber="149">8.08±0.48 (7.68.83)</td>
</tr>
<tr id="35409D6DFFB5007450D4FCD0FAA7101E" box="[151,1360,806,857]" gridrow="10" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FCD0FF1F101E" box="[151,232,806,857]" gridcol="0" gridrow="10" pageId="6" pageNumber="149">LGFS</th>
<td id="7691F411FFB50074510EFCD0FE22101E" box="[333,469,806,857]" gridcol="1" gridrow="10" pageId="6" pageNumber="149">10.99±1.19 (8.4612.17)</td>
<td id="7691F411FFB50074524DFCD0FD61101E" box="[526,662,806,857]" gridcol="2" gridrow="10" pageId="6" pageNumber="149">10.75±1.56 (9.0612.9)</td>
<td id="7691F411FFB50074528EFCD0FCDC101E" box="[717,811,806,857]" gridcol="3" gridrow="10" pageId="6" pageNumber="149">10.2</td>
<td id="7691F411FFB500745326FCD0FC22101E" box="[869,981,806,857]" gridcol="4" gridrow="10" pageId="6" pageNumber="149">8.95±0.21 (8.89.1)</td>
<td id="7691F411FFB50074545FFCD0FB7A101E" box="[1052,1165,806,857]" gridcol="5" gridrow="10" pageId="6" pageNumber="149">11.84±0.65 (11.212.9)</td>
<td id="7691F411FFB500745490FCD0FAA7101E" box="[1235,1360,806,857]" gridcol="6" gridrow="10" pageId="6" pageNumber="149">11.94±0.63 (11.212.96)</td>
</tr>
<tr id="35409D6DFFB5007450D4FC9FFAA710DA" box="[151,1360,873,925]" gridrow="11" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FC9FFF1F10DA" box="[151,232,873,925]" gridcol="0" gridrow="11" pageId="6" pageNumber="149">IND</th>
<td id="7691F411FFB50074510EFC9FFE2210DA" box="[333,469,873,925]" gridcol="1" gridrow="11" pageId="6" pageNumber="149">1.94±0.14 (1.742.12)</td>
<td id="7691F411FFB50074524DFC9FFD6110DA" box="[526,662,873,925]" gridcol="2" gridrow="11" pageId="6" pageNumber="149">2.25±0.56 (1.573.11)</td>
<td id="7691F411FFB50074528EFC9FFCDC10DA" box="[717,811,873,925]" gridcol="3" gridrow="11" pageId="6" pageNumber="149">1.8</td>
<td id="7691F411FFB500745326FC9FFC2210DA" box="[869,981,873,925]" gridcol="4" gridrow="11" pageId="6" pageNumber="149">1.2</td>
<td id="7691F411FFB50074545FFC9FFB7A10DA" box="[1052,1165,873,925]" gridcol="5" gridrow="11" pageId="6" pageNumber="149">2.27±0.19 (22.5)</td>
<td id="7691F411FFB500745490FC9FFAA710DA" box="[1235,1360,873,925]" gridcol="6" gridrow="11" pageId="6" pageNumber="149">2.61±0.36 (23.16)</td>
</tr>
<tr id="35409D6DFFB5007450D4FC5AFAA710A7" box="[151,1360,940,992]" gridrow="12" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FC5AFF1F10A7" box="[151,232,940,992]" gridcol="0" gridrow="12" pageId="6" pageNumber="149">NLP</th>
<td id="7691F411FFB50074510EFC5AFE2210A7" box="[333,469,940,992]" gridcol="1" gridrow="12" pageId="6" pageNumber="149">0.9±0.09 (0.741.01)</td>
<td id="7691F411FFB50074524DFC5AFD6110A7" box="[526,662,940,992]" gridcol="2" gridrow="12" pageId="6" pageNumber="149">0.87±0.15 (0.741.11)</td>
<td id="7691F411FFB50074528EFC5AFCDC10A7" box="[717,811,940,992]" gridcol="3" gridrow="12" pageId="6" pageNumber="149">1</td>
<td id="7691F411FFB500745326FC5AFC2210A7" box="[869,981,940,992]" gridcol="4" gridrow="12" pageId="6" pageNumber="149">0.75±0.07 (0.70.8)</td>
<td id="7691F411FFB50074545FFC5AFB7A10A7" box="[1052,1165,940,992]" gridcol="5" gridrow="12" pageId="6" pageNumber="149">1.07±0.13 (0.91.3)</td>
<td id="7691F411FFB500745490FC5AFAA710A7" box="[1235,1360,940,992]" gridcol="6" gridrow="12" pageId="6" pageNumber="149">1.05±0.16 (0.91.28)</td>
</tr>
<tr id="35409D6DFFB5007450D4FC06FAA71764" box="[151,1360,1008,1059]" gridrow="13" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FC06FF1F1764" box="[151,232,1008,1059]" gridcol="0" gridrow="13" pageId="6" pageNumber="149">ICD</th>
<td id="7691F411FFB50074510EFC06FE221764" box="[333,469,1008,1059]" gridcol="1" gridrow="13" pageId="6" pageNumber="149">3.3±0.34 (2.883.8)</td>
<td id="7691F411FFB50074524DFC06FD611764" box="[526,662,1008,1059]" gridcol="2" gridrow="13" pageId="6" pageNumber="149">3.27±0.38 (2.933.88)</td>
<td id="7691F411FFB50074528EFC06FCDC1764" box="[717,811,1008,1059]" gridcol="3" gridrow="13" pageId="6" pageNumber="149">2.8</td>
<td id="7691F411FFB500745326FC06FC221764" box="[869,981,1008,1059]" gridcol="4" gridrow="13" pageId="6" pageNumber="149">2.45±0.07 (2.42.5)</td>
<td id="7691F411FFB50074545FFC06FB7A1764" box="[1052,1165,1008,1059]" gridcol="5" gridrow="13" pageId="6" pageNumber="149">3.14±0.57 (1.93.6)</td>
<td id="7691F411FFB500745490FC06FAA71764" box="[1235,1360,1008,1059]" gridcol="6" gridrow="13" pageId="6" pageNumber="149">3.51±0.28 (1.93.98)</td>
</tr>
<tr id="35409D6DFFB5007450D4FBC5FAA71720" box="[151,1360,1075,1127]" gridrow="14" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FBC5FF1F1720" box="[151,232,1075,1127]" gridcol="0" gridrow="14" pageId="6" pageNumber="149">HLL</th>
<td id="7691F411FFB50074510EFBC5FE221720" box="[333,469,1075,1127]" gridcol="1" gridrow="14" pageId="6" pageNumber="149">11±1.62 (8.0312.79)</td>
<td id="7691F411FFB50074524DFBC5FD611720" box="[526,662,1075,1127]" gridcol="2" gridrow="14" pageId="6" pageNumber="149">11.07±1.54 (9.4713.09)</td>
<td id="7691F411FFB50074528EFBC5FCDC1720" box="[717,811,1075,1127]" gridcol="3" gridrow="14" pageId="6" pageNumber="149">10.4</td>
<td id="7691F411FFB500745326FBC5FC221720" box="[869,981,1075,1127]" gridcol="4" gridrow="14" pageId="6" pageNumber="149">8.3±0.28 (8.18.5)</td>
<td id="7691F411FFB50074545FFBC5FB7A1720" box="[1052,1165,1075,1127]" gridcol="5" gridrow="14" pageId="6" pageNumber="149">12.26±0.85 (11.213.6)</td>
<td id="7691F411FFB500745490FBC5FAA71720" box="[1235,1360,1075,1127]" gridcol="6" gridrow="14" pageId="6" pageNumber="149">12.71±1.36 (11.214.16)</td>
</tr>
<tr id="35409D6DFFB5007450D4FB80FAA717ED" box="[151,1360,1142,1194]" gridrow="15" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FB80FF1F17ED" box="[151,232,1142,1194]" gridcol="0" gridrow="15" pageId="6" pageNumber="149">TW</th>
<td id="7691F411FFB50074510EFB80FE2217ED" box="[333,469,1142,1194]" gridcol="1" gridrow="15" pageId="6" pageNumber="149">6.62±1.39 (5.169.2)</td>
<td id="7691F411FFB50074524DFB80FD6117ED" box="[526,662,1142,1194]" gridcol="2" gridrow="15" pageId="6" pageNumber="149">5.26±0.63 (4.36.02)</td>
<td id="7691F411FFB50074528EFB80FCDC17ED" box="[717,811,1142,1194]" gridcol="3" gridrow="15" pageId="6" pageNumber="149">5.4</td>
<td id="7691F411FFB500745326FB80FC2217ED" box="[869,981,1142,1194]" gridcol="4" gridrow="15" pageId="6" pageNumber="149">3.9±0.14 (3.84)</td>
<td id="7691F411FFB50074545FFB80FB7A17ED" box="[1052,1165,1142,1194]" gridcol="5" gridrow="15" pageId="6" pageNumber="149">6.93±0.96 (5.88.4)</td>
<td id="7691F411FFB500745490FB80FAA717ED" box="[1235,1360,1142,1194]" gridcol="6" gridrow="15" pageId="6" pageNumber="149">6.1±0.46 (5.86.9)</td>
</tr>
<tr id="35409D6DFFB5007450D4FB4CFAA717AA" box="[151,1360,1210,1261]" gridrow="16" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FB4CFF1F17AA" box="[151,232,1210,1261]" gridcol="0" gridrow="16" pageId="6" pageNumber="149">FLL</th>
<td id="7691F411FFB50074510EFB4CFE2217AA" box="[333,469,1210,1261]" gridcol="1" gridrow="16" pageId="6" pageNumber="149">10.38±1.54 (7.7511.91)</td>
<td id="7691F411FFB50074524DFB4CFD6117AA" box="[526,662,1210,1261]" gridcol="2" gridrow="16" pageId="6" pageNumber="149">11.28±3.87 (8.0617.62)</td>
<td id="7691F411FFB50074528EFB4CFCDC17AA" box="[717,811,1210,1261]" gridcol="3" gridrow="16" pageId="6" pageNumber="149">8.9</td>
<td id="7691F411FFB500745326FB4CFC2217AA" box="[869,981,1210,1261]" gridcol="4" gridrow="16" pageId="6" pageNumber="149">8.00</td>
<td id="7691F411FFB50074545FFB4CFB7A17AA" box="[1052,1165,1210,1261]" gridcol="5" gridrow="16" pageId="6" pageNumber="149">11.27±0.74 (10.112.4)</td>
<td id="7691F411FFB500745490FB4CFAA717AA" box="[1235,1360,1210,1261]" gridcol="6" gridrow="16" pageId="6" pageNumber="149">11.51±1.22 (10.113.61)</td>
</tr>
<tr id="35409D6DFFB5007450D4FB0BFAA71677" box="[151,1360,1277,1328]" gridrow="17" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FB0BFF1F1677" box="[151,232,1277,1328]" gridcol="0" gridrow="17" pageId="6" pageNumber="149">VGS</th>
<td id="7691F411FFB50074510EFB0BFE221677" box="[333,469,1277,1328]" gridcol="1" gridrow="17" pageId="6" pageNumber="149">10.49±1.13 (8.1811.68)</td>
<td id="7691F411FFB50074524DFB0BFD611677" box="[526,662,1277,1328]" gridcol="2" gridrow="17" pageId="6" pageNumber="149">10.47±1.81 (8.7213.2)</td>
<td id="7691F411FFB50074528EFB0BFCDC1677" box="[717,811,1277,1328]" gridcol="3" gridrow="17" pageId="6" pageNumber="149">9.2</td>
<td id="7691F411FFB500745326FB0BFC221677" box="[869,981,1277,1328]" gridcol="4" gridrow="17" pageId="6" pageNumber="149">8.45±0.35 (8.28.7)</td>
<td id="7691F411FFB50074545FFB0BFB7A1677" box="[1052,1165,1277,1328]" gridcol="5" gridrow="17" pageId="6" pageNumber="149">11.66±0.52 (11.112.4)</td>
<td id="7691F411FFB500745490FB0BFAA71677" box="[1235,1360,1277,1328]" gridcol="6" gridrow="17" pageId="6" pageNumber="149">11.4±0.56 (11.112.2)</td>
</tr>
<tr id="35409D6DFFB5007450D4FAB6FAA71633" box="[151,1360,1344,1396]" gridrow="18" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FAB6FF1F1633" box="[151,232,1344,1396]" gridcol="0" gridrow="18" pageId="6" pageNumber="149">FSL</th>
<td id="7691F411FFB50074510EFAB6FE221633" box="[333,469,1344,1396]" gridcol="1" gridrow="18" pageId="6" pageNumber="149">12.75±1.42 (10.1614.8)</td>
<td id="7691F411FFB50074524DFAB6FD611633" box="[526,662,1344,1396]" gridcol="2" gridrow="18" pageId="6" pageNumber="149">12.59±1.77 (10.9115.17)</td>
<td id="7691F411FFB50074528EFAB6FCDC1633" box="[717,811,1344,1396]" gridcol="3" gridrow="18" pageId="6" pageNumber="149">11.9</td>
<td id="7691F411FFB500745326FAB6FC221633" box="[869,981,1344,1396]" gridcol="4" gridrow="18" pageId="6" pageNumber="149">10.3±0.28 (10.110.5)</td>
<td id="7691F411FFB50074545FFAB6FB7A1633" box="[1052,1165,1344,1396]" gridcol="5" gridrow="18" pageId="6" pageNumber="149">14.46±0.79 (13.515.6)</td>
<td id="7691F411FFB500745490FAB6FAA71633" box="[1235,1360,1344,1396]" gridcol="6" gridrow="18" pageId="6" pageNumber="149">14.37±0.85 (13.515.59)</td>
</tr>
<tr id="35409D6DFFB5007450D4FA75FAA716F0" box="[151,1360,1411,1463]" gridrow="19" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FA75FF1F16F0" box="[151,232,1411,1463]" gridcol="0" gridrow="19" pageId="6" pageNumber="149">UHL</th>
