treatments-xml/data/03/B7/C7/03B7C77AFFCAFFFEBCF5FE4D0E6105FD.xml
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<document id="13D419E4E3DD610EE4F61F88ECD4C7CA" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFCAFFFEBCF5FE4D0E6105FD" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus halimunsalakensis Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="33" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="06D299182D82B2A3200D8F9A0641ACEE">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="6FA4EAF6EE46523A6CF0EDC5468D5B4C">Frisch, Johannes</mods:namePart>
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</mods:role>
<mods:namePart id="C70C9A21D2985B1317DF558F6F8642E7">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="466602559039D0CAC08E5EECE1FBBCAB">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="59C67A744D730F670675485189E0D098">Soil Organisms</mods:title>
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<mods:date id="64812F3B30397BC39DB7AF3B11A90EB3">2022</mods:date>
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<mods:number id="E9A4D49C423E19CF3DF62C7120FF724C">2022-04-01</mods:number>
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<mods:number id="7B57A840D550D96ACD88249FECE0279B">95</mods:number>
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<mods:number id="332F170457BF9F89432CC4224A703B71">1</mods:number>
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<mods:identifier id="63021F2B034533DCFF547EFBE4714025" type="DOI">10.25674/so95iss1id311</mods:identifier>
<mods:identifier id="6A25D41A1FD0B09ADC1960DDE99C6144" type="ISSN">2509-9523</mods:identifier>
<mods:identifier id="C230D6EF5AE649EED5832E17D69EF43B" type="Zenodo-Dep">10724236</mods:identifier>
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<treatment id="03B7C77AFFCAFFFEBCF5FE4D0E6105FD" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFFEBCF5FE4D0E6105FD" httpUri="http://treatment.plazi.org/id/03B7C77AFFCAFFFEBCF5FE4D0E6105FD" lastPageNumber="33" pageId="10" pageNumber="33">
<subSubSection id="C30425E7FFCAFFFEBCF5FE4D0FF505DD" box="[174,744,436,464]" pageId="10" pageNumber="33" type="nomenclature">
<paragraph id="8BA1766CFFCAFFFEBCF5FE4D0FF505DD" blockId="10.[174,744,436,497]" box="[174,744,436,464]" pageId="10" pageNumber="33">
<heading id="D0E9C100FFCAFFFEBCF5FE4D0FF505DD" box="[174,744,436,464]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="33" reason="5">
<taxonomicName id="4C1E0DEFFFCAFFFEBCF5FE4D0F7A05C2" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[174,615,436,463]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="halimunsalakensis" status="sp. nov.">
<emphasis id="B96AAA7EFFCAFFFEBCF5FE4D0F7A05C2" bold="true" box="[174,615,436,463]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBCF5FE4D0F1305C2" bold="true" box="[174,526,436,463]" italics="true" pageId="10" pageNumber="33">Scopaeus halimunsalakensis</emphasis>
Frisch
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2591705FFCAFFFEBE2EFE4E0FF505DD" box="[629,744,438,464]" pageId="10" pageNumber="33" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCAFFFEBCF5FE2E0CE305FC" box="[174,510,470,497]" pageId="10" pageNumber="33" type="description">
<paragraph id="8BA1766CFFCAFFFEBCF5FE2E0CE305FC" blockId="10.[174,744,436,497]" box="[174,510,470,497]" pageId="10" pageNumber="33">
(
<figureCitation id="13256AE9FFCAFFFEBCEDFE2E0C1D05FD" box="[182,256,470,496]" captionStart="Figures 17" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 17. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="10" pageNumber="33">Figs 7</figureCitation>
,
<figureCitation id="13256AE9FFCAFFFEBD57FE2E0C3605FD" box="[268,299,470,496]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="10" pageNumber="33">32</figureCitation>
,
<figureCitation id="13256AE9FFCAFFFEBD6CFE2E0C9A05FD" box="[311,391,470,497]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">65 67</figureCitation>
,
<figureCitation id="13256AE9FFCAFFFEBDC8FE2E0CDD05FD" box="[403,448,470,496]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="10" pageNumber="33">124</figureCitation>
,
<figureCitation id="13256AE9FFCAFFFEBD97FE2E0CEB05FC" box="[460,502,470,497]" captionStart="Figures 146154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146154. Sperm pump (146154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="10" pageNumber="33">152</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCAFFFEBCF5FDE20C7C0734" pageId="10" pageNumber="33" type="materials_examined">
<paragraph id="8BA1766CFFCAFFFEBCF5FDE20C7C0734" blockId="10.[151,764,537,1976]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBCF5FDE20C71063C" bold="true" box="[174,364,537,562]" pageId="10" pageNumber="33">Type specimens:</emphasis>
<materialsCitation id="3B767C31FFCAFFFEBD2EFDE10DFB0699" collectingDate="2015-09-21" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="10" pageNumber="33" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
<collectingCountry id="F30936FCFFCAFFFEBD2EFDE10CFC063C" box="[373,481,537,561]" name="Indonesia" pageId="10" pageNumber="33">Indonesia</collectingCountry>
,
<collectingRegion id="49DAB88EFFCAFFFEBDB5FDE20F77063C" box="[494,618,538,561]" country="Indonesia" name="Jawa Barat" pageId="10" pageNumber="33">Jawa Barat</collectingRegion>
:
<typeStatus id="54A5C8CEFFCAFFFEBE2DFDE10FC7063C" box="[630,730,537,561]" pageId="10" pageNumber="33" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFCAFFFEBEB8FDE20FEA063C" box="[739,759,538,561]" count="1" pageId="10" pageNumber="33" type="male"></specimenCount>
,
<collectingCounty id="62C00EE0FFCAFFFEBCCCFDC20C15065F" box="[151,264,570,594]" pageId="10" pageNumber="33">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFCAFFFEBD4AFDC20C66065F" box="[273,379,570,594]" pageId="10" pageNumber="33">Sirnarasa</collectingMunicipality>
:
<location id="8EC120B7FFCAFFFEBDD8FDC20F0D065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFFEBCF5FE4D0E6105FD:8EC120B7FFCAFFFEBDD8FDC20F0D065F" box="[387,528,570,594]" country="Indonesia" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Cimaja River" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Cimaja River</location>
(S-slope
<location id="8EC120B7FFCAFFFEBE2AFDC30FE8065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFFEBCF5FE4D0E6105FD:8EC120B7FFCAFFFEBE2AFDC30FE8065F" box="[625,757,570,594]" country="Indonesia" county="Sukabumi" latitude="-6.8588886" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" name="Mt Halimun" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Mt Halimun</location>
) (
<geoCoordinate id="EE2A10ABFFCAFFFEBCC4FDA30C00067E" box="[159,285,603,627]" degrees="06" direction="south" minutes="51" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="32" value="-6.8588886">06°5132S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFCAFFFEBD72FDA30CA8067E" box="[297,437,603,627]" degrees="106" direction="east" minutes="31" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="06" value="106.51833">106°3106E</geoCoordinate>
),
<quantity id="4CE6DB89FFCAFFFEBD92FDA30F13067E" box="[457,526,603,627]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="10" pageNumber="33" unit="m" value="670.0">
<elevation id="0033915FFFCAFFFEBD92FDA30F13067E" box="[457,526,603,627]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="10" pageNumber="33" unit="m" value="670.0">670 m</elevation>
</quantity>
,
<date id="FFA050ACFFCAFFFEBE41FDA30F62067E" box="[538,639,603,627]" pageId="10" pageNumber="33" value="2015-09-21">
<collectingDate id="EFE4A944FFCAFFFEBE41FDA30F62067E" box="[538,639,603,627]" pageId="10" pageNumber="33" value="2015-09-21">21.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCAFFFEBEE3FDA30FE6067E" box="[696,763,603,627]" pageId="10" pageNumber="33">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCAFFFEBCC4FD840DC30699" box="[159,222,636,660]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54A5C8CEFFCAFFFEBCAFFD850C420698" box="[244,351,637,661]" pageId="10" pageNumber="33" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D18BDE5FFCAFFFEBD29FD840CE20699" box="[370,511,636,660]" count="7" pageId="10" pageNumber="33" type="generic">7 specimens</specimenCount>
):
<materialsCitation id="3B767C31FFCAFFFEBE4FFD840CD206B8" collectingDate="2015-09-21" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="670" latitude="-6.8588886" location="Cimaja River" longLatPrecision="21" longitude="106.51833" municipality="Sirnarasa" pageId="10" pageNumber="33" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCAFFFEBE4FFD840F5D0699" box="[532,576,636,660]" count="1" pageId="10" pageNumber="33" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFCAFFFEBE14FD840F660699" box="[591,635,636,660]" count="4" pageId="10" pageNumber="33" type="female">4 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED0FEEA9FFCAFFFEBD72FD650C6606B8" box="[297,379,669,693]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="33">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFCAFFFEBDD2FD650CDA06B8" box="[393,455,669,693]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFCAFFFEBD86FD650C5A06FA" collectingDate="2015-09-13" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" elevation="640" latitude="-6.665278" location="Mt Salak" longLatPrecision="21" longitude="106.76" municipality="Bogor" pageId="10" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCAFFFEBD86FD650F1A06B9" box="[477,519,669,693]" count="1" pageId="10" pageNumber="33" type="female">1 ♀</specimenCount>
,
<collectingMunicipality id="6BC5EC16FFCAFFFEBE43FD650F4106B8" box="[536,604,669,693]" pageId="10" pageNumber="33">Bogor</collectingMunicipality>
, NO-slope
<location id="8EC120B7FFCAFFFEBE86FD650DCE06DB" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFFEBCF5FE4D0E6105FD:8EC120B7FFCAFFFEBE86FD650DCE06DB" country="Indonesia" latitude="-6.665278" longLatPrecision="21" longitude="106.76" municipality="Bogor" name="Mt Salak" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Mt Salak</location>
(
<geoCoordinate id="EE2A10ABFFCAFFFEBCB8FD460C7F06DB" box="[227,354,702,726]" degrees="06" direction="south" minutes="39" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="55" value="-6.665278">06°3955S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFCAFFFEBD35FD460CE606DB" box="[366,507,702,726]" degrees="106" direction="east" minutes="45" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="36" value="106.76">106°4536E</geoCoordinate>
),
<quantity id="4CE6DB89FFCAFFFEBE54FD460F4B06D8" box="[527,598,702,726]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="33" unit="m" value="640.0">
<elevation id="0033915FFFCAFFFEBE54FD460F4B06D8" box="[527,598,702,726]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="33" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="FFA050ACFFCAFFFEBE38FD460FD406DB" box="[611,713,702,726]" pageId="10" pageNumber="33" value="2015-09-13">
<collectingDate id="EFE4A944FFCAFFFEBE38FD460FD406DB" box="[611,713,702,726]" pageId="10" pageNumber="33" value="2015-09-13">13.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCAFFFEBCCCFD270DC706FA" box="[151,218,735,759]" pageId="10" pageNumber="33">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCAFFFEBCB6FD270C2206FA" box="[237,319,735,759]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="33">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFCAFFFEBD0DFD270C400734" collectingDate="2015-09-22" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="1000" latitude="-6.817222" location="Cisareno River" longLatPrecision="21" longitude="106.5025" municipality="Sirnaresmi" pageId="10" pageNumber="33" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCAFFFEBD0DFD270C9C06FB" box="[342,385,735,759]" count="1" pageId="10" pageNumber="33" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62C00EE0FFCAFFFEBDC8FD270F1906FA" box="[403,516,735,759]" pageId="10" pageNumber="33">Sukabumi</collectingCounty>
,
<collectingMunicipality id="6BC5EC16FFCAFFFEBE48FD270F9306FA" box="[531,654,735,759]" pageId="10" pageNumber="33">Sirnaresmi</collectingMunicipality>
:
<location id="8EC120B7FFCAFFFEBEC7FD270DCE0715" LSID="urn:lsid:plazi:treatment:03B7C77AFFCAFFFEBCF5FE4D0E6105FD:8EC120B7FFCAFFFEBEC7FD270DCE0715" country="Indonesia" county="Sukabumi" latitude="-6.817222" longLatPrecision="21" longitude="106.5025" municipality="Sirnaresmi" name="Cisareno River" pageId="10" pageNumber="33" stateProvince="Jawa Barat">Cisareno River</location>
(
<geoCoordinate id="EE2A10ABFFCAFFFEBCB3FD070C75071A" box="[232,360,767,791]" degrees="06" direction="south" minutes="49" orientation="latitude" pageId="10" pageNumber="33" precision="15" seconds="02" value="-6.817222">06°4902S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFCAFFFEBD21FD070F17071A" box="[378,522,767,791]" degrees="106" direction="east" minutes="30" orientation="longitude" pageId="10" pageNumber="33" precision="15" seconds="09" value="106.5025">106°3009E</geoCoordinate>
),
<quantity id="4CE6DB89FFCAFFFEBE7FFCF80F60071A" box="[548,637,768,792]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="33" unit="m" value="1000.0">
<elevation id="0033915FFFCAFFFEBE7FFCF80F60071A" box="[548,637,768,792]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="33" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFA050ACFFCAFFFEBED4FD070FE5071A" box="[655,760,767,791]" pageId="10" pageNumber="33" value="2015-09-22">
<collectingDate id="EFE4A944FFCAFFFEBED4FD070FE5071A" box="[655,760,767,791]" pageId="10" pageNumber="33" value="2015-09-22">22.9.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCAFFFEBC9EFCD80C150735" box="[197,264,800,824]" pageId="10" pageNumber="33">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCAFFFEBD4DFCD90C480734" box="[278,341,801,825]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="33" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCAFFFEBCF5FCB90FE103BA" pageId="10" pageNumber="33" type="description">
<paragraph id="8BA1766CFFCAFFFEBCF5FCB90CCB00CE" blockId="10.[151,764,537,1976]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBCF5FCB90C260754" bold="true" box="[174,315,833,857]" pageId="10" pageNumber="33">Description:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFCAFFFEBE29FCB90FAB0754" box="[626,694,833,857]" captionStart="Figures 17" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 17. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="10" pageNumber="33">Fig. 7</figureCitation>
. Head subquadrate. Antennae with penultimate segment quadrate. Mesotibia slender. Forebody surface matt due to dense, rugulose, setose punctation; abdomen somewhat shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown except for yellow-brown posterior sixth of elytra and tip of abdomen; maxillary palpi and antennae medium brown, latter gradually lighter towards light brown, distal segments; legs yellow-brown with femora dark brown except for distal ends. Total body length
<quantity id="4CE6DB89FFCAFFFEBE36FB720FE500AF" box="[621,760,1162,1186]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="10" pageNumber="33" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4 2.9 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFCAFFFEBD1CFB530CCF00CE" box="[327,466,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="10" pageNumber="33" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5 1.6 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFCAFFFEBCF5FB340FB90339" blockId="10.[151,764,537,1976]" pageId="10" pageNumber="33">
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII with posterior margin bisinuate with strongly convex median third; emargination covering about distal fifth of sternite length (
<figureCitation id="13256AE9FFCAFFFEBEC9FAD60FEC014A" box="[658,753,1326,1351]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="10" pageNumber="33">Fig. 124</figureCitation>
). Aedeagus (
<figureCitation id="13256AE9FFCAFFFEBD4FFAB70C760165" box="[276,363,1359,1384]" captionStart="Figures 2734" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 2734. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="10" pageNumber="33">Figs 32</figureCitation>
,
<figureCitation id="13256AE9FFCAFFFEBD20FAB70CDE016A" box="[379,451,1359,1383]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">65 67</figureCitation>
) about
<quantity id="4CE6DB89FFCAFFFEBE45FAA80F72016A" box="[542,623,1360,1384]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="33" unit="mm" value="0.8">0.8 mm</quantity>
long; apical lobes slender, in ventral and dorsal view subparallel with mediad curved ends (
<figureCitation id="13256AE9FFCAFFFEBD2CFA690CF301A4" box="[375,494,1425,1449]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Figs 66, 67</figureCitation>
); dorsal lobe distally with truncate, finely serrate ventral margin; flagellum long, strongly projecting from apical lobes apically; apical lobes, dorsal lobe and flagellum strongly curved ventrally in about longitudinal middle (
<figureCitation id="13256AE9FFCAFFFEBD23F9ED0CD80220" box="[376,453,1557,1581]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Fig. 65</figureCitation>
: arrow I); lateral lobes short, subacute, little projecting from phallobase, each bearing apical group of apicad pointing setae and widely separate, proximal group of ventromediad pointing setae (
<figureCitation id="13256AE9FFCAFFFEBEFFF98F0DA902BD" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Figs 66, 67</figureCitation>
); ventral lobe in lateral view elongate with straight ventral margin and round apex (
<figureCitation id="13256AE9FFCAFFFEBDAEF9410F5E02DC" box="[501,579,1721,1745]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Fig. 65</figureCitation>
), in ventral view bilobate with deep, narrow, median emargination (
<figureCitation id="13256AE9FFCAFFFEBEF0F9220FE502FF" box="[683,760,1754,1778]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Fig. 66</figureCitation>
: arrow
<collectionCode id="ED0FEEA9FFCAFFFEBC80F9030DED031E" box="[219,240,1787,1811]" pageId="10" pageNumber="33">II</collectionCode>
); median foramen large, round distally; transverse ridge weakly sclerotized, inconspicuous (
<figureCitation id="13256AE9FFCAFFFEBE11F8E40F850339" box="[586,664,1820,1844]" captionStart="Figures 6267" captionStartId="22.[151,224,1885,1907]" captionTargetBox="[220,1321,396,1832]" captionTargetId="figure-119@22.[151,1398,322,1867]" captionTargetPageId="22" captionText="Figures 6267. Apex of aedeagus in lateral (62, 65), ventral (63, 66), dorsal view (64, 67) of Scopaeus mixtus, holotype, Malaysia, Sabah: Sandakan (6264); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (6567). Abbreviations: al apical lobes, dl dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, s groups of setae, vl ventral lobe. Arrows: dorsal emargination of apical lobes, dorsal lobe and flagellum (I); apical emargination of dorsal lobe (II)." figureDoi="http://doi.org/10.5281/zenodo.10724270" httpUri="https://zenodo.org/record/10724270/files/figure.png" pageId="10" pageNumber="33">Fig. 66</figureCitation>
).
</paragraph>
<paragraph id="8BA1766CFFCAFFFEBCF5F8C50FFC0378" blockId="10.[151,764,537,1976]" pageId="10" pageNumber="33">
Female: Sperm pump with strongly widened ends and short, circular chamber (
<figureCitation id="13256AE9FFCAFFFEBDFBF8A50CE60378" box="[416,507,1885,1909]" captionStart="Figures 146154" captionStartId="40.[151,224,1862,1884]" captionTargetBox="[200,1323,249,1795]" captionTargetId="figure-141@40.[151,1398,229,1843]" captionTargetPageId="40" captionText="Figures 146154. Sperm pump (146154) and bursa (148, 149) of ♀ of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (146); S. batukaruensis, paratype, Bali, Tabanan:Batukaru (147); S. jacobsoni, Jawa Barat, Sukabumi:Cidahu (148, with lateral gonocoxal plates IX; 150); S. mixtus, Malaysia, Sabah: Lokan (149, with lateral gonocoxal plates IX; 151); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (152); S. tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (153, with bursal duct and bursa; 154). Long scale bar: 146, 147, 150152, 154; short scale bar: 148, 149, 153." figureDoi="http://doi.org/10.5281/zenodo.10724309" httpUri="https://zenodo.org/record/10724309/files/figure.png" pageId="10" pageNumber="33">Fig. 152</figureCitation>
); bursa membranous.
</paragraph>
<paragraph id="8BA1766CFFCAFFFEBCF5F8860FE103BA" blockId="10.[151,764,537,1976]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBCF5F8860C37039B" bold="true" box="[174,298,1918,1942]" pageId="10" pageNumber="33">Phylogeny:</emphasis>
<taxonomicName id="4C1E0DEFFFCAFFFEBD1AF8870F65039B" authorityName="Frisch" authorityYear="2022" box="[321,632,1918,1942]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="halimunsalakensis">
<emphasis id="B96AAA7EFFCAFFFEBD1AF8870F65039B" box="[321,632,1918,1942]" italics="true" pageId="10" pageNumber="33">Scopaeus halimunsalakensis</emphasis>
</taxonomicName>
is to be classified in the
<taxonomicName id="4C1E0DEFFFCAFFFEBD60F8580CB003BA" authorityName="Kraatz" authorityYear="1859" box="[315,429,1951,1975]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B96AAA7EFFCAFFFEBD60F8580CB003BA" box="[315,429,1951,1975]" italics="true" pageId="10" pageNumber="33">S. limbatus</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF8F0B9DFFCAFFFEBE10F8670FED03BA" author="Frisch, J." box="[587,752,1951,1975]" pageId="10" pageNumber="33" pagination="73 - 96" refId="ref31539" refString="Frisch, J. (2005): &quot; Scopaeus limbatus Kraatz &quot; of ancient authors - a monophyletic species group distributed in the Mascarene Islands, the South-East Palaearctic, the Oriental and the Australian regions (Coleoptera, Staphylinidae, Paederinae). - Mitteilungen aus dem Museum fur Naturkunde Berlin, Deutsche Entomologische Zeitschrift 52: 73 - 96." type="journal article" year="2005">Frisch 2005: 75</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCAFFFEBF72FF0909690546" pageId="10" pageNumber="33" type="distribution">
<paragraph id="8BA1766CFFCAFFFEBF72FF0909690546" blockId="10.[786,1399,241,496]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBF72FF090EA20504" bold="true" box="[809,959,241,265]" pageId="10" pageNumber="33">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFCAFFFEBF9CFF0A09E30504" authorityName="Frisch" authorityYear="2022" box="[967,1278,241,265]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="halimunsalakensis">
<emphasis id="B96AAA7EFFCAFFFEBF9CFF0A09E30504" box="[967,1278,241,265]" italics="true" pageId="10" pageNumber="33">Scopaeus halimunsalakensis</emphasis>
</taxonomicName>
is hitherto known only from the surroundings of Mount Halimun-Salak National Park,
<collectingRegion id="49DAB88EFFCAFFFEBFACFECC096D0546" box="[1015,1136,308,331]" country="Indonesia" name="Jawa Barat" pageId="10" pageNumber="33">Jawa Barat</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCAFFFEBF72FEAC0E6105FD" pageId="10" pageNumber="33" type="etymology">
<paragraph id="8BA1766CFFCAFFFEBF72FEAC0E6105FD" blockId="10.[786,1399,241,496]" pageId="10" pageNumber="33">
<emphasis id="B96AAA7EFFCAFFFEBF72FEAC0EAD0561" bold="true" box="[809,944,340,364]" pageId="10" pageNumber="33">Etymology:</emphasis>
The epithet
<taxonomicName id="4C1E0DEFFFCAFFFEB86DFEAC081D0561" authorityName="Frisch" authorityYear="2022" box="[1078,1280,340,364]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="halimunsalakensis">
<emphasis id="B96AAA7EFFCAFFFEB86DFEAC081D0561" box="[1078,1280,340,364]" italics="true" pageId="10" pageNumber="33">halimunsalakensis</emphasis>
</taxonomicName>
(adjective, Latin,withsuffix
<emphasis id="B96AAA7EFFCAFFFEBF87FE8E090E0580" box="[988,1043,374,397]" italics="true" pageId="10" pageNumber="33">ensis</emphasis>
, derived from the noun Halimun-Salak) refers to the Mount Halimun-Salak National Park, in the vicinity of which the
<typeStatus id="54A5C8CEFFCAFFFEB80AFE4F099C05C2" box="[1105,1153,439,463]" pageId="10" pageNumber="33">type</typeStatus>
specimens have been collected.
</paragraph>
</subSubSection>
</treatment>
</document>