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<document id="E8D6C7FDE85B1013C05C0057818AED60" ID-CLB-Dataset="115951" ID-DOI="10.1093/zoolinnean/zlab033" ID-GBIF-Dataset="75b10efe-a17d-4d10-b46e-094c6a7ebf01" ID-ISSN="0024-4082" ID-Zenodo-Dep="6967774" ID-ZooBank="BE2B3859-8F6A-4543-8A69-91840F82377B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1653991067759" checkinUser="plazi" docAuthor="Lin, Shou-Wang, Lopardo, Lara &amp; Uhl, Gabriele" docDate="2022" docId="03B787C7FF5576A5FC92D49C4E78F319" docLanguage="en" docName="zlab033.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Oedothorax sexmaculatus SAITO &amp; ONO 2001, INCERTAE SEDIS" docType="treatment" docVersion="7" lastPageNumber="553" masterDocId="FF8EFFBFFFDD762DFFABD4584A34F51A" masterDocTitle="Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)" masterLastPageNumber="584" masterPageNumber="417" pageNumber="553" updateTime="1699338434129" updateUser="ExternalLinkService">
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<mods:title id="74377FD70126B02F5703D090746846EB">Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae)</mods:title>
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<mods:namePart id="BCA50EA08C0EC02711A6A9F5F7B30D22">Lin, Shou-Wang</mods:namePart>
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<mods:namePart id="6A0F85DA37C6D8C4B0EF5EB0A9070FAF">Lopardo, Lara</mods:namePart>
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<mods:namePart id="7154F1DBAA6A5CD198A5BF21F03BF6D8">Uhl, Gabriele</mods:namePart>
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<mods:title id="AE9C45CAA5DD0071D75AC67097D83050">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="4DEDC95CCFCB638E801D60C1CA176861">2022</mods:date>
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<treatment id="03B787C7FF5576A5FC92D49C4E78F319" ID-DOI="http://doi.org/10.5281/zenodo.6968052" ID-GBIF-Taxon="198618725" ID-Zenodo-Dep="6968052" LSID="urn:lsid:plazi:treatment:03B787C7FF5576A5FC92D49C4E78F319" httpUri="http://treatment.plazi.org/id/03B787C7FF5576A5FC92D49C4E78F319" lastPageNumber="553" pageId="136" pageNumber="553">
<subSubSection id="C304655AFF5576A5FC92D49C4E8DF5E0" pageId="136" pageNumber="553" type="nomenclature">
<paragraph id="8BA136D1FF5576A5FC92D49C4E8DF5E0" blockId="136.[825,1408,196,250]" pageId="136" pageNumber="553">
<heading id="D0E981BDFF5576A5FC92D49C4E8DF5E0" centered="true" fontSize="9" level="2" pageId="136" pageNumber="553" reason="2">
<emphasis id="B96AEAC3FF5576A5FC92D49C490AF5C1" box="[825,830,196,219]" italics="true" pageId="136" pageNumber="553"></emphasis>
<taxonomicName id="4C1E4D52FF5576A5FC95D49C4FB4F5C7" ID-CoL="48WVF" authority="SAITO &amp; ONO, 2001 INCERTAE" authorityName="SAITO &amp; ONO" authorityYear="2001" box="[830,1408,196,221]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="136" pageNumber="553" phylum="Arthropoda" rank="species" species="sexmaculatus" status="INCERTAE SEDIS">
<emphasis id="B96AEAC3FF5576A5FC95D49C4EADF5C1" box="[830,1177,196,219]" italics="true" pageId="136" pageNumber="553">OEDOTHORAX SEXMACULATUS</emphasis>
<bibRefCitation id="EF8F4B20FF5576A5FB0BD49C4FB4F5C7" author="Saito H &amp; Ono H" box="[1184,1408,196,221]" pageId="136" pageNumber="553" pagination="1 - 59" refId="ref94881" refString="Saito H, Ono H. 2001. New genera and species of the spider family Linyphiidae (Arachnida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo (A) 27: 1 - 59." type="journal article" year="2001">SAITO &amp; ONO, 2001</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A25957B8FF5576A5FBABD4BE4E8DF5E0" box="[1024,1209,230,250]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FBABD4BE4E8DF5E0" box="[1024,1209,230,250]" italics="true" pageId="136" pageNumber="553">INCERTAE SEDIS</emphasis>
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5576A5FC82D55349D2F467" pageId="136" pageNumber="553" type="reference_group">
<paragraph id="8BA136D1FF5576A5FC82D55349F6F45A" blockId="136.[809,1425,267,381]" pageId="136" pageNumber="553">
<treatmentCitationGroup id="AB0E11FFFF5576A5FC82D55349F6F45A" pageId="136" pageNumber="553">
<treatmentCitation id="0ABF10C0FF5576A5FC82D5534F6DF43B" author="Saito H &amp; Ono H" box="[809,1369,267,289]" page="5" pageId="136" pageNumber="553" year="2001">
<taxonomicName id="4C1E4D52FF5576A5FC82D5534F6DF43B" ID-CoL="48WVF" authority="Saito &amp; Ono, 2001: 5" authorityName="SAITO &amp; ONO" authorityPageNumber="5" authorityYear="2001" box="[809,1369,267,289]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="136" pageNumber="553" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B96AEAC3FF5576A5FC82D5534E6CF43A" box="[809,1112,267,288]" italics="true" pageId="136" pageNumber="553">Oedothorax sexmaculatus</emphasis>
<bibRefCitation id="EF8F4B20FF5576A5FBCAD5534F6DF43B" author="Saito H &amp; Ono H" box="[1121,1369,267,289]" pageId="136" pageNumber="553" pagination="1 - 59" refId="ref94881" refString="Saito H, Ono H. 2001. New genera and species of the spider family Linyphiidae (Arachnida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo (A) 27: 1 - 59." type="journal article" year="2001">Saito &amp; Ono, 2001: 5</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 59 (Dmf).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136D1FF5576A5FC82D51049D2F467" blockId="136.[809,1425,267,381]" pageId="136" pageNumber="553">
<treatmentCitationGroup id="AB0E11FFFF5576A5FC82D51049D2F467" pageId="136" pageNumber="553">
<treatmentCitation id="0ABF10C0FF5576A5FC82D5104F6DF447" author="Ono H &amp; Matsuda M &amp; Saito H" box="[809,1369,328,350]" page="284" pageId="136" pageNumber="553" year="2009">
<taxonomicName id="4C1E4D52FF5576A5FC82D5104F6DF447" ID-CoL="48WVF" authority="Ono et al., 2009: 284" authorityName="Ono" authorityPageNumber="284" authorityYear="2009" box="[809,1369,328,350]" class="Arachnida" family="Linyphiidae" genus="Oedothorax" kingdom="Animalia" order="Araneae" pageId="136" pageNumber="553" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B96AEAC3FF5576A5FC82D5104E6DF447" box="[809,1113,328,349]" italics="true" pageId="136" pageNumber="553">Oedothorax sexmaculatus</emphasis>
<bibRefCitation id="EF8F4B20FF5576A5FBC9D5104F6DF447" author="Ono H &amp; Matsuda M &amp; Saito H" box="[1122,1369,328,350]" pageId="136" pageNumber="553" pagination="253 - 344" refId="ref93932" refString="Ono H, Matsuda M, Saito H. 2009. Linyphiidae, Pimoidae. In: Ono H, ed. The spiders of Japan with keys to the families and genera and illustrations of the species. Kanagawa: Tokai University Press, 253 - 344." type="book chapter" year="2009">
Ono
<emphasis id="B96AEAC3FF5576A5FB31D5114EE2F447" box="[1178,1238,328,350]" italics="true" pageId="136" pageNumber="553">et al.</emphasis>
, 2009: 284
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 343347 (mf).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5576A5FC82D5F94E2AF797" pageId="136" pageNumber="553" type="materials_examined">
<paragraph id="8BA136D1FF5576A5FC82D5F94E2AF797" blockId="136.[809,1424,416,653]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D5F949CEF4AC" box="[809,1018,416,438]" italics="true" pageId="136" pageNumber="553">Type material:</emphasis>
<materialsCitation id="3B763C8CFF5576A5FBA6D5F84E81F708" ID-GBIF-Occurrence="3880945395" collectingDate="1986-06-11" collectorName="K. Kumada" country="Japan" county="Jododaria" location="Hunshu" municipality="Azuma-yama Mts." pageId="136" pageNumber="553" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A58873FF5576A5FBA6D5F84EBDF4AC" box="[1037,1161,416,438]" pageId="136" pageNumber="553" type="holotype">Holotype</typeStatus>
:
<emphasis id="B96AEAC3FF5576A5FB0FD5F84F3EF4AF" bold="true" box="[1188,1290,416,437]" pageId="136" pageNumber="553">
<collectingCountry id="F3097641FF5576A5FB0FD5F84F34F4AF" box="[1188,1280,416,437]" name="Japan" pageId="136" pageNumber="553">Japan</collectingCountry>
:
</emphasis>
<location id="8EC1600AFF5576A5FAB7D5F84FBCF4AC" LSID="urn:lsid:plazi:treatment:03B787C7FF5576A5FC92D49C4E78F319:8EC1600AFF5576A5FAB7D5F84FBCF4AC" box="[1308,1416,416,438]" country="Japan" county="Jododaria" municipality="Azuma-yama Mts." name="Hunshu" pageId="136" pageNumber="553">Hunshu</location>
, Fukuschima Pref.,
<location id="8EC1600AFF5576A5FB96D5E74EF2F4CE" LSID="urn:lsid:plazi:treatment:03B787C7FF5576A5FC92D49C4E78F319:8EC1600AFF5576A5FB96D5E74EF2F4CE" box="[1085,1222,447,468]" country="Japan" county="Jododaria" municipality="Azuma-yama Mts." name="Yama-gun" pageId="136" pageNumber="553">Yama-gun</location>
,
<collectingMunicipality id="6BC5ACABFF5576A5FB74D5E74957F4E9" pageId="136" pageNumber="553">Azuma-yama Mts.</collectingMunicipality>
,
<collectingCounty id="62C04E5DFF5576A5FCDED58649C2F4EE" box="[885,1014,478,500]" pageId="136" pageNumber="553">Jododaria</collectingCounty>
,
<specimenCount id="9D18FD58FF5576A5FBA3D5B84E2CF4E8" box="[1032,1048,480,498]" pageId="136" pageNumber="553" type="male"></specimenCount>
,
<date id="FFA01011FF5576A5FB87D5864E80F4EE" box="[1068,1204,478,500]" pageId="136" pageNumber="553" value="1986-06-11">
<collectingDate id="EFE4E9F9FF5576A5FB87D5864E80F4EE" box="[1068,1204,478,500]" pageId="136" pageNumber="553" value="1986-06-11">11.vi.1986</collectingDate>
</date>
, leg.
<collectorName id="26EB5307FF5576A5FB54D5864FBBF4EE" box="[1279,1423,478,500]" pageId="136" pageNumber="553">K. Kumada</collectorName>
(NSMT-Ar 4572, not examined)
</materialsCitation>
.
<materialsCitation id="3B763C8CFF5576A5FB68D5A54E2EF797" ID-GBIF-Occurrence="3880945411" collectingDate="1983-06-01" collectorName="H. Saito" country="Japan" county="Tone-gun" location="Marunuma" municipality="Katashina-mura" pageId="136" pageNumber="553" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Gunma" typeStatus="paratype">
<typeStatus id="54A58873FF5576A5FB68D5A54F77F708" box="[1219,1347,509,530]" pageId="136" pageNumber="553" type="paratype">Paratypes</typeStatus>
: same data as holotype,
<specimenCount id="9D18FD58FF5576A5FBADD6434E11F72A" box="[1030,1061,539,561]" pageId="136" pageNumber="553" type="male">5♂</specimenCount>
<specimenCount id="9D18FD58FF5576A5FB8DD6434E70F72A" box="[1062,1092,539,561]" pageId="136" pageNumber="553" type="female">6♀</specimenCount>
(NSMT-Ar 45734574, not examined);
<collectingRegion id="49DAF833FF5576A5FC04D6624E72F74A" box="[943,1094,570,592]" country="Japan" name="Gunma" pageId="136" pageNumber="553">Gunma Pref.</collectingRegion>
,
<collectingCounty id="62C04E5DFF5576A5FBE4D6624E8AF755" box="[1103,1214,570,591]" pageId="136" pageNumber="553">Tone-gun</collectingCounty>
,
<collectingMunicipality id="6BC5ACABFF5576A5FB62D6624FB9F74A" box="[1225,1421,570,592]" pageId="136" pageNumber="553">Katashina-mura</collectingMunicipality>
,
<location id="8EC1600AFF5576A5FC82D6014980F774" LSID="urn:lsid:plazi:treatment:03B787C7FF5576A5FC92D49C4E78F319:8EC1600AFF5576A5FC82D6014980F774" box="[809,948,601,622]" county="Tone-gun" municipality="Katashina-mura" name="Marunuma" pageId="136" pageNumber="553" stateProvince="Gunma">Marunuma</location>
,
<specimenCount id="9D18FD58FF5576A5FC68D60049D6F777" box="[963,994,600,621]" pageId="136" pageNumber="553" type="male">1♂</specimenCount>
<date id="FFA01011FF5576A5FC47D6004E6AF774" box="[1004,1118,600,622]" pageId="136" pageNumber="553" value="1983-06-01">
<collectingDate id="EFE4E9F9FF5576A5FC47D6004E6AF774" box="[1004,1118,600,622]" pageId="136" pageNumber="553" value="1983-06-01">1.vi.1983</collectingDate>
</date>
, leg.
<collectorName id="26EB5307FF5576A5FB08D6014F3CF774" box="[1187,1288,600,622]" pageId="136" pageNumber="553">H. Saito</collectorName>
(NSMT-Ar 4575, not examined)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5576A5FC82D6F64E3DF13C" pageId="136" pageNumber="553" type="diagnosis">
<paragraph id="8BA136D1FF5576A5FC82D6F64994F7D9" blockId="136.[809,928,686,707]" box="[809,928,686,707]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D6F64994F7D9" box="[809,928,686,707]" italics="true" pageId="136" pageNumber="553">Diagnosis:</emphasis>
</paragraph>
<paragraph id="8BA136D1FF5576A5FC82D6BC498AF68E" blockId="136.[809,1425,740,916]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D6BC494EF7E3" box="[809,890,740,761]" italics="true" pageId="136" pageNumber="553">Males:</emphasis>
This species can be distinguished by the variegated opisthosoma, the palpal apophysis and the depiction of the bulbal structure in figures 8 and
<quantity id="4CE69B34FF5576A5FACFD77A4FA5F622" box="[1380,1425,802,824]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="136" pageNumber="553" unit="in" value="9.0">9 in</quantity>
<bibRefCitation id="EF8F4B20FF5576A5FC82D7184E35F64C" author="Saito H &amp; Ono H" box="[809,1025,832,854]" pageId="136" pageNumber="553" pagination="1 - 59" refId="ref94881" refString="Saito H, Ono H. 2001. New genera and species of the spider family Linyphiidae (Arachnida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo (A) 27: 1 - 59." type="journal article" year="2001">Saito &amp; Ono (2001)</bibRefCitation>
, in which the individual structures of the embolic division are unfortunately not clearly recognizable.
</paragraph>
<paragraph id="8BA136D1FF5576A5FC82D7EC4E3DF13C" blockId="136.[809,1425,948,1062]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D7EC49BAF6D3" box="[809,910,948,969]" italics="true" pageId="136" pageNumber="553">Females:</emphasis>
This species can be identified by the dorsal pattern of opisthosoma with six dark patches, and the epigynal external morphology illustrated in figure
<quantity id="4CE69B34FF5576A5FACDD7AA4FA5F112" box="[1382,1425,1010,1032]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="136" pageNumber="553" unit="in" value="9.0">9 in</quantity>
<bibRefCitation id="EF8F4B20FF5576A5FC82D0484E37F13C" author="Saito H &amp; Ono H" box="[809,1027,1040,1062]" pageId="136" pageNumber="553" pagination="1 - 59" refId="ref94881" refString="Saito H, Ono H. 2001. New genera and species of the spider family Linyphiidae (Arachnida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo (A) 27: 1 - 59." type="journal article" year="2001">Saito &amp; Ono (2001)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5576A5FC82D01F4E4FF147" box="[809,1147,1095,1117]" pageId="136" pageNumber="553" type="distribution">
<paragraph id="8BA136D1FF5576A5FC82D01F4E4FF147" blockId="136.[809,1147,1095,1117]" box="[809,1147,1095,1117]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D01F498FF146" box="[809,955,1095,1116]" italics="true" pageId="136" pageNumber="553">Distribution:</emphasis>
<collectingCountry id="F3097641FF5576A5FC6CD01F4E3BF146" box="[967,1039,1095,1116]" name="Japan" pageId="136" pageNumber="553">Japan</collectingCountry>
: Honshu.
</paragraph>
</subSubSection>
<subSubSection id="C304655AFF5576A5FC82D0254E78F319" pageId="136" pageNumber="553" type="discussion">
<paragraph id="8BA136D1FF5576A5FC82D0254E78F319" blockId="136.[809,1425,1149,1539]" pageId="136" pageNumber="553">
<emphasis id="B96AEAC3FF5576A5FC82D02549A1F188" box="[809,917,1149,1170]" italics="true" pageId="136" pageNumber="553">Remarks:</emphasis>
The description of
<bibRefCitation id="EF8F4B20FF5576A5FBD1D0254F6CF18E" author="Saito H &amp; Ono H" box="[1146,1368,1149,1172]" pageId="136" pageNumber="553" pagination="1 - 59" refId="ref94881" refString="Saito H, Ono H. 2001. New genera and species of the spider family Linyphiidae (Arachnida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo (A) 27: 1 - 59." type="journal article" year="2001">Saito &amp; Ono (2001)</bibRefCitation>
is in accordance with the general combination of somatic characters of
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<emphasis id="B96AEAC3FF5576A5FC6DD0E34E7FF1CA" box="[966,1099,1211,1232]" italics="true" pageId="136" pageNumber="553">Oedothorax</emphasis>
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,
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<emphasis id="B96AEAC3FF5576A5FBF2D0E34E8BF1CA" box="[1113,1215,1211,1232]" italics="true" pageId="136" pageNumber="553">Mitrager</emphasis>
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and other related taxa. Although the configuration of the embolic division is not fully recognizable in their drawings, the embolus seems to be prolaterally spiralled (fig.
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), but the ARP tip is not pointed at tip and not spiral as in
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<emphasis id="B96AEAC3FF5576A5FB80D10C4E9AF073" box="[1067,1198,1364,1385]" italics="true" pageId="136" pageNumber="553">Oedothorax</emphasis>
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; its PC has a clearly broader distal part (fig.
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) than in
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. The six distinct spots on the opisthosoma is seen neither in
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</emphasis>
nor in
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.
</emphasis>
Due to these dissimilarities, this species most probably belongs to another genus.
</paragraph>
</subSubSection>
</treatment>
</document>