226 lines
43 KiB
XML
226 lines
43 KiB
XML
<document id="F2FC1987876595CA2226089AC9B0A478" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFD9FFEDBC8FFBDE0861004D" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus cuspilobatus Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="48" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="48" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3E9851117D6EDC7F84F43286C8BF12FD">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="1C7180D483F130B98A1DAEC51562D429" type="personal">
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<mods:roleTerm id="7DBAD2A55102C2BFA61169A19A94EABE">Author</mods:roleTerm>
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<mods:namePart id="127241D69CBFD0D3A38833E8C785A38A">Frisch, Johannes</mods:namePart>
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<mods:roleTerm id="44B03D42483AAFF9E410BDF0342D6766">Author</mods:roleTerm>
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<mods:namePart id="6FA4DB91B8FFA2BB74AA5473E512A186">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="E18862C4E4DAAF15C69F4767149F4DC5">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:typeOfResource id="FCF23B1FF57BB216684AD21E9E28834A">text</mods:typeOfResource>
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<mods:title id="6A07A9FCB112091CD9984DBABE25EFA3">Soil Organisms</mods:title>
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<mods:part id="AEA8330C676031753636717654AF765F">
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<mods:date id="BAF712EB7478F4054108B542019062C7">2022</mods:date>
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<mods:number id="B8CF2B90FBF49F33A3B76E2F8B6920D5">2022-04-01</mods:number>
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<mods:number id="1B22DC75CD811C6EC6850C39D8514384">95</mods:number>
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<mods:url id="91E0793F40E06A3AD7F78BBB811BEF09">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="EE38313973983B086B44ACCDEF70442F">journal article</mods:classification>
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<mods:identifier id="2604A58D7A1144004D3CCA2305488625" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFD9FFEDBC8FFBDE0861004D" ID-DOI="http://doi.org/10.5281/zenodo.10724313" ID-Zenodo-Dep="10724313" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFEDBC8FFBDE0861004D" httpUri="http://treatment.plazi.org/id/03B7C77AFFD9FFEDBC8FFBDE0861004D" lastPageNumber="48" pageId="25" pageNumber="48">
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<subSubSection id="C30425E7FFD9FFEDBC8FFBDE0FDC004D" box="[212,705,1061,1088]" pageId="25" pageNumber="48" type="nomenclature">
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<paragraph id="8BA1766CFFD9FFEDBC8FFBDE0FDC004D" blockId="25.[212,705,1061,1121]" box="[212,705,1061,1088]" pageId="25" pageNumber="48">
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<heading id="D0E9C100FFD9FFEDBC8FFBDE0FDC004D" box="[212,705,1061,1088]" fontSize="11" level="2" pageId="25" pageNumber="48" reason="5">
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<taxonomicName id="4C1E0DEFFFD9FFEDBC8FFBDE0F5D004D" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[212,576,1061,1088]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="cuspilobatus" status="sp. nov.">
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<emphasis id="B96AAA7EFFD9FFEDBC8FFBDE0CF6004D" bold="true" box="[212,491,1061,1088]" italics="true" pageId="25" pageNumber="48">Scopaeus cuspilobatus</emphasis>
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Frisch
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFD9FFEDBE17FBDE0FDC004D" box="[588,705,1062,1088]" pageId="25" pageNumber="48" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30425E7FFD9FFEDBC8FFBBE0F6D006C" box="[212,624,1094,1121]" pageId="25" pageNumber="48" type="description">
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<paragraph id="8BA1766CFFD9FFEDBC8FFBBE0F6D006C" blockId="25.[212,705,1061,1121]" box="[212,624,1094,1121]" pageId="25" pageNumber="48">
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(
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<figureCitation id="13256AE9FFD9FFEDBC87FBBE0C28006D" box="[220,309,1094,1120]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="25" pageNumber="48">Figs 16</figureCitation>
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,
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<figureCitation id="13256AE9FFD9FFEDBD19FBBF0C7C006C" box="[322,353,1095,1121]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">38</figureCitation>
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,
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<figureCitation id="13256AE9FFD9FFEDBD35FBBF0CDF006C" box="[366,450,1095,1121]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92 – 94</figureCitation>
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, 134, 167, 168)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFD9FFEDBC8FFB720FA20108" pageId="25" pageNumber="48" type="materials_examined">
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<paragraph id="8BA1766CFFD9FFEDBC8FFB720FA20108" blockId="25.[189,800,1162,1975]" pageId="25" pageNumber="48">
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<emphasis id="B96AAA7EFFD9FFEDBC8FFB720C8E00AF" bold="true" box="[212,403,1162,1186]" pageId="25" pageNumber="48">Type specimens:</emphasis>
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<materialsCitation id="3B767C31FFD9FFEDBDC1FB720E0100E9" collectingDate="2015-09-21" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="25" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
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<collectingCountry id="F30936FCFFD9FFEDBDC1FB720F1500AF" box="[410,520,1162,1186]" name="Indonesia" pageId="25" pageNumber="48">Indonesia</collectingCountry>
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:
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<typeStatus id="54A5C8CEFFD9FFEDBE48FB720F6500AF" box="[531,632,1162,1186]" pageId="25" pageNumber="48" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFD9FFEDBEDBFB730F8900AF" box="[640,660,1163,1186]" count="1" pageId="25" pageNumber="48" type="male">♂</specimenCount>
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,
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<collectingRegion id="49DAB88EFFD9FFEDBEFBFB730E0600AF" box="[672,795,1163,1186]" country="Indonesia" name="Jawa Barat" pageId="25" pageNumber="48">Jawa Barat</collectingRegion>
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,
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<collectingCounty id="62C00EE0FFD9FFEDBCE6FB530C2D00CE" box="[189,304,1195,1219]" pageId="25" pageNumber="48">Sukabumi</collectingCounty>
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:
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<collectingMunicipality id="6BC5EC16FFD9FFEDBD11FB530CAC00CE" box="[330,433,1195,1219]" pageId="25" pageNumber="48">Sirnarasa</collectingMunicipality>
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(
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<location id="8EC120B7FFD9FFEDBD95FB530F6F00CE" LSID="urn:lsid:plazi:treatment:03B7C77AFFD9FFEDBC8FFBDE0861004D:8EC120B7FFD9FFEDBD95FB530F6F00CE" box="[462,626,1195,1219]" country="Indonesia" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Cimaja River" pageId="25" pageNumber="48" stateProvince="Jawa Barat">Cimaja River</location>
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), (
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<geoCoordinate id="EE2A10ABFFD9FFEDBEC7FB530E0700CE" box="[668,794,1195,1219]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="25" pageNumber="48" precision="15" seconds="32" value="-6.8588886">06°51’32’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFD9FFEDBCE6FB340C5600E9" box="[189,331,1228,1252]" degrees="106" direction="east" minutes="31" orientation="longitude" pageId="25" pageNumber="48" precision="15" seconds="06" value="106.51833">106°31’06’’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFD9FFEDBD3CFB340CAB00EE" box="[359,438,1228,1252]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="25" pageNumber="48" unit="m" value="670.0">
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<elevation id="0033915FFFD9FFEDBD3CFB340CAB00EE" box="[359,438,1228,1252]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="25" pageNumber="48" unit="m" value="670.0">670 m</elevation>
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</quantity>
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,
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<date id="FFA050ACFFD9FFEDBD92FB340F2D00E9" box="[457,560,1228,1252]" pageId="25" pageNumber="48" value="2015-09-21">
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<collectingDate id="EFE4A944FFD9FFEDBD92FB340F2D00E9" box="[457,560,1228,1252]" pageId="25" pageNumber="48" value="2015-09-21">21.9.2015</collectingDate>
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</date>
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, leg.
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<collectorName id="26EB13BAFFD9FFEDBE22FB340FA000E9" box="[633,701,1228,1252]" pageId="25" pageNumber="48">Frisch</collectorName>
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(
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||
<collectionCode id="ED0FEEA9FFD9FFEDBE8FFB340E0900E9" box="[724,788,1228,1252]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="25" pageNumber="48" type="Museum">MZB</collectionCode>
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)
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</materialsCitation>
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.
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||
<materialsCitation id="3B767C31FFD9FFEDBCE6FB150FA60108" box="[189,699,1261,1285]" collectingDate="2015-09-21" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="25" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
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<typeStatus id="54A5C8CEFFD9FFEDBCE6FB150C3F0108" box="[189,290,1261,1285]" pageId="25" pageNumber="48" type="paratype">Paratype</typeStatus>
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:
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<specimenCount id="9D18BDE5FFD9FFEDBD77FB150C4E0109" box="[300,339,1261,1285]" count="1" pageId="25" pageNumber="48" type="female">1 ♀</specimenCount>
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||
, same data as holotype (
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<collectionCode id="ED0FEEA9FFD9FFEDBE3BFB150FAE0108" box="[608,691,1261,1285]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="25" pageNumber="48">MFNB</collectionCode>
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||
)
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||
</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFD9FFEDBC8FFAF508540754" pageId="25" pageNumber="48" type="description">
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<paragraph id="8BA1766CFFD9FFEDBC8FFAF50FB102BD" blockId="25.[189,800,1162,1975]" pageId="25" pageNumber="48">
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<emphasis id="B96AAA7EFFD9FFEDBC8FFAF50C790128" bold="true" box="[212,356,1293,1317]" pageId="25" pageNumber="48">Description:</emphasis>
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||
Habitus and coloring as in
|
||
<figureCitation id="13256AE9FFD9FFEDBED4FAF60FC1012B" box="[655,732,1294,1318]" captionStart="Figures 14–16" captionStartId="9.[189,262,1930,1952]" captionTargetBox="[262,1384,910,1880]" captionTargetId="figure-57@9.[189,1436,894,1911]" captionTargetPageId="9" captionText="Figures 14–16. Habitus of Scopaeus heronifer, paratype, Jawa Barat: Mt Salak (14); S. diversilobatus, paratype, Jawa Barat, Sukabumi: Djampang Kulon (15); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (16)." figureDoi="http://doi.org/10.5281/zenodo.10724246" httpUri="https://zenodo.org/record/10724246/files/figure.png" pageId="25" pageNumber="48">Fig. 16</figureCitation>
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. Head subquadrate. Penultimate antennal segment quadrate. Mesotibia moderately thickened. Setose punctation of body surface fine, dense on subnitid head, rugulose on matt elytra, more spacious on shiny pronotum; microreticulation absent. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown except for narrow, light brown posterior margin of elytra and tip of abdomen; antennae medium brown with darker segments 2 – 6; maxillary palpi light brown with darker penultimate segment; legs light brown with medium brown femora. Total body length
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||
<quantity id="4CE6DB89FFD9FFEDBD53F9600C4302BD" box="[264,350,1688,1712]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="25" pageNumber="48" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; forebody length 2.0 –
|
||
<quantity id="4CE6DB89FFD9FFEDBE08F9600FB502BD" box="[595,680,1688,1712]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="25" pageNumber="48" unit="mm" value="2.1">2.1 mm</quantity>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD9FFEDBC8FF94109290699" blockId="25.[189,800,1162,1975]" lastBlockId="25.[826,1437,241,1088]" pageId="25" pageNumber="48">
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||
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with posterior margin with two very short, lateral emarginations and barely recognizably concave middle (Fig. 134). Aedeagus (
|
||
<figureCitation id="13256AE9FFD9FFEDBD65F8C50C850358" box="[318,408,1853,1877]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
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||
,
|
||
<figureCitation id="13256AE9FFD9FFEDBDF2F8C50CE80358" box="[425,501,1853,1877]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92 – 94</figureCitation>
|
||
) about
|
||
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||
long with asymmetrical distal lobes; apical lobes curved ventrad with round ends (
|
||
<figureCitation id="13256AE9FFD9FFEDBDD6F8860CFB039B" box="[397,486,1918,1942]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDBDACF8860F09039B" box="[503,532,1918,1942]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92</figureCitation>
|
||
), in ventral and dorsal view stout with broad, subtruncate apices (
|
||
<figureCitation id="13256AE9FFD9FFEDBECDF8670E0E03BA" box="[662,787,1951,1975]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Figs 93, 94</figureCitation>
|
||
); dextral apical lobe with little, ventroproximal tooth (
|
||
<figureCitation id="13256AE9FFD9FFEDBF19FEEA0E8B0527" box="[834,918,274,298]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 92</figureCitation>
|
||
: arrow I), convexly widened dextrad distally (
|
||
<figureCitation id="13256AE9FFD9FFEDBF1AFECB0EC80546" box="[833,981,307,331]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Figs 93, 94</figureCitation>
|
||
); sinistral apical lobe evenly convex distolaterally, somewhat overlapping dextral apical lobe ventrally (
|
||
<figureCitation id="13256AE9FFD9FFEDBFBFFE8D09280580" box="[996,1077,373,397]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 93</figureCitation>
|
||
); dorsal lobe broad with convex apex (
|
||
<figureCitation id="13256AE9FFD9FFEDBFD8FE6E0EC705A3" box="[899,986,406,430]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 94</figureCitation>
|
||
); flagellum winding dextrad (
|
||
<figureCitation id="13256AE9FFD9FFEDB961FE6E089205A3" box="[1338,1423,406,430]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 93</figureCitation>
|
||
), strongly projecting from apical lobes ventrally (
|
||
<figureCitation id="13256AE9FFD9FFEDB91FFE4F088505C2" box="[1348,1432,439,463]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDBF61FE200E4505FD" box="[826,856,472,496]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92</figureCitation>
|
||
); lateral lobes reduced, marked by unequal groups of setae (
|
||
<figureCitation id="13256AE9FFD9FFEDBFFBFE000902061C" box="[928,1055,504,529]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Figs 93, 94</figureCitation>
|
||
); ventral lobe broad, subquadrate, with truncate apex (
|
||
<figureCitation id="13256AE9FFD9FFEDB848FDE1097B063F" box="[1043,1126,537,562]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDB82BFDE20990063F" box="[1136,1165,538,562]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92</figureCitation>
|
||
), in ventral view narrow, strongly shifted sinistrad (
|
||
<figureCitation id="13256AE9FFD9FFEDB838FDC209A8065F" box="[1123,1205,570,594]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 93</figureCitation>
|
||
); transverse ridge of median foramen concave (
|
||
<figureCitation id="13256AE9FFD9FFEDB839FDA309AE067E" box="[1122,1203,603,627]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">Fig. 93</figureCitation>
|
||
), strongly projecting ventrally (
|
||
<figureCitation id="13256AE9FFD9FFEDBFF0FD840EE20699" box="[939,1023,636,660]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDB850FD8409340699" box="[1035,1065,636,660]" captionStart="Figures 92–97" captionStartId="28.[151,224,1886,1908]" captionTargetBox="[223,1329,734,1834]" captionTargetId="figure-235@28.[151,1398,685,1867]" captionTargetPageId="28" captionText="Figures 92–97. Apex of aedeagus in lateral (92, 95), ventral (93, 96), dorsal view (94, 97) of Scopaeus cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (92–94); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (95– 97). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral tooth of dextral apical lobe (I), lateral ridges of median foramen (II, III)." figureDoi="http://doi.org/10.5281/zenodo.10724282" httpUri="https://zenodo.org/record/10724282/files/figure.png" pageId="25" pageNumber="48">92</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD9FFEDBF0AFD650920071A" blockId="25.[826,1437,241,1088]" pageId="25" pageNumber="48">
|
||
Female: Sperm pump with chamber abruptly narrowed at transition point towards bursal duct (
|
||
<figureCitation id="13256AE9FFD9FFEDB8B1FD46085806DB" box="[1258,1349,702,726]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="25" pageNumber="48">Fig. 168</figureCitation>
|
||
); bursal duct about three times as long as sperm pump; bursa membranous (
|
||
<figureCitation id="13256AE9FFD9FFEDBF8EFCF8092C071A" box="[981,1073,767,792]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="25" pageNumber="48">Fig. 167</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD9FFEDBF0AFCD908540754" blockId="25.[826,1437,241,1088]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBF0AFCD90ECD0734" bold="true" box="[849,976,801,825]" pageId="25" pageNumber="48">Phylogeny:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBF8DFCD909CD0735" authorityName="Frisch" authorityYear="2022" box="[982,1232,800,824]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="cuspilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDBF8DFCD909CD0735" box="[982,1232,800,824]" italics="true" pageId="25" pageNumber="48">Scopaeus cuspilobatus</emphasis>
|
||
</taxonomicName>
|
||
is a member of the
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBF61FCBA0EB70754" authorityName="Oke" authorityYear="1933" box="[826,938,833,857]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96AAA7EFFD9FFEDBF61FCBA0EB70754" box="[826,938,833,857]" italics="true" pageId="25" pageNumber="48">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="EF8F0B9DFFD9FFEDB808FCB908200754" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[1107,1341,833,857]" pageId="25" pageNumber="48" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD9FFEDBF0BFC9A084F0796" pageId="25" pageNumber="48" type="distribution">
|
||
<paragraph id="8BA1766CFFD9FFEDBF0BFC9A084F0796" blockId="25.[826,1437,241,1088]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBF0BFC9A0EF50777" bold="true" box="[848,1000,866,890]" pageId="25" pageNumber="48">Distribution:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDBFA5FC9B08150777" authorityName="Frisch" authorityYear="2022" box="[1022,1288,866,890]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="cuspilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDBFA5FC9B08150777" box="[1022,1288,866,890]" italics="true" pageId="25" pageNumber="48">Scopaeus cuspilobatus</emphasis>
|
||
</taxonomicName>
|
||
is hitherto known only from the
|
||
<typeStatus id="54A5C8CEFFD9FFEDB87CFC7C094A0791" box="[1063,1111,900,924]" pageId="25" pageNumber="48">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="49DAB88EFFD9FFEDB888FC7C08530796" box="[1235,1358,900,923]" country="Indonesia" name="Jawa Barat" pageId="25" pageNumber="48">Jawa Barat</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD9FFEDBF0BFC5C0861004D" pageId="25" pageNumber="48" type="etymology">
|
||
<paragraph id="8BA1766CFFD9FFEDBF0BFC5C0861004D" blockId="25.[826,1437,241,1088]" pageId="25" pageNumber="48">
|
||
<emphasis id="B96AAA7EFFD9FFEDBF0BFC5C0ECB07B1" bold="true" box="[848,982,932,956]" pageId="25" pageNumber="48">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFD9FFEDB802FC5C09FF07B1" authorityName="Frisch" authorityYear="2022" box="[1113,1250,932,956]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="48" phylum="Arthropoda" rank="species" species="cuspilobatus">
|
||
<emphasis id="B96AAA7EFFD9FFEDB802FC5C09FF07B1" box="[1113,1250,932,956]" italics="true" pageId="25" pageNumber="48">cuspilobatus</emphasis>
|
||
</taxonomicName>
|
||
[adjective, Latin, composed of the noun
|
||
<emphasis id="B96AAA7EFFD9FFEDB81BFC3E099907D0" box="[1088,1156,966,989]" italics="true" pageId="25" pageNumber="48">cuspis</emphasis>
|
||
(sting, thorn, spine) and
|
||
<emphasis id="B96AAA7EFFD9FFEDBF61FC1E0E9707F3" box="[826,906,998,1022]" italics="true" pageId="25" pageNumber="48">lobatus</emphasis>
|
||
(indicating a characteristic/feature of the lobe)] refers to the distinct, ventral spine of the dextral apical lobe of the aedeagus of this new species (
|
||
<figureCitation id="13256AE9FFD9FFEDB8ACFBDF0855004D" box="[1271,1352,1063,1088]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="25" pageNumber="48">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD9FFEDB90FFBD0086D004D" box="[1364,1392,1064,1088]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="25" pageNumber="48">83</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |