246 lines
32 KiB
XML
246 lines
32 KiB
XML
<document id="35C29DFDA95EB1A1D8FCF5D79F41AE7F" ID-DOI="10.11646/zootaxa.5370.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10147814" ID-ZooBank="48903495-7C6C-46E4-9B1B-D6A2F2781873" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1700202702953" checkinUser="plazi" docAuthor="Harzhauser, Mathias & Landau, Bernard" docDate="2023" docId="03BEE17BFFDDC078FF0EFD5578D030C9" docLanguage="en" docName="zootaxa.5370.1.1.pdf" docOrigin="Zootaxa 5370 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5370.1.1/52270" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Heliacus globosus Harzhauser & Landau 2023, nov. sp." docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FF879903FFC1C064FF99FFF27C023738" masterDocTitle="The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition" masterLastPageNumber="74" masterPageNumber="1" pageNumber="29" updateTime="1706534291121" updateUser="carolina">
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<mods:title id="BFBD9BA3EA9AF172B733022D43746A5E">The Architectonicidae and Mathildidae (Gastropoda, Heterobranchia) of the Miocene Paratethys Sea-victims of the Miocene Climatic Transition</mods:title>
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<mods:namePart id="E37221E42B89EB5E61A3BC31B63690D4">Harzhauser, Mathias</mods:namePart>
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<mods:affiliation id="D95F63B31F7EC08B347B2BC2F296D395">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<mods:nameIdentifier id="1DCDBA1E7723E19A539A79A0F9E9B520" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:namePart id="136976D16271F7BB1C36A6CEF33C81F2">Landau, Bernard</mods:namePart>
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<mods:affiliation id="65A5823CA7F60C76F632523A8A49E7FA">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655; & Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 & Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655;</mods:affiliation>
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<mods:nameIdentifier id="E3023E78FDB04056C13E8E4752D98EFC" type="email">mathias.harzhauser@nhm-wien.ac.at</mods:nameIdentifier>
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<mods:date id="BFBF6A7B13F7098E880AE2107B4F850B">2023</mods:date>
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<mods:number id="8FA5CB7343A3D69F248BDBAABCB24872">2023-11-14</mods:number>
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<heading id="D0E0E701FFDDC078FF0EFD557DD835FA" bold="true" box="[151,474,679,706]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
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<taxonomicName id="4C172BEEFFDDC078FF0EFD557D7635FA" ID-CoL="BXSL8" authority="Harzhauser & Landau, 2023" authorityName="Harzhauser & Landau" authorityYear="2023" box="[151,372,679,706]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="globosus" status="sp. nov.">
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<emphasis id="B9638C7FFFDDC078FF0EFD557D7635FA" bold="true" box="[151,372,679,706]" italics="true" pageId="28" pageNumber="29">Heliacus globosus</emphasis>
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<taxonomicNameLabel id="A2503104FFDDC078FEE2FD5A7DD835FA" box="[379,474,680,706]" pageId="28" pageNumber="29" rank="species">nov. sp.</taxonomicNameLabel>
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<paragraph id="8BA8506DFFDDC078FF0EFD3E7D5835D4" blockId="28.[151,474,679,748]" box="[151,346,716,748]" pageId="28" pageNumber="29">
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<figureCitation id="132C4CE8FFDDC078FF0EFD3E7CF235DE" box="[151,240,716,742]" captionStart="FIGURE 4" captionStartId="9.[151,250,1590,1614]" captionTargetBox="[151,1436,151,1566]" captionTargetId="figure-233@9.[151,1436,151,1569]" captionTargetPageId="9" captionText="FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale." figureDoi="http://doi.org/10.5281/zenodo.10147820" httpUri="https://zenodo.org/record/10147820/files/figure.png" pageId="28" pageNumber="29">Figs 4L</figureCitation>
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,
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<figureCitation id="132C4CE8FFDDC078FF65FD3E7D2B35DE" box="[252,297,716,742]" captionStart="FIGURE 11" captionStartId="22.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-30@22.[151,1436,181,1185]" captionTargetPageId="22" captionText="FIGURE 11 A1–A3. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Romania). B1–B3. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). C1–C3. Nipteraxis pozsgayae (Kovács Leél-Őssy & Vicián, 2023), holotype, PAL 2022.52, Ottnangian, Parádfürdö, Ilona Valley (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.10147834" httpUri="https://zenodo.org/record/10147834/files/figure.png" pageId="28" pageNumber="29">11B</figureCitation>