<td id="7691F411FFB50074510EFA75FE2216F0" box="[333,469,1411,1463]" gridcol="1" gridrow="19" pageId="6" pageNumber="149">7.22±2 (2.638.58)</td>
<td id="7691F411FFB50074524DFA75FD6116F0" box="[526,662,1411,1463]" gridcol="2" gridrow="19" pageId="6" pageNumber="149">6.51±2.65 (3.29.01)</td>
<td id="7691F411FFB50074528EFA75FCDC16F0" box="[717,811,1411,1463]" gridcol="3" gridrow="19" pageId="6" pageNumber="149">6.5</td>
<td id="7691F411FFB500745326FA75FC2216F0" box="[869,981,1411,1463]" gridcol="4" gridrow="19" pageId="6" pageNumber="149">5.9</td>
<td id="7691F411FFB50074545FFA75FB7A16F0" box="[1052,1165,1411,1463]" gridcol="5" gridrow="19" pageId="6" pageNumber="149">8.1±0.64 (7.49.4)</td>
<td id="7691F411FFB500745490FA75FAA716F0" box="[1235,1360,1411,1463]" gridcol="6" gridrow="19" pageId="6" pageNumber="149">6.16±2 (7.49.2)</td>
</tr>
<tr id="35409D6DFFB5007450D4FA31FAA716BD" box="[151,1360,1479,1530]" gridrow="20" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4FA31FF1F16BD" box="[151,232,1479,1530]" gridcol="0" gridrow="20" pageId="6" pageNumber="149">AGL</th>
<td id="7691F411FFB50074510EFA31FE2216BD" box="[333,469,1479,1530]" gridcol="1" gridrow="20" pageId="6" pageNumber="149">21.94±3.75 (16.1426.4)</td>
<td id="7691F411FFB50074524DFA31FD6116BD" box="[526,662,1479,1530]" gridcol="2" gridrow="20" pageId="6" pageNumber="149">21.2±3.05 (16.1824.13)</td>
<td id="7691F411FFB50074528EFA31FCDC16BD" box="[717,811,1479,1530]" gridcol="3" gridrow="20" pageId="6" pageNumber="149">19.3</td>
<td id="7691F411FFB500745326FA31FC2216BD" box="[869,981,1479,1530]" gridcol="4" gridrow="20" pageId="6" pageNumber="149">17.9±0.28 (17.718.1)</td>
<td id="7691F411FFB50074545FFA31FB7A16BD" box="[1052,1165,1479,1530]" gridcol="5" gridrow="20" pageId="6" pageNumber="149">27.37±1.89 (25.230.1)</td>
<td id="7691F411FFB500745490FA31FAA716BD" box="[1235,1360,1479,1530]" gridcol="6" gridrow="20" pageId="6" pageNumber="149">24.71±1.96 (25.228.35)</td>
</tr>
<tr id="35409D6DFFB5007450D4F9FCFAA71579" box="[151,1360,1546,1598]" gridrow="21" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F9FCFF1F1579" box="[151,232,1546,1598]" gridcol="0" gridrow="21" pageId="6" pageNumber="149">VL</th>
<td id="7691F411FFB50074510EF9FCFE221579" box="[333,469,1546,1598]" gridcol="1" gridrow="21" pageId="6" pageNumber="149">4.34±0.7 (3.055.09)</td>
<td id="7691F411FFB50074524DF9FCFD611579" box="[526,662,1546,1598]" gridcol="2" gridrow="21" pageId="6" pageNumber="149">4.06±0.85 (2.934.94)</td>
<td id="7691F411FFB50074528EF9FCFCDC1579" box="[717,811,1546,1598]" gridcol="3" gridrow="21" pageId="6" pageNumber="149">4</td>
<td id="7691F411FFB500745326F9FCFC221579" box="[869,981,1546,1598]" gridcol="4" gridrow="21" pageId="6" pageNumber="149">2.56±0.62 (2.123)</td>
<td id="7691F411FFB50074545FF9FCFB7A1579" box="[1052,1165,1546,1598]" gridcol="5" gridrow="21" pageId="6" pageNumber="149">4.86±0.53 (4.45.8)</td>
<td id="7691F411FFB500745490F9FCFAA71579" box="[1235,1360,1546,1598]" gridcol="6" gridrow="21" pageId="6" pageNumber="149">4.36±0.64 (4.45.78)</td>
</tr>
<tr id="35409D6DFFB5007450D4F9BBFAA715C6" box="[151,1360,1613,1665]" gridrow="22" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F9BBFF1F15C6" box="[151,232,1613,1665]" gridcol="0" gridrow="22" pageId="6" pageNumber="149">HaW</th>
<td id="7691F411FFB50074510EF9BBFE2215C6" box="[333,469,1613,1665]" gridcol="1" gridrow="22" pageId="6" pageNumber="149">3.66±0.5 (3.044.41)</td>
<td id="7691F411FFB50074524DF9BBFD6115C6" box="[526,662,1613,1665]" gridcol="2" gridrow="22" pageId="6" pageNumber="149">3.79±0.84 (2.684.84)</td>
<td id="7691F411FFB50074528EF9BBFCDC15C6" box="[717,811,1613,1665]" gridcol="3" gridrow="22" pageId="6" pageNumber="149">3.3</td>
<td id="7691F411FFB500745326F9BBFC2215C6" box="[869,981,1613,1665]" gridcol="4" gridrow="22" pageId="6" pageNumber="149">2.65±0.21 (2.52.8)</td>
<td id="7691F411FFB50074545FF9BBFB7A15C6" box="[1052,1165,1613,1665]" gridcol="5" gridrow="22" pageId="6" pageNumber="149">4.24±0.44 (3.85.1)</td>
<td id="7691F411FFB500745490F9BBFAA715C6" box="[1235,1360,1613,1665]" gridcol="6" gridrow="22" pageId="6" pageNumber="149">4.36±0.57 (3.85.1)</td>
</tr>
<tr id="35409D6DFFB5007450D4F967FAA71583" box="[151,1360,1681,1732]" gridrow="23" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F967FF1F1583" box="[151,232,1681,1732]" gridcol="0" gridrow="23" pageId="6" pageNumber="149">HaL</th>
<td id="7691F411FFB50074510EF967FE221583" box="[333,469,1681,1732]" gridcol="1" gridrow="23" pageId="6" pageNumber="149">3.85±0.64 (2.754.64)</td>
<td id="7691F411FFB50074524DF967FD611583" box="[526,662,1681,1732]" gridcol="2" gridrow="23" pageId="6" pageNumber="149">3.81±1 (2.915.35)</td>
<td id="7691F411FFB50074528EF967FCDC1583" box="[717,811,1681,1732]" gridcol="3" gridrow="23" pageId="6" pageNumber="149">3.7</td>
<td id="7691F411FFB500745326F967FC221583" box="[869,981,1681,1732]" gridcol="4" gridrow="23" pageId="6" pageNumber="149">2.6±0.14 (2.52.7)</td>
<td id="7691F411FFB50074545FF967FB7A1583" box="[1052,1165,1681,1732]" gridcol="5" gridrow="23" pageId="6" pageNumber="149">4.04±0.43 (3.85)</td>
<td id="7691F411FFB500745490F967FAA71583" box="[1235,1360,1681,1732]" gridcol="6" gridrow="23" pageId="6" pageNumber="149">4.2±0.49 (3.84.9)</td>
</tr>
<tr id="35409D6DFFB5007450D4F922FAA7144F" box="[151,1360,1748,1800]" gridrow="24" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F922FF1F144F" box="[151,232,1748,1800]" gridcol="0" gridrow="24" pageId="6" pageNumber="149">WF3</th>
<td id="7691F411FFB50074510EF922FE22144F" box="[333,469,1748,1800]" gridcol="1" gridrow="24" pageId="6" pageNumber="149">0.86±0.09 (0.731.05)</td>
<td id="7691F411FFB50074524DF922FD61144F" box="[526,662,1748,1800]" gridcol="2" gridrow="24" pageId="6" pageNumber="149">0.79±0.15 (0.590.98)</td>
<td id="7691F411FFB50074528EF922FCDC144F" box="[717,811,1748,1800]" gridcol="3" gridrow="24" pageId="6" pageNumber="149">0.9</td>
<td id="7691F411FFB500745326F922FC22144F" box="[869,981,1748,1800]" gridcol="4" gridrow="24" pageId="6" pageNumber="149">0.6</td>
<td id="7691F411FFB50074545FF922FB7A144F" box="[1052,1165,1748,1800]" gridcol="5" gridrow="24" pageId="6" pageNumber="149">0.77±0.08 (0.70.9)</td>
<td id="7691F411FFB500745490F922FAA7144F" box="[1235,1360,1748,1800]" gridcol="6" gridrow="24" pageId="6" pageNumber="149">0.82±0.11 (0.70.99)</td>
</tr>
<tr id="35409D6DFFB5007450D4F8E1FAA7140C" box="[151,1360,1815,1867]" gridrow="25" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F8E1FF1F140C" box="[151,232,1815,1867]" gridcol="0" gridrow="25" pageId="6" pageNumber="149">LF2</th>
<td id="7691F411FFB50074510EF8E1FE22140C" box="[333,469,1815,1867]" gridcol="1" gridrow="25" pageId="6" pageNumber="149">1.61±0.22 (1.331.92)</td>
<td id="7691F411FFB50074524DF8E1FD61140C" box="[526,662,1815,1867]" gridcol="2" gridrow="25" pageId="6" pageNumber="149">1.81±0.29 (1.592.31)</td>
<td id="7691F411FFB50074528EF8E1FCDC140C" box="[717,811,1815,1867]" gridcol="3" gridrow="25" pageId="6" pageNumber="149">1.6</td>
<td id="7691F411FFB500745326F8E1FC22140C" box="[869,981,1815,1867]" gridcol="4" gridrow="25" pageId="6" pageNumber="149">1.45±0.07 (1.41.5)</td>
<td id="7691F411FFB50074545FF8E1FB7A140C" box="[1052,1165,1815,1867]" gridcol="5" gridrow="25" pageId="6" pageNumber="149">2.01±0.22 (1.82.4)</td>
<td id="7691F411FFB500745490F8E1FAA7140C" box="[1235,1360,1815,1867]" gridcol="6" gridrow="25" pageId="6" pageNumber="149">2.24±0.4 (1.82.7)</td>
</tr>
<tr id="35409D6DFFB5007450D4F8ADFAA714C9" box="[151,1360,1883,1934]" gridrow="26" pageId="6" pageNumber="149">
<th id="7691F411FFB5007450D4F8ADFF1F14C9" box="[151,232,1883,1934]" gridcol="0" gridrow="26" pageId="6" pageNumber="149">LF3</th>
<td id="7691F411FFB50074510EF8ADFE2214C9" box="[333,469,1883,1934]" gridcol="1" gridrow="26" pageId="6" pageNumber="149">2.02±0.25 (1.752.39)</td>
<td id="7691F411FFB50074524DF8ADFD6114C9" box="[526,662,1883,1934]" gridcol="2" gridrow="26" pageId="6" pageNumber="149">2.29±0.45 (1.863)</td>
<td id="7691F411FFB50074528EF8ADFCDC14C9" box="[717,811,1883,1934]" gridcol="3" gridrow="26" pageId="6" pageNumber="149">2.3</td>
<td id="7691F411FFB500745326F8ADFC2214C9" box="[869,981,1883,1934]" gridcol="4" gridrow="26" pageId="6" pageNumber="149">1.75±0.21 (1.61.9)</td>
<td id="7691F411FFB50074545FF8ADFB7A14C9" box="[1052,1165,1883,1934]" gridcol="5" gridrow="26" pageId="6" pageNumber="149">2.56±0.24 (2.43.1)</td>
<td id="7691F411FFB500745490F8ADFAA714C9" box="[1235,1360,1883,1934]" gridcol="6" gridrow="26" pageId="6" pageNumber="149">2.76±0.42 (2.43.45)</td>
</tr>
</table>
</paragraph>
<caption id="DF0FCFA7FFB4FF8C50CEFF78FE261387" isContinuationCaption="true" pageId="7" pageNumber="150" startId="7.[141,197,142,163]" targetType="table">
<paragraph id="8BCF9F2FFFB4FF8C50CEFF78FE261387" blockId="7.[141,1395,141,192]" pageId="7" pageNumber="150">
<emphasis id="B904433DFFB4FF8C50CEFF78FEAE13E4" bold="true" box="[141,345,141,163]" pageId="7" pageNumber="150">Table 2 (continued).</emphasis>
Morphometric data for
<taxonomicName id="4C70E4ACFFB4FF8C5202FF78FCE513E4" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[577,786,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB4FF8C5202FF78FCE513E4" box="[577,786,142,163]" italics="true" pageId="7" pageNumber="150">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB4FF8C535BFF78FC9213E4" bold="true" box="[792,869,142,163]" pageId="7" pageNumber="150">
<taxonomicNameLabel id="A237FE46FFB4FF8C535BFF78FC9213E4" box="[792,869,142,163]" pageId="7" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B904433DFFB4FF8C5332FF78FC2613E4" box="[881,977,142,163]" italics="true" pageId="7" pageNumber="150">
<taxonomicName id="4C70E4ACFFB4FF8C5332FF78FC3A13E4" box="[881,973,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="kamuk">B. kamuk</taxonomicName>
,
</emphasis>
and
<emphasis id="B904433DFFB4FF8C5443FF78FB8F13E4" box="[1024,1144,142,163]" italics="true" pageId="7" pageNumber="150">
<taxonomicName id="4C70E4ACFFB4FF8C5443FF78FB8213E4" box="[1024,1141,142,163]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="pesrubra">B. pesrubra</taxonomicName>
.
</emphasis>
These data were obtained prior to morphometric correction.
</paragraph>
</caption>
<paragraph id="8BCF9F2FFFB4FF8C50DAFF21FABC1401" pageId="7" pageNumber="150">
<table id="F9706D8FFFB4007450DAFF21FABC1401" box="[153,1355,215,1862]" colsContinueFrom="6.[151,1360,220,1934]" gridcols="7" gridrows="26" pageId="7" pageNumber="150">
<tr id="35409D6DFFB4007450DAFF21FABC13AA" box="[153,1355,215,237]" gridrow="0" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFF21FEF913AA" box="[153,270,215,237]" gridcol="0" gridrow="0" pageId="7" pageNumber="150">Variable</th>
<th id="7691F411FFB400745119FF21FD7B13AA" box="[346,652,215,237]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="7" pageNumber="150">
<taxonomicName id="4C70E4ACFFB4FF8C51D3FF2EFE0913AB" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[400,510,215,237]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB4FF8C51D3FF2EFE0913AB" box="[400,510,215,237]" italics="true" pageId="7" pageNumber="150">B. bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB4FF8C5247FF2EFDA713AA" bold="true" box="[516,592,216,237]" pageId="7" pageNumber="150">
<taxonomicNameLabel id="A237FE46FFB4FF8C5247FF2EFDA713AA" box="[516,592,216,237]" pageId="7" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="7691F411FFB40074528DFF21FC2113AA" box="[718,982,215,237]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="7" pageNumber="150">
<taxonomicName id="4C70E4ACFFB4FF8C5365FF2EFC8813AB" box="[806,895,215,237]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="kamuk">
<emphasis id="B904433DFFB4FF8C5365FF2EFC8813AB" box="[806,895,215,237]" italics="true" pageId="7" pageNumber="150">B. kamuk</emphasis>
</taxonomicName>
</th>
<th id="7691F411FFB40074545EFF21FABC13AA" box="[1053,1355,215,237]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="7" pageNumber="150">
<taxonomicName id="4C70E4ACFFB4FF8C5434FF2EFB1E13AB" authority="Mean" authorityName="Mean" box="[1143,1257,215,237]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="150" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B904433DFFB4FF8C5434FF2EFB1E13AB" box="[1143,1257,215,237]" italics="true" pageId="7" pageNumber="150">B. pesrubra</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="35409D6DFFB4007450DAFF08FABC1253" box="[153,1355,254,276]" gridrow="1" pageId="7" pageNumber="150" rowspan-0="1">
<td id="7691F411FFB400745119FF08FD7B1253" box="[346,652,254,276]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="7" pageNumber="150">Mean±S.D. (Range)</td>
<td id="7691F411FFB40074528DFF08FC211253" box="[718,982,254,276]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="7" pageNumber="150">Mean±S.D. (Range)</td>
<td id="7691F411FFB40074545EFF08FABC1253" box="[1053,1355,254,276]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="7" pageNumber="150">Mean±S.D. (Range)</td>
</tr>
<tr id="35409D6DFFB4007450DAFED2FABC127D" box="[153,1355,292,314]" gridrow="2" pageId="7" pageNumber="150" rowspan-0="1">
<td id="7691F411FFB400745119FED2FE3D127D" box="[346,458,292,314]" gridcol="1" gridrow="2" pageId="7" pageNumber="150">♀ (N = 8)</td>
<td id="7691F411FFB400745258FED2FD7B127D" box="[539,652,292,314]" gridcol="2" gridrow="2" pageId="7" pageNumber="150">♂ (N = 5)</td>
<td id="7691F411FFB40074528DFED2FCDB127D" box="[718,812,292,314]" gridcol="3" gridrow="2" pageId="7" pageNumber="150">♀ (N = 1)</td>
<td id="7691F411FFB400745325FED2FC21127D" box="[870,982,292,314]" gridcol="4" gridrow="2" pageId="7" pageNumber="150">♂ (N = 2)</td>
<td id="7691F411FFB40074545EFED2FB79127D" box="[1053,1166,292,314]" gridcol="5" gridrow="2" pageId="7" pageNumber="150">♀ (N = 7)</td>
<td id="7691F411FFB400745499FED2FABC127D" box="[1242,1355,292,314]" gridcol="6" gridrow="2" pageId="7" pageNumber="150">♂ (N = 8)</td>
</tr>
<tr id="35409D6DFFB4007450DAFEBCFABC123A" box="[153,1355,330,381]" gridrow="3" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFEBCFEF9123A" box="[153,270,330,381]" gridcol="0" gridrow="3" pageId="7" pageNumber="150">FoW</th>
<td id="7691F411FFB400745119FEBCFE3D123A" box="[346,458,330,381]" gridcol="1" gridrow="3" pageId="7" pageNumber="150">4.44±0.76 (3.445.42)</td>
<td id="7691F411FFB400745258FEBCFD7B123A" box="[539,652,330,381]" gridcol="2" gridrow="3" pageId="7" pageNumber="150">4.56±0.87 (3.635.74)</td>
<td id="7691F411FFB40074528DFEBCFCDB123A" box="[718,812,330,381]" gridcol="3" gridrow="3" pageId="7" pageNumber="150">3.9</td>
<td id="7691F411FFB400745325FEBCFC21123A" box="[870,982,330,381]" gridcol="4" gridrow="3" pageId="7" pageNumber="150">3.45±0.21 (3.33.6)</td>
<td id="7691F411FFB40074545EFEBCFB79123A" box="[1053,1166,330,381]" gridcol="5" gridrow="3" pageId="7" pageNumber="150">4.96±0.4 (4.55.6)</td>
<td id="7691F411FFB400745499FEBCFABC123A" box="[1242,1355,330,381]" gridcol="6" gridrow="3" pageId="7" pageNumber="150">5.17±0.77 (4.56.02)</td>
</tr>
<tr id="35409D6DFFB4007450DAFE7BFABC1286" box="[153,1355,397,449]" gridrow="4" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFE7BFEF91286" box="[153,270,397,449]" gridcol="0" gridrow="4" pageId="7" pageNumber="150">FoL</th>
<td id="7691F411FFB400745119FE7BFE3D1286" box="[346,458,397,449]" gridcol="1" gridrow="4" pageId="7" pageNumber="150">4.34±0.58 (3.425.03)</td>
<td id="7691F411FFB400745258FE7BFD7B1286" box="[539,652,397,449]" gridcol="2" gridrow="4" pageId="7" pageNumber="150">4.23±0.91 (3.275.53)</td>
<td id="7691F411FFB40074528DFE7BFCDB1286" box="[718,812,397,449]" gridcol="3" gridrow="4" pageId="7" pageNumber="150">4.3</td>
<td id="7691F411FFB400745325FE7BFC211286" box="[870,982,397,449]" gridcol="4" gridrow="4" pageId="7" pageNumber="150">3.00</td>
<td id="7691F411FFB40074545EFE7BFB791286" box="[1053,1166,397,449]" gridcol="5" gridrow="4" pageId="7" pageNumber="150">4.6±0.35 (4.15.2)</td>
<td id="7691F411FFB400745499FE7BFABC1286" box="[1242,1355,397,449]" gridcol="6" gridrow="4" pageId="7" pageNumber="150">4.78±0.39 (4.15.3)</td>
</tr>
<tr id="35409D6DFFB4007450DAFE27FABC1143" box="[153,1355,465,516]" gridrow="5" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFE27FEF91143" box="[153,270,465,516]" gridcol="0" gridrow="5" pageId="7" pageNumber="150">WT3</th>
<td id="7691F411FFB400745119FE27FE3D1143" box="[346,458,465,516]" gridcol="1" gridrow="5" pageId="7" pageNumber="150">0.9±0.16 (0.681.24)</td>
<td id="7691F411FFB400745258FE27FD7B1143" box="[539,652,465,516]" gridcol="2" gridrow="5" pageId="7" pageNumber="150">0.77±0.1 (0.670.92)</td>
<td id="7691F411FFB40074528DFE27FCDB1143" box="[718,812,465,516]" gridcol="3" gridrow="5" pageId="7" pageNumber="150">0.9</td>
<td id="7691F411FFB400745325FE27FC211143" box="[870,982,465,516]" gridcol="4" gridrow="5" pageId="7" pageNumber="150">0.65±0.07 (0.60.7)</td>
<td id="7691F411FFB40074545EFE27FB791143" box="[1053,1166,465,516]" gridcol="5" gridrow="5" pageId="7" pageNumber="150">0.8±0.1 (0.71)</td>
<td id="7691F411FFB400745499FE27FABC1143" box="[1242,1355,465,516]" gridcol="6" gridrow="5" pageId="7" pageNumber="150">0.86±0.13 (0.71.1)</td>
</tr>
<tr id="35409D6DFFB4007450DAFDE2FABC110F" box="[153,1355,532,584]" gridrow="6" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFDE2FEF9110F" box="[153,270,532,584]" gridcol="0" gridrow="6" pageId="7" pageNumber="150">LT2</th>
<td id="7691F411FFB400745119FDE2FE3D110F" box="[346,458,532,584]" gridcol="1" gridrow="6" pageId="7" pageNumber="150">1.82±0.19 (1.522.05)</td>
<td id="7691F411FFB400745258FDE2FD7B110F" box="[539,652,532,584]" gridcol="2" gridrow="6" pageId="7" pageNumber="150">1.93±0.16 (1.772.15)</td>
<td id="7691F411FFB40074528DFDE2FCDB110F" box="[718,812,532,584]" gridcol="3" gridrow="6" pageId="7" pageNumber="150">1.5</td>
<td id="7691F411FFB400745325FDE2FC21110F" box="[870,982,532,584]" gridcol="4" gridrow="6" pageId="7" pageNumber="150">1.2±0.14 (1.11.3)</td>
<td id="7691F411FFB40074545EFDE2FB79110F" box="[1053,1166,532,584]" gridcol="5" gridrow="6" pageId="7" pageNumber="150">2.31±0.22 (22.7)</td>
<td id="7691F411FFB400745499FDE2FABC110F" box="[1242,1355,532,584]" gridcol="6" gridrow="6" pageId="7" pageNumber="150">2.44±0.5 (23.09)</td>
</tr>
<tr id="35409D6DFFB4007450DAFDA1FABC11CC" box="[153,1355,599,651]" gridrow="7" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFDA1FEF911CC" box="[153,270,599,651]" gridcol="0" gridrow="7" pageId="7" pageNumber="150">LT3</th>
<td id="7691F411FFB400745119FDA1FE3D11CC" box="[346,458,599,651]" gridcol="1" gridrow="7" pageId="7" pageNumber="150">2.44±0.44 (1.662.94)</td>
<td id="7691F411FFB400745258FDA1FD7B11CC" box="[539,652,599,651]" gridcol="2" gridrow="7" pageId="7" pageNumber="150">2.46±0.22 (2.152.68)</td>
<td id="7691F411FFB40074528DFDA1FCDB11CC" box="[718,812,599,651]" gridcol="3" gridrow="7" pageId="7" pageNumber="150">2.3</td>
<td id="7691F411FFB400745325FDA1FC2111CC" box="[870,982,599,651]" gridcol="4" gridrow="7" pageId="7" pageNumber="150">1.65±0.07 (1.61.7)</td>
<td id="7691F411FFB40074545EFDA1FB7911CC" box="[1053,1166,599,651]" gridcol="5" gridrow="7" pageId="7" pageNumber="150">2.79±0.17 (2.63)</td>
<td id="7691F411FFB400745499FDA1FABC11CC" box="[1242,1355,599,651]" gridcol="6" gridrow="7" pageId="7" pageNumber="150">3.05±0.42 (2.63.73)</td>
</tr>
<tr id="35409D6DFFB4007450DAFD6DFABC1189" box="[153,1355,667,718]" gridrow="8" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFD6DFEF91189" box="[153,270,667,718]" gridcol="0" gridrow="8" pageId="7" pageNumber="150">VGS/SL</th>
<td id="7691F411FFB400745119FD6DFE3D1189" box="[346,458,667,718]" gridcol="1" gridrow="8" pageId="7" pageNumber="150">0.24±0.01 (0.230.26)</td>
<td id="7691F411FFB400745258FD6DFD7B1189" box="[539,652,667,718]" gridcol="2" gridrow="8" pageId="7" pageNumber="150">0.25±0.01 (0.230.27)</td>
<td id="7691F411FFB40074528DFD6DFCDB1189" box="[718,812,667,718]" gridcol="3" gridrow="8" pageId="7" pageNumber="150">0.24</td>
<td id="7691F411FFB400745325FD6DFC211189" box="[870,982,667,718]" gridcol="4" gridrow="8" pageId="7" pageNumber="150">0.24±0 (0.240.24)</td>
<td id="7691F411FFB40074545EFD6DFB791189" box="[1053,1166,667,718]" gridcol="5" gridrow="8" pageId="7" pageNumber="150">0.23±0.01 (0.220.24)</td>
<td id="7691F411FFB400745499FD6DFABC1189" box="[1242,1355,667,718]" gridcol="6" gridrow="8" pageId="7" pageNumber="150">0.24±0.01 (0.220.25)</td>
</tr>
<tr id="35409D6DFFB4007450DAFD28FABC1056" box="[153,1355,734,785]" gridrow="9" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFD28FEF91056" box="[153,270,734,785]" gridcol="0" gridrow="9" pageId="7" pageNumber="150">IND/HEW</th>
<td id="7691F411FFB400745119FD28FE3D1056" box="[346,458,734,785]" gridcol="1" gridrow="9" pageId="7" pageNumber="150">0.29±0.02 (0.260.32)</td>
<td id="7691F411FFB400745258FD28FD7B1056" box="[539,652,734,785]" gridcol="2" gridrow="9" pageId="7" pageNumber="150">0.35±0.05 (0.290.43)</td>
<td id="7691F411FFB40074528DFD28FCDB1056" box="[718,812,734,785]" gridcol="3" gridrow="9" pageId="7" pageNumber="150">0.3</td>
<td id="7691F411FFB400745325FD28FC211056" box="[870,982,734,785]" gridcol="4" gridrow="9" pageId="7" pageNumber="150">0.23</td>
<td id="7691F411FFB40074545EFD28FB791056" box="[1053,1166,734,785]" gridcol="5" gridrow="9" pageId="7" pageNumber="150">0.32±0.03 (0.290.38)</td>
<td id="7691F411FFB400745499FD28FABC1056" box="[1242,1355,734,785]" gridcol="6" gridrow="9" pageId="7" pageNumber="150">0.37±0.04 (0.290.42)</td>
</tr>
<tr id="35409D6DFFB4007450DAFCD7FABC1013" box="[153,1355,801,852]" gridrow="10" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFCD7FEF91013" box="[153,270,801,852]" gridcol="0" gridrow="10" pageId="7" pageNumber="150">AGL/SL</th>
<td id="7691F411FFB400745119FCD7FE3D1013" box="[346,458,801,852]" gridcol="1" gridrow="10" pageId="7" pageNumber="150">0.51±0.03 (0.470.58)</td>
<td id="7691F411FFB400745258FCD7FD7B1013" box="[539,652,801,852]" gridcol="2" gridrow="10" pageId="7" pageNumber="150">0.51±0.05 (0.460.58)</td>
<td id="7691F411FFB40074528DFCD7FCDB1013" box="[718,812,801,852]" gridcol="3" gridrow="10" pageId="7" pageNumber="150">0.503</td>
<td id="7691F411FFB400745325FCD7FC211013" box="[870,982,801,852]" gridcol="4" gridrow="10" pageId="7" pageNumber="150">0.51</td>
<td id="7691F411FFB40074545EFCD7FB791013" box="[1053,1166,801,852]" gridcol="5" gridrow="10" pageId="7" pageNumber="150">0.54±0.01 (0.530.55)</td>
<td id="7691F411FFB400745499FCD7FABC1013" box="[1242,1355,801,852]" gridcol="6" gridrow="10" pageId="7" pageNumber="150">0.51±0.02 (0.530.54)</td>
</tr>
<tr id="35409D6DFFB4007450DAFC92FABC10DF" box="[153,1355,868,920]" gridrow="11" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFC92FEF910DF" box="[153,270,868,920]" gridcol="0" gridrow="11" pageId="7" pageNumber="150">HEW/SL</th>
<td id="7691F411FFB400745119FC92FE3D10DF" box="[346,458,868,920]" gridcol="1" gridrow="11" pageId="7" pageNumber="150">0.16±0.01 (0.150.17)</td>
<td id="7691F411FFB400745258FC92FD7B10DF" box="[539,652,868,920]" gridcol="2" gridrow="11" pageId="7" pageNumber="150">0.15±0.01 (0.150.17)</td>
<td id="7691F411FFB40074528DFC92FCDB10DF" box="[718,812,868,920]" gridcol="3" gridrow="11" pageId="7" pageNumber="150">0.156</td>
<td id="7691F411FFB400745325FC92FC2110DF" box="[870,982,868,920]" gridcol="4" gridrow="11" pageId="7" pageNumber="150">0.15±0.01 (0.140.15)</td>
<td id="7691F411FFB40074545EFC92FB7910DF" box="[1053,1166,868,920]" gridcol="5" gridrow="11" pageId="7" pageNumber="150">0.14±0.01 (0.120.16)</td>
<td id="7691F411FFB400745499FC92FABC10DF" box="[1242,1355,868,920]" gridcol="6" gridrow="11" pageId="7" pageNumber="150">0.15±0.01 (0.120.16)</td>
</tr>
<tr id="35409D6DFFB4007450DAFC5EFABC109C" box="[153,1355,936,987]" gridrow="12" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFC5EFEF9109C" box="[153,270,936,987]" gridcol="0" gridrow="12" pageId="7" pageNumber="150">HEW/AGL</th>
<td id="7691F411FFB400745119FC5EFE3D109C" box="[346,458,936,987]" gridcol="1" gridrow="12" pageId="7" pageNumber="150">0.31±0.03 (0.250.36)</td>
<td id="7691F411FFB400745258FC5EFD7B109C" box="[539,652,936,987]" gridcol="2" gridrow="12" pageId="7" pageNumber="150">0.31±0.02 (0.280.34)</td>
<td id="7691F411FFB40074528DFC5EFCDB109C" box="[718,812,936,987]" gridcol="3" gridrow="12" pageId="7" pageNumber="150">0.311</td>
<td id="7691F411FFB400745325FC5EFC21109C" box="[870,982,936,987]" gridcol="4" gridrow="12" pageId="7" pageNumber="150">0.29±0.01 (0.280.29)</td>
<td id="7691F411FFB40074545EFC5EFB79109C" box="[1053,1166,936,987]" gridcol="5" gridrow="12" pageId="7" pageNumber="150">0.26±0.02 (0.230.29)</td>
<td id="7691F411FFB400745499FC5EFABC109C" box="[1242,1355,936,987]" gridcol="6" gridrow="12" pageId="7" pageNumber="150">0.28±0.01 (0.230.3)</td>
</tr>
<tr id="35409D6DFFB4007450DAFC1DFABC1758" box="[153,1355,1003,1055]" gridrow="13" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFC1DFEF91758" box="[153,270,1003,1055]" gridcol="0" gridrow="13" pageId="7" pageNumber="150">SNL/HEW</th>
<td id="7691F411FFB400745119FC1DFE3D1758" box="[346,458,1003,1055]" gridcol="1" gridrow="13" pageId="7" pageNumber="150">0.41±0.04 (0.360.45)</td>
<td id="7691F411FFB400745258FC1DFD7B1758" box="[539,652,1003,1055]" gridcol="2" gridrow="13" pageId="7" pageNumber="150">0.45±0.03 (0.410.48)</td>
<td id="7691F411FFB40074528DFC1DFCDB1758" box="[718,812,1003,1055]" gridcol="3" gridrow="13" pageId="7" pageNumber="150">0.383</td>
<td id="7691F411FFB400745325FC1DFC211758" box="[870,982,1003,1055]" gridcol="4" gridrow="13" pageId="7" pageNumber="150">0.38±0.03 (0.360.4)</td>
<td id="7691F411FFB40074545EFC1DFB791758" box="[1053,1166,1003,1055]" gridcol="5" gridrow="13" pageId="7" pageNumber="150">0.42±0.03 (0.370.46)</td>
<td id="7691F411FFB400745499FC1DFABC1758" box="[1242,1355,1003,1055]" gridcol="6" gridrow="13" pageId="7" pageNumber="150">0.43±0.1 (0.370.49)</td>
</tr>
<tr id="35409D6DFFB4007450DAFBD8FABC1725" box="[153,1355,1070,1122]" gridrow="14" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFBD8FEF91725" box="[153,270,1070,1122]" gridcol="0" gridrow="14" pageId="7" pageNumber="150">HLL/SL</th>
<td id="7691F411FFB400745119FBD8FE3D1725" box="[346,458,1070,1122]" gridcol="1" gridrow="14" pageId="7" pageNumber="150">0.26±0.01 (0.250.27)</td>
<td id="7691F411FFB400745258FBD8FD7B1725" box="[539,652,1070,1122]" gridcol="2" gridrow="14" pageId="7" pageNumber="150">0.27±0.01 (0.260.28)</td>
<td id="7691F411FFB40074528DFBD8FCDB1725" box="[718,812,1070,1122]" gridcol="3" gridrow="14" pageId="7" pageNumber="150">0.271</td>
<td id="7691F411FFB400745325FBD8FC211725" box="[870,982,1070,1122]" gridcol="4" gridrow="14" pageId="7" pageNumber="150">0.24±0.01 (0.230.24)</td>
<td id="7691F411FFB40074545EFBD8FB791725" box="[1053,1166,1070,1122]" gridcol="5" gridrow="14" pageId="7" pageNumber="150">0.24±0.01 (0.220.26)</td>
<td id="7691F411FFB400745499FBD8FABC1725" box="[1242,1355,1070,1122]" gridcol="6" gridrow="14" pageId="7" pageNumber="150">0.26±0.02 (0.220.3)</td>
</tr>
<tr id="35409D6DFFB4007450DAFB84FABC17E2" box="[153,1355,1138,1189]" gridrow="15" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFB84FEF917E2" box="[153,270,1138,1189]" gridcol="0" gridrow="15" pageId="7" pageNumber="150">FLL/SL</th>
<td id="7691F411FFB400745119FB84FE3D17E2" box="[346,458,1138,1189]" gridcol="1" gridrow="15" pageId="7" pageNumber="150">0.24±0.01 (0.220.26)</td>
<td id="7691F411FFB400745258FB84FD7B17E2" box="[539,652,1138,1189]" gridcol="2" gridrow="15" pageId="7" pageNumber="150">0.26±0.05 (0.230.26)</td>
<td id="7691F411FFB40074528DFB84FCDB17E2" box="[718,812,1138,1189]" gridcol="3" gridrow="15" pageId="7" pageNumber="150">0.232</td>
<td id="7691F411FFB400745325FB84FC2117E2" box="[870,982,1138,1189]" gridcol="4" gridrow="15" pageId="7" pageNumber="150">0.23±0.01 (0.220.23)</td>
<td id="7691F411FFB40074545EFB84FB7917E2" box="[1053,1166,1138,1189]" gridcol="5" gridrow="15" pageId="7" pageNumber="150">0.22±0.02 (0.190.24)</td>
<td id="7691F411FFB400745499FB84FABC17E2" box="[1242,1355,1138,1189]" gridcol="6" gridrow="15" pageId="7" pageNumber="150">0.24±0.02 (0.190.27)</td>
</tr>
<tr id="35409D6DFFB4007450DAFB43FABC17AE" box="[153,1355,1205,1257]" gridrow="16" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFB43FEF917AE" box="[153,270,1205,1257]" gridcol="0" gridrow="16" pageId="7" pageNumber="150">HAL/VGS</th>
<td id="7691F411FFB400745119FB43FE3D17AE" box="[346,458,1205,1257]" gridcol="1" gridrow="16" pageId="7" pageNumber="150">0.37±0.03 (0.330.4)</td>
<td id="7691F411FFB400745258FB43FD7B17AE" box="[539,652,1205,1257]" gridcol="2" gridrow="16" pageId="7" pageNumber="150">0.36±0.03 (0.330.41)</td>
<td id="7691F411FFB40074528DFB43FCDB17AE" box="[718,812,1205,1257]" gridcol="3" gridrow="16" pageId="7" pageNumber="150">0.402</td>
<td id="7691F411FFB400745325FB43FC2117AE" box="[870,982,1205,1257]" gridcol="4" gridrow="16" pageId="7" pageNumber="150">0.31±0.03 (0.290.33)</td>
<td id="7691F411FFB40074545EFB43FB7917AE" box="[1053,1166,1205,1257]" gridcol="5" gridrow="16" pageId="7" pageNumber="150">0.35±0.03 (0.310.4)</td>
<td id="7691F411FFB400745499FB43FABC17AE" box="[1242,1355,1205,1257]" gridcol="6" gridrow="16" pageId="7" pageNumber="150">0.37±0.03 (0.310.43)</td>
</tr>
<tr id="35409D6DFFB4007450DAFB0EFABC166B" box="[153,1355,1272,1324]" gridrow="17" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFB0EFEF9166B" box="[153,270,1272,1324]" gridcol="0" gridrow="17" pageId="7" pageNumber="150">FOL/VGS</th>
<td id="7691F411FFB400745119FB0EFE3D166B" box="[346,458,1272,1324]" gridcol="1" gridrow="17" pageId="7" pageNumber="150">0.41±0.02 (0.370.44)</td>
<td id="7691F411FFB400745258FB0EFD7B166B" box="[539,652,1272,1324]" gridcol="2" gridrow="17" pageId="7" pageNumber="150">0.4±0.03 (0.370.43)</td>
<td id="7691F411FFB40074528DFB0EFCDB166B" box="[718,812,1272,1324]" gridcol="3" gridrow="17" pageId="7" pageNumber="150">0.467</td>
<td id="7691F411FFB400745325FB0EFC21166B" box="[870,982,1272,1324]" gridcol="4" gridrow="17" pageId="7" pageNumber="150">0.36±0.01 (0.340.37)</td>
<td id="7691F411FFB40074545EFB0EFB79166B" box="[1053,1166,1272,1324]" gridcol="5" gridrow="17" pageId="7" pageNumber="150">0.39±0.02 (0.370.42)</td>
<td id="7691F411FFB400745499FB0EFABC166B" box="[1242,1355,1272,1324]" gridcol="6" gridrow="17" pageId="7" pageNumber="150">0.42±0.03 (0.370.45)</td>
</tr>
<tr id="35409D6DFFB4007450DAFACAFABC1628" box="[153,1355,1340,1391]" gridrow="18" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFACAFEF91628" box="[153,270,1340,1391]" gridcol="0" gridrow="18" pageId="7" pageNumber="150">HAW/HEW</th>
<td id="7691F411FFB400745119FACAFE3D1628" box="[346,458,1340,1391]" gridcol="1" gridrow="18" pageId="7" pageNumber="150">0.54±0.04 (0.480.59)</td>
<td id="7691F411FFB400745258FACAFD7B1628" box="[539,652,1340,1391]" gridcol="2" gridrow="18" pageId="7" pageNumber="150">0.58±0.07 (0.490.66)</td>
<td id="7691F411FFB40074528DFACAFCDB1628" box="[718,812,1340,1391]" gridcol="3" gridrow="18" pageId="7" pageNumber="150">0.55</td>
<td id="7691F411FFB400745325FACAFC211628" box="[870,982,1340,1391]" gridcol="4" gridrow="18" pageId="7" pageNumber="150">0.52±0.06 (0.470.56)</td>
<td id="7691F411FFB40074545EFACAFB791628" box="[1053,1166,1340,1391]" gridcol="5" gridrow="18" pageId="7" pageNumber="150">0.59±0.04 (0.540.66)</td>
<td id="7691F411FFB400745499FACAFABC1628" box="[1242,1355,1340,1391]" gridcol="6" gridrow="18" pageId="7" pageNumber="150">0.62±0.07 (0.540.76)</td>
</tr>
<tr id="35409D6DFFB4007450DAFA89FABC16F5" box="[153,1355,1407,1458]" gridrow="19" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFA89FEF916F5" box="[153,270,1407,1458]" gridcol="0" gridrow="19" pageId="7" pageNumber="150">FOW/HEW</th>
<td id="7691F411FFB400745119FA89FE3D16F5" box="[346,458,1407,1458]" gridcol="1" gridrow="19" pageId="7" pageNumber="150">0.66±0.06 (0.550.73)</td>
<td id="7691F411FFB400745258FA89FD7B16F5" box="[539,652,1407,1458]" gridcol="2" gridrow="19" pageId="7" pageNumber="150">0.7±0.06 (0.640.79)</td>
<td id="7691F411FFB40074528DFA89FCDB16F5" box="[718,812,1407,1458]" gridcol="3" gridrow="19" pageId="7" pageNumber="150">0.65</td>
<td id="7691F411FFB400745325FA89FC2116F5" box="[870,982,1407,1458]" gridcol="4" gridrow="19" pageId="7" pageNumber="150">0.67±0.07 (0.620.72)</td>
<td id="7691F411FFB40074545EFA89FB7916F5" box="[1053,1166,1407,1458]" gridcol="5" gridrow="19" pageId="7" pageNumber="150">0.7±0.07 (0.630.84)</td>
<td id="7691F411FFB400745499FA89FABC16F5" box="[1242,1355,1407,1458]" gridcol="6" gridrow="19" pageId="7" pageNumber="150">0.73±0.09 (0.630.9)</td>
</tr>
<tr id="35409D6DFFB4007450DAFA34FABC16B1" box="[153,1355,1474,1526]" gridrow="20" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAFA34FEF916B1" box="[153,270,1474,1526]" gridcol="0" gridrow="20" pageId="7" pageNumber="150">LT2/HEW</th>
<td id="7691F411FFB400745119FA34FE3D16B1" box="[346,458,1474,1526]" gridcol="1" gridrow="20" pageId="7" pageNumber="150">0.27±0.03 (0.240.3)</td>
<td id="7691F411FFB400745258FA34FD7B16B1" box="[539,652,1474,1526]" gridcol="2" gridrow="20" pageId="7" pageNumber="150">0.31±0.03 (0.260.32)</td>
<td id="7691F411FFB40074528DFA34FCDB16B1" box="[718,812,1474,1526]" gridcol="3" gridrow="20" pageId="7" pageNumber="150">0.25</td>
<td id="7691F411FFB400745325FA34FC2116B1" box="[870,982,1474,1526]" gridcol="4" gridrow="20" pageId="7" pageNumber="150">0.23±0.02 (0.220.25)</td>
<td id="7691F411FFB40074545EFA34FB7916B1" box="[1053,1166,1474,1526]" gridcol="5" gridrow="20" pageId="7" pageNumber="150">0.33±0.04 (0.270.39)</td>
<td id="7691F411FFB400745499FA34FABC16B1" box="[1242,1355,1474,1526]" gridcol="6" gridrow="20" pageId="7" pageNumber="150">0.35±0.07 (0.270.45)</td>
</tr>
<tr id="35409D6DFFB4007450DAF9F3FABC157E" box="[153,1355,1541,1593]" gridrow="21" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAF9F3FEF9157E" box="[153,270,1541,1593]" gridcol="0" gridrow="21" pageId="7" pageNumber="150">LT3/FOW</th>
<td id="7691F411FFB400745119F9F3FE3D157E" box="[346,458,1541,1593]" gridcol="1" gridrow="21" pageId="7" pageNumber="150">0.55±0.05 (0.480.63)</td>
<td id="7691F411FFB400745258F9F3FD7B157E" box="[539,652,1541,1593]" gridcol="2" gridrow="21" pageId="7" pageNumber="150">0.56±0.08 (0.470.64)</td>
<td id="7691F411FFB40074528DF9F3FCDB157E" box="[718,812,1541,1593]" gridcol="3" gridrow="21" pageId="7" pageNumber="150">0.59</td>
<td id="7691F411FFB400745325F9F3FC21157E" box="[870,982,1541,1593]" gridcol="4" gridrow="21" pageId="7" pageNumber="150">0.48±0.05 (0.440.52)</td>
<td id="7691F411FFB40074545EF9F3FB79157E" box="[1053,1166,1541,1593]" gridcol="5" gridrow="21" pageId="7" pageNumber="150">0.56±0.05 (0.510.65)</td>
<td id="7691F411FFB400745499F9F3FABC157E" box="[1242,1355,1541,1593]" gridcol="6" gridrow="21" pageId="7" pageNumber="150">0.6±0.08 (0.510.71)</td>
</tr>
<tr id="35409D6DFFB4007450DAF9BFFABC153B" box="[153,1355,1609,1660]" gridrow="22" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAF9BFFEF9153B" box="[153,270,1609,1660]" gridcol="0" gridrow="22" pageId="7" pageNumber="150">LT2/FOL</th>
<td id="7691F411FFB400745119F9BFFE3D153B" box="[346,458,1609,1660]" gridcol="1" gridrow="22" pageId="7" pageNumber="150">0.42±0.03 (0.370.48)</td>
<td id="7691F411FFB400745258F9BFFD7B153B" box="[539,652,1609,1660]" gridcol="2" gridrow="22" pageId="7" pageNumber="150">0.47±0.09 (0.350.54)</td>
<td id="7691F411FFB40074528DF9BFFCDB153B" box="[718,812,1609,1660]" gridcol="3" gridrow="22" pageId="7" pageNumber="150">0.349</td>
<td id="7691F411FFB400745325F9BFFC21153B" box="[870,982,1609,1660]" gridcol="4" gridrow="22" pageId="7" pageNumber="150">0.4±0.05 (0.370.43)</td>
<td id="7691F411FFB40074545EF9BFFB79153B" box="[1053,1166,1609,1660]" gridcol="5" gridrow="22" pageId="7" pageNumber="150">0.5±0.05 (0.420.57)</td>
<td id="7691F411FFB400745499F9BFFABC153B" box="[1242,1355,1609,1660]" gridcol="6" gridrow="22" pageId="7" pageNumber="150">0.51±0.1 (0.420.68)</td>
</tr>
<tr id="35409D6DFFB4007450DAF97AFABC1587" box="[153,1355,1676,1728]" gridrow="23" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAF97AFEF91587" box="[153,270,1676,1728]" gridcol="0" gridrow="23" pageId="7" pageNumber="150">LF2/HAL</th>
<td id="7691F411FFB400745119F97AFE3D1587" box="[346,458,1676,1728]" gridcol="1" gridrow="23" pageId="7" pageNumber="150">0.43±0.08 (0.290.58)</td>
<td id="7691F411FFB400745258F97AFD7B1587" box="[539,652,1676,1728]" gridcol="2" gridrow="23" pageId="7" pageNumber="150">0.49±0.08 (0.420.6)</td>
<td id="7691F411FFB40074528DF97AFCDB1587" box="[718,812,1676,1728]" gridcol="3" gridrow="23" pageId="7" pageNumber="150">0.432</td>
<td id="7691F411FFB400745325F97AFC211587" box="[870,982,1676,1728]" gridcol="4" gridrow="23" pageId="7" pageNumber="150">0.56±0.06 (0.520.6)</td>
<td id="7691F411FFB40074545EF97AFB791587" box="[1053,1166,1676,1728]" gridcol="5" gridrow="23" pageId="7" pageNumber="150">0.5±0.05 (0.450.58)</td>
<td id="7691F411FFB400745499F97AFABC1587" box="[1242,1355,1676,1728]" gridcol="6" gridrow="23" pageId="7" pageNumber="150">0.53±0.08 (0.450.63)</td>
</tr>
<tr id="35409D6DFFB4007450DAF926FABC1444" box="[153,1355,1744,1795]" gridrow="24" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAF926FEF91444" box="[153,270,1744,1795]" gridcol="0" gridrow="24" pageId="7" pageNumber="150">WT3/FOW</th>
<td id="7691F411FFB400745119F926FE3D1444" box="[346,458,1744,1795]" gridcol="1" gridrow="24" pageId="7" pageNumber="150">0.2±0.03 (0.160.24)</td>
<td id="7691F411FFB400745258F926FD7B1444" box="[539,652,1744,1795]" gridcol="2" gridrow="24" pageId="7" pageNumber="150">0.17±0.01 (0.160.18)</td>
<td id="7691F411FFB40074528DF926FCDB1444" box="[718,812,1744,1795]" gridcol="3" gridrow="24" pageId="7" pageNumber="150">0.231</td>
<td id="7691F411FFB400745325F926FC211444" box="[870,982,1744,1795]" gridcol="4" gridrow="24" pageId="7" pageNumber="150">0.19±0.03 (0.170.21)</td>
<td id="7691F411FFB40074545EF926FB791444" box="[1053,1166,1744,1795]" gridcol="5" gridrow="24" pageId="7" pageNumber="150">0.16±0.03 (0.140.22)</td>
<td id="7691F411FFB400745499F926FABC1444" box="[1242,1355,1744,1795]" gridcol="6" gridrow="24" pageId="7" pageNumber="150">0.17±0.02 (0.140.21)</td>
</tr>
<tr id="35409D6DFFB4007450DAF8E5FABC1401" box="[153,1355,1811,1862]" gridrow="25" pageId="7" pageNumber="150">
<th id="7691F411FFB4007450DAF8E5FEF91401" box="[153,270,1811,1862]" gridcol="0" gridrow="25" pageId="7" pageNumber="150">WF3/HAW</th>
<td id="7691F411FFB400745119F8E5FE3D1401" box="[346,458,1811,1862]" gridcol="1" gridrow="25" pageId="7" pageNumber="150">0.24±0.03 (0.210.28)</td>
<td id="7691F411FFB400745258F8E5FD7B1401" box="[539,652,1811,1862]" gridcol="2" gridrow="25" pageId="7" pageNumber="150">0.21±0.03 (0.160.26)</td>
<td id="7691F411FFB40074528DF8E5FCDB1401" box="[718,812,1811,1862]" gridcol="3" gridrow="25" pageId="7" pageNumber="150">0.273</td>
<td id="7691F411FFB400745325F8E5FC211401" box="[870,982,1811,1862]" gridcol="4" gridrow="25" pageId="7" pageNumber="150">0.23±0.02 (0.210.24)</td>
<td id="7691F411FFB40074545EF8E5FB791401" box="[1053,1166,1811,1862]" gridcol="5" gridrow="25" pageId="7" pageNumber="150">0.18±0.01 (0.170.21)</td>
<td id="7691F411FFB400745499F8E5FABC1401" box="[1242,1355,1811,1862]" gridcol="6" gridrow="25" pageId="7" pageNumber="150">0.19±0.03 (0.170.24)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C36ACCA4FFBBFF8250D3FE1BFD1F1428" lastPageId="9" lastPageNumber="152" pageId="8" pageNumber="151" type="description">
<paragraph id="8BCF9F2FFFBBFF8350D3FE1BFD7517EF" blockId="8.[141,749,493,1955]" pageId="8" pageNumber="151">
<emphasis id="B904433DFFBBFF8350D3FE1BFE5C1141" bold="true" box="[144,427,493,518]" pageId="8" pageNumber="151">
Description of
<typeStatus id="54CB218DFFBBFF835101FE18FE511141" box="[322,422,494,518]" pageId="8" pageNumber="151" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male having a SL of
<quantity id="4C8832CAFFBBFF8350D3FDFDFF081165" box="[144,255,523,547]" metricMagnitude="-2" metricUnit="m" metricValue="4.904999999999999" pageId="8" pageNumber="151" unit="mm" value="49.05">49.05 mm</quantity>
(
<figureCitation id="134B83AAFFBBFF835152FDFDFEA11164" box="[273,342,523,547]" captionStart="Fig" captionStartId="10.[141,176,688,710]" captionTargetBox="[159,1364,165,650]" captionTargetId="figure-21@10.[138,1397,138,675]" captionTargetPageId="10" captionText="Fig. 4. Holotype of Bolitoglossa bolanosi sp. nov. (UCR 22965) in life on a white background. Photograph by Erick Arias." figureDoi="http://doi.org/10.5281/zenodo.11390653" httpUri="https://zenodo.org/record/11390653/files/figure.png" pageId="8" pageNumber="151">Fig. 4</figureCitation>
). Head slightly wider than neck and shoulders (HeW
<quantity id="4C8832CAFFBBFF835104FDDEFE6A1107" box="[327,413,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="8" pageNumber="151" unit="mm" value="7.3">7.3 mm</quantity>
, NeW
<quantity id="4C8832CAFFBBFF8351AAFDDEFDC81107" box="[489,575,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="151" unit="mm" value="6.2">6.2 mm</quantity>
, ShW
<quantity id="4C8832CAFFBBFF8352CBFDDEFD281107" box="[648,735,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="8" pageNumber="151" unit="mm" value="7.1">7.1 mm</quantity>
), with the greatest width of the head just posterior to the articulation of the jaws; truncate in dorsal outline and rounded to truncate in profile; snout moderate (SnL
<quantity id="4C8832CAFFBBFF83528AFD76FF4B11F2" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="8" pageNumber="151" unit="mm" value="3.4">3.4 mm</quantity>
, 6.9% of SL), with nearly terminal non-protruding small nostrils (LNH
<quantity id="4C8832CAFFBBFF835139FD4DFE251195" box="[378,466,699,723]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="151" unit="mm" value="0.6">0.6 mm</quantity>
, RNW
<quantity id="4C8832CAFFBBFF835268FD4DFD751194" box="[555,642,699,723]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="151" unit="mm" value="0.4">0.4 mm</quantity>
) directed anterolaterally; internarial area convex in dorsal outline. Snout protruding beyond the anterior margin of the lower lip in lateral view. Eyes relatively large (EW = 92% of SnL), weakly protruding beyond the dorsal and ventral outline of the head, directed anterolaterally, with a distinct suborbital groove. Top of head flat and smooth, tapering slightly toward anterior terminus, lacking contrasting interorbital or other dermal structures. Canthus rostralis weakly rounded; intercanthal area flat to slightly convex; and loreal region slightly concave. Evident cirri (nasolabial protuberances) on tip of snout; nasolabial grooves start at ventrolateral margins of nares and terminate in a cirrus. Gular fold is well-defined, starting on the dorsolateral portion of the neck, below the postorbital groove. Evident mental gland is visible under the skin of the anterior intermandibular region.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF8350EEFB58FE41168A" blockId="8.[141,749,493,1955]" pageId="8" pageNumber="151">
Arms relatively long and slender (FLL =
<quantity id="4C8832CAFFBBFF8352C2FB58FD101782" box="[641,743,1198,1222]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="8" pageNumber="151" unit="mm" value="12.6">12.6 mm</quantity>
, 25.7% of SL), without noticeable hypertrophied forearm compared to the upper arm. Hands well-developed and slender (HaL =
<quantity id="4C8832CAFFBBFF835177FAF3FE7F165A" box="[308,392,1285,1309]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="151" unit="mm" value="5.4">5.4 mm</quantity>
, 40.9% of VGS; HaW =
<quantity id="4C8832CAFFBBFF8352D1FAF0FD11165A" box="[658,742,1286,1310]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="151" unit="mm" value="4.8">4.8 mm</quantity>
, 65.8% of HeW). Fingers II, III, and IV protrude freely, with at least the ultimate phalange free beyond interdigital tissue margin (LF2
<quantity id="4C8832CAFFBBFF83511EFAA8FE461632" box="[349,433,1374,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="151" unit="mm" value="2.3">2.3 mm</quantity>
, LF3 3.0 mm;
<figureCitation id="134B83AAFFBBFF83520EFAA8FD7B1632" box="[589,652,1373,1398]" captionStart="Fig" captionStartId="10.[141,176,1873,1895]" captionTargetBox="[141,1394,786,1860]" captionTargetId="figure-43@10.[141,1394,786,1860]" captionTargetPageId="10" captionText="Fig. 5. Dorsal views of the hand and foot of the holotypes of (AB) Bolitoglossa bolanosi sp. nov. (UCR 22965) and (CD) B. pesrubra in preservation. Photograph by Erick Arias." figureDoi="http://doi.org/10.5281/zenodo.11390655" httpUri="https://zenodo.org/record/11390655/files/figure.png" pageId="8" pageNumber="151">Fig. 5</figureCitation>
). Tips of fingers rounded; terminal pads weakly discernible on the ventral surfaces of fingers. Relative lengths of fingers on right hand: I &lt;IV &lt;II &lt;III.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF8350EEFA25FEA81457" blockId="8.[141,749,493,1955]" pageId="8" pageNumber="151">
Legs moderately long and slender (HLL
<quantity id="4C8832CAFFBBFF83523EFA25FD1416AD" box="[637,739,1491,1515]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="151" unit="mm" value="13.1">13.1 mm</quantity>
, 26.7% of SL). Feet well-developed and slender (FoL
<quantity id="4C8832CAFFBBFF8350CEF9FBFF1B1562" box="[141,236,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="151" unit="mm" value="5.5">5.5 mm</quantity>
, 41.7% of VGS; FoW
<quantity id="4C8832CAFFBBFF835248F9FBFD9E1562" box="[523,617,1549,1573]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="8" pageNumber="151" unit="mm" value="5.7">5.7 mm</quantity>
, 78.1% of HeW). Toes II, III, IV, and V protruding freely beyond interdigital tissue margin (LT2
<quantity id="4C8832CAFFBBFF8351AAF9BEFDB61527" box="[489,577,1608,1632]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="151" unit="mm" value="1.9">1.9 mm</quantity>
, LT3
<quantity id="4C8832CAFFBBFF8352C0F9BEFD2B1527" box="[643,732,1608,1632]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="151" unit="mm" value="2.7">2.7 mm</quantity>
), toe I with minimal indentation at interdigital spaces. Toe III is most free of interdigital tissue, with about the entire distal phalanx protruding. Tips of toes rounded; terminal pads weakly discernible on ventral distal surface of toes. Relative lengths of toes on right foot: I &lt;V &lt;II &lt;IV &lt;III.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF8350EEF8E0FC9A1211" blockId="8.[141,749,493,1955]" lastBlockId="8.[791,1396,142,342]" pageId="8" pageNumber="151">Body subcylindrical (slightly wider than high) in cross-section, and relatively slender (TW = 6.0 mm; TW = 26.5% of AGL). Between the axilla and groin, 11 costal grooves are visible, 13 if counting axillary and inguinal grooves; costal grooves are most visible on ventral and lateral portions of the body. Adpressed limbs separated by one costal fold; 12 costal folds total between axilla and groin. The tail is long, cylindrical in cross-section, with an evident constriction at the base, and some caudal grooves discernible on the anterior portion of the tail. The skin on the surfaces of the head, body, limbs, and tail is smooth.</paragraph>
<paragraph id="8BCF9F2FFFBBFF835354FE8EFB0B1021" blockId="8.[791,1397,376,1163]" pageId="8" pageNumber="151">
<emphasis id="B904433DFFBBFF835354FE8EFC2A12D7" bold="true" box="[791,989,376,400]" pageId="8" pageNumber="151">Coloration in life.</emphasis>
The ground color of dorsal surfaces of the head, trunk, tail, hind limbs, and forelimbs is brownish black to brownish violet with numerous fine lighter patches of chromatophores scattered throughout the dorsal surface, especially concentrated on the head. The head and hindlimbs are lighter than the trunk. A pair of irregular dorsolateral stripes, which run from the tail base to the intercanthal area crossing the superior eyelids, are formed by yellowish-bronze blotches. The iris is bright dark bronze with a dark brownish-black reticulation. The upper surfaces of the arms are lighter than the trunk, pinkish brown, with orange blotches in the proximal portion of the humerus and on the ventrolateral surface of the ulna. The upper surface of the legs is similar in color to the trunk, uniform brownish black, except in the foot which is paler. The dorsal and dorsolateral surfaces of the tail are nearly uniform brownish black.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF835374FC9DFC3D17CC" blockId="8.[791,1397,376,1163]" pageId="8" pageNumber="151">The ventrolateral surfaces of the body, tail, hindlimbs, and forelimbs are lighter than the dorsal surface. The gular surfaces are paler than the venter, consisting of lighter brown with a paler blotch on the anterior part, the mental gland. The ventral surface of the trunk and the tail are slightly lighter than the dorsal surface. The ventral surfaces of the arms and legs are lighter than the dorsal surfaces with irregular orange blotches. The palmar and plantar surfaces are light brown with black chromatophores.</paragraph>
<paragraph id="8BCF9F2FFFBBFF835354FB5BFC0E165A" blockId="8.[791,1394,1197,1310]" pageId="8" pageNumber="151">
<emphasis id="B904433DFFBBFF835354FB5BFBE21781" bold="true" box="[791,1045,1197,1222]" pageId="8" pageNumber="151">Coloration in ethanol.</emphasis>
After more than seven years in ethanol (70%), the overall coloration of the
<typeStatus id="54CB218DFFBBFF835557FB3DFA8617A4" box="[1300,1393,1227,1251]" pageId="8" pageNumber="151" type="holotype">holotype</typeStatus>
has darkened throughout and contains a principal dark brown-blackish tone.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF835354FAB6FC5615B4" blockId="8.[791,1394,1344,1779]" pageId="8" pageNumber="151">
<emphasis id="B904433DFFBBFF835354FAB6FC371631" bold="true" pageId="8" pageNumber="151">
Measurements (in mm), limb interval, and percentages of the
<typeStatus id="54CB218DFFBBFF83531DFAA8FC371631" box="[862,960,1374,1398]" pageId="8" pageNumber="151" type="holotype">holotype</typeStatus>
</emphasis>
. SL 49.05; TL 57.5; ShW 7.1; HeW 7.3; NeW 6.2; EW 3.2; SnL 3.4; JSL 8.2; LGFS 12.9; LNH 0.6; RNW 0.4; IND 3.1; NLP 1.1; ICD 3.9; HLL 13.1; FLL 12.6; TW 6.0; VGS 13.2; FSL 15.2; UHL 9.0; AGL 22.8; VL 4.9; HaW 4.8; HaL 5.4; LF2 2.3; LF3 3.0; WF3 1.0; FoW 5.7; FoL 5.5; LT2 1.9; LT3 2.7; WT3 0.9. Limb interval 1. Measurements in relative percentages: VGS/ SL 26.9%; IND/HeW 42.5%; AGL/SL 46.5%; HeW/SL 14.9%; HeW/AGL 32.0%; SnL/ HeW 46.6%; LNH/HeW 8.2%; LNH/SL 1.2%; RNW/HeW 5.5%; RNW/SL 0.8%; HLL/SL 26.7%; FLL/SL 25.7%; HaL/VGS 40.9%; FoL/ VGS 41.7%; HaW/HeW 65.7%; FoW/HeW 78.1%; LT2/ FoL 34,5%; LF2/HaL 42.6%; WT3/FoW 15.8%; WF3/ HaW 20.8%.
</paragraph>
<paragraph id="8BCF9F2FFFBBFF825354F8E0FE9D1606" blockId="8.[791,1394,1813,1955]" lastBlockId="9.[141,745,793,1345]" lastPageId="9" lastPageNumber="152" pageId="8" pageNumber="151">
<emphasis id="B904433DFFBBFF835354F8E0FBEB146A" bold="true" box="[791,1052,1813,1838]" pageId="8" pageNumber="151">Noteworthy variation.</emphasis>
The female
<typeStatus id="54CB218DFFBBFF8354FFF8E0FAD51469" box="[1212,1314,1814,1838]" pageId="8" pageNumber="151" type="paratype">paratypes</typeStatus>
(UCR 22421 and UCR 24245) have more rounded snouts, and much less defined and protruding cirri or nasolabial protuberances, which are sexually dimorphic features. In coloration, this species is extremely polymorphic, some individuals are almost uniform black, which is common in juveniles (UCR 22247,
<figureCitation id="134B83AAFFBAFF8251FAFCC1FDE71008" box="[441,528,823,847]" captionStart="Fig" captionStartId="11.[141,176,1002,1024]" captionTargetBox="[189,1347,161,975]" captionTargetId="figure-495@11.[186,1350,158,978]" captionTargetPageId="11" captionText="Fig. 6. In-life photographs of variations within Bolitoglossa bolanosi sp. nov. (A) Paratopotype male subadult UCR 22966, (B) paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and (D) paratype adult female UCR 24245. Photographs A and C by Erick Arias, B and D by Omar Zúñiga." figureDoi="http://doi.org/10.5281/zenodo.11390657" httpUri="https://zenodo.org/record/11390657/files/figure.png" pageId="9" pageNumber="152">Fig. 6B</figureCitation>
). The
<typeStatus id="54CB218DFFBAFF82521EFCC1FD1F1008" box="[605,744,823,847]" pageId="9" pageNumber="152" type="paratopotype">paratopotype</typeStatus>
(UCR 22964,
<figureCitation id="134B83AAFFBAFF825160FCA2FE82102B" box="[291,373,852,876]" captionStart="Fig" captionStartId="11.[141,176,1002,1024]" captionTargetBox="[189,1347,161,975]" captionTargetId="figure-495@11.[186,1350,158,978]" captionTargetPageId="11" captionText="Fig. 6. In-life photographs of variations within Bolitoglossa bolanosi sp. nov. (A) Paratopotype male subadult UCR 22966, (B) paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and (D) paratype adult female UCR 24245. Photographs A and C by Erick Arias, B and D by Omar Zúñiga." figureDoi="http://doi.org/10.5281/zenodo.11390657" httpUri="https://zenodo.org/record/11390657/files/figure.png" pageId="9" pageNumber="152">Fig. 6A</figureCitation>
) has the pair of dorsolateral stripes more vivid and continuous, the blotches in the upper eyelid are iridescent green; in addition, it has a band of pale orange from posterior head to tail, where it is suffused with the red of the tail. The presence of red on the tail is common; generally the red color in the proximal portion is only on the dorsal surface, but the tail becomes completely red in the distal portion. Several specimens showed a similar coloration morpho “mottled” (
<figureCitation id="134B83AAFFBAFF8252CBFBC9FD2C1710" box="[648,731,1087,1111]" captionStart="Fig" captionStartId="11.[141,176,1002,1024]" captionTargetBox="[189,1347,161,975]" captionTargetId="figure-495@11.[186,1350,158,978]" captionTargetPageId="11" captionText="Fig. 6. In-life photographs of variations within Bolitoglossa bolanosi sp. nov. (A) Paratopotype male subadult UCR 22966, (B) paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and (D) paratype adult female UCR 24245. Photographs A and C by Erick Arias, B and D by Omar Zúñiga." figureDoi="http://doi.org/10.5281/zenodo.11390657" httpUri="https://zenodo.org/record/11390657/files/figure.png" pageId="9" pageNumber="152">Fig. 6C</figureCitation>
), with ground color brownish black to reddish brown with numerous and irregular yellow to red blotches or spots; these specimens lack the dorsolateral stripes. The female UCR 24245 (
<figureCitation id="134B83AAFFBAFF825161FB42FE8F178B" box="[290,376,1204,1228]" captionStart="Fig" captionStartId="11.[141,176,1002,1024]" captionTargetBox="[189,1347,161,975]" captionTargetId="figure-495@11.[186,1350,158,978]" captionTargetPageId="11" captionText="Fig. 6. In-life photographs of variations within Bolitoglossa bolanosi sp. nov. (A) Paratopotype male subadult UCR 22966, (B) paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and (D) paratype adult female UCR 24245. Photographs A and C by Erick Arias, B and D by Omar Zúñiga." figureDoi="http://doi.org/10.5281/zenodo.11390657" httpUri="https://zenodo.org/record/11390657/files/figure.png" pageId="9" pageNumber="152">Fig. 6D</figureCitation>
) has a striking coloration pattern formed by large yellow blotches about a brownish-black ground color; although less evident this specimen has the pair of dorsolateral stripes but the blotches are larger and suffused in the back.
</paragraph>
<caption id="DF0FCFA7FFBAFF8250CCFD28FA9611B3" ID-DOI="http://doi.org/10.5281/zenodo.11390649" ID-Zenodo-Dep="11390649" box="[143,1377,734,756]" httpUri="https://zenodo.org/record/11390649/files/figure.png" pageId="9" pageNumber="152" startId="9.[143,178,734,756]" targetBox="[272,1218,148,713]" targetPageId="9" targetType="figure">
<paragraph id="8BCF9F2FFFBAFF8250CCFD28FA9611B3" blockId="9.[143,1377,734,756]" box="[143,1377,734,756]" pageId="9" pageNumber="152">
<emphasis id="B904433DFFBAFF8250CCFD28FF3811B4" bold="true" box="[143,207,734,756]" pageId="9" pageNumber="152">Fig. 3.</emphasis>
Linear discriminant analysis shows the morphological separation between the new species and the two species close to it.
</paragraph>
</caption>
<paragraph id="8BCF9F2FFFBAFF8250D3FA92FD1F1428" blockId="9.[141,744,1380,1903]" pageId="9" pageNumber="152">
<emphasis id="B904433DFFBAFF8250D3FA92FE0316DE" bold="true" pageId="9" pageNumber="152">
Measurements (in mm), limb intervals, and percentages of the
<typeStatus id="54CB218DFFBAFF82513EFA77FE0716DE" box="[381,496,1409,1433]" pageId="9" pageNumber="152" type="paratype">paratypes</typeStatus>
.
</emphasis>
SL 39.450.0; ShW 4.97.5; HeW 6.37.7; NeW 5.06.7; EW 1.83.2; SnL 2.43.4; JSL 6.08.2; LGFS 10.212.9; IND 1.73.1; NLP 0.71.1; ICD 2.93.9; HLL 9.913.1; FLL 8.9 17.6; TW 5.29.2; VGS 9.613.2; FSL 11.415.2; UHL 6.89.0; AGL 18.626.4; VL 3.55.1; HaW 3.04.8; HaL 3.45.4; LF2 1.32.3; LF3 1.83.0; WF3 0.71.1; FoW 3.45.7; FoL 3.85.5; LT2 1.52.1; LT3 1.72.9; WT3 0.71.2. Limb intervals 35. Measurements in relative percentages: VGS/SL 23.426.9%; IND/HeW 25.8 42.6%; AGL/SL 46.457.6%; HeW/SL 14.516.9%; HeW/AGL 25.235.7%; SnL/HeW 35.747.6%; HLL/ SL 24.626.8%; FLL/SL 22.326.1%; HaL/VGS 32.7 40.5%; FoL/VGS 37.144.4%; HaW/HeW 48.066.3%; FoW/HeW 54.578.6%; LT2/HeW 24.131.9%; LT3/ FoW 46.763.0%; LT2/FoL 34.945.8%; LF2/HaL 29.3 46.9%; WT3/FoW 15.524.1%; WF3/HaW 16.228.3%.
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFBAFF8250CEF867FA8610A5" pageId="9" pageNumber="152" type="etymology">
<paragraph id="8BCF9F2FFFBAFF8250CEF867FA8610A5" blockId="9.[141,744,1937,1962]" lastBlockId="9.[791,1396,793,994]" pageId="9" pageNumber="152">
<emphasis id="B904433DFFBAFF8250CEF867FEF914EE" bold="true" box="[141,270,1937,1961]" pageId="9" pageNumber="152">Etymology.</emphasis>
The name
<taxonomicName id="4C70E4ACFFBAFF82513DF867FE1114EE" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[382,486,1937,1961]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF8251C9F867FE1114EE" box="[394,486,1937,1961]" italics="true" pageId="9" pageNumber="152">bolanosi</emphasis>
</taxonomicName>
” is a patronym honoring the Costa Rican herpetologist Federico Bolaños, and is used as a noun in the genitive case. We name this species after our dear friend in recognition of his scientific contributions to the knowledge of the herpetology of
<collectingCountry id="F367DFBFFFBAFF825354FC79FC6210E0" box="[791,917,911,935]" name="Costa Rica" pageId="9" pageNumber="152">Costa Rica</collectingCountry>
, as curator of the Herpetology section at Museo de Zoología of Universidad de
<collectingCountry id="F367DFBFFFBAFF8254F3FC5AFAD01083" box="[1200,1319,940,964]" name="Costa Rica" pageId="9" pageNumber="152">Costa Rica</collectingCountry>
, and as the mentor of most herpetologists present in the country.
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFBAFF805354FBF2FEF917D1" lastPageId="11" lastPageNumber="154" pageId="9" pageNumber="152" type="biology_ecology">
<paragraph id="8BCF9F2FFFBAFF825354FBF2FAF21693" blockId="9.[791,1399,1028,1962]" pageId="9" pageNumber="152">
<emphasis id="B904433DFFBAFF825354FBF2FB04175B" bold="true" box="[791,1267,1028,1052]" pageId="9" pageNumber="152">Habitat and natural history observations.</emphasis>
The habitat of
<taxonomicName id="4C70E4ACFFBAFF825376FBD7FBE9177E" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[821,1054,1057,1081]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF825376FBD7FBE9177E" box="[821,1054,1057,1081]" italics="true" pageId="9" pageNumber="152">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
in the subalpine rain páramo is characterized by having a very short dry season (one to two months), annual precipitation ranging from
<quantity id="4C8832CAFFBAFF825578FBAAFC5217D6" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="152" unit="mm" value="1500.0" valueMax="2000.0" valueMin="1000.0">1,000 to 2,000 mm</quantity>
, and annual temperatures between 3 and 6 ºC. The páramo vegetation consists of extensive, deep moss mats, spongy soil, ferns, and small isolated trees with arboreal bromeliads.
<taxonomicName id="4C70E4ACFFBAFF825408FB27FACA17AE" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[1099,1341,1233,1257]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF825408FB27FACA17AE" box="[1099,1341,1233,1257]" italics="true" pageId="9" pageNumber="152">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
was found within both moss and bromeliads. It also occurs in Montane rain forest which is dominated by
<taxonomicName id="4C70E4ACFFBAFF82555EFAFAFA811664" box="[1309,1398,1292,1315]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="152" phylum="Tracheophyta" rank="genus">
<emphasis id="B904433DFFBAFF82555EFAFAFA811664" box="[1309,1398,1292,1315]" italics="true" pageId="9" pageNumber="152">Quercus</emphasis>
</taxonomicName>
covered with moss, with a great abundance of bryophytes and epiphytes. The
<typeStatus id="54CB218DFFBAFF8253ACFAB1FBEA1618" box="[1007,1053,1351,1375]" pageId="9" pageNumber="152">type</typeStatus>
locality occurs in the Montane rain forest (
<bibRefCitation id="EFE1E2DEFFBAFF8253EBFA92FB69163B" author="Bolanos R &amp; Watson V &amp; Tosi J." box="[936,1182,1380,1404]" pageId="9" pageNumber="152" pagination="750" refId="ref10978" refString="Bolanos R, Watson V, Tosi J. 2005. Mapa Ecologico de Costa Rica (Zonas de Vida), Segun el Sistema de Clasificacion de Zonas de Vida del Mundo de L. R. Holdridge, Escala 1: 750 000. Centro Cientifico Tropical, San Jose, Costa Rica." type="journal article" year="2005">Bolaños et al. 2005</bibRefCitation>
;
<bibRefCitation id="EFE1E2DEFFBAFF8254F7FA92FA9C163B" author="Holdridge LR" box="[1204,1387,1380,1404]" pageId="9" pageNumber="152" refId="ref11344" refString="Holdridge LR. 1967. Life Zone Ecology. Revised edition. Tropical Science Center, San Jose, Costa Rica. 206 p." type="book" year="1967">Holdridge 1967</bibRefCitation>
), characterized by a very short dry season (one to two months), an annual precipitation range of
<quantity id="4C8832CAFFBAFF825498FA69FCB41694" metricMagnitude="0" metricUnit="m" metricValue="3.35" metricValueMax="4.5" metricValueMin="2.2" pageId="9" pageNumber="152" unit="mm" value="3350.0" valueMax="4500.0" valueMin="2200.0">2,200 to 4,500 mm</quantity>
, and annual temperatures from 6 to 12 ºC.
</paragraph>
<paragraph id="8BCF9F2FFFBAFF805374FA2CFEF917D1" blockId="9.[791,1399,1028,1962]" lastBlockId="11.[141,745,1120,1174]" lastPageId="11" lastPageNumber="154" pageId="9" pageNumber="152">
Very little is known about the natural history of
<taxonomicName id="4C70E4ACFFBAFF825523FA2CFC841548" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF825523FA2CFC841548" italics="true" pageId="9" pageNumber="152">B. bolanosi</emphasis>
</taxonomicName>
, but it is important to note that females guarding a clutch of eggs were found on Cerro Hakú (
<date id="FFCEB9EFFFBAFF825544F9E2FCB9150E" pageId="9" pageNumber="152" value="2015-12">December 2015</date>
) and Cerro Dúrika (
<date id="FFCEB9EFFFBAFF825461F9C7FB42150E" box="[1058,1205,1585,1609]" pageId="9" pageNumber="152" value="2016-01">January 2016</date>
), both under moss on the floor. All specimens from Cerro Utyum were found within bromeliads at heights of&gt;
<quantity id="4C8832CAFFBAFF825431F99AFB6E15C4" box="[1138,1177,1644,1668]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="152" unit="m" value="3.0">3 m</quantity>
over the soil, despite active searches in the moss of the páramo. Unlike in the peaks Dúrika, Arbolado, and Hakú, the specimens were mainly found in the moss at &lt;
<quantity id="4C8832CAFFBAFF825413F932FB8F159C" box="[1104,1144,1732,1756]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="152" unit="m" value="2.0">2 m</quantity>
over soil, despite active searches in the bromeliads at&gt;
<quantity id="4C8832CAFFBAFF825438F917FB5F15BE" box="[1147,1192,1761,1785]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="152" unit="m" value="3.0">3 m</quantity>
above the ground (especially on Cerro Arbolado and Cerro Hakú). In the summit of Cerro Dúrika,
<taxonomicName id="4C70E4ACFFBAFF825461F8EBFB6C1473" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[1058,1179,1820,1844]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF825461F8EBFB6C1473" box="[1058,1179,1820,1844]" italics="true" pageId="9" pageNumber="152">B. bolanosi</emphasis>
</taxonomicName>
is very near (~
<quantity id="4C8832CAFFBAFF825574F8EAFA871473" box="[1335,1392,1820,1844]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="152" unit="km" value="4.0">4 km</quantity>
) to a site with
<taxonomicName id="4C70E4ACFFBAFF8253E7F8CCFBD61416" box="[932,1057,1849,1873]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B904433DFFBAFF8253E7F8CCFBD61416" box="[932,1057,1849,1873]" italics="true" pageId="9" pageNumber="152">B. pesrubra</emphasis>
</taxonomicName>
. In the summit of Cerro Utyum, it is to only ~
<quantity id="4C8832CAFFBAFF8253DCF8A1FC231428" box="[927,980,1879,1903]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="152" unit="km" value="7.0">7 km</quantity>
to a site with
<emphasis id="B904433DFFBAFF825427F8A1FB3A1428" box="[1124,1229,1879,1903]" italics="true" pageId="9" pageNumber="152">
<taxonomicName id="4C70E4ACFFBAFF825427F8A1FB3E1428" box="[1124,1225,1879,1903]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="kamuk">B. kamuk</taxonomicName>
,
</emphasis>
and in this peak
<taxonomicName id="4C70E4ACFFBAFF82535AF883FC6214CB" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[793,917,1908,1932]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFBAFF82535AF883FC6214CB" box="[793,917,1908,1932]" italics="true" pageId="9" pageNumber="152">B. bolanosi</emphasis>
</taxonomicName>
is sympatric with an unnamed miniaturized salamander related to
<emphasis id="B904433DFFBAFF825449F864FB6514EE" box="[1034,1170,1938,1961]" italics="true" pageId="9" pageNumber="152">
<taxonomicName id="4C70E4ACFFBAFF825449F864FB7914EE" authorityName="Bolanos &amp; Wake" authorityYear="2009" box="[1034,1166,1938,1961]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="152" phylum="Chordata" rank="species" species="pygmaea">B. pygmaea</taxonomicName>
.
</emphasis>
In addition, the
<typeStatus id="54CB218DFFBAFF82550AF864FA8014ED" box="[1353,1399,1938,1962]" pageId="9" pageNumber="152">type</typeStatus>
locality of
<taxonomicName id="4C70E4ACFFB8FF805141FB97FE7F173F" box="[258,392,1120,1144]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="splendida">
<emphasis id="B904433DFFB8FF805141FB97FE7F173F" box="[258,392,1120,1144]" italics="true" pageId="11" pageNumber="154">B. splendida</emphasis>
</taxonomicName>
is only ~
<quantity id="4C8832CAFFB8FF8051AEFB96FDD3173F" box="[493,548,1120,1144]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="154" unit="km" value="4.0">4 km</quantity>
from the site with
<emphasis id="B904433DFFB8FF8050CEFB88FEF917D1" box="[141,270,1150,1174]" italics="true" pageId="11" pageNumber="154">
<taxonomicName id="4C70E4ACFFB8FF8050CEFB88FEFD17D1" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[141,266,1150,1174]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="bolanosi">B. bolanosi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF0FCFA7FFB9FF8150CEFD46FADA1181" ID-DOI="http://doi.org/10.5281/zenodo.11390653" ID-Zenodo-Dep="11390653" box="[141,1325,688,710]" httpUri="https://zenodo.org/record/11390653/files/figure.png" pageId="10" pageNumber="153" startId="10.[141,176,688,710]" targetBox="[159,1364,165,650]" targetPageId="10" targetType="figure">
<paragraph id="8BCF9F2FFFB9FF8150CEFD46FADA1181" blockId="10.[141,1325,688,710]" box="[141,1325,688,710]" pageId="10" pageNumber="153">
<emphasis id="B904433DFFB9FF8150CEFD46FF3A1182" bold="true" box="[141,205,688,710]" pageId="10" pageNumber="153">Fig. 4.</emphasis>
Holotype of
<taxonomicName id="4C70E4ACFFB9FF81510FFD46FDEA1182" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[332,541,688,709]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="153" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB9FF81510FFD46FDEA1182" box="[332,541,688,709]" italics="true" pageId="10" pageNumber="153">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB9FF815260FD46FD981182" bold="true" box="[547,623,688,709]" pageId="10" pageNumber="153">
<taxonomicNameLabel id="A237FE46FFB9FF815260FD46FD981182" box="[547,623,688,709]" pageId="10" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UCR 22965) in life on a white background.
<emphasis id="B904433DFFB9FF815466FD46FAD01181" box="[1061,1319,688,710]" italics="true" pageId="10" pageNumber="153">Photograph by Erick Arias</emphasis>
.
</paragraph>
</caption>
<caption id="DF0FCFA7FFB9FF8150CEF8A7FD7914C4" ID-DOI="http://doi.org/10.5281/zenodo.11390655" ID-Zenodo-Dep="11390655" httpUri="https://zenodo.org/record/11390655/files/figure.png" pageId="10" pageNumber="153" startId="10.[141,176,1873,1895]" targetBox="[141,1394,786,1860]" targetPageId="10" targetType="figure">
<paragraph id="8BCF9F2FFFB9FF8150CEF8A7FD7914C4" blockId="10.[141,1395,1873,1924]" pageId="10" pageNumber="153">
<emphasis id="B904433DFFB9FF8150CEF8A7FF391421" bold="true" box="[141,206,1873,1895]" pageId="10" pageNumber="153">Fig. 5.</emphasis>
Dorsal views of the hand and foot of the holotypes of
<emphasis id="B904433DFFB9FF8152AEF8A7FCDD1420" bold="true" box="[749,810,1873,1895]" pageId="10" pageNumber="153">(AB)</emphasis>
<taxonomicName id="4C70E4ACFFB9FF815371F8A7FBF31421" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[818,1028,1873,1894]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="153" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB9FF815371F8A7FBF31421" box="[818,1028,1873,1894]" italics="true" pageId="10" pageNumber="153">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<emphasis id="B904433DFFB9FF815448F8A7FBAE1421" bold="true" box="[1035,1113,1873,1894]" pageId="10" pageNumber="153">
<taxonomicNameLabel id="A237FE46FFB9FF815448F8A7FBAE1421" box="[1035,1113,1873,1894]" pageId="10" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(UCR 22965) and
<emphasis id="B904433DFFB9FF81555BF8A7FAA11420" bold="true" box="[1304,1366,1873,1895]" pageId="10" pageNumber="153">(CD)</emphasis>
<taxonomicName id="4C70E4ACFFB9FF81551DF8A4FF1314C4" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="10" pageNumber="153" phylum="Chordata" rank="species" species="pesrubra">
<emphasis id="B904433DFFB9FF81551DF8A4FF1314C4" italics="true" pageId="10" pageNumber="153">B. pesrubra</emphasis>
</taxonomicName>
in preservation.
<emphasis id="B904433DFFB9FF8151C6F898FD7F14C4" box="[389,648,1902,1923]" italics="true" pageId="10" pageNumber="153">Photograph by Erick Arias</emphasis>
.
</paragraph>
</caption>
<caption id="DF0FCFA7FFB8FF8050CEFC1CFDA9177E" ID-DOI="http://doi.org/10.5281/zenodo.11390657" ID-Zenodo-Dep="11390657" httpUri="https://zenodo.org/record/11390657/files/figure.png" pageId="11" pageNumber="154" startId="11.[141,176,1002,1024]" targetBox="[189,1347,161,975]" targetPageId="11" targetType="figure">
<paragraph id="8BCF9F2FFFB8FF8050CEFC1CFDA9177E" blockId="11.[141,1394,1002,1082]" pageId="11" pageNumber="154">
<emphasis id="B904433DFFB8FF8050CEFC1CFF3910B8" bold="true" box="[141,206,1002,1024]" pageId="11" pageNumber="154">Fig. 6.</emphasis>
In-life photographs of variations within
<taxonomicName id="4C70E4ACFFB8FF80521CFC1CFCC510B8" authority="Arias &amp; Chaves &amp; Parra-Olea, 2023" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" box="[607,818,1002,1023]" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="bolanosi" status="sp. nov.">
<emphasis id="B904433DFFB8FF80521CFC1CFCC510B8" box="[607,818,1002,1023]" italics="true" pageId="11" pageNumber="154">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237FE46FFB8FF80537AFC1CFC7E1747" box="[825,905,1002,1024]" pageId="11" pageNumber="154" rank="species">
<emphasis id="B904433DFFB8FF80537AFC1CFC7510B8" bold="true" box="[825,898,1002,1023]" pageId="11" pageNumber="154">sp. nov</emphasis>
.
</taxonomicNameLabel>
<emphasis id="B904433DFFB8FF8053D2FC1CFC451747" bold="true" box="[913,946,1002,1024]" pageId="11" pageNumber="154">(A)</emphasis>
Paratopotype male subadult UCR 22966,
<emphasis id="B904433DFFB8FF805510FC1CFA851747" bold="true" box="[1363,1394,1002,1024]" pageId="11" pageNumber="154">(B)</emphasis>
paratype juvenile UCR 24247, (C) paratype adult female UCR 22421, and
<emphasis id="B904433DFFB8FF805321FBF1FC73175A" bold="true" box="[866,900,1031,1053]" pageId="11" pageNumber="154">(D)</emphasis>
paratype adult female UCR 24245.
<emphasis id="B904433DFFB8FF8054A1FBF1FDAF177E" italics="true" pageId="11" pageNumber="154">Photographs A and C by Erick Arias, B and D by Omar Zúñiga</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C36ACCA4FFB8FF8050CEFB4EFD7A16D9" pageId="11" pageNumber="154" type="distribution">
<paragraph id="8BCF9F2FFFB8FF8050CEFB4EFD7A16D9" blockId="11.[141,746,1208,1438]" pageId="11" pageNumber="154">
<emphasis id="B904433DFFB8FF8050CEFB4EFED71797" bold="true" box="[141,288,1208,1232]" pageId="11" pageNumber="154">Distribution.</emphasis>
The known distribution area of
<taxonomicName id="4C70E4ACFFB8FF805227FB4EFF1E17A9" authorityName="Arias &amp; Chaves &amp; Parra-Olea" authorityYear="2023" class="Amphibia" family="Plethodontidae" genus="Bolitoglossa" kingdom="Animalia" order="Caudata" pageId="11" pageNumber="154" phylum="Chordata" rank="species" species="bolanosi">
<emphasis id="B904433DFFB8FF805227FB4EFF1E17A9" italics="true" pageId="11" pageNumber="154">Bolitoglossa bolanosi</emphasis>
</taxonomicName>
is very small, restricted throughout to ~
<quantity id="4C8832CAFFB8FF8052E7FB20FD1E17A9" box="[676,745,1238,1262]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="154" unit="km" value="15.0">15 km</quantity>
on the summits of the peaks Dúrika, Arbolado, Hakú, and Utyum on the Cordillera de Talamanca (
<figureCitation id="134B83AAFFB8FF805229FAE6FD5E166F" box="[618,681,1296,1320]" captionStart="Fig" captionStartId="1.[139,174,894,916]" captionTargetBox="[142,1392,142,876]" captionTargetId="figure-526@1.[139,1392,141,878]" captionTargetPageId="1" captionText="Fig. 1. (A) Map showing the known populations of the species within the Bolitoglossa subpalmata species group. (B) Map showing the localities for B. bolanosi sp. nov. and the species close to it in the páramo of the Cordillera de Talamanca. The open shapes indicate the historical data and the solid shapes indicate the localities with molecular data included herein." figureDoi="http://doi.org/10.5281/zenodo.11390645" httpUri="https://zenodo.org/record/11390645/files/figure.png" pageId="11" pageNumber="154">Fig. 1</figureCitation>
). The altitudinal range of the new species is
<quantity id="4C8832CAFFB8FF805207FAD8FD1D1602" box="[580,746,1326,1350]" metricMagnitude="3" metricUnit="m" metricValue="2.895" metricValueMax="3.2399999999999998" metricValueMin="2.55" pageId="11" pageNumber="154" unit="m" value="2895.0" valueMax="3240.0" valueMin="2550.0">
<geoCoordinate id="EE44F9E8FFB8FF805207FAD8FD771601" box="[580,640,1326,1350]" degrees="2,550" direction="north" orientation="latitude" pageId="11" pageNumber="154" precision="55" value="2.55">2,550</geoCoordinate>
<geoCoordinate id="EE44F9E8FFB8FF8052C3FAD8FD3D1601" box="[640,714,1326,1350]" degrees="3,240" direction="west" orientation="longitude" pageId="11" pageNumber="154" precision="55" value="-3.24">3,240</geoCoordinate>
m
</quantity>
asl. All the populations of the new species were found in primary vegetation (páramo and forest) and all are within a protected area, La Amistad International Park.
</paragraph>
</subSubSection>
<subSubSection id="C36ACCA4FFB8FF8050CEFA36FE6615E1" pageId="11" pageNumber="154" type="conservation">
<paragraph id="8BCF9F2FFFB8FF8050CEFA36FE6615E1" blockId="11.[141,746,1472,1702]" pageId="11" pageNumber="154">
<emphasis id="B904433DFFB8FF8050CEFA36FE8A169F" bold="true" box="[141,381,1472,1496]" pageId="11" pageNumber="154">Conservation status.</emphasis>
The conservation status of this species is uncertain; however, its known distribution range is small (&lt;
<quantity id="4C8832CAFFB8FF805106FA0DFE4B1554" box="[325,444,1531,1555]" metricMagnitude="7" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="154" unit="km" value="20000.0">20,000 km</quantity>
<superScript id="7C053267FFB8FF8051FFFA0FFE331540" attach="left" box="[444,452,1529,1543]" fontSize="6" pageId="11" pageNumber="154">2</superScript>
) and all know populations are restricted to summits of peaks, sites that are highly vulnerable to forest fires and other effects of climate change. We suggest that it should be tentatively considered as Vulnerable (VU) following the IUCN (2017) category criteria.
</paragraph>
</subSubSection>
</treatment>
</document>