236 lines
37 KiB
XML
236 lines
37 KiB
XML
<document id="453F78D42D63D405A039363D330559A8" ID-DOI="10.11646/zootaxa.4742.3.11" ID-GBIF-Dataset="a1b47413-2f76-404c-86e4-571b06c6a1cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677917" ID-ZooBank="B5B11009-059B-4DC1-85B6-831A67B72166" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272996897" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="03BE87E8FF9E7F1C9CF7FB4D521FF828" docLanguage="en" docName="zootaxa.4742.3.11.pdf" docOrigin="Zootaxa 4742 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cloeon Leach 1815" docType="treatment" docVersion="4" lastPageNumber="576" masterDocId="FF87FF90FF9F7F1F9C60FFB85021FFCB" masterDocTitle="New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon)" masterLastPageNumber="587" masterPageNumber="573" pageNumber="574" updateTime="1698720838785" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C2E34FB7784F0945607440C6499F7AF3">New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon)</mods:title>
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<mods:namePart id="39BBBCB156286965E4774C97B37DDBA9">Kluge, Nikita J.</mods:namePart>
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<mods:title id="7AABE0FAE79AB153D826AE71CC1CEECB">Zootaxa</mods:title>
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<mods:part id="0E41D6AC86CC12A6C9462E9EB4A06E07">
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<mods:date id="F945D420915FE394E2BC178403BE245F">2020</mods:date>
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<mods:number id="53C27D1DE1CBE54CD3BB4978DDC2F70F">2020-02-21</mods:number>
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<treatment id="03BE87E8FF9E7F1C9CF7FB4D521FF828" ID-DOI="http://doi.org/10.5281/zenodo.3684998" ID-GBIF-Taxon="163883513" ID-Zenodo-Dep="3684998" LSID="urn:lsid:plazi:treatment:03BE87E8FF9E7F1C9CF7FB4D521FF828" httpUri="http://treatment.plazi.org/id/03BE87E8FF9E7F1C9CF7FB4D521FF828" lastPageId="3" lastPageNumber="576" pageId="1" pageNumber="574">
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<subSubSection id="C30D6575FF9E7F1E9CF7FB4D5251FAE5" pageId="1" pageNumber="574" type="nomenclature">
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<paragraph id="8BA836FEFF9E7F1E9CF7FB4D513FFAC4" blockId="1.[151,649,1269,1387]" box="[151,286,1269,1295]" pageId="1" pageNumber="574">
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<heading id="D0E08192FF9E7F1E9CF7FB4D513FFAC4" bold="true" box="[151,286,1269,1295]" fontSize="11" level="1" pageId="1" pageNumber="574" reason="1">
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<emphasis id="B963EAECFF9E7F1E9CF7FB4D513FFAC4" bold="true" box="[151,286,1269,1295]" pageId="1" pageNumber="574">
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<taxonomicName id="4C174D7DFF9E7F1E9CF7FB4D50CEFAC4" ID-CoL="3R5Y" ID-ENA="197151" authorityName="Leach" authorityYear="1815" box="[151,239,1269,1295]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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/fg1
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA836FEFF9E7F1E9CF7FAAD5251FAE5" blockId="1.[151,649,1269,1387]" box="[151,624,1301,1326]" pageId="1" pageNumber="574">
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Branch «6»:
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<bibRefCitation id="EF864B0FFF9E7F1E9D7AFAAD5238FAE5" author="Kluge, N. J. & Novikova, E. A." box="[282,537,1301,1326]" pageId="1" pageNumber="574" pagination="29 - 54" refId="ref6849" refString="Kluge, N. J. & Novikova, E. A. (1992) Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR. [Entomologicheskoe Obozrenie] (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83 (in Russian). English translation: Entomological Review, 71 (9), 29 - 54." type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
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:
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<figureCitation id="132C2A7BFF9E7F1E9E43FAAD524DFAE5" box="[547,620,1301,1326]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="1" pageNumber="574">Fig. 11</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30D6575FF9E7F1E9CF7FA8C5223FAA0" pageId="1" pageNumber="574" type="description">
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<paragraph id="8BA836FEFF9E7F1E9CF7FA8C52A8FA86" blockId="1.[151,649,1269,1387]" box="[151,649,1332,1357]" pageId="1" pageNumber="574">
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Hierarchica name:
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<taxonomicName id="4C174D7DFF9E7F1E9D38FA8C5186FA86" authorityName="Leach" authorityYear="1815" box="[344,423,1332,1357]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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/fg1:
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<bibRefCitation id="EF864B0FFF9E7F1E9DB7FA8C52A4FA86" author="Kluge" box="[471,645,1332,1357]" pageId="1" pageNumber="574" pagination="361 - 376" refId="ref6645" refString="Kluge N. J. (2012) Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal, 20 (4), 361 - 376. https: // doi. org / 10.15298 / rusentj. 20.4.02" type="journal article" year="2012">Kluge 2012: 362</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BA836FEFF9E7F1E9CF7FAEA5223FAA0" blockId="1.[151,649,1269,1387]" box="[151,514,1362,1387]" pageId="1" pageNumber="574">Tribus Cloeonini: Kluge 2016: 492.</paragraph>
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</subSubSection>
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<subSubSection id="C30D6575FF9E7F1C9CF7FA21521FF828" lastPageId="3" lastPageNumber="576" pageId="1" pageNumber="574" type="diagnosis">
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<paragraph id="8BA836FEFF9E7F1E9CF7FA215199FA7F" blockId="1.[151,1437,1433,2036]" box="[151,440,1433,1460]" pageId="1" pageNumber="574">
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<heading id="D0E08192FF9E7F1E9CF7FA215199FA7F" bold="true" box="[151,440,1433,1460]" fontSize="11" level="1" pageId="1" pageNumber="574" reason="1">
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<emphasis id="B963EAECFF9E7F1E9CF7FA215199FA7F" bold="true" box="[151,440,1433,1460]" pageId="1" pageNumber="574">
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Diagnosis of
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<taxonomicName id="4C174D7DFF9E7F1E9D50FA2151A7FA7F" authorityName="Leach" authorityYear="1815" box="[304,390,1433,1460]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="574" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
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/fg1.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA836FEFF9E7F1E9CA7FA06530EF93C" blockId="1.[151,1437,1433,2036]" pageId="1" pageNumber="574">
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<emphasis id="B963EAECFF9E7F1E9CA7FA065133FA13" bold="true" box="[199,274,1470,1496]" italics="true" pageId="1" pageNumber="574">Larva.</emphasis>
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Both mandibles with dense setae between prostheca and mola (
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<figureCitation id="132C2A7BFF9E7F1E9FB3FA05543AFA1C" box="[979,1051,1469,1495]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1897,1922]" captionTargetBox="[151,1436,270,1869]" captionTargetId="figure@2.[151,1436,270,1869]" captionTargetPageId="2" captionText="FIGURES 1–8. Larval exuviae. 1–2, Procloeon (Monilistylus) monilistylus: 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8, Procloeon (Monilistylus) ornatipennis sp. n.: 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3677921" httpUri="https://zenodo.org/record/3677921/files/figure.png" pageId="1" pageNumber="574">Figs 4</figureCitation>
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,
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<figureCitation id="132C2A7BFF9E7F1E9846FA05544EFA1C" box="[1062,1135,1469,1495]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="1" pageNumber="574">18, 20</figureCitation>
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). Labial palp with 3rd (distalmost) segment truncate (
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<figureCitation id="132C2A7BFF9E7F1E9DA8FA59526AFA37" box="[456,587,1505,1532]" captionStart="FIGURES 26–29" captionStartId="6.[151,264,1811,1836]" captionTargetBox="[151,1436,236,1786]" captionTargetId="figure@6.[151,1436,236,1786]" captionTargetPageId="6" captionText="FIGURES 26–29. Procloeon (Monilistylus) ornatipennis sp. n., exuviae of labium: 26, general view with focus on ventral side; 27, labial palp with focus on dorsal side; 28, glossa and paraglossa with focus on ventral side; 29, the same with focus on dorsal side (26, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3677925" httpUri="https://zenodo.org/record/3677925/files/figure.png" pageId="1" pageNumber="574">Figs 26–27</figureCitation>
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). On anterior side of tibia, proximal end of patella-tibial suture bent distally and connected under acute angle with row of long thin hairs (
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<figureCitation id="132C2A7BFF9E7F1E9F2EF9BD53EEF9D4" box="[846,975,1541,1567]" captionStart="FIGURES 30–32" captionStartId="8.[151,264,1781,1806]" captionTargetBox="[151,1436,182,1755]" captionTargetId="figure@8.[151,1436,181,1755]" captionTargetPageId="8" captionText="FIGURES 30–32. Procloeon (Monilistylus) ornatipennis sp. n., legs: 30, foreleg in anterior view (holotype); 31, base of tibia in anterior view; 32, claw." figureDoi="http://doi.org/10.5281/zenodo.3677927" httpUri="https://zenodo.org/record/3677927/files/figure.png" pageId="1" pageNumber="574">Figs 30–31</figureCitation>
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) (patella-tibial suture has equal structure on all legs – see Anteropatellata). Claw symmetric, slightly curved, with 2 similar rows of denticles (
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<figureCitation id="132C2A7BFF9E7F1E997BF9915550F988" box="[1307,1393,1577,1603]" captionStart="FIGURES 30–32" captionStartId="8.[151,264,1781,1806]" captionTargetBox="[151,1436,182,1755]" captionTargetId="figure@8.[151,1436,181,1755]" captionTargetPageId="8" captionText="FIGURES 30–32. Procloeon (Monilistylus) ornatipennis sp. n., legs: 30, foreleg in anterior view (holotype); 31, base of tibia in anterior view; 32, claw." figureDoi="http://doi.org/10.5281/zenodo.3677927" httpUri="https://zenodo.org/record/3677927/files/figure.png" pageId="1" pageNumber="574">Fig. 32</figureCitation>
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); in some taxa without denticles. Abdominal segment IX and often one or several previous uromeres with longitudinal row of spines on each lateral margin (
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<figureCitation id="132C2A7BFF9E7F1E9E2FF9C95284F947" box="[591,677,1649,1676]" captionStart="FIGURES 33–38" captionStartId="9.[151,264,1848,1873]" captionTargetBox="[164,1423,273,1823]" captionTargetId="figure@9.[164,1423,273,1823]" captionTargetPageId="9" captionText="FIGURES 33–38. Procloeon (Monilistylus) ornatipennis sp. n.: 33, larval exuviae of posterior margin of 9th abdominal sternum with protogonostyli; 34, subimaginal tissues with gonostyli, extracted from it; 35, exuviae of paraprocts and protogonostyli of another specimen; 36, exuviae of 5th abdominal sternum (Figs 33–36 under the same magnification); 37,middle of posterior margin of 9th abdominal tergum; 38, portion of mesonotum (magnification as in Fig. 37).Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.3677929" httpUri="https://zenodo.org/record/3677929/files/figure.png" pageId="1" pageNumber="574">Fig. 35</figureCitation>
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). Tergalii often with anal-proximal expansion curved dorsally and forming a second (dorsal) lamella of tergalius; this lamella is absent in selected species of
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<taxonomicName id="4C174D7DFF9E7F1E98EFF92D5529F964" authorityName="Bengtsson" authorityYear="1915" box="[1167,1288,1685,1711]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
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/g1, including both species of
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<emphasis id="B963EAECFF9E7F1E9D2BF90151F4F918" box="[331,469,1721,1747]" italics="true" pageId="1" pageNumber="574">Monilistylus</emphasis>
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(
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<figureCitation id="132C2A7BFF9E7F1E9D86F901527DF91F" box="[486,604,1721,1748]" captionStart="FIGURES 9–25" captionStartId="3.[151,264,1705,1730]" captionTargetBox="[189,1402,191,1671]" captionTargetId="figure@3.[177,1411,181,1679]" captionTargetPageId="3" captionText="FIGURES 9–25. Details of larvae. 9–23, Procloeon (Monilistylus) ornatipennis sp. n.: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25, Procloeon (Monilistylus) monilistylus, kinetodontium and prostheca rotated after pressure of slide." figureDoi="http://doi.org/10.5281/zenodo.3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="1" pageNumber="574">Figs 9–15</figureCitation>
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). Tergalii able to intensive rhythmic respiratory movements. Larval cerci and paracercus with dark rings on each 4th joining (
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<figureCitation id="132C2A7BFF9E7F1E9EBDF9655300F93C" box="[733,801,1757,1783]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1897,1922]" captionTargetBox="[151,1436,270,1869]" captionTargetId="figure@2.[151,1436,270,1869]" captionTargetPageId="2" captionText="FIGURES 1–8. Larval exuviae. 1–2, Procloeon (Monilistylus) monilistylus: 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8, Procloeon (Monilistylus) ornatipennis sp. n.: 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged." figureDoi="http://doi.org/10.5281/zenodo.3677921" httpUri="https://zenodo.org/record/3677921/files/figure.png" pageId="1" pageNumber="574">Fig. 8</figureCitation>
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).
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</paragraph>
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<paragraph id="8BA836FEFF9E7F1E9CA7F8B95469F8AF" blockId="1.[151,1437,1433,2036]" pageId="1" pageNumber="574">
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<emphasis id="B963EAECFF9E7F1E9CA7F8B95162F8D7" bold="true" box="[199,323,1793,1820]" italics="true" pageId="1" pageNumber="574">Subimago.</emphasis>
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All tarsomeres covered with pointed microlepides, without blunt microlepides. Process of posterodorsal angle of postsubalar sclerite diminished, pointed, with concave antero-dorsal margin (
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<figureCitation id="132C2A7BFF9E7F1E98CAF89D54DDF8F4" box="[1194,1276,1829,1855]" captionStart="FIGURES 39–46" captionStartId="10.[151,264,1885,1910]" captionTargetBox="[151,1436,181,1860]" captionTargetId="figure@10.[151,1436,181,1860]" captionTargetPageId="10" captionText="FIGURES 39–46. Procloeon (Monilistylus) ornatipennis sp. n.: 39–40, male imagines; 41, male imaginal genitals; 42, exuviae of right half of subimaginal mesonotum; 43, exuviae of left half of subimaginal mesopleuron; 44, wing; 45, abdominal terga II–VII; 46, tibia and tarsus of middle leg (41–43 and 45–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3677931" httpUri="https://zenodo.org/record/3677931/files/figure.png" pageId="1" pageNumber="574">Fig. 43</figureCitation>
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) (unlike other Turbanoculata, in which this process is larger and has convex antero-dorsal margin).
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</paragraph>
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<paragraph id="8BA836FEFF9E7F1E9CA7F8D55292F81B" blockId="1.[151,1437,1433,2036]" pageId="1" pageNumber="574">
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<emphasis id="B963EAECFF9E7F1E9CA7F8D5525AF843" bold="true" box="[199,635,1901,1928]" italics="true" pageId="1" pageNumber="574">Winged stages (imago and subimago).</emphasis>
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On fore wing each space with no more than 1 marginal intercalary (
|
||
<figureCitation id="132C2A7BFF9E7F1E990FF8D55096F860" captionStart="FIGURES 39–46" captionStartId="10.[151,264,1885,1910]" captionTargetBox="[151,1436,181,1860]" captionTargetId="figure@10.[151,1436,181,1860]" captionTargetPageId="10" captionText="FIGURES 39–46. Procloeon (Monilistylus) ornatipennis sp. n.: 39–40, male imagines; 41, male imaginal genitals; 42, exuviae of right half of subimaginal mesonotum; 43, exuviae of left half of subimaginal mesopleuron; 44, wing; 45, abdominal terga II–VII; 46, tibia and tarsus of middle leg (41–43 and 45–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3677931" httpUri="https://zenodo.org/record/3677931/files/figure.png" pageId="1" pageNumber="574">Fig. 44</figureCitation>
|
||
). Hind wing, if present, of «
|
||
<emphasis id="B963EAECFF9E7F1E9D9CF82952B1F860" box="[508,656,1937,1963]" italics="true" pageId="1" pageNumber="574">Centroptilum</emphasis>
|
||
-
|
||
<typeStatus id="54AC885CFF9E7F1E9EF8F82A52EFF867" box="[664,718,1938,1964]" pageId="1" pageNumber="574">type</typeStatus>
|
||
»: relatively narrow and long, with hooked costal projection (in both species of
|
||
<emphasis id="B963EAECFF9E7F1E9D26F80D51F1F804" box="[326,464,1973,1999]" italics="true" pageId="1" pageNumber="574">Monilistylus</emphasis>
|
||
hind wings absent).
|
||
</paragraph>
|
||
<paragraph id="8BA836FEFF9E7F1D9CA7F8615418FF30" blockId="1.[151,1437,1433,2036]" lastBlockId="2.[151,1436,152,251]" lastPageId="2" lastPageNumber="575" pageId="1" pageNumber="574">
|
||
<emphasis id="B963EAECFF9E7F1E9CA7F861514FF83F" bold="true" box="[199,366,2009,2036]" pageId="1" pageNumber="574">
|
||
<emphasis id="B963EAECFF9E7F1E9CA7F8615147F83F" bold="true" box="[199,358,2009,2036]" italics="true" pageId="1" pageNumber="574">Male genitals</emphasis>
|
||
.
|
||
</emphasis>
|
||
Gonovectes fused by their median ends with penial bridge, unable for protraction (
|
||
<figureCitation id="132C2A7BFF9E7F1E9959F86155B1F83F" box="[1337,1424,2009,2036]" captionStart="FIGURES 52–54" captionStartId="12.[151,264,1768,1793]" captionTargetBox="[165,1436,188,1732]" captionTargetId="figure@12.[151,1436,181,1743]" captionTargetPageId="12" captionText="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677935" httpUri="https://zenodo.org/record/3677935/files/figure.png" pageId="1" pageNumber="574">Fig. 52</figureCitation>
|
||
). Gonostylus with 2nd segment thickened apically, 3rd (distalmost) segment clavate (
|
||
<figureCitation id="132C2A7BFF9D7F1D9831FF21548AFF78" box="[1105,1195,153,179]" captionStart="FIGURES 47–51" captionStartId="11.[151,264,1579,1604]" captionTargetBox="[164,1423,181,1549]" captionTargetId="figure@11.[164,1423,181,1553]" captionTargetPageId="11" captionText="FIGURES 47–51. Procloeon (Monilistylus) ornatipennis sp. n. (holotype): 47, median projection of penis; 48, exuviae of subimaginal gonostyli; 49, genitals with all visible muscles (muscles and hidden parts of penis shown by interrupted lines); 50, the same with penis, its muscles and gonoducts (shown by dotted line); 51, the same with muscles of styliger (penis not shown). Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.IX-X, area of attachment of intersegmental paraproctal muscle; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677933" httpUri="https://zenodo.org/record/3677933/files/figure.png" pageId="2" pageNumber="575">Figs 49</figureCitation>
|
||
,
|
||
<figureCitation id="132C2A7BFF9D7F1D98D8FF2054F6FF78" box="[1208,1239,152,179]" captionStart="FIGURES 52–54" captionStartId="12.[151,264,1768,1793]" captionTargetBox="[165,1436,188,1732]" captionTargetId="figure@12.[151,1436,181,1743]" captionTargetPageId="12" captionText="FIGURES 52–54. Procloeon (Monilistylus) monilistylus: 52–53, genital of male imago (corrected from Kluge et al. 2014); 54, exuviae of subimaginal gonostyli. Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., median and lateral portions of styligeral muscle; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.3677935" httpUri="https://zenodo.org/record/3677935/files/figure.png" pageId="2" pageNumber="575">53</figureCitation>
|
||
). In mature larva ready to molt to subimago, subimaginal gonostyli folded under larval cuticle by «
|
||
<taxonomicName id="4C174D7DFF9D7F1D985CFF0554ACFF1C" authorityName="Leach" authorityYear="1815" box="[1084,1165,189,215]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9D7F1D985CFF0554ACFF1C" box="[1084,1165,189,215]" italics="true" pageId="2" pageNumber="575">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<typeStatus id="54AC885CFF9D7F1D98F5FF0554EAFF1C" box="[1173,1227,189,215]" pageId="2" pageNumber="575">type</typeStatus>
|
||
»: 2nd segment directed laterally, 3rd segment curved either caudally (
|
||
<figureCitation id="132C2A7BFF9D7F1D9E83FF595316FF30" box="[739,823,225,251]" captionStart="FIGURES 33–38" captionStartId="9.[151,264,1848,1873]" captionTargetBox="[164,1423,273,1823]" captionTargetId="figure@9.[164,1423,273,1823]" captionTargetPageId="9" captionText="FIGURES 33–38. Procloeon (Monilistylus) ornatipennis sp. n.: 33, larval exuviae of posterior margin of 9th abdominal sternum with protogonostyli; 34, subimaginal tissues with gonostyli, extracted from it; 35, exuviae of paraprocts and protogonostyli of another specimen; 36, exuviae of 5th abdominal sternum (Figs 33–36 under the same magnification); 37,middle of posterior margin of 9th abdominal tergum; 38, portion of mesonotum (magnification as in Fig. 37).Abbreviations: gs1, gs2, gs3, segments of gonostylus; m.gs, gonostylar muscle; m.gv, gonovectal (penial) muscle; m.s., lateral portion of styligeral muscle." figureDoi="http://doi.org/10.5281/zenodo.3677929" httpUri="https://zenodo.org/record/3677929/files/figure.png" pageId="2" pageNumber="575">Fig. 34</figureCitation>
|
||
), or caudally-medially.
|
||
</paragraph>
|
||
<caption id="DF686676FF9D7F1D9CF7F8D1524FF825" ID-DOI="http://doi.org/10.5281/zenodo.3677921" ID-Zenodo-Dep="3677921" httpUri="https://zenodo.org/record/3677921/files/figure.png" pageId="2" pageNumber="575" startId="2.[151,264,1897,1922]" targetBox="[151,1436,270,1869]" targetPageId="2">
|
||
<paragraph id="8BA836FEFF9D7F1D9CF7F8D1524FF825" blockId="2.[151,1437,1897,2030]" pageId="2" pageNumber="575">
|
||
<emphasis id="B963EAECFF9D7F1D9CF7F8D1511DF849" bold="true" box="[151,316,1897,1922]" pageId="2" pageNumber="575">FIGURES 1–8.</emphasis>
|
||
Larval exuviae. 1–2,
|
||
<taxonomicName id="4C174D7DFF9D7F1D9E7FF8D153BEF84A" authority="monilistylus" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[543,927,1897,1921]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="monilistylus" subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9D7F1D9E7FF8D152A9F84A" box="[543,648,1897,1921]" italics="true" pageId="2" pageNumber="575">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF9D7F1D9EF7F8D15335F84A" box="[663,788,1897,1921]" italics="true" pageId="2" pageNumber="575">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF9D7F1D9F45F8D153BEF84A" box="[805,927,1897,1921]" italics="true" pageId="2" pageNumber="575">monilistylus</emphasis>
|
||
</taxonomicName>
|
||
: 1, right half of pro- and mesonotum; 2, thoracic sterna, pleura, metanotum, abdominal sterna and terga. 3–8,
|
||
<taxonomicName id="4C174D7DFF9D7F1D9F63F83554A8F86E" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[771,1161,1933,1957]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="575" phylum="Arthropoda" rank="species" species="ornatipennis" status="sp. nov." subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9D7F1D9F63F835534DF86E" box="[771,876,1933,1957]" italics="true" pageId="2" pageNumber="575">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF9D7F1D9F1CF83553D8F86E" box="[892,1017,1933,1957]" italics="true" pageId="2" pageNumber="575">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF9D7F1D9869F83654A8F86E" box="[1033,1161,1934,1957]" italics="true" pageId="2" pageNumber="575">ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2505797FF9D7F1D98F0F83654EBF86E" box="[1168,1226,1934,1957]" pageId="2" pageNumber="575" rank="species">sp. n.</taxonomicNameLabel>
|
||
: 3, right half of pro- and mesonotum; 4, head; 5, thoracic sterna, pleura, metanotum and two legs; 6, abdominal sterna and terga (with caudalii of holotype); 7–8, caudalii of holotype, enlarged.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF686676FF9C7F1C9CF7F91152D3F8E5" ID-DOI="http://doi.org/10.5281/zenodo.3677923" ID-Zenodo-Dep="3677923" httpUri="https://zenodo.org/record/3677923/files/figure.png" pageId="3" pageNumber="576" startId="3.[151,264,1705,1730]" targetBox="[189,1402,191,1671]" targetPageId="3">
|
||
<paragraph id="8BA836FEFF9C7F1C9CF7F91152D3F8E5" blockId="3.[151,1437,1705,1838]" pageId="3" pageNumber="576">
|
||
<emphasis id="B963EAECFF9C7F1C9CF7F911516BF909" bold="true" box="[151,330,1705,1730]" pageId="3" pageNumber="576">FIGURES 9–25.</emphasis>
|
||
Details of larvae. 9–23,
|
||
<taxonomicName id="4C174D7DFF9C7F1C9E32F91153FDF90A" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[594,988,1705,1730]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="ornatipennis" status="sp. nov." subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9C7F1C9E32F911529AF90A" box="[594,699,1705,1729]" italics="true" pageId="3" pageNumber="576">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF9C7F1C9EADF911536BF90A" box="[717,842,1705,1729]" italics="true" pageId="3" pageNumber="576">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF9C7F1C9F3CF91253FDF90A" box="[860,988,1706,1729]" italics="true" pageId="3" pageNumber="576">ornatipennis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2505797FF9C7F1C9F85F9125400F90A" box="[997,1057,1706,1729]" pageId="3" pageNumber="576" rank="species">sp. n.</taxonomicNameLabel>
|
||
: 9–15, tergalii I–VII (holotype); 16, labrum; 17, maxilla (dorsal view); 18, right mandible; 19, its kinetodontium rotated after pressure of slide; 20, left mandible; 21–23, its incisor, kinetodontium and prostheca rotated after pressure of slide. 24–25,
|
||
<taxonomicName id="4C174D7DFF9C7F1C9874F94955B4F8C2" authority="monilistylus" authorityName="monilistylus Kluge, Tiunova & Novikova" authorityYear="2014" box="[1044,1429,1777,1802]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="monilistylus" subGenus="Monilistylus">
|
||
<emphasis id="B963EAECFF9C7F1C9874F949545CF8C2" box="[1044,1149,1777,1801]" italics="true" pageId="3" pageNumber="576">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B963EAECFF9C7F1C98EEF949552AF8C2" box="[1166,1291,1777,1801]" italics="true" pageId="3" pageNumber="576">Monilistylus</emphasis>
|
||
)
|
||
<emphasis id="B963EAECFF9C7F1C997BF94955B4F8C2" box="[1307,1429,1777,1801]" italics="true" pageId="3" pageNumber="576">monilistylus</emphasis>
|
||
</taxonomicName>
|
||
, kinetodontium and prostheca rotated after pressure of slide.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA836FEFF9C7F1C9CA7F8E3521FF828" blockId="3.[151,1436,1883,2019]" pageId="3" pageNumber="576">
|
||
<emphasis id="B963EAECFF9C7F1C9CA7F8E35146F8BD" bold="true" box="[199,359,1883,1910]" pageId="3" pageNumber="576">Composition.</emphasis>
|
||
Taxon
|
||
<taxonomicName id="4C174D7DFF9C7F1C9DA5F8E45238F8BD" authorityName="Leach" authorityYear="1815" box="[453,537,1884,1910]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg1 includes
|
||
<taxonomicName id="4C174D7DFF9C7F1C9ED9F8E45456F8BD" authority="Kluge & Novikova 1992" authorityName="Kluge & Novikova" authorityYear="1992" box="[697,1143,1884,1910]" class="Insecta" family="Baetidae" genus="Similicloeon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9C7F1C9ED9F8E45364F8BD" box="[697,837,1884,1910]" italics="true" pageId="3" pageNumber="576">Similicloeon</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9C7F1C9F31F8E45456F8BD" author="Kluge, N. J. & Novikova, E. A." box="[849,1143,1884,1910]" pageId="3" pageNumber="576" pagination="29 - 54" refId="ref6849" refString="Kluge, N. J. & Novikova, E. A. (1992) Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR. [Entomologicheskoe Obozrenie] (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83 (in Russian). English translation: Entomological Review, 71 (9), 29 - 54." type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C174D7DFF9C7F1C98C9F8E45169F851" authority="Kluge & Novikova 1992" authorityName="Kluge & Novikova" authorityYear="1992" class="Insecta" family="Baetidae" genus="Intercloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9C7F1C98C9F8E45508F8BD" box="[1193,1321,1884,1910]" italics="true" pageId="3" pageNumber="576">Intercloeon</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9C7F1C9956F8E45169F851" author="Kluge, N. J. & Novikova, E. A." pageId="3" pageNumber="576" pagination="29 - 54" refId="ref6849" refString="Kluge, N. J. & Novikova, E. A. (1992) Revision of the Palaearctic genera and subgenera of mayflies of the subfamily Cloeoninae (Ephemeroptera, Baetidae) with description of new species from the USSR. [Entomologicheskoe Obozrenie] (Revue d'Entomologie de l'URSS), 71 (1), 60 - 83 (in Russian). English translation: Entomological Review, 71 (9), 29 - 54." type="journal article" year="1992">Kluge & Novikova 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C174D7DFF9C7F1C9D3AF838518AF851" authorityName="Leach" authorityYear="1815" box="[346,427,1920,1946]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
|
||
/fg2 (or genus
|
||
<taxonomicName id="4C174D7DFF9C7F1C9E2DF8385307F851" authority="Leach 1815" authorityName="Leach" authorityYear="1815" box="[589,806,1919,1946]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B963EAECFF9C7F1C9E2DF83852BFF851" box="[589,670,1920,1946]" italics="true" pageId="3" pageNumber="576">Cloeon</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9C7F1C9EC4F8385307F851" author="Leach, W. E." box="[676,806,1919,1946]" pageId="3" pageNumber="576" pagination="57 - 172" refId="ref7186" refString="Leach, W. E. (1815) Entomology. Brewster's Edinburg Encyclopedia, 1 st Edition, 9 (1), 57 - 172." type="journal article" year="1815">Leach 1815</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2505797FF9C7F1C9F4CF8385348F851" box="[812,873,1920,1946]" pageId="3" pageNumber="576" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
),
|
||
<taxonomicName id="4C174D7DFF9C7F1C9F1CF8385573F851" authority="Hill, Pfeiffer & Jacobus 2010" authorityName="Hill, Pfeiffer & Jacobus" authorityYear="2010" box="[892,1362,1920,1946]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9C7F1C9F1CF8385423F851" box="[892,1026,1920,1946]" italics="true" pageId="3" pageNumber="576">Waynokiops</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9C7F1C9869F8385573F851" author="Hill, M. A. & Pfeiffer, J. & Jacobus, L. M." box="[1033,1362,1920,1946]" pageId="3" pageNumber="576" pagination="61 - 68" refId="ref6119" refString="Hill, M. A., Pfeiffer, J. & Jacobus, L. M. (2010) A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America. Zootaxa, 2481, 61 - 68." type="journal article" year="2010">Hill, Pfeiffer & Jacobus 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C174D7DFF9C7F1C993CF83951DBF875" authority="Bogoescu 1947" authorityName="Bogoescu" authorityYear="1947" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B963EAECFF9C7F1C993CF8395160F875" italics="true" pageId="3" pageNumber="576">Pseudocentroptilum</emphasis>
|
||
Bogoescu 1947
|
||
</taxonomicName>
|
||
[with the single species
|
||
<taxonomicName id="4C174D7DFF9C7F1C9F7AF81D54B1F875" authority="Tshernova 1941" authorityName="Tshernova" authorityYear="1941" box="[794,1168,1956,1982]" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="unguiculatum">
|
||
<emphasis id="B963EAECFF9C7F1C9F7AF81D53F3F875" box="[794,978,1956,1982]" italics="true" pageId="3" pageNumber="576">P. unguiculatum</emphasis>
|
||
<bibRefCitation id="EF864B0FFF9C7F1C9FBBF81C54B1F875" author="Tshernova, O. A." box="[987,1168,1956,1982]" pageId="3" pageNumber="576" pagination="239 - 244" refId="ref7214" refString="Tshernova, O. A. (1941) Ephemeroptera collected on the river Ili by the Moscow Zoological Museum's Expedition. Archives du Musee Zoollogique de l'Universite de Moscou, 6, 239 - 244. [in Russian]" type="journal article" year="1941">Tshernova 1941</bibRefCitation>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C174D7DFF9C7F1C98DCF81D50F2F829" authority="Bogoescu 1947" authorityName="Bogoescu" authorityYear="1947" class="Insecta" family="Baetidae" genus="Pseudocentroptilum" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="motasi">
|
||
<emphasis id="B963EAECFF9C7F1C98DCF81D5504F875" box="[1212,1317,1957,1982]" italics="true" pageId="3" pageNumber="576">P. motasi</emphasis>
|
||
Bogoescu 1947
|
||
</taxonomicName>
|
||
)] and
|
||
<taxonomicName id="4C174D7DFF9C7F1C9D78F87051AEF829" authorityName="Bengtsson" authorityYear="1915" box="[280,399,1992,2018]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="587" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
|
||
/g1 (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |