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<document id="23A09A823B918FC2E75C5C2E1A0DCE6E" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FFC2760DFC2FD2C049C2F226" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax tingitanus" docType="treatment" docVersion="8" lastPageNumber="449" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="448" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="3E387D39CDB36E81AA493844E91EC99D">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<treatment id="03B787C7FFC2760DFC2FD2C049C2F226" ID-DOI="http://doi.org/10.5281/zenodo.6967824" ID-GBIF-Taxon="198618715" ID-Zenodo-Dep="6967824" LSID="urn:lsid:plazi:treatment:03B787C7FFC2760DFC2FD2C049C2F226" httpUri="http://treatment.plazi.org/id/03B787C7FFC2760DFC2FD2C049C2F226" lastPageId="32" lastPageNumber="449" pageId="31" pageNumber="448">
<subSubSection id="C304655AFFC27632FC2FD2C04F6EF3AA" box="[900,1370,1687,1712]" pageId="31" pageNumber="448" type="nomenclature">
<paragraph id="8BA136D1FFC27632FC2FD2C04F6EF3AA" blockId="31.[900,1370,1687,1712]" box="[900,1370,1687,1712]" pageId="31" pageNumber="448">
<heading id="D0E981BDFFC27632FC2FD2C04F6EF3AA" box="[900,1370,1687,1712]" centered="true" fontSize="9" level="2" pageId="31" pageNumber="448" reason="2">
<taxonomicName id="4C1E4D52FFC27632FC2FD2C04F6EF3AA" ID-CoL="48WVN" authority="(SIMON, 1884)" baseAuthorityName="Simon" baseAuthorityYear="1884" box="[900,1370,1687,1712]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="448" phylum="Arthropoda" rank="species" species="tingitanus">
<emphasis id="B96AEAC3FFC27632FC2FD2C04E84F3B5" box="[900,1200,1688,1711]" italics="true" pageId="31" pageNumber="448">OEDOTHORAX TINGITANUS</emphasis>
(
<bibRefCitation id="EF8F4B20FFC27632FB6BD2CF4F66F3AA" author="Simon E" box="[1216,1362,1687,1712]" pageId="31" pageNumber="448" pagination="180 - 885" refId="ref95278" refString="Simon E. 1884. Les arachnides de France. Tome cinquieme, deuxieme et troisieme partie. Roret Paris, 180 - 885." type="book chapter" year="1884">SIMON, 1884</bibRefCitation>
)
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<subSubSection id="C304655AFFC27632FC94D2E64EB3F3EC" pageId="31" pageNumber="448" type="description">
<paragraph id="8BA136D1FFC27632FC94D2E64EB3F3EC" blockId="31.[831,1438,1726,1782]" pageId="31" pageNumber="448">
(
<figureCitation id="13252A54FFC27632FCECD2E64990F3CD" box="[839,932,1726,1751]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="31" pageNumber="448">FIGS 7X</figureCitation>
,
<figureCitation id="13252A54FFC27632FC1AD2E649F8F3CD" box="[945,972,1726,1751]" captionStart="Figure 8" captionStartId="14.[146,225,1007,1029]" captionTargetBox="[147,1428,197,966]" captionTargetId="figure-341@14.[145,1428,195,967]" captionTargetPageId="14" captionText="Figure 8. Male chelicerae, stridulatory striae on lateral side. A, Oedothorax gibbosus (Blackwall, 1841). B, Oe. trilobatus (Banks, 1896). C, Oe. gibbifer (Kulczyński, 1882). D, Oe. apicatus (Blackwall, 1850). E, Oe. retusus (Westring, 1851). F, Oe. paludigena Simon, 1926. G, Oe. meridionalis Tanasevitch, 1987. H, Oe. agrestis (Blackwall, 1853). I, Oe. fuscus (Blackwall, 1834). J, Oe. tingitanus (Simon, 1884). Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967804" httpUri="https://zenodo.org/record/6967804/files/figure.png" pageId="31" pageNumber="448">8J</figureCitation>
,
<figureCitation id="13252A54FFC27632FC72D2E649C7F3CD" box="[985,1011,1726,1751]" captionStart="Figure 9" captionStartId="15.[163,241,931,953]" captionTargetBox="[164,1444,196,890]" captionTargetId="figure-419@15.[163,1446,195,892]" captionTargetPageId="15" captionText="Figure 9. Male opisthosoma, dorsal view. A, Oedothorax gibbosus (Blackwall, 1841). B, Oe. trilobatus (Banks, 1896). C, Oe. gibbifer (Kulczyński, 1882). D, Oe. apicatus (Blackwall, 1850). E, Oe. retusus (Westring, 1851). F, Oe. paludigena Simon, 1926. G, Oe. meridionalis Tanasevitch, 1987. H, Oe. agrestis (Blackwall, 1853). I, Oe. fuscus (Blackwall, 1834). J, Oe. tingitanus (Simon, 1884). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967808" httpUri="https://zenodo.org/record/6967808/files/figure.png" pageId="31" pageNumber="448">9J</figureCitation>
,
<figureCitation id="13252A54FFC27632FBABD2E64E28F3CD" box="[1024,1052,1726,1751]" captionStart="Figure 1" captionStartId="6.[145,222,1872,1894]" captionTargetBox="[209,1361,198,1830]" captionTargetId="figure-25@6.[206,1364,195,1833]" captionTargetPageId="6" captionText="Figure 1. The consensus tree of 296 most-parsimonious trees from the analysis of the expanded matrix of Miller &amp; Hormiga (2004) and Frick et al.(2010) (Matrix I), with Bremer/Jackknife support values shown beside nodes. Newly appended taxa in bold." figureDoi="http://doi.org/10.5281/zenodo.6967780" httpUri="https://zenodo.org/record/6967780/files/figure.png" pageId="31" pageNumber="448">11</figureCitation>
; SUPPORTING INFORMATION,
<figureCitation id="13252A54FFC27632FAD8D29A4E4AF3EC" captionStart="Figure 1" captionStartId="6.[145,222,1872,1894]" captionTargetBox="[209,1361,198,1830]" captionTargetId="figure-25@6.[206,1364,195,1833]" captionTargetPageId="6" captionText="Figure 1. The consensus tree of 296 most-parsimonious trees from the analysis of the expanded matrix of Miller &amp; Hormiga (2004) and Frick et al.(2010) (Matrix I), with Bremer/Jackknife support values shown beside nodes. Newly appended taxa in bold." figureDoi="http://doi.org/10.5281/zenodo.6967780" httpUri="https://zenodo.org/record/6967780/files/figure.png" pageId="31" pageNumber="448">FIG. S1I</figureCitation>
)
</paragraph>
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<paragraph id="8BA136D1FFC27632FC90D35D4F31F220" blockId="31.[827,1384,1797,1880]" pageId="31" pageNumber="448">
<treatmentCitationGroup id="AB0E11FFFFC27632FC90D35D4F31F220" pageId="31" pageNumber="448">
<treatmentCitation id="0ABF10C0FFC27632FC90D35D4F50F201" author="Simon E" box="[827,1380,1797,1819]" page="483" pageId="31" pageNumber="448" year="1884">
<taxonomicName id="4C1E4D52FFC27632FC90D35D4F50F201" authority="Simon, 1884: 483 (Dm)" authorityName="Simon" authorityYear="1884" box="[827,1380,1797,1819]" class="Arachnida" family="Linyphiidae" genus="Gongylidium" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="448" phylum="Arthropoda" rank="species" species="tingitanum">
<emphasis id="B96AEAC3FFC27632FC90D35D4E62F201" box="[827,1110,1797,1819]" italics="true" pageId="31" pageNumber="448">Gongylidium tingitanum</emphasis>
<bibRefCitation id="EF8F4B20FFC27632FBF5D35D4F14F200" author="Simon E" box="[1118,1312,1797,1819]" pageId="31" pageNumber="448" pagination="180 - 885" refId="ref95278" refString="Simon E. 1884. Les arachnides de France. Tome cinquieme, deuxieme et troisieme partie. Roret Paris, 180 - 885." type="book chapter" year="1884">Simon, 1884: 483</bibRefCitation>
(Dm)
</taxonomicName>
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.
<treatmentCitation id="0ABF10C0FFC27632FC90D37C4F36F220" author="Simon E" box="[827,1282,1828,1850]" page="633" pageId="31" pageNumber="448" year="1894">
<taxonomicName id="4C1E4D52FFC27632FC90D37C4F36F220" authority="Simon, 1894: 633 (Dm)" authorityName="Simon" authorityYear="1894" box="[827,1282,1828,1850]" class="Arachnida" family="Linyphiidae" genus="Neriene" kingdom="Animalia" order="Araneae" pageId="31" pageNumber="448" phylum="Arthropoda" rank="species" species="insolens">
<emphasis id="B96AEAC3FFC27632FC90D37C49C1F223" box="[827,1013,1828,1849]" italics="true" pageId="31" pageNumber="448">Neriene insolens</emphasis>
<bibRefCitation id="EF8F4B20FFC27632FC50D37C4E89F220" author="Simon E" box="[1019,1213,1828,1850]" pageId="31" pageNumber="448" pagination="489 - 760" refId="ref95303" refString="Simon E. 1894. Histoire naturelle des araignees, Vol. 1. Paris: Roret, 489 - 760." type="book chapter" year="1894">Simon, 1894: 633</bibRefCitation>
(Dm)
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BA136D1FFC27632FC90D31A4F25F242" blockId="31.[827,1384,1797,1880]" box="[827,1297,1858,1880]" pageId="31" pageNumber="448">
<emphasis id="B96AEAC3FFC27632FC90D31A4E1AF242" box="[827,1070,1858,1880]" italics="true" pageId="31" pageNumber="448">Stylothorax tingitana</emphasis>
<bibRefCitation id="EF8F4B20FFC27632FB9ED31A4F39F242" author="Reimoser E" box="[1077,1293,1858,1880]" pageId="31" pageNumber="448" pagination="1 - 280" refId="ref94586" refString="Reimoser E. 1919. Katolog der echten Spinnen (Araneae) des Palaarktischen Gebietes. Abhandlungen der Zoologisch- Botanischen Gesellschaft in Wien 10: 1 - 280." type="journal article" year="1919">Reimoser, 1919: 73</bibRefCitation>
.
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD49D4AF3F5E0" blockId="32.[145,760,197,403]" pageId="32" pageNumber="449">
<treatmentCitationGroup id="AB0E11FFFFFD760DFF3AD49D4AF3F5E0" pageId="32" pageNumber="449">
<treatmentCitation id="0ABF10C0FFFD760DFF3AD49D485BF5C1" author="Denis J" box="[145,623,197,219]" page="151" pageId="32" pageNumber="449" year="1968">
<taxonomicName id="4C1E4D52FFFD760DFF3AD49D485BF5C1" ID-CoL="48WVN" authority="Denis, 1968: 151" authorityName="Denis" authorityPageNumber="151" authorityYear="1968" box="[145,623,197,219]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="tingitanus">
<emphasis id="B96AEAC3FFFD760DFF3AD49D4BAAF5C1" box="[145,414,197,219]" italics="true" pageId="32" pageNumber="449">Oedothorax tingitanus</emphasis>
<bibRefCitation id="EF8F4B20FFFD760DFE0CD49D485BF5C1" author="Denis J" box="[423,623,197,219]" pageId="32" pageNumber="449" pagination="137 - 164" refId="ref90547" refString="Denis J. 1968. Notes d'araneologie marocaine. X, Les erigonides du Maroc. Bulletin de la Societe des Sciences Naturelles du Maroc 47: 137 - 164." type="journal article" year="1968">Denis, 1968: 151</bibRefCitation>
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, figs 1115 (mf).
</treatmentCitationGroup>
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<paragraph id="8BA136D1FFFD760DFF3AD55A4ADDF42D" blockId="32.[145,760,197,403]" pageId="32" pageNumber="449">
<treatmentCitationGroup id="AB0E11FFFFFD760DFF3AD55A4ADDF42D" pageId="32" pageNumber="449">
<treatmentCitation id="0ABF10C0FFFD760DFF3AD55A485DF402" author="Bosmans R" box="[145,617,258,280]" page="59" pageId="32" pageNumber="449" year="1985">
<taxonomicName id="4C1E4D52FFFD760DFF3AD55A485DF402" ID-CoL="48WVN" authority="Bosmans, 1985: 59" authorityName="Bosmans" authorityPageNumber="59" authorityYear="1985" box="[145,617,258,280]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="tingitanus">
<emphasis id="B96AEAC3FFFD760DFF3AD55A4BBBF402" box="[145,399,258,280]" italics="true" pageId="32" pageNumber="449">Oedothorax tingitanus</emphasis>
<bibRefCitation id="EF8F4B20FFFD760DFE3DD55B485DF402" author="Bosmans R" box="[406,617,258,280]" pageId="32" pageNumber="449" pagination="58 - 75" refId="ref89322" refString="Bosmans R. 1985. Etudes sur les Linyphiidae nord-africains. II. Le genre Oedothorax Bertkau en Africa du Nord, avec une revision des caracteres diagnostiques des males des especes ouest-palearctiques. Biologisch Jaarboek Dodonaea 53: 58 - 75." type="journal article" year="1985">Bosmans, 1985: 59</bibRefCitation>
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, figs 13, 22, 36 (mf).
</treatmentCitationGroup>
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<paragraph id="8BA136D1FFFD760DFF3AD518489CF489" blockId="32.[145,760,197,403]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD5184B76F44F" box="[145,322,320,341]" italics="true" pageId="32" pageNumber="449">Type material:</emphasis>
<materialsCitation id="3B763C8CFFFD760DFEE4D51B489CF489" ID-GBIF-Occurrence="3880945329" collectionCode="MNHN" collectorName="Simon" country="Morocco" location="Morocco" pageId="32" pageNumber="449" specimenCode="MNHN 4881" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="lectotype">
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<typeStatus id="54A58873FFFD760DFEC1D5184BD4F44F" box="[362,480,320,341]" pageId="32" pageNumber="449" type="lectotype">Lectotype</typeStatus>
and
<specimenCount id="9D18FD58FFFD760DFD8BD518480AF44F" box="[544,574,320,341]" pageId="32" pageNumber="449" type="female">2♀</specimenCount>
<typeStatus id="54A58873FFFD760DFDECD51848CCF44C" box="[583,760,320,342]" pageId="32" pageNumber="449" type="paralectotype">paralectotypes</typeStatus>
based on
<bibRefCitation id="EF8F4B20FFFD760DFF54D5074B8DF46E" author="Bosmans R" box="[255,441,350,372]" pageId="32" pageNumber="449" pagination="58 - 75" refId="ref89322" refString="Bosmans R. 1985. Etudes sur les Linyphiidae nord-africains. II. Le genre Oedothorax Bertkau en Africa du Nord, avec une revision des caracteres diagnostiques des males des especes ouest-palearctiques. Biologisch Jaarboek Dodonaea 53: 58 - 75." type="journal article" year="1985">Bosmans (1985)</bibRefCitation>
:
<emphasis id="B96AEAC3FFFD760DFE6CD507480AF46E" bold="true" box="[455,574,351,372]" pageId="32" pageNumber="449">
<collectingCountry id="F3097641FFFD760DFE6CD507480CF46E" box="[455,568,351,372]" name="Morocco" pageId="32" pageNumber="449">Morocco</collectingCountry>
:
</emphasis>
close to Tanger, 1868, coll.
<collectorName id="26EB5307FFFD760DFEA3D5254B65F489" box="[264,337,381,403]" pageId="32" pageNumber="449">Simon</collectorName>
(
<specimenCode id="DBB89EAAFFFD760DFEF4D5254BC0F488" box="[351,500,381,402]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="32" pageNumber="449">MNHN 4881</specimenCode>
, not examined).
</materialsCitation>
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD5E448CEF757" blockId="32.[145,762,444,589]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD5E44B43F4CB" box="[145,375,444,465]" italics="true" pageId="32" pageNumber="449">Examined material:</emphasis>
<materialsCitation id="3B763C8CFFFD760DFE2FD5E448C3F757" ID-GBIF-Occurrence="3880945386" collectingDate="1977-05-16" collectingDateMax="1977-05-17" collectingDateMin="1977-05-16" collectionCode="NHM" collectorName="P. Hillyard &amp; P. D. Hillyard" country="Morocco" determinerName="A. F. Millidge" location="Mediterranean" pageId="32" pageNumber="449" specimenCount="33" specimenCount-female="9" specimenCount-male="24">
<emphasis id="B96AEAC3FFFD760DFE2FD5E44BCEF4CB" bold="true" box="[388,506,444,465]" pageId="32" pageNumber="449">
<collectingCountry id="F3097641FFFD760DFE2FD5E44BC1F4CB" box="[388,501,444,465]" name="Morocco" pageId="32" pageNumber="449">Morocco</collectingCountry>
:
</emphasis>
<location id="8EC1600AFFFD760DFDA9D5E44884F4C8" LSID="urn:lsid:plazi:treatment:03B787C7FFC2760DFC2FD2C049C2F226:8EC1600AFFFD760DFDA9D5E44884F4C8" box="[514,688,444,466]" country="Morocco" name="Mediterranean" pageId="32" pageNumber="449">Mediterranean</location>
coast, salt marsh, west of Mdiq (Tekouan),
<specimenCount id="9D18FD58FFFD760DFDE9D5834854F4F5" box="[578,608,475,496]" pageId="32" pageNumber="449" type="male">8♂</specimenCount>
<specimenCount id="9D18FD58FFFD760DFDCAD583484AF4F5" box="[609,638,475,496]" pageId="32" pageNumber="449" type="female">8♀</specimenCount>
<date id="FFA01011FFFD760DFD2DD58348C0F4EA" box="[646,756,475,496]" pageId="32" pageNumber="449" value="1977-05-17">
<collectingDate id="EFE4E9F9FFFD760DFD2DD58348C0F4EA" box="[646,756,475,496]" pageId="32" pageNumber="449" value="1977-05-17">17.v.1977</collectingDate>
</date>
, coll.
<collectorName id="26EB5307FFFD760DFF61D5A24B7BF715" box="[202,335,505,527]" pageId="32" pageNumber="449">P. Hillyard</collectorName>
, det.
<determinerName id="7E7DDA15FFFD760DFE39D5A14801F715" box="[402,565,505,527]" pageId="32" pageNumber="449">A. F. Millidge</determinerName>
(
<collectionCode id="ED0FAE14FFFD760DFDECD5A148B9F715" box="[583,653,505,527]" pageId="32" pageNumber="449">NHM</collectionCode>
); Merdia herga, west coast, salt marsh vegetation,
<specimenCount id="9D18FD58FFFD760DFD00D64048EEF737" box="[683,730,536,558]" pageId="32" pageNumber="449" type="male">16♂</specimenCount>
<specimenCount id="9D18FD58FFFD760DFD70D64048CDF737" box="[731,761,536,557]" pageId="32" pageNumber="449" type="female">1♀</specimenCount>
<date id="FFA01011FFFD760DFF3AD66E4AC9F756" box="[145,253,566,588]" pageId="32" pageNumber="449" value="1977-05-16">
<collectingDate id="EFE4E9F9FFFD760DFF3AD66E4AC9F756" box="[145,253,566,588]" pageId="32" pageNumber="449" value="1977-05-16">16.v.1977</collectingDate>
</date>
, coll.
<collectorName id="26EB5307FFFD760DFE93D66F4BE5F756" box="[312,465,566,588]" pageId="32" pageNumber="449">P. D. Hillyard</collectorName>
, det.
<determinerName id="7E7DDA15FFFD760DFDA2D66F48A9F756" box="[521,669,566,588]" pageId="32" pageNumber="449">A. F. Millidge</determinerName>
(
<collectionCode id="ED0FAE14FFFD760DFD01D66F48DBF757" box="[682,751,567,589]" pageId="32" pageNumber="449">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD62E4B3CF791" blockId="32.[145,264,630,651]" box="[145,264,630,651]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD62E4B3CF791" box="[145,264,630,651]" italics="true" pageId="32" pageNumber="449">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD6EC4B78F101" blockId="32.[145,761,692,1051]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD6EC4AD6F7D3" box="[145,226,692,713]" italics="true" pageId="32" pageNumber="449">Males:</emphasis>
The lack of male prosomal modifications and the presence of an embolic base protuberance distinguishes the males of this species from the
<taxonomicName id="4C1E4D52FFFD760DFF3DD7484ACFF63F" box="[150,251,784,805]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B96AEAC3FFFD760DFF3DD7484ACFF63F" box="[150,251,784,805]" italics="true" pageId="32" pageNumber="449">gibbosus</emphasis>
</taxonomicName>
-like species group (Clade 28). Males share with
<emphasis id="B96AEAC3FFFD760DFF64D7774B7CF65E" box="[207,328,815,836]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFF56D7774B7CF65E" box="[253,328,815,836]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="fuscus">fuscus</taxonomicName>
</emphasis>
the bifurcated protegulum, but can be distinguished by the extended distribution of papillae (from the distal part of the mesal branch of protegulum until the base of TS) and the presence of a tooth prolateral to the palpal tibia prolateral apophysis (in
<emphasis id="B96AEAC3FFFD760DFE98D7904B92F6C7" box="[307,422,968,989]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFEF5D7904B92F6C7" box="[350,422,968,989]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="fuscus">fuscus</taxonomicName>
</emphasis>
the papillae is limited to the distal part of the mesal branch and the prolateral tooth is absent).
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD01C4876F35E" blockId="32.[145,762,1092,1604]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD01C4AC0F143" box="[145,244,1092,1113]" italics="true" pageId="32" pageNumber="449">Females:</emphasis>
Can be distinguished from other species by the epigynal configuration and number of sub-AME setae (two; one in
<emphasis id="B96AEAC3FFFD760DFEDFD0D94BAEF18C" box="[372,410,1153,1174]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
<taxonomicName id="4C1E4D52FFFD760DFE08D0D94876F18C" authority=", Oe." authorityName="Oe." box="[419,578,1153,1175]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="gibbosus">
<emphasis id="B96AEAC3FFFD760DFE08D0D9483FF18C" box="[419,523,1153,1174]" italics="true" pageId="32" pageNumber="449">gibbosus</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFDB6D0D94876F18C" box="[541,578,1153,1174]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFDE7D0D948CEF18C" authority=", Oe." authorityName="Oe." box="[588,762,1153,1175]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="trilobatus">
<emphasis id="B96AEAC3FFFD760DFDE7D0D948F7F18C" box="[588,707,1153,1174]" italics="true" pageId="32" pageNumber="449">trilobatus</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFD7FD0D948CEF18C" box="[724,762,1153,1174]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFF3AD0F84B15F1AF" authority=", Oe." authorityName="Oe." box="[145,289,1184,1205]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="apicatus">
<emphasis id="B96AEAC3FFFD760DFF3AD0F84AC5F1AF" box="[145,241,1184,1205]" italics="true" pageId="32" pageNumber="449">apicatus</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFF55D0F84B15F1AF" box="[254,289,1184,1205]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFE83D0F84B9FF1AF" authority=", Oe." authorityName="Oe." box="[296,427,1184,1205]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="retusus">
<emphasis id="B96AEAC3FFFD760DFE83D0F84B4FF1AF" box="[296,379,1184,1205]" italics="true" pageId="32" pageNumber="449">retusus</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFE2CD0F84B9FF1AF" box="[391,427,1184,1205]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFE1AD0F8483CF1AF" baseAuthorityName="Kulczynski" baseAuthorityYear="1882" box="[433,520,1184,1205]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="gibbifer">
<emphasis id="B96AEAC3FFFD760DFE1AD0F8483CF1AF" box="[433,520,1184,1205]" italics="true" pageId="32" pageNumber="449">gibbifer</emphasis>
</taxonomicName>
and
<emphasis id="B96AEAC3FFFD760DFDEBD0F848DEF1AF" box="[576,746,1184,1205]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFDC1D0F848DEF1AF" authorityName="Pantini &amp; Mazzoleni" authorityYear="2018" box="[618,746,1184,1205]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="paludigena">paludigena</taxonomicName>
</emphasis>
). Distinguished from
<emphasis id="B96AEAC3FFFD760DFEDAD0E7483DF1C9" box="[369,521,1214,1236]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFE31D0E6483DF1C9" box="[410,521,1214,1235]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="trilobatus">trilobatus</taxonomicName>
</emphasis>
and
<emphasis id="B96AEAC3FFFD760DFDEBD0E748CCF1C9" box="[576,760,1214,1236]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFDC3D0E648CCF1C9" authorityName="Tanasevitch" authorityYear="2015" box="[616,760,1214,1235]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="meridionalis">meridionalis</taxonomicName>
</emphasis>
by the much more anteriorly located copulatory openings (
<figureCitation id="13252A54FFFD760DFEA7D0A44B5BF00B" box="[268,367,1276,1298]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11E</figureCitation>
); from
<emphasis id="B96AEAC3FFFD760DFE63D0A44BD8F00B" box="[456,492,1276,1297]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
<taxonomicName id="4C1E4D52FFFD760DFE5ED0A448B1F00B" authority=", Oe." authorityName="Oe." box="[501,645,1276,1297]" class="Arachnida" family="Linyphiidae" genus="Neriene" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="agrestis">
<emphasis id="B96AEAC3FFFD760DFE5ED0A44865F00B" box="[501,593,1276,1297]" italics="true" pageId="32" pageNumber="449">agrestis</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFDCAD0A448B1F00B" box="[609,645,1276,1297]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFD25D0A44A81F02A" authority=", Oe." authorityName="Oe." class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="apicatus">
<emphasis id="B96AEAC3FFFD760DFD25D0A448C6F00B" box="[654,754,1276,1297]" italics="true" pageId="32" pageNumber="449">apicatus</emphasis>
,
<emphasis id="B96AEAC3FFFD760DFF3AD1434A81F02A" box="[145,181,1307,1328]" italics="true" pageId="32" pageNumber="449">Oe.</emphasis>
</taxonomicName>
<taxonomicName id="4C1E4D52FFFD760DFF16D1424B23F035" baseAuthorityName="Kulczynski" baseAuthorityYear="1882" box="[189,279,1306,1327]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="gibbifer">
<emphasis id="B96AEAC3FFFD760DFF16D1424B23F035" box="[189,279,1306,1327]" italics="true" pageId="32" pageNumber="449">gibbifer</emphasis>
</taxonomicName>
and
<emphasis id="B96AEAC3FFFD760DFEFFD1434BE1F02A" box="[340,469,1307,1328]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFE2BD1434BE1F02A" box="[384,469,1307,1328]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="retusus">retusus</taxonomicName>
</emphasis>
by the mesal copulatory openings (
<figureCitation id="13252A54FFFD760DFEA1D1614B59F054" box="[266,365,1337,1359]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11E</figureCitation>
; lateral in the first three species,
<figureCitation id="13252A54FFFD760DFF3AD1004AD2F074" box="[145,230,1368,1390]" captionStart="Figure 5" captionStartId="10.[144,222,1846,1868]" captionTargetBox="[148,1428,353,1804]" captionTargetId="figure-82@10.[145,1431,349,1807]" captionTargetPageId="10" captionText="Figure 5. Oedothorax agrestis (Blackwall, 1853). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. EH, epigyne. E, ventral view. F, dorsal view. G, anterior view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967794" httpUri="https://zenodo.org/record/6967794/files/figure.png" pageId="32" pageNumber="449">Figs 5E</figureCitation>
,
<figureCitation id="13252A54FFFD760DFF5AD1004B2EF077" box="[241,282,1368,1389]" captionStart="Figure 12" captionStartId="20.[145,224,1684,1706]" captionTargetBox="[147,1415,214,1637]" captionTargetId="figure-24@20.[145,1429,195,1645]" captionTargetPageId="20" captionText="Figure 12. Oedothorax apicatus (Blackwall, 1850). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, male right palp, expanded, image flipped horizontally. FH, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967816" httpUri="https://zenodo.org/record/6967816/files/figure.png" pageId="32" pageNumber="449">12F</figureCitation>
,
<figureCitation id="13252A54FFFD760DFE8FD1004B67F074" box="[292,339,1368,1390]" captionStart="Figure 15" captionStartId="26.[145,224,1672,1694]" captionTargetBox="[149,1428,198,1630]" captionTargetId="figure-24@26.[145,1431,195,1633]" captionTargetPageId="26" captionText="Figure 15. Oedothorax gibbifer (Kulczyński, 1882). AF, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, copulatory bulb, apical view. F, embolic division, ventral view. GJ, epigyne. G, ventral view. H, dorsal view. I, anterior view. J, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967826" httpUri="https://zenodo.org/record/6967826/files/figure.png" pageId="32" pageNumber="449">15G</figureCitation>
; posterior in
<emphasis id="B96AEAC3FFFD760DFE4FD100486BF077" box="[484,607,1368,1389]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFDA7D100486BF077" box="[524,607,1368,1389]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="retusus">retusus</taxonomicName>
</emphasis>
<figureCitation id="13252A54FFFD760DFDCFD10048F4F074" box="[612,704,1368,1390]" captionStart="Figure 16" captionStartId="28.[145,223,1712,1734]" captionTargetBox="[161,1425,197,1658]" captionTargetId="figure-47@28.[145,1428,195,1673]" captionTargetPageId="28" captionText="Figure 16. Oedothorax retusus (Westring, 1851). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. EG, epigyne. E, ventral view. F, dorsal view. G, external morphology. H, male right spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967828" httpUri="https://zenodo.org/record/6967828/files/figure.png" pageId="32" pageNumber="449">Fig. 16E</figureCitation>
) and the more anteriorly extended copulatory ducts; from
<emphasis id="B96AEAC3FFFD760DFF3AD1CD4B14F0B0" box="[145,288,1429,1450]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFF17D1CD4B14F0B0" box="[188,288,1429,1450]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="gibbosus">gibbosus</taxonomicName>
</emphasis>
by the more curved and less convergent borders between the dorsal and ventral plates (
<figureCitation id="13252A54FFFD760DFD67D1EC4A89F0FD" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11E</figureCitation>
, in contrast to
<figureCitation id="13252A54FFFD760DFEC2D18A4B8CF0F2" box="[361,440,1490,1512]" captionStart="Figure 6" captionStartId="12.[145,223,1762,1784]" captionTargetBox="[200,1374,198,1719]" captionTargetId="figure-24@12.[190,1380,195,1722]" captionTargetPageId="12" captionText="Figure 6. Oedothorax gibbosus (Blackwall, 1841).AE, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. FH, epigyne. F, ventral view. G, dorsal view. H, external morphology. I, male spinnerets. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967798" httpUri="https://zenodo.org/record/6967798/files/figure.png" pageId="32" pageNumber="449">Fig. 6F</figureCitation>
); from
<emphasis id="B96AEAC3FFFD760DFDA2D18B484EF0FD" box="[521,634,1490,1512]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFD98D18A484EF0FD" box="[563,634,1490,1511]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="fuscus">fuscus</taxonomicName>
</emphasis>
by the less convoluted copulatory ducts compared to
<emphasis id="B96AEAC3FFFD760DFD29D1A948CDF31C" box="[642,761,1521,1542]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFD04D1A948CDF31C" box="[687,761,1521,1542]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="fuscus">fuscus</taxonomicName>
</emphasis>
(
<figureCitation id="13252A54FFFD760DFF32D2484AC8F33F" box="[153,252,1552,1574]" captionStart="Figure 14" captionStartId="24.[145,223,1846,1868]" captionTargetBox="[152,1425,551,1803]" captionTargetId="figure-193@24.[145,1431,547,1807]" captionTargetPageId="24" captionText="Figure 14. Oedothorax fuscus (Blackwall, 1834). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. EG, epigyne. E, ventral view. F, dorsal view. G, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967822" httpUri="https://zenodo.org/record/6967822/files/figure.png" pageId="32" pageNumber="449">Fig. 14E</figureCitation>
) and the narrower separation between the copulatory ducts (wider in
<emphasis id="B96AEAC3FFFD760DFE6AD2774806F359" box="[449,562,1582,1604]" italics="true" pageId="32" pageNumber="449">
Oe.
<taxonomicName id="4C1E4D52FFFD760DFE40D2764806F359" box="[491,562,1582,1603]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="449" phylum="Arthropoda" rank="species" species="fuscus">fuscus</taxonomicName>
</emphasis>
).
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD2364B2DF399" blockId="32.[145,281,1646,1667]" box="[145,281,1646,1667]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD2364B2DF399" box="[145,281,1646,1667]" italics="true" pageId="32" pageNumber="449">Description:</emphasis>
</paragraph>
<paragraph id="8BA136D1FFFD760DFF3AD2F448CCF241" blockId="32.[145,761,1708,1883]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFF3AD2F44BBBF3DB" box="[145,399,1708,1729]" italics="true" pageId="32" pageNumber="449">Male (Merdia herga):</emphasis>
Total length: 1.96. Prosoma: 0.90 long, 0.68 wide, unmodified (
<figureCitation id="13252A54FFFD760DFDA6D2924869F3FA" box="[525,605,1738,1760]" captionStart="Figure 7" captionStartId="13.[163,240,1315,1337]" captionTargetBox="[166,1440,198,1272]" captionTargetId="figure-25@13.[163,1443,195,1275]" captionTargetPageId="13" captionText="Figure 7. Male prosomal morphology, lateral.A, Emertongone montifera (Emerton, 1882) (Paquin &amp; Dupérré, 2003, fig. 1187). B, Shaanxinus mingchihensis Lin, 2019 (traced from photograph). C, Oedothorax paracymbialis Tanasevitch, 2015 (traced from photograph). D, Oe. kodaikanal Tanasevitch, 2015 (traced from photograph). E, Oe. cunur Tanasevitch, 2015 (traced from photograph). F, Oe. stylus Tanasevitch, 2015 (traced from photograph). G, Atypena cirrifrons (Heimer, 1984) (traced from photograph). H, A. formosana (Oi, 1977) (Oi, 1977, fig. 2). I, Oe. nazareti Scharff, 1989 (Scharff, 1989: fig. 7). J, Gongylidium rufipes (Linnaeus, 1758) (traced from photograph). K, Ummeliata insecticeps (Bösenberg &amp; Strand, 1906) (Tu &amp; Li 2004: fig. 8A). L, U. esyunini (Zhang, Zhang &amp; Yu, 2003) (Zhang et al., 2003, fig. 2A). M, Hylyphantes graminicola (Sundevall, 1830) (traced from photograph). N, Tmeticus tolli Kulczyński, 1908 (traced from photograph). O, Oedothorax gibbosus (Blackwall, 1841) [Roberts, 1987: fig. 21 (e)]. P, Oe. trilobatus (Banks, 1896) (Bishop &amp; Crosby, 1935: fig. 79). Q, Oe. gibbifer (Kulczyński, 1882) (Heimer &amp; Nentwig, 1991: fig.604.5).R, Oe.apicatus (Blackwall, 1850) [Roberts, 1987: fig.23 (d)].S, Oe. retusus (Westring, 1851) [Roberts, 1987: fig. 23 (c)]. T, Oe. paludigena Simon, 1926 [Millidge, 1975: fig. (5)]. U, Oe. meridionalis Tanasevitch, 1987 (Tanasevitch, 1987: fig. 107).V, Oe. agrestis (Blackwall, 1853) (Roberts, 1987: fig. 23B).W, Oe. fuscus (Blackwall, 1834) (Roberts, 1987: fig. 23A). X, Oe. tingitanus (Simon, 1884) (Denis, 1968: fig. 11)." figureDoi="http://doi.org/10.5281/zenodo.6967802" httpUri="https://zenodo.org/record/6967802/files/figure.png" pageId="32" pageNumber="449">Fig. 7X</figureCitation>
). Eyes: AME- AME: 0.06, AME width: 0.07, AME-ALE: 0.04, ALE width: 0.07, ALE-PLE: 0.00, PLE width: 0.08, PLE- PME: 0.02, PME width: 0.05, PME-PME: 0.03. Clypeus: not hirsute, two sub-AME setae. Sternum: 0.52 long,
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D49D4EADF7BC" blockId="32.[809,1425,197,679]" pageId="32" pageNumber="449">
0.5 wide. Chelicerae: mastidia absent; stridulatory striae imbricated, rows widely and evenly spaced (
<figureCitation id="13252A54FFFD760DFC9AD55A49B5F40D" box="[817,897,258,280]" captionStart="Figure 8" captionStartId="14.[146,225,1007,1029]" captionTargetBox="[147,1428,197,966]" captionTargetId="figure-341@14.[145,1428,195,967]" captionTargetPageId="14" captionText="Figure 8. Male chelicerae, stridulatory striae on lateral side. A, Oedothorax gibbosus (Blackwall, 1841). B, Oe. trilobatus (Banks, 1896). C, Oe. gibbifer (Kulczyński, 1882). D, Oe. apicatus (Blackwall, 1850). E, Oe. retusus (Westring, 1851). F, Oe. paludigena Simon, 1926. G, Oe. meridionalis Tanasevitch, 1987. H, Oe. agrestis (Blackwall, 1853). I, Oe. fuscus (Blackwall, 1834). J, Oe. tingitanus (Simon, 1884). Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6967804" httpUri="https://zenodo.org/record/6967804/files/figure.png" pageId="32" pageNumber="449">Fig. 8J</figureCitation>
). Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.60, 1.51, 2.02 and 2.34 times diameter of tibia, respectively; Tm I: 0.66. Pedipalp: TPA rodlike, distal part scaly; BT short, situated close to tip of tibial prolateral apophysis, pointed anteriorly; tooth prolateral to TPA; PC distal setae at median position (
<figureCitation id="13252A54FFFD760DFC9AD5E249A5F4D5" box="[817,913,442,464]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11A</figureCitation>
); T without papillae, PT bifurcate, papillaebearing region extending from distal part of mesal branch until base of TS, ectal branch without papillae; TS short (
<figureCitation id="13252A54FFFD760DFC0FD64E4E3FF731" box="[932,1035,534,556]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11D</figureCitation>
); DSA tip round (
<figureCitation id="13252A54FFFD760DFB43D64E4F79F731" box="[1256,1357,534,556]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11A</figureCitation>
); EM median-long, cylindrical, distally oriented, with small papillae at tip; TP with several small protuberances (
<figureCitation id="13252A54FFFD760DFC9AD62A49B9F79D" box="[817,909,626,648]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11B</figureCitation>
); E not broadened at basal part. Opisthosoma: brown, evenly coloured (
<figureCitation id="13252A54FFFD760DFB96D6C94EBEF7BC" box="[1085,1162,657,679]" captionStart="Figure 9" captionStartId="15.[163,241,931,953]" captionTargetBox="[164,1444,196,890]" captionTargetId="figure-419@15.[163,1446,195,892]" captionTargetPageId="15" captionText="Figure 9. Male opisthosoma, dorsal view. A, Oedothorax gibbosus (Blackwall, 1841). B, Oe. trilobatus (Banks, 1896). C, Oe. gibbifer (Kulczyński, 1882). D, Oe. apicatus (Blackwall, 1850). E, Oe. retusus (Westring, 1851). F, Oe. paludigena Simon, 1926. G, Oe. meridionalis Tanasevitch, 1987. H, Oe. agrestis (Blackwall, 1853). I, Oe. fuscus (Blackwall, 1834). J, Oe. tingitanus (Simon, 1884). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967808" httpUri="https://zenodo.org/record/6967808/files/figure.png" pageId="32" pageNumber="449">Fig. 9J</figureCitation>
).
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D68849D7F16E" blockId="32.[809,1425,720,1140]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D6884E03F7FF" box="[809,1079,720,741]" italics="true" pageId="32" pageNumber="449">Female (Merdia herga):</emphasis>
Total length: 2.30. Prosoma: 0.97 long, 0.77 wide. Eyes: AME-AME: 0.03, AME width: 0.06, AME-ALE: 0.02, ALE width: 0.08, ALE- PLE: 0.01, PLE width: 0.07, PLE-PME: 0.06, PME width: 0.07, PME-PME: 0.05. Clypeus: not hirsute, two sub-AME setae. Sternum: 0.61 long; 0.58 wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.68, 1.59, 2.05 and 2.27 times diameter of tibia, respectively; Tm I: 0.66. Chelicerae: stridulatory striae similar to male. Epigyne: Clade 13 characteristic morphology, borders between dorsal and ventral plates converging anteriorly, copulatory duct of median length (
<figureCitation id="13252A54FFFD760DFC46D0674E42F14F" box="[1005,1142,1087,1109]" captionStart="Figure 11" captionStartId="19.[163,241,1726,1748]" captionTargetBox="[180,1442,196,1686]" captionTargetId="figure-26@19.[163,1446,195,1687]" captionTargetPageId="19" captionText="Figure 11. Oedothorax tingitanus (Simon, 1884). AD, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, distal view. F, G, epigyne. F, ventral view. G, dorsal view. H, external morphology. Scale bar 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6967814" httpUri="https://zenodo.org/record/6967814/files/figure.png" pageId="32" pageNumber="449">Fig. 11FH</figureCitation>
). Opisthosoma: brown, evenly coloured.
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D0C54E7CF1CB" blockId="32.[809,1425,1181,1233]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D0C549AFF1A8" box="[809,923,1181,1202]" italics="true" pageId="32" pageNumber="449">Variation:</emphasis>
Measurements are based on
<specimenCount id="9D18FD58FFFD760DFB42D0C54F21F1A8" box="[1257,1301,1181,1202]" pageId="32" pageNumber="449" type="male">10♂</specimenCount>
<specimenCount id="9D18FD58FFFD760DFABED0C54F06F1A8" box="[1301,1330,1181,1202]" pageId="32" pageNumber="449" type="female">1♀</specimenCount>
(Merdia herga) and
<specimenCount id="9D18FD58FFFD760DFC00D0E449FCF1CA" box="[939,968,1212,1233]" pageId="32" pageNumber="449" type="female">8♀</specimenCount>
(Tekouan).
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D0A24944F0D2" blockId="32.[809,1425,1274,1480]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D0A24EDFF015" box="[809,1259,1274,1296]" italics="true" pageId="32" pageNumber="449">Males (N = 10, means in parentheses):</emphasis>
Total length 1.842.06 (1.93). Prosoma: 0.820.91 (0.87) long, 0.66 0.73 (0.69) wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.241.78 (1.47,
<emphasis id="B96AEAC3FFFD760DFB64D10F4ED7F076" box="[1231,1251,1367,1388]" italics="true" pageId="32" pageNumber="449">N</emphasis>
= 9), 1.461.76 (1.59), 1.902.30 (2.11,
<emphasis id="B96AEAC3FFFD760DFBEFD12E4E6CF091" box="[1092,1112,1398,1419]" italics="true" pageId="32" pageNumber="449">N</emphasis>
= 9) and 1.982.44 (2.21) times diameter of tibia, respectively; Tm I: 0.600.68 (0.64).
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D1A94F2EF3BA" blockId="32.[809,1425,1521,1696]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D1A94EDAF31C" box="[809,1262,1521,1543]" italics="true" pageId="32" pageNumber="449">Females (N = 9, means in parentheses):</emphasis>
Total length 2.102.69 (2.44). Prosoma: 0.971.07 (1.04) long, 0.77 0.88 (0.83) wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 1.581.92 (1.70), 1.411.90 (1.71), 1.862.20 (2.02) and 1.902.37 (2.13) times diameter of tibia, respectively; Tm I: 0.560.71 (0.65).
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D2914F09F3C5" blockId="32.[809,1341,1737,1759]" box="[809,1341,1737,1759]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D291498EF3C4" box="[809,954,1737,1758]" italics="true" pageId="32" pageNumber="449">Distribution:</emphasis>
<collectingCountry id="F3097641FFFD760DFC6CD2914E3FF3C5" box="[967,1035,1737,1759]" name="Spain" pageId="32" pageNumber="449">Spain</collectingCountry>
,
<collectingCountry id="F3097641FFFD760DFBBDD2914E41F3C4" box="[1046,1141,1737,1758]" name="Morocco" pageId="32" pageNumber="449">Morocco</collectingCountry>
,
<collectingCountry id="F3097641FFFD760DFBD4D2914EE0F3C5" box="[1151,1236,1737,1759]" name="Algeria" pageId="32" pageNumber="449">Algeria</collectingCountry>
,
<collectingCountry id="F3097641FFFD760DFB75D2914F0DF3C5" box="[1246,1337,1737,1759]" name="Tunisia" pageId="32" pageNumber="449">Tunisia</collectingCountry>
.
</paragraph>
<paragraph id="8BA136D1FFFD760DFC82D35F49C2F226" blockId="32.[809,1425,1799,1852]" pageId="32" pageNumber="449">
<emphasis id="B96AEAC3FFFD760DFC82D35F49BDF206" box="[809,905,1799,1820]" italics="true" pageId="32" pageNumber="449">Habitat:</emphasis>
Marshes, salt meadows, wet meadows, lake shores, riversides.
</paragraph>
</subSubSection>
</treatment>
</document>