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–B
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<figureCitation id="132C4CE8FFDDC078FECBFD2C7D5835D4" box="[338,346,734,748]" captionStart="FIGURE 3" captionStartId="6.[151,250,2002,2026]" captionTargetId="figure-99@6.[337,1250,1242,1978]" captionTargetPageId="6" captionText="FIGURE 3. Descriptive terms used in descriptions of Mathildidae." figureDoi="http://doi.org/10.5281/zenodo.10307672" httpUri="https://zenodo.org/record/10307672/files/figure.png" pageId="28" pageNumber="29">
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<emphasis id="B9638C7FFFDDC078FF0EFCFD7D443410" bold="true" box="[151,326,782,809]" pageId="28" pageNumber="29">Type material.</emphasis>
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<specimenCode id="DBB1F816FFDDC078FE27FCFD7E5B3411" box="[446,601,782,809]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="29" type="Museum">NHMW 1907</specimenCode>
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/0328/1959a, SL:
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, MD:
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<quantity id="4CEFFD88FFDDC078FC51FCFD782D3411" box="[968,1071,783,809]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="28" pageNumber="29" unit="mm" value="10.7">10.7 mm</quantity>
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, M̂llersdorf (
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<collectingCountry id="F30010FDFFDDC078FB48FCFD792B3411" box="[1233,1321,783,809]" name="Austria" pageId="28" pageNumber="29">Austria</collectingCountry>
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),
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<figureCitation id="132C4CE8FFDDC078FAA7FCFD799A3411" box="[1342,1432,783,809]" captionStart="FIGURE 4" captionStartId="9.[151,250,1590,1614]" captionTargetBox="[151,1436,151,1566]" captionTargetId="figure-233@9.[151,1436,151,1569]" captionTargetPageId="9" captionText="FIGURE 4 Protoconchs of Paratethyan Architectonicidae. A. Ammotectonica gregorovae nov. sp., paratype, NHMW 1862/0029/0078c. B. Simplexollata anticollata nov. sp., paratype, NHMW 1854/0035/0331, Lăpugiu de Sus (Romania). C. Simplexollata carocollata (Lamarck, 1822), NHMW 1850/0018/0115, Léognan (France), Burdigalian. D. Simplexollata simplex (Bronn, 1831), NHMW 2013/0078/0109, Baden (Austria).E. Nipteraxis deformatusnov. sp., paratype, NHMW 2010/0004/0456a, Bad Vöslau (Austria). F. Nipteraxis exmoniliferus (Sacco, 1892), NHMW 1907/0323/1959, M̂llersdorf (Austria). G. Solatectonica kostejana (Boettger, 1907), NHMW 2012/0229/0049b, Traiskirchen (Austria). H. Nipteraxis monilifer (Bronn, 1831), NHMW 1846/0037/0427, M̂llersdorf (Austria). I. Solatisonax tavianii nov. sp., NHMW2016/0266/0002, SL: 8.0 mm, MD: 15.0 mm, Lăpugiu des Sus (Romania). J. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Roma- nia). K. Granosolarium semilaevis (Sacco, 1892), NHMW 1846/0037/0428d, M̂llersdorf (Austria). L. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). M. Heliacus miser (Dujardin, 1837), NHMW 2023/0267/0001, Steinebrunn (Austria). N. Pseudotorinia grasemanni nov. sp., paratype, NHMW 2023/0267/0010, Steinebrunn (Austria). O. Pseudotorinia marthae (Boettger, 1902), NHMW 2012/0229/0049, Traiskirchen (Austria). All figures same scale." figureDoi="http://doi.org/10.5281/zenodo.10147820" httpUri="https://zenodo.org/record/10147820/files/figure.png" pageId="28" pageNumber="29">Figs 4L</figureCitation>
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,
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<figureCitation id="132C4CE8FFDDC078FF0EFCC17CC63475" box="[151,196,819,845]" captionStart="FIGURE 11" captionStartId="22.[151,250,1209,1233]" captionTargetBox="[151,1436,181,1185]" captionTargetId="figure-30@22.[151,1436,181,1185]" captionTargetPageId="22" captionText="FIGURE 11 A1–A3. Solatisonax transversa nov. sp., holotype, NHMW 2016/0177/0341, Lăpugiu de Sus (Romania). B1–B3. Heliacus globosus nov. sp., holotype NHMW 1907/0328/1959a, M̂llersdorf (Austria). C1–C3. Nipteraxis pozsgayae (Kovács Leél-Őssy & Vicián, 2023), holotype, PAL 2022.52, Ottnangian, Parádfürdö, Ilona Valley (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.10147834" httpUri="https://zenodo.org/record/10147834/files/figure.png" pageId="28" pageNumber="29">11B</figureCitation>
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–B
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.
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locality.
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(
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||
),
|
||
<collectingRegion id="49D39E8FFFDDC078FDF5FCA57EB93449" box="[620,699,855,881]" country="Austria" name="Wien" pageId="28" pageNumber="29">Vienna</collectingRegion>
|
||
<location id="8EC806B6FFDDC078FD5BFCA57F073449" LSID="urn:lsid:plazi:treatment:03BEE17BFFDDC078FF0EFD5578D030C9:8EC806B6FFDDC078FD5BFCA57F073449" box="[706,773,855,881]" country="Austria" municipality="Mollersdorf" name="Basin" pageId="28" pageNumber="29" stateProvince="Wien">Basin</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA8506DFFDDC078FF5EFC897EF834AD" blockId="28.[151,1437,782,2035]" box="[199,762,891,917]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFC897D6D34AD" bold="true" box="[199,367,891,917]" pageId="28" pageNumber="29">
|
||
<typeStatus id="54ACEECFFFDDC078FF5EFC897D0034AD" box="[199,258,891,917]" pageId="28" pageNumber="29">Type</typeStatus>
|
||
stratum.
|
||
</emphasis>
|
||
Silty clay of the Baden Formation.
|
||
</paragraph>
|
||
<paragraph id="8BA8506DFFDDC078FF5EFC6C7F173481" blockId="28.[151,1437,782,2035]" box="[199,789,926,953]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFC6C7CFF3480" bold="true" box="[199,253,926,952]" pageId="28" pageNumber="29">Age.</emphasis>
|
||
Middle Miocene, middle Badenian (Langhian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D03E6FFDDC078FF5EFC31788934E5" box="[199,1163,963,989]" pageId="28" pageNumber="29" type="etymology">
|
||
<paragraph id="8BA8506DFFDDC078FF5EFC31788934E5" blockId="28.[151,1437,782,2035]" box="[199,1163,963,989]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFC317D4C34E5" bold="true" box="[199,334,963,989]" pageId="28" pageNumber="29">Etymology.</emphasis>
|
||
Latin
|
||
<taxonomicName id="4C172BEEFFDDC078FE0EFC317DF934E5" authorityName="Harzhauser & Landau" authorityYear="2023" box="[407,507,963,989]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="globosus">
|
||
<emphasis id="B9638C7FFFDDC078FE0EFC317DF934E5" box="[407,507,963,989]" italics="true" pageId="28" pageNumber="29">globosus</emphasis>
|
||
</taxonomicName>
|
||
[= globular, spherical], referring to the rounded periphery.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D03E6FFDDC078FF5EFC14791A331D" pageId="28" pageNumber="29" type="diagnosis">
|
||
<paragraph id="8BA8506DFFDDC078FF5EFC14791A331D" blockId="28.[151,1437,782,2035]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFC147D403338" bold="true" box="[199,322,998,1024]" pageId="28" pageNumber="29">Diagnosis.</emphasis>
|
||
Medium-sized, solid shell with convex spire whorls and rounded periphery, characterized by relatively broad adapical MC, deep position of LPC and IPC forming weak angulation closely below LPC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D03E6FFDDC078FF5EFBDC7F5632ED" pageId="28" pageNumber="29" type="description">
|
||
<paragraph id="8BA8506DFFDDC078FF5EFBDC7F5632ED" blockId="28.[151,1437,782,2035]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFBDC7D5B3370" bold="true" box="[199,345,1070,1096]" pageId="28" pageNumber="29">Description.</emphasis>
|
||
Medium-sized, solid, turbinid shell with moderately elevated, gradate spire and rounded periphery. Protoconch heterostrophic, medium-sized of 1.2 visible whorls (diameter
|
||
<quantity id="4CEFFD88FFDDC078FC66FBA1785B3355" box="[1023,1113,1107,1133]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="29" unit="mm" value="1.2">1.2 mm</quantity>
|
||
). Teleoconch of 3.25 whorls. First teleoconch whorl with finely beaded SSC, beaded UPC and LPC and two poorly defined beaded MCs. SSC and MCs increasing in strength on subsequent whorl; adapical MC nearly as strong as SSC. Weaker spiral cord of beads intercalated between MCs. On second whorl secondary spiral row of beads intercalated between UPC and LPC. Abapically, beads becoming lower, subquadratic on last two whorls, and indistinct close to aperture. Periphery rounded without angulation or keel. UPC slightly more prominent forming periphery, LPC just below forming weak angulation. Whorls attached to LPC. Base weakly convex. LPC placed just medial to periphery in basal view. IPC distinct, followed by seven faintly beaded spiral cords on basal field, widening towards umbilicus. UC moderately prominent, slightly overhanging umbilicus, delimited from basal field by shallow narrow groove. Umbilicus cylindrical, moderately narrow (25% of total shell diameter). Umbilical wall with delicate opisthocyrt growth lines. Aperture moderately large, circular, no CG and PG developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D03E6FFDDC078FF5EFA2C7D893091" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="8BA8506DFFDDC078FF5EFA2C7F053059" blockId="28.[151,1437,782,2035]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EFA2C7D4E32C0" bold="true" box="[199,332,1502,1528]" pageId="28" pageNumber="29">Discussion.</emphasis>
|
||
Although represented by a single specimen, it is distinct enough to warrant formal description.
|
||
<taxonomicName id="4C172BEEFFDDC078FAEBFA2D7D463125" authority="Harzhauser & Landau, 2023" authorityName="Harzhauser & Landau" authorityYear="2023" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="globosus" status="sp. nov.">
|
||
<emphasis id="B9638C7FFFDDC078FAEBFA2D7D463125" italics="true" pageId="28" pageNumber="29">Heliacus globosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9638C7FFFDDC078FED3F9F17DA13125" bold="true" box="[330,419,1539,1565]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A2503104FFDDC078FED3F9F17DA13125" box="[330,419,1539,1565]" pageId="28" pageNumber="29" rank="species">nov. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is characterized by the position of the LPC slightly below the UPC and the strongly convex last whorl.
|
||
<taxonomicName id="4C172BEEFFDDC078FE8BF9D57E8A3179" authority="(Borson, 1821)" baseAuthorityName="Borson" baseAuthorityYear="1821" box="[274,648,1575,1601]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="radiatus">
|
||
<emphasis id="B9638C7FFFDDC078FE8BF9D57DDB3179" box="[274,473,1575,1601]" italics="true" pageId="28" pageNumber="29">Heliacus radiatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF862D9CFFDDC078FE7EF9D57E823179" author="Borson, S." box="[487,640,1575,1601]" pageId="28" pageNumber="29" pagination="297 - 364" refId="ref49541" refString="Borson, S. (1821) Continuazione del saggio di orittografia Piemontese. Memorie della reale accademia delle scienze di Torino, Series 1, 26, 297 - 364, pls. 5 - 6. [https: // www. biodiversitylibrary. org / page / 7653833]" type="journal article" year="1821">Borson, 1821</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Oligocene of the Colli Torinesi might be a related species and agrees especially in the sculpture of the spire whorls. It is distinguished from
|
||
<taxonomicName id="4C172BEEFFDDC078FB98F9B978D2315D" authorityName="Harzhauser & Landau" authorityYear="2023" box="[1025,1232,1611,1637]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="globosus">
|
||
<emphasis id="B9638C7FFFDDC078FB98F9B978D2315D" box="[1025,1232,1611,1637]" italics="true" pageId="28" pageNumber="29">Heliacus globosus</emphasis>
|
||
</taxonomicName>
|
||
in its much wider umbilicus and the prominent axial folds on the basal field (see Pavia 1976: pl. 1, figs 6a–c). Among extant
|
||
<taxonomicName id="4C172BEEFFDDC078FAA0F99D7CE93195" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FAA0F99D799F31B1" box="[1337,1437,1647,1673]" italics="true" pageId="28" pageNumber="29">Heliacus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C172BEEFFDDC078FF61F9617EBF3195" authority="(Schepman, 1909)" baseAuthorityName="Schepman" baseAuthorityYear="1909" box="[248,701,1683,1709]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maduernsis">
|
||
<emphasis id="B9638C7FFFDDC078FF61F9617DE73195" box="[248,485,1683,1709]" italics="true" pageId="28" pageNumber="29">Heliacus maduernsis</emphasis>
|
||
(Schepman, 1909)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C172BEEFFDDC078FD6EF96178973195" authority="(Gmelin, 1791)" baseAuthorityName="Gmelin" baseAuthorityYear="1791" box="[759,1173,1683,1709]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="stramineus">
|
||
<emphasis id="B9638C7FFFDDC078FD6EF9617FDC3194" box="[759,990,1683,1709]" italics="true" pageId="28" pageNumber="29">Heliacus stramineus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF862D9CFFDDC078FC76F961788F3195" author="Gmelin, J. F." box="[1007,1165,1683,1709]" pageId="28" pageNumber="29" pagination="3021 - 3910" refId="ref52952" refString="Gmelin, J. F. (1791) Vermes. In: Gmelin, J. F. (Ed.), Caroli a Linnaei systema naturae per regna tria naturae. Tome 1 (6). 13 th Edition. G. E. Beer, Lipsiae [Leipzig], pp. 3021 - 3910. [http: // www. biodiversitylibrary. org / item / 83098 # 5]" type="book chapter" year="1791">Gmelin, 1791</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
have comparable shell outlines with rather reduced peripheral keels, rounded periphery, convex dorsal side of the last whorl and circular aperture. These Indo-West Pacific species both differ from
|
||
<taxonomicName id="4C172BEEFFDDC078FCACF9297FBE31CD" authorityName="Harzhauser & Landau" authorityYear="2023" box="[821,956,1755,1781]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="globosus">
|
||
<emphasis id="B9638C7FFFDDC078FCACF9297FBE31CD" box="[821,956,1755,1781]" italics="true" pageId="28" pageNumber="29">H. globosus</emphasis>
|
||
</taxonomicName>
|
||
in their more delicate sculpture. The Pliocene to Recent Mediterranean
|
||
<taxonomicName id="4C172BEEFFDDC078FE6EF90D7F843021" authority="(Tiberi, 1872)" baseAuthorityName="Tiberi" baseAuthorityYear="1872" box="[503,902,1791,1817]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fallaciosus">
|
||
<emphasis id="B9638C7FFFDDC078FE6EF90D7EDF3021" box="[503,733,1791,1817]" italics="true" pageId="28" pageNumber="29">Heliacus fallaciosus</emphasis>
|
||
(Tiberi, 1872)
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="4C172BEEFFDDC078FC28F90D79973021" authority="(d'Orbigny, 1852)" baseAuthorityName="d'Orbigny" baseAuthorityYear="1852" box="[945,1429,1790,1817]" class="Gastropoda" family="Architectonicidae" genus="Heliacus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heterostropha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="subvariegatus">
|
||
<emphasis id="B9638C7FFFDDC078FC28F90D78B93021" box="[945,1211,1791,1817]" italics="true" pageId="28" pageNumber="29">Heliacus subvariegatus</emphasis>
|
||
(d’Orbigny, 1852)
|
||
</taxonomicName>
|
||
] differs in its more angulated periphery, the higher position of the LPC and the funnel-shaped umbilicus (
|
||
<bibRefCitation id="EF862D9CFFDDC078FAB1F8D17D2C3059" author="Melone, G. & Taviani, M." pageId="28" pageNumber="29" pagination="149 - 192" refId="ref57012" refString="Melone, G. & Taviani, M. (1985) Revisione delle Architectonicidae del Mediterraneo. Lavori S. I. M., 21, 149 - 192." type="journal article" year="1985">Melone & Taviani 1985</bibRefCitation>
|
||
: figs 31–33;
|
||
<bibRefCitation id="EF862D9CFFDDC078FE27F8B57E413059" author="Chirli, C." box="[446,579,1863,1889]" pageId="28" pageNumber="29" refId="ref50127" refString="Chirli, C. (2013) Malacofauna pliocenica Toscana. Vol. 9. Gastropoda fine Scaphopoda. Chirli, C., Firenze, 169 pp., 24 pls." type="book" year="2013">Chirli 2013</bibRefCitation>
|
||
: pl. 5, figs 7–12).
|
||
</paragraph>
|
||
<paragraph id="8BA8506DFFDDC078FF5EF8987D893091" blockId="28.[151,1437,782,2035]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EF8987DAA30BC" bold="true" box="[199,424,1898,1924]" pageId="28" pageNumber="29">Paleoenvironment.</emphasis>
|
||
The occurrence in the Baden Formation suggests middle to outer neritic water depths (
|
||
<bibRefCitation id="EF862D9CFFDDC078FF06F87D7D7C3091" author="Kranner, M. & Harzhauser, M. & Mandic, O. & Strauss, P. & Siedl, W. & Piller, W. E." box="[159,382,1935,1961]" pageId="28" pageNumber="29" pagination="105187" refId="ref55630" refString="Kranner, M., Harzhauser, M., Mandic, O., Strauss, P., Siedl, W. & Piller, W. E. (2021) Early and middle Miocene paleobathymetry of the Vienna Basin (Austria). Marine and Petroleum Geology, 132, 105187. https: // doi. org / 10.1016 / j. marpetgeo. 2021.105187" type="journal article" year="2021">
|
||
Kranner
|
||
<emphasis id="B9638C7FFFDDC078FE9AF87D7D3E3091" box="[259,316,1935,1961]" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30D03E6FFDDC078FF5EF84078D030C9" pageId="28" pageNumber="29" type="distribution">
|
||
<paragraph id="8BA8506DFFDDC078FF5EF840795030F5" blockId="28.[151,1437,782,2035]" box="[199,1362,1970,1997]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EF8407D6330F4" bold="true" box="[199,353,1970,1996]" pageId="28" pageNumber="29">Distribution.</emphasis>
|
||
Only known from the Langhian equivalent of the Badenian of the Central Paratethys Sea.
|
||
</paragraph>
|
||
<paragraph id="8BA8506DFFDDC078FF5EF82478D030C9" blockId="28.[151,1437,782,2035]" box="[199,1234,2006,2035]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9638C7FFFDDC078FF5EF8247DE130C8" bold="true" box="[199,483,2006,2033]" pageId="28" pageNumber="29">Central Paratethys Sea.</emphasis>
|
||
Middle Miocene (Badenian):
|
||
<emphasis id="B9638C7FFFDDC078FCAFF8257FC830C8" box="[822,970,2007,2032]" italics="true" pageId="28" pageNumber="29">
|
||
<collectingRegion id="49D39E8FFFDDC078FCAFF8257F8130C8" box="[822,899,2007,2032]" country="Austria" name="Wien" pageId="28" pageNumber="29">Vienna</collectingRegion>
|
||
Basin
|
||
</emphasis>
|
||
: M̂llersdorf (
|
||
<collectingCountry id="F30010FDFFDDC078FBF4F82578C430C9" box="[1133,1222,2007,2033]" name="Austria" pageId="28" pageNumber="29">Austria</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |