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<document id="D6347104BC1D5F067A565A204F256D13" ID-CLB-Dataset="302400" ID-DOI="10.11646/phytotaxa.365.3.7" ID-GBIF-Dataset="ccea10f5-10cf-4f30-8daf-625fb5513d82" ID-ISSN="1179-3163" ID-Zenodo-Dep="13704454" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725583221623" checkinUser="felipe" docAuthor="Lin, Che-Wei &amp; Lee, Chi-Hung" docDate="2018" docId="03BE8790C80B66552C9BF90B91EA6E9C" docLanguage="en" docName="phytotaxa.365.3.7.pdf" docOrigin="Phytotaxa 365 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.365.3.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phyllagathis stellata C. W. Lin &amp; C. H. Lee. A. Habit 2018, sp. nov." docType="treatment" docVersion="4" lastPageNumber="299" masterDocId="FF87FFE8C80B66512C13FF8695096C11" masterDocTitle="Phyllagathis stellata (Sonerileae, Melastomataceae), a new species from southwestern Sarawak, Borneo" masterLastPageNumber="300" masterPageNumber="295" pageNumber="295" updateTime="1725597381276" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="583AA5DA5A818058F045BDE9F43DBE49">Phyllagathis stellata (Sonerileae, Melastomataceae), a new species from southwestern Sarawak, Borneo</mods:title>
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<mods:namePart id="163FDB9A8CC2B75567F0ECE87F3489E9">Lin, Che-Wei</mods:namePart>
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<treatment id="03BE8790C80B66552C9BF90B91EA6E9C" ID-DOI="http://doi.org/10.5281/zenodo.13704537" ID-GBIF-Taxon="240418466" ID-Zenodo-Dep="13704537" LSID="urn:lsid:plazi:treatment:03BE8790C80B66552C9BF90B91EA6E9C" httpUri="http://treatment.plazi.org/id/03BE8790C80B66552C9BF90B91EA6E9C" lastPageId="4" lastPageNumber="299" pageId="0" pageNumber="295">
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<paragraph id="8BA83686C80B66512C9BF90B964A6AB9" blockId="0.[136,835,1677,1704]" box="[136,835,1677,1704]" pageId="0" pageNumber="295">
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<taxonomicName id="4C174D05C80B66512C9BF90B97656AB9" ID-CoL="4H33F" authority="C.W. Lin &amp; C.H. Lee" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[136,620,1677,1704]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="0" pageNumber="295" phylum="Tracheophyta" rank="species" species="stellata" status="sp. nov.">
<emphasis id="B963EA94C80B66512C9BF90B947B6AB9" bold="true" box="[136,370,1677,1704]" italics="true" pageId="0" pageNumber="295">Phyllagathis stellata</emphasis>
C.W. Lin &amp; C.H. Lee
</taxonomicName>
,
<emphasis id="B963EA94C80B66512E64F90997C56AB9" box="[631,716,1679,1704]" italics="true" pageId="0" pageNumber="295">
<taxonomicNameLabel id="A25057EFC80B66512E64F90997C56AB9" box="[631,716,1679,1704]" pageId="0" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132C2A03C80B66512EC8F90996146AB9" box="[731,797,1679,1704]" captionStart="FIGURE 1" captionStartId="1.[136,229,1926,1947]" captionTargetBox="[158,1430,243,1837]" captionTargetId="figure-14@1.[151,1436,214,1855]" captionTargetPageId="1" captionText="FIGURE 1. Phyllagathis stellata C.W. Lin &amp; C.H. Lee. A. Habit; B. Portion of leaf abaxial surface; C, C. Flower, front and side views; D, D'. Petal, inside and side views; E, E', E''. Stamens, side, dorsal and ventral views; F. Hypanthium; G. Longitudinal section of ovary; H. Portion of crown lobe, showing margins denticulate with gland-trichomes; I. Capsules, side view. J. Infructescences.All from C.W. Lin 643 (TAIF)." figureDoi="http://doi.org/10.5281/zenodo.13704456" httpUri="https://zenodo.org/record/13704456/files/figure.png" pageId="0" pageNumber="295">Fig. 1</figureCitation>
,
<figureCitation id="132C2A03C80B66512F38F90996336AB9" box="[811,826,1679,1704]" captionStart="FIGURE 2" captionStartId="2.[136,229,1936,1957]" captionTargetBox="[151,1436,187,1908]" captionTargetId="figure-16@2.[151,1436,187,1908]" captionTargetPageId="2" captionText="FIGURE 2. Phyllagathis stellata C.W. Lin &amp; C.H. Lee. A. Habit and habitats; B. Young plant; C. Portion of leaf adaxial surface; D. Portion of leaf abaxial surface; E. Immature inflorescence; F. Inflorescence; G. Flower, front view; H. Hypanthium; I. Anthers, dorsal, side and ventral views; J. Appendage of sepals; K. Vertical section of an ovary; L. Capsules, apical view; M. Old-age capsules.All from C.W. Lin 643 (TAIF)." figureDoi="http://doi.org/10.5281/zenodo.13704458" httpUri="https://zenodo.org/record/13704458/files/figure.png" pageId="0" pageNumber="295">2</figureCitation>
)
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<paragraph id="8BA83686C80B66512C9BF93396C56AFF" blockId="0.[136,1452,1717,1774]" pageId="0" pageNumber="295">
Type:—
<collectingCountry id="F3007616C80B66512CC6F933945F6ADB" box="[213,342,1717,1738]" name="Malaysia" pageId="0" pageNumber="295">MALAYSIA</collectingCountry>
. Borneo,
<collectingRegion id="49D3F864C80B66512DA2F933970F6ADB" box="[433,518,1717,1738]" country="Malaysia" name="Sarawak" pageId="0" pageNumber="295">Sarawak</collectingRegion>
, Sri Aman Division, Batang Ai,
<emphasis id="B963EA94C80B66512F2FF933965A6ADB" box="[828,851,1717,1738]" italics="true" pageId="0" pageNumber="295">ca</emphasis>
.
<quantity id="4CEF9B63C80B66512F4CF93396956ADB" box="[863,924,1717,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="295" unit="m" value="100.0">100 m</quantity>
elev. Type specimens pressed from plants cultivated in a nursery in Taiwan,
<date id="FFA91046C80B66512D94F95F94FC6AFF" box="[391,501,1753,1774]" pageId="0" pageNumber="295" value="2017-09-05">5 Sep 2017</date>
,
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<collectionCode id="ED06AE43C80B66512DECF95F97196AFF" box="[511,528,1753,1774]" country="Denmark" name="University of Copenhagen" pageId="0" pageNumber="295" type="Herbarium">C</collectionCode>
.
<collectionCode id="ED06AE43C80B66512E00F95F97236AFF" box="[531,554,1753,1774]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="0" pageNumber="295" type="Herbarium">W</collectionCode>
. Lin 643
</emphasis>
(
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<collectionCode id="ED06AE43C80B66512EF7F95F96166AFF" box="[740,799,1753,1774]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15478" lsid="urn:lsid:biocol.org:col:15478" name="Taiwan Forestry Research Institute" pageId="0" pageNumber="295" type="Herbarium">TAIF</collectionCode>
!,
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<collectionCode id="ED06AE43C80B66512F64F95F96B76AFF" box="[887,958,1753,1774]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15074" lsid="urn:lsid:biocol.org:col:15074" name="Research Center for Biodiversity, Academia Sinica" pageId="0" pageNumber="295" type="Herbarium">HAST</collectionCode>
!).
</paragraph>
<paragraph id="8BA83686C80B66512C9BF89991676BB5" blockId="0.[136,1452,1823,2064]" pageId="0" pageNumber="295">
Diagnosis:
<taxonomicName id="4C174D05C80B66512D17F89994E16B29" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[260,488,1823,1848]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="0" pageNumber="295" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80B66512D17F89994E16B29" box="[260,488,1823,1848]" italics="true" pageId="0" pageNumber="295">Phyllagathis stellata</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C174D05C80B66512E72F89991EA6B29" authority="(M. P. Nayar, 1965: 502) Cellinese (2003: 85)" authorityName="Cellinese" authorityPageNumber="85" authorityYear="2003" baseAuthorityName="M. P. Nayar" baseAuthorityPageNumber="502" baseAuthorityYear="1965" box="[609,1251,1823,1848]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="0" pageNumber="295" phylum="Tracheophyta" rank="species" species="jacobsiana">
<emphasis id="B963EA94C80B66512E72F89997FC6B29" box="[609,757,1823,1848]" italics="true" pageId="0" pageNumber="295">P. jacobsiana</emphasis>
(M.P.
<bibRefCitation id="EF864B77C80B66512F29F89996F06B29" author="Nayar, M. P." box="[826,1017,1823,1848]" pageId="0" pageNumber="295" pagination="501 - 506" refId="ref2440" refString="Nayar, M. P. (1965) Contributions to the knowledge of Indo-Malaysian and other Asiatic Melastomataceae. Kew Bulletin 19: 501 - 506. https: // doi. org / 10.2307 / 4108209" type="journal article" year="1965">Nayar, 1965: 502</bibRefCitation>
)
<bibRefCitation id="EF864B77C80B66512817F89991EA6B29" author="Cellinese, N." box="[1028,1251,1823,1848]" pageId="0" pageNumber="295" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 85)</bibRefCitation>
</taxonomicName>
in its hairy leaves, pentamerous flowers and fleshy finger appendiculate sepals. The new species is different in its lamina with 11 or 13 basal veins (
<emphasis id="B963EA94C80B66512D00F8E194386B91" box="[275,305,1895,1920]" italics="true" pageId="0" pageNumber="295">vs.</emphasis>
7 or 9-veins), inflorescence shorter (
<emphasis id="B963EA94C80B66512EDFF8E197E26B91" box="[716,747,1895,1920]" italics="true" pageId="0" pageNumber="295">vs.</emphasis>
longer) than petiole, hypanthium ribbed and densely verrucose (
<emphasis id="B963EA94C80B66512C82F80D95B96BB5" box="[145,176,1931,1956]" italics="true" pageId="0" pageNumber="295">vs.</emphasis>
smooth), stamens with a dorsal tuberculate appendage (
<emphasis id="B963EA94C80B66512F36F80D964A6BB5" box="[805,835,1931,1956]" italics="true" pageId="0" pageNumber="295">vs.</emphasis>
dorsally inappendiculate).
</paragraph>
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Caulescent herbs, ascending or creeping, terrestrial. Stems unbranched, rooting at nodes and creeping up soil slope, ascending or suberect upper part; olive green to reddish brown, to
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or longer,
<quantity id="4CEF9B63C80B6651284EF85591D06BFD" box="[1117,1241,2003,2028]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="9.0" metricValueMin="4.0" pageId="0" pageNumber="295" unit="cm" value="0.65" valueMax="0.9" valueMin="0.4">0.40.9 cm</quantity>
in diameter, terete, stout, slightly woody at base, covered with densely gland-tipped trichomes; internodes
<quantity id="4CEF9B63C80B66512871F87191E96401" box="[1122,1248,2039,2064]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.8" metricValueMin="0.5" pageId="0" pageNumber="295" unit="cm" value="1.65" valueMax="2.8" valueMin="0.5">0.52.8 cm</quantity>
long. Leaf blades 25, decussate, 1018 ×
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,, isophyllous, very widely to depressed ovate, base cordate, margins denticulate and sparsely incised, with rows of long gland-tipped trichomes up to
<quantity id="4CEF9B63C80866522F9AFF4296C06CCC" box="[905,969,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="298" unit="mm" value="5.0">5 mm</quantity>
long, apex shortly acuminate; chartaceous; venation acrodromous, 11 or 13 veined, basal; veins slightly depressed on the adaxial surface and prominent on the abaxial surface, secondary and tertiary veins numerous and conspicuous, reticulate; adaxially emerald green, slightly bullate and long gland-tipped trichomes between veins; abaxially pale green, sparsely gland-tipped trichomes on all veins. Petioles
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long, flat terete and slightly grooved adaxially, densely covered with gland-tipped trichomes and minutely translucent puberulous. Bracts inconspicuous,
<emphasis id="B963EA94C80866522F29FEFE965D6D80" box="[826,852,376,401]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
.
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long, linear. Inflorescences in the upper leaf axils, very congested pleiochasium (umbelliform in young plants), peduncle
<quantity id="4CEF9B63C80866522FE0FE1A915F6DA4" box="[1011,1110,412,437]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="3" pageNumber="298" unit="cm" value="9.5" valueMax="13.0" valueMin="6.0">613 cm</quantity>
long, reddish brown, covered with gland-tipped trichomes, densely glandular and minutely translucent puberulous. Flowers pentamerous, pedicels
<emphasis id="B963EA94C80866522C9BFE6295AB6DEC" box="[136,162,484,509]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
.
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long, densely glandular and minutely translucent puberulous. Hypanthium campanulate,
<emphasis id="B963EA94C808665228FFFE62900F6DEC" box="[1260,1286,484,509]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
.
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long,
<quantity id="4CEF9B63C8086652298EFE6295BD6E30" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="298" unit="mm" value="3.0">3 mm</quantity>
in diameter, covered with densely minutely translucent puberulous and gland-tipped trichomes &lt;
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long; 10- ribbed, with undertint verrucose, giving hypanthium a rugose appearance. Sepals 5, persistent, very widely triangular, apex rounded, connate into a torus, each lobe with an acute-angled triangular external tooth; lobes alternating with fleshy finger appendages, appendage unbranched or multi-branched, tipped by a gland. Petals 5, oblique, glabrous, widely elliptic to oblong,
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×
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wide, white, sometimes pale pinkish, apex mucronate-apiculate. Stamens 10, isomorphic, filaments slightly flat,
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long, white, pinkish toward apex, anthers subulate, apex attenuate, slightly curved ventrally,
<quantity id="4CEF9B63C80866522E32FD66979C6EE8" box="[545,661,736,761]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.0" metricValueMin="4.5" pageId="3" pageNumber="298" unit="mm" value="5.25" valueMax="6.0" valueMin="4.5">4.56 mm</quantity>
long, purplish to bluish, pore 1, connective distinct, with one pair of ventral auricular appendages on the base, dorsal appendage
<emphasis id="B963EA94C80866522F3CFC8296406F0C" box="[815,841,772,797]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
.
<quantity id="4CEF9B63C80866522F4AFC8296BB6F0C" box="[857,946,772,797]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="298" unit="mm" value="0.3">0.3 mm</quantity>
, tuberculate. Style
<emphasis id="B963EA94C80866522898FC8291AC6F0C" box="[1163,1189,772,797]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
.
<quantity id="4CEF9B63C808665228A6FC82900C6F0C" box="[1205,1285,772,797]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="298" unit="mm" value="15.0">15 mm</quantity>
long, filiform, glabrous, stigma capitate. Ovary
<emphasis id="B963EA94C80866522DE7FCAE971F6F50" box="[500,534,808,833]" italics="true" pageId="3" pageNumber="298">ca.</emphasis>
4/5 as long as the hypanthium, crown with fully connate lobes, margins denticulate with each tooth tipped by a glandular trichome, anther pockets shallow, placentae stalked. Capsules on pedicels
<quantity id="4CEF9B63C80866522967FCCA95BD6F98" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="3" pageNumber="298" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">610 mm</quantity>
long, hypanthium cup-shaped,
<emphasis id="B963EA94C80866522E06FCF697266F98" box="[533,559,880,905]" italics="true" pageId="3" pageNumber="298">ca</emphasis>
. 6 ×
<quantity id="4CEF9B63C80866522E7AFCF697A56F98" box="[617,684,880,905]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="298" unit="mm" value="6.0">6 mm</quantity>
, placentae disintegrating after seed dehiscence.
</paragraph>
<caption id="DF68660EC80A66502C9BF80095F46416" ID-DOI="http://doi.org/10.5281/zenodo.13704456" ID-Zenodo-Dep="13704456" httpUri="https://zenodo.org/record/13704456/files/figure.png" pageId="1" pageNumber="296" startId="1.[136,229,1926,1947]" targetBox="[158,1430,243,1837]" targetPageId="1" targetType="figure">
<paragraph id="8BA83686C80A66502C9BF80095F46416" blockId="1.[136,1452,1925,2055]" pageId="1" pageNumber="296">
<emphasis id="B963EA94C80A66502C9BF80095F76B8A" bold="true" box="[136,254,1926,1947]" pageId="1" pageNumber="296">FIGURE 1.</emphasis>
<taxonomicName id="4C174D05C80A66502D10F80396086B8A" authority="C. W. Lin &amp; C. H. Lee. A. Habit" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[259,769,1925,1947]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="296" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80A66502D10F80394C56B8A" bold="true" box="[259,460,1925,1947]" italics="true" pageId="1" pageNumber="296">Phyllagathis stellata</emphasis>
C.W. Lin &amp; C.H. Lee. A. Habit
</taxonomicName>
; B. Portion of leaf abaxial surface; C, C. Flower, front and side views; D, D'. Petal, inside and side views; E, E', E''. Stamens, side, dorsal and ventral views; F. Hypanthium; G. Longitudinal section of ovary; H. Portion of crown lobe, showing margins denticulate with gland-trichomes; I. Capsules, side view. J. Infructescences. All from
<emphasis id="B963EA94C80A6650294BF84890A26BF2" box="[1368,1451,1998,2019]" italics="true" pageId="1" pageNumber="296">C.W. Lin</emphasis>
643 (TAIF).
</paragraph>
</caption>
<caption id="DF68660EC80966532C9BF816945F6400" ID-DOI="http://doi.org/10.5281/zenodo.13704458" ID-Zenodo-Dep="13704458" httpUri="https://zenodo.org/record/13704458/files/figure.png" pageId="2" pageNumber="297" startId="2.[136,229,1936,1957]" targetBox="[151,1436,187,1908]" targetPageId="2" targetType="figure">
<paragraph id="8BA83686C80966532C9BF816945F6400" blockId="2.[136,1452,1935,2065]" pageId="2" pageNumber="297">
<emphasis id="B963EA94C80966532C9BF81695F66BB4" bold="true" box="[136,255,1936,1957]" pageId="2" pageNumber="297">FIGURE 2.</emphasis>
<taxonomicName id="4C174D05C80966532D14F80996186BB4" authority="C. W. Lin &amp; C. H. Lee. A. Habit" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[263,785,1935,1957]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="297" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80966532D14F80994DB6BB4" bold="true" box="[263,466,1935,1957]" italics="true" pageId="2" pageNumber="297">Phyllagathis stellata</emphasis>
C.W. Lin &amp; C.H. Lee. A. Habit
</taxonomicName>
and habitats; B. Young plant; C. Portion of leaf adaxial surface; D. Portion of leaf abaxial surface; E. Immature inflorescence; F. Inflorescence; G. Flower, front view; H. Hypanthium; I. Anthers, dorsal, side and ventral views; J. Appendage of sepals; K. Vertical section of an ovary; L. Capsules, apical view; M. Old-age capsules. All from
<emphasis id="B963EA94C80966532C9BF87A95D56400" box="[136,220,2044,2065]" italics="true" pageId="2" pageNumber="297">C.W. Lin</emphasis>
643 (TAIF).
</paragraph>
</caption>
<caption id="DF68660EC80866522C9BF78A97F06430" box="[136,761,2060,2081]" pageId="3" pageNumber="298" startId="3.[136,229,2060,2081]" targetType="figure">
<paragraph id="8BA83686C80866522C9BF78A97F06430" blockId="3.[136,761,2060,2081]" box="[136,761,2060,2081]" pageId="3" pageNumber="298">
<emphasis id="B963EA94C80866522C9BF78A95F76430" bold="true" box="[136,254,2060,2081]" pageId="3" pageNumber="298">FIGURE 3.</emphasis>
Distribution of
<taxonomicName id="4C174D05C80866522D84F78A94FF6430" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[407,502,2060,2081]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="298" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80866522D84F78A94FF6430" box="[407,502,2060,2081]" italics="true" pageId="3" pageNumber="298">P. stellata</emphasis>
</taxonomicName>
(star) in Sarawak, Borneo.
</paragraph>
</caption>
<paragraph id="8BA83686C80F66552CAEFF26906F6CCC" blockId="4.[136,1455,160,653]" pageId="4" pageNumber="299">
<emphasis id="B963EA94C80F66552CAEFF26975E6CA8" bold="true" box="[189,599,160,185]" pageId="4" pageNumber="299">Additional specimen examined:—</emphasis>
<collectingCountry id="F3007616C80F66552E44FF2697E56CA8" box="[599,748,160,185]" name="Malaysia" pageId="4" pageNumber="299">MALAYSIA</collectingCountry>
. Borneo,
<collectingRegion id="49D3F864C80F66552F48FF2696B76CA8" box="[859,958,160,185]" country="Malaysia" name="Sarawak" pageId="4" pageNumber="299">Sarawak</collectingRegion>
, Sri Aman Division, Batang Ai,
<emphasis id="B963EA94C80F66552924FF2690586CA8" box="[1335,1361,160,185]" italics="true" pageId="4" pageNumber="299">ca</emphasis>
.
<quantity id="4CEF9B63C80F66552971FF2690A56CA8" box="[1378,1452,160,185]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="299" unit="m" value="100.0">100 m</quantity>
elev.
<typeStatus id="54AC8824C80F66552CD1FF4295F36CCC" box="[194,250,196,221]" pageId="4" pageNumber="299">Type</typeStatus>
specimens pressed from plants cultivated in a nursery in Taiwan,
<date id="FFA91046C80F66552FCFFF4291626CCC" box="[988,1131,196,221]" pageId="4" pageNumber="299" value="2017-09-10">10 Sep 2017</date>
,
<emphasis id="B963EA94C80F66552865FF4290006CCC" box="[1142,1289,196,221]" italics="true" pageId="4" pageNumber="299">
<collectionCode id="ED06AE43C80F66552865FF4291836CCC" box="[1142,1162,196,221]" country="Denmark" name="University of Copenhagen" pageId="4" pageNumber="299" type="Herbarium">C</collectionCode>
.
<collectionCode id="ED06AE43C80F6655289EFF4291A16CCC" box="[1165,1192,196,221]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="4" pageNumber="299" type="Herbarium">W</collectionCode>
. Lin 646
</emphasis>
(
<collectionCode id="ED06AE43C80F66552904FF4290536CCC" box="[1303,1370,196,221]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15478" lsid="urn:lsid:biocol.org:col:15478" name="Taiwan Forestry Research Institute" pageId="4" pageNumber="299" type="Herbarium">TAIF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D650DC80F66552CAEFF6E91D36D34" pageId="4" pageNumber="299" type="distribution">
<paragraph id="8BA83686C80F66552CAEFF6E91D36D34" blockId="4.[136,1455,160,653]" pageId="4" pageNumber="299">
<emphasis id="B963EA94C80F66552CAEFF6E94E36D10" bold="true" box="[189,490,232,257]" pageId="4" pageNumber="299">Distribution and ecology:</emphasis>
—This new species is endemic to
<collectingCountry id="F3007616C80F66552F71FF6E96CC6D10" box="[866,965,232,257]" name="Malaysia" pageId="4" pageNumber="299">Sarawak</collectingCountry>
, currently only known from Batang Ai (
<figureCitation id="132C2A03C80F6655296DFF6E95916D34" captionStart="FIGURE 3" captionStartId="3.[136,229,2060,2081]" captionText="FIGURE 3. Distribution of P. stellata (star) in Sarawak, Borneo." pageId="4" pageNumber="299">Fig. 3</figureCitation>
). Growing on riparian forest on soil steep slope in deeply shaded areas, at
<quantity id="4CEF9B63C80F66552FC8FE8A91536D34" box="[987,1114,268,293]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="299" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100200 m</quantity>
elevations.
</paragraph>
</subSubSection>
<subSubSection id="C30D650DC80F66552CAEFEB691EA6E9C" pageId="4" pageNumber="299" type="etymology">
<paragraph id="8BA83686C80F66552CAEFEB6977C6D7C" blockId="4.[136,1455,160,653]" pageId="4" pageNumber="299">
<emphasis id="B963EA94C80F66552CAEFEB694416D58" bold="true" box="[189,328,304,329]" pageId="4" pageNumber="299">Etymology:</emphasis>
—The specific epithet refers to the resemblance of this new species capsule from apical view that appears star-like with radiating appendages.
</paragraph>
<paragraph id="8BA83686C80F66552CAEFEFE97BA6DA4" blockId="4.[136,1455,160,653]" pageId="4" pageNumber="299">
<emphasis id="B963EA94C80F66552CAEFEFE94F06D80" bold="true" box="[189,505,376,401]" pageId="4" pageNumber="299">Proposed IUCN Category:</emphasis>
—DD-Data Deficient, according to
<bibRefCitation id="EF864B77C80F66552F99FEFE912A6D80" author="IUCN Standards &amp; Petitions Subcommittee" box="[906,1059,376,401]" pageId="4" pageNumber="299" refId="ref1975" refString="IUCN Standards and Petitions Subcommittee (2016) Guidelines for using the IUCN Red List categories and criteria. Version 11. Available from: http: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 9 September 2016)" type="book" year="2016">IUCN (2016)</bibRefCitation>
: the species is rare but possibly not threatened as it grows in an inaccessible location.
</paragraph>
<paragraph id="8BA83686C80F66552CAEFE4691EA6E9C" blockId="4.[136,1455,160,653]" pageId="4" pageNumber="299">
<emphasis id="B963EA94C80F66552CAEFE4695F66DC8" bold="true" box="[189,255,448,473]" pageId="4" pageNumber="299">Note:</emphasis>
<taxonomicName id="4C174D05C80F66552D14FE4694E66DC8" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" box="[263,495,448,473]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80F66552D14FE4694E66DC8" box="[263,495,448,473]" italics="true" pageId="4" pageNumber="299">Phyllagathis stellata</emphasis>
</taxonomicName>
belongs to a distinct group of Bornean
<taxonomicName id="4C174D05C80F66552FA7FE46914B6DC8" box="[948,1090,448,473]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="genus">
<emphasis id="B963EA94C80F66552FA7FE46914B6DC8" box="[948,1090,448,473]" italics="true" pageId="4" pageNumber="299">Phyllagathis</emphasis>
</taxonomicName>
that have pentamerous flowers. Among this group that includes
<taxonomicName id="4C174D05C80F66552DEAFE6296A76DEC" authority="(Stapf) Cellinese (2003: 83)" authorityName="Cellinese" authorityPageNumber="83" authorityYear="2003" baseAuthorityName="Stapf" box="[505,942,484,509]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="guttata">
<emphasis id="B963EA94C80F66552DEAFE62976C6DEC" box="[505,613,484,509]" italics="true" pageId="4" pageNumber="299">P. guttata</emphasis>
(Stapf)
<bibRefCitation id="EF864B77C80F66552ED7FE6296A76DEC" author="Cellinese, N." box="[708,942,484,509]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 83)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C174D05C80F66552FAFFE6290AE6DEC" authority="(Nayar) Cellinese (2003: 85)" authorityName="Cellinese" authorityPageNumber="85" authorityYear="2003" baseAuthorityName="M. P. Nayar" baseAuthorityYear="1965" box="[956,1447,484,509]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="jacobsiana">
<emphasis id="B963EA94C80F66552FAFFE62915D6DEC" box="[956,1108,484,509]" italics="true" pageId="4" pageNumber="299">P. jacobsiana</emphasis>
(Nayar)
<bibRefCitation id="EF864B77C80F665528AFFE6290AE6DEC" author="Cellinese, N." box="[1212,1447,484,509]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 85)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C174D05C80F66552C9BFD8E97606E30" authority="(Nayar) Cellinese (2003: 88)" authorityName="Cellinese" authorityPageNumber="88" authorityYear="2003" baseAuthorityName="Nayar" box="[136,617,520,545]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="osmantha">
<emphasis id="B963EA94C80F66552C9BFD8E941A6E30" box="[136,275,520,545]" italics="true" pageId="4" pageNumber="299">P. osmantha</emphasis>
(Nayar)
<bibRefCitation id="EF864B77C80F66552D6EFD8E97606E30" author="Cellinese, N." box="[381,617,520,545]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 88)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C174D05C80F66552E64FD8E91276E30" authority="Cellinese (2003: 89)" authorityName="Cellinese" authorityPageNumber="89" authorityYear="2003" box="[631,1070,520,545]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="penrissenensis">
<emphasis id="B963EA94C80F66552E64FD8E96316E30" box="[631,824,520,545]" italics="true" pageId="4" pageNumber="299">P. penrissenensis</emphasis>
<bibRefCitation id="EF864B77C80F66552F51FD8E91276E30" author="Cellinese, N." box="[834,1070,520,545]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 89)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C174D05C80F6655282FFD8E94B36E54" authority="C. W. Lin, Chien F. Chen &amp; T. Y. A. Yang (2015: 123)" authorityName="C. W. Lin, Chien F. Chen &amp; T. Y. A. Yang" authorityYear="2015" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="rajah">
<emphasis id="B963EA94C80F6655282FFD8E919E6E30" box="[1084,1175,520,545]" italics="true" pageId="4" pageNumber="299">P. rajah</emphasis>
C.W.Lin, Chien F.Chen &amp; T.Y.A.Yang (2015: 123)
</taxonomicName>
,
<taxonomicName id="4C174D05C80F66552DD4FDAA96556E54" authority="(Stapf) Cellinese (2003: 91)" authorityName="Cellinese" authorityPageNumber="91" authorityYear="2003" baseAuthorityName="Stapf" box="[455,860,556,581]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="rufa">
<emphasis id="B963EA94C80F66552DD4FDAA971A6E54" box="[455,531,556,581]" italics="true" pageId="4" pageNumber="299">P. rufa</emphasis>
(Stapf)
<bibRefCitation id="EF864B77C80F66552E60FDAA96556E54" author="Cellinese, N." box="[627,860,556,581]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 91)</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C174D05C80F66552F86FDAA90AE6E54" authority="(Cogniaux) Cellinese (2003: 93)" authorityName="Cellinese" authorityPageNumber="93" authorityYear="2003" baseAuthorityName="Cogniaux" box="[917,1447,556,581]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="subacaulis">
<emphasis id="B963EA94C80F66552F86FDAA91226E54" box="[917,1067,556,581]" italics="true" pageId="4" pageNumber="299">P. subacaulis</emphasis>
(Cogniaux)
<bibRefCitation id="EF864B77C80F665528AFFDAA90AE6E54" author="Cellinese, N." box="[1212,1447,556,581]" pageId="4" pageNumber="299" pagination="69 - 97" refId="ref1926" refString="Cellinese, N. (2003) Revision of the genus Phyllagathis (Melastomataceae: Sonerileae) II. The species in Borneo and Natuna Island. Blumea 48 (1): 69 - 97. https: // doi. org / 10.3767 / 000651903 X 686060" type="journal article" year="2003">Cellinese (2003: 93)</bibRefCitation>
</taxonomicName>
, it most resembles
<taxonomicName id="4C174D05C80F66552D45FDD694E76E78" authorityName="Cellinese" authorityYear="2003" baseAuthorityName="M. P. Nayar" baseAuthorityYear="1965" box="[342,494,592,617]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="jacobsiana">
<emphasis id="B963EA94C80F66552D45FDD694E76E78" box="[342,494,592,617]" italics="true" pageId="4" pageNumber="299">P. jacobsiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C174D05C80F66552E34FDD6979B6E78" authorityName="Cellinese" authorityYear="2003" baseAuthorityName="Stapf" box="[551,658,592,617]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="guttata">
<emphasis id="B963EA94C80F66552E34FDD6979B6E78" box="[551,658,592,617]" italics="true" pageId="4" pageNumber="299">P. guttata</emphasis>
</taxonomicName>
in its fleshy finger appendiculate sepals. However, in other aspects
<taxonomicName id="4C174D05C80F66552984FDD695D36E9C" authorityName="C. W. Lin &amp; C. H. Lee. A. Habit" authorityYear="2018" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="299" phylum="Tracheophyta" rank="species" species="stellata">
<emphasis id="B963EA94C80F66552984FDD695D36E9C" italics="true" pageId="4" pageNumber="299">P. stellata</emphasis>
</taxonomicName>
is sharply distinct. A comparison of salient features of the three species is shown in
<tableCitation id="C695033DC80F6655289AFDF291D46E9C" box="[1161,1245,628,653]" captionStart="TABLE 1" captionStartId="4.[136,224,701,724]" captionText="TABLE 1. Comparison of Phyllagathis stellata with P. jacobsiana and P. guttata" pageId="4" pageNumber="299">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>