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<document id="4FA6CCC5B5D73B5FAF27DE8FC31A0F31" ID-DOI="10.1649/0010-065X-73.2.343" ID-ISSN="1938-4394" ID-Zenodo-Dep="10085324" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622212308669" checkinUser="carolina" docAuthor="Clark, Shawn M. &amp; Anderson, E. Russell" docDate="2019" docId="03B787BF553D401CFE96DA04AD68CAD0" docLanguage="en" docName="TheColeopBull.73.2.343-357.pdf" docOrigin="The Coleopterists Bulletin 73 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-73.2.343" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Triarius Jacoby 1887" docType="treatment" docVersion="6" lastPageNumber="350" masterDocId="FF8EFFC7553B401BFFA4DB21AC60CE7E" masterDocTitle="A Review of Triarius Jacoby, 1887 (Coleoptera: Chrysomelidae: Galerucinae: Luperini), with Descriptions of a New Genus and Four New Species" masterLastPageNumber="357" masterPageNumber="343" pageNumber="349" updateTime="1699916141952" updateUser="ExternalLinkService">
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<mods:title id="A62FBCD98918E1693F3DAD0380F9A1EF">A Review of Triarius Jacoby, 1887 (Coleoptera: Chrysomelidae: Galerucinae: Luperini), with Descriptions of a New Genus and Four New Species</mods:title>
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<subSubSection id="C3046522553D401DFE96DA04AE6ECF44" box="[306,526,293,314]" pageId="6" pageNumber="349" type="nomenclature">
<paragraph id="8BA136A9553D401DFE96DA04AE6ECF44" blockId="6.[306,526,293,341]" box="[306,526,293,314]" pageId="6" pageNumber="349">
<heading id="D0E981C5553D401DFE96DA04AE6ECF44" box="[306,526,293,314]" centered="true" fontSize="9" level="2" pageId="6" pageNumber="349" reason="2">
<taxonomicName id="4C1E4D2A553D401DFE96DA04AE6ECF44" ID-CoL="7YX2" authority="Jacoby, 1887" authorityName="Jacoby" authorityYear="1887" box="[306,526,293,314]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553D401DFE96DA04AE6ECF44" bold="true" box="[306,526,293,314]" pageId="6" pageNumber="349">
<emphasis id="B96AEABB553D401DFE96DA04ADE3CF44" bold="true" box="[306,387,293,314]" italics="true" pageId="6" pageNumber="349">Triarius</emphasis>
<bibRefCitation id="EF8F4B58553D401DFE2EDA04AE6ECF44" author="Jacoby, M." box="[394,526,293,314]" pageId="6" pageNumber="349" refId="ref12144" refString="Jacoby, M. 1887. Biologia Centrali-Americana, Insecta, Coleoptera, Galerucidae, vol. 6, pt. 1. London, UK." type="book" year="1887">Jacoby, 1887</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C3046522553D401DFEF6DA1EAD8ECF2A" box="[338,494,319,341]" pageId="6" pageNumber="349" type="description">
<paragraph id="8BA136A9553D401DFEF6DA1EAD8ECF2A" blockId="6.[306,526,293,341]" box="[338,494,319,341]" pageId="6" pageNumber="349">
<figureCitation id="13252A2C553D401DFEF6DA1EADD3CF2A" box="[338,435,319,341]" captionStart="Fig" captionStartId="6.[197,228,1687,1706]" captionTargetBox="[179,1188,443,1668]" captionTargetId="figure-76@6.[175,1190,441,1670]" captionTargetPageId="6" captionText="Fig. 2. Triarius species, dorsal habitus. A) T. pini, male, B) T. pini, female, C) T. trivittatus, D) T. nigroflavus, male, E) T. nigroflavus, female, F) T. melanolomatus, G) T. lividus, H) T. texanus, I) T. novoleonis." figureDoi="http://doi.org/10.5281/zenodo.10120531" httpUri="https://zenodo.org/record/10120531/files/figure.png" pageId="6" pageNumber="349">Figs. 2ai</figureCitation>
,
<figureCitation id="13252A2C553D401DFE1BDA1EAD8ECF2A" box="[447,494,319,341]" captionStart="Fig" captionStartId="8.[197,228,1301,1320]" captionTargetBox="[177,1180,196,1283]" captionTargetId="figure-200@8.[175,1190,194,1285]" captionTargetPageId="8" captionText="Fig. 3. Triarius species, aedeagi, dorsal and lateral views. A) T. pini, B) T. nigroflavus, C) T. melanolomatus, D) T. lividus, E) T. trivittatus, F) T. texanus, G) T. novoleonis." figureDoi="http://doi.org/10.5281/zenodo.10120533" httpUri="https://zenodo.org/record/10120533/files/figure.png" pageId="6" pageNumber="349">3ag</figureCitation>
</paragraph>
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<subSubSection id="C3046522553D401DFF08DA4BADCCCF01" box="[172,428,362,383]" pageId="6" pageNumber="349" type="reference_group">
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<treatmentCitationGroup id="AB0E1187553D401DFF08DA4BADCCCF01" box="[172,428,362,383]" pageId="6" pageNumber="349">
<treatmentCitation id="0ABF10B8553D401DFF08DA4BADCCCF01" author="Jacoby, M." box="[172,428,362,383]" page="571" pageId="6" pageNumber="349" year="1887">
<taxonomicName id="4C1E4D2A553D401DFF08DA4BADCCCF01" ID-CoL="7YX2" authority="Jacoby 1887: 571" authorityName="Jacoby" authorityPageNumber="571" authorityYear="1887" box="[172,428,362,383]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553D401DFF08DA4BAC99CF01" box="[172,249,362,383]" italics="true" pageId="6" pageNumber="349">Triarius</emphasis>
<bibRefCitation id="EF8F4B58553D401DFEA5DA4BADCCCF01" author="Jacoby, M." box="[257,428,362,383]" pageId="6" pageNumber="349" refId="ref12144" refString="Jacoby, M. 1887. Biologia Centrali-Americana, Insecta, Coleoptera, Galerucidae, vol. 6, pt. 1. London, UK." type="book" year="1887">Jacoby 1887: 571</bibRefCitation>
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</paragraph>
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<subSubSection id="C3046522553D401DFD7FDBE3AF39CE8F" pageId="6" pageNumber="349" type="type_taxon">
<paragraph id="8BA136A9553D401DFD7FDBE3AF39CE8F" blockId="6.[707,1193,194,382]" pageId="6" pageNumber="349">
<emphasis id="B96AEABB553D401DFD7FDBE3AF03CEA9" bold="true" box="[731,867,194,215]" pageId="6" pageNumber="349">
<typeStatus id="54A5880B553D401DFD7FDBE3AF6DCEA9" box="[731,781,194,215]" pageId="6" pageNumber="349">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C1E4D2A553D401DFCCFDBE3A8C9CEA9" authority="Jacoby, 1887" authorityName="Jacoby. Just" authorityYear="1887" box="[875,1193,194,215]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B96AEABB553D401DFCCFDBE3A843CEA9" box="[875,1059,194,215]" italics="true" pageId="6" pageNumber="349">Triarius mexicanus</emphasis>
<bibRefCitation id="EF8F4B58553D401DFB8FDBE3A8C9CEA9" author="Jacoby, M." box="[1067,1193,194,215]" pageId="6" pageNumber="349" refId="ref12144" refString="Jacoby, M. 1887. Biologia Centrali-Americana, Insecta, Coleoptera, Galerucidae, vol. 6, pt. 1. London, UK." type="book" year="1887">Jacoby, 1887</bibRefCitation>
</taxonomicName>
(by monotypy).
</paragraph>
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<subSubSection id="C3046522553D401CFD7FDBD6AD87CC34" lastPageId="7" lastPageNumber="350" pageId="6" pageNumber="349" type="diagnosis">
<paragraph id="8BA136A9553D401CFD7FDBD6AD87CC34" blockId="6.[707,1193,194,382]" lastBlockId="7.[151,639,194,1198]" lastPageId="7" lastPageNumber="350" pageId="6" pageNumber="349">
<emphasis id="B96AEABB553D401DFD7FDBD6AF22CF72" bold="true" box="[731,834,247,268]" pageId="6" pageNumber="349">Diagnosis.</emphasis>
The basal carina of the pronotum is complete. Although fine, it extends without interruption along the entire posterior margin. Each eye is narrowly separated from the base of the mandible by a distance subequal to the width of the basal maxillary palpomere. The length of the basal antennomere is subequal to the maximum diameter of the eye. The epipleuron is broad in the basal portion and very narrow in the distal portion. Each tibia is armed with a terminal spur. The rectangular lobe at the apex of the male abdomen is conspicuous. In these characters,
<taxonomicName id="4C1E4D2A553C401CFEE4DA40ADEBCF08" authorityName="Jacoby" authorityYear="1887" box="[320,395,353,374]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFEE4DA40ADEBCF08" box="[320,395,353,374]" italics="true" pageId="7" pageNumber="350">Triarius</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C1E4D2A553C401CFDB3DA40AE17CF08" authorityName="Clark and Anderson" authorityYear="2019" box="[535,631,353,374]" class="Insecta" family="Chrysomelidae" genus="Triariodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFDB3DA40AE17CF08" box="[535,631,353,374]" italics="true" pageId="7" pageNumber="350">Triariodes</emphasis>
</taxonomicName>
. However, the frontal tubercles of
<taxonomicName id="4C1E4D2A553C401CFE78DA5DAE47CFEF" authorityName="Jacoby" authorityYear="1887" box="[476,551,380,401]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFE78DA5DAE47CFEF" box="[476,551,380,401]" italics="true" pageId="7" pageNumber="350">Triarius</emphasis>
</taxonomicName>
are only about as broad as long, being limited laterad by a shallow (sometimes rather inconspicuous) depression. In
<taxonomicName id="4C1E4D2A553C401CFEA8DAEAAD0CCF9E" authorityName="Clark and Anderson" authorityYear="2019" box="[268,364,459,480]" class="Insecta" family="Chrysomelidae" genus="Triariodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFEA8DAEAAD0CCF9E" box="[268,364,459,480]" italics="true" pageId="7" pageNumber="350">Triariodes</emphasis>
</taxonomicName>
, the depression is nearly absent, such that the tubercles are strongly transverse, closely approaching the eye. Moreover, the aedeagus of
<taxonomicName id="4C1E4D2A553C401CFF11D93AAD60CC4E" authorityName="Jacoby" authorityYear="1887" box="[181,256,539,560]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFF11D93AAD60CC4E" box="[181,256,539,560]" italics="true" pageId="7" pageNumber="350">Triarius</emphasis>
</taxonomicName>
is symmetrical, with the distal portion only weakly spatulate, or not at all.
</paragraph>
</subSubSection>
<caption id="DF616621553D401DFF61DDB6A860C8BC" ID-DOI="http://doi.org/10.5281/zenodo.10120531" ID-Zenodo-Dep="10120531" httpUri="https://zenodo.org/record/10120531/files/figure.png" pageId="6" pageNumber="349" startId="6.[197,228,1687,1706]" targetBox="[179,1188,443,1668]" targetPageId="6">
<paragraph id="8BA136A9553D401DFF61DDB6A860C8BC" blockId="6.[173,1193,1687,1730]" pageId="6" pageNumber="349">
<emphasis id="B96AEABB553D401DFF61DDB6AC9FC8D4" bold="true" box="[197,255,1687,1706]" pageId="6" pageNumber="349">Fig. 2.</emphasis>
<taxonomicName id="4C1E4D2A553D401DFEB1DDB6ADC0C8D4" box="[277,416,1687,1706]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96AEABB553D401DFEB1DDB6AD39C8D4" box="[277,345,1687,1706]" italics="true" pageId="6" pageNumber="349">Triarius</emphasis>
species
</taxonomicName>
, dorsal habitus. A)
<taxonomicName id="4C1E4D2A553D401DFDF5DDB6AEEAC8D4" baseAuthorityName="Schaeffer" baseAuthorityYear="1906" box="[593,650,1687,1706]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="B96AEABB553D401DFDF5DDB6AEEAC8D4" box="[593,650,1687,1706]" italics="true" pageId="6" pageNumber="349">T. pini</emphasis>
</taxonomicName>
, male, B)
<taxonomicName id="4C1E4D2A553D401DFD4FDDB6AF44C8D4" baseAuthorityName="Schaeffer" baseAuthorityYear="1906" box="[747,804,1687,1706]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="pini">
<emphasis id="B96AEABB553D401DFD4FDDB6AF44C8D4" box="[747,804,1687,1706]" italics="true" pageId="6" pageNumber="349">T. pini</emphasis>
</taxonomicName>
, female, C)
<taxonomicName id="4C1E4D2A553D401DFC32DDB6AF9FC8D4" authorityName="Horn" authorityYear="1893" box="[918,1023,1687,1706]" class="Insecta" family="Chrysomelidae" genus="Ttiarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="trivittatus">
<emphasis id="B96AEABB553D401DFC32DDB6AF9FC8D4" box="[918,1023,1687,1706]" italics="true" pageId="6" pageNumber="349">T. trivittatus</emphasis>
</taxonomicName>
, D)
<taxonomicName id="4C1E4D2A553D401DFB88DDB6A8C3C8D4" box="[1068,1187,1687,1706]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="nigroflavus">
<emphasis id="B96AEABB553D401DFB88DDB6A8C3C8D4" box="[1068,1187,1687,1706]" italics="true" pageId="6" pageNumber="349">T. nigroflavus</emphasis>
</taxonomicName>
, male, E)
<taxonomicName id="4C1E4D2A553D401DFF58DD8EAD11C8BC" box="[252,369,1711,1730]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="nigroflavus">
<emphasis id="B96AEABB553D401DFF58DD8EAD11C8BC" box="[252,369,1711,1730]" italics="true" pageId="6" pageNumber="349">T. nigroflavus</emphasis>
</taxonomicName>
, female, F)
<taxonomicName id="4C1E4D2A553D401DFE7FDD8EAE13C8BC" authorityName="Blake" authorityYear="1942" box="[475,627,1711,1730]" class="Insecta" family="Chrysomelidae" genus="Luperodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="melanolomatus">
<emphasis id="B96AEABB553D401DFE7FDD8EAE13C8BC" box="[475,627,1711,1730]" italics="true" pageId="6" pageNumber="349">T. melanolomatus</emphasis>
</taxonomicName>
, G)
<taxonomicName id="4C1E4D2A553D401DFD3FDD8EAE8AC8BC" box="[667,746,1711,1730]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="lividus">
<emphasis id="B96AEABB553D401DFD3FDD8EAE8AC8BC" box="[667,746,1711,1730]" italics="true" pageId="6" pageNumber="349">T. lividus</emphasis>
</taxonomicName>
, H)
<taxonomicName id="4C1E4D2A553D401DFCB7DD8EAF09C8BC" authorityName="Clark and Anderson" authorityYear="2019" box="[787,873,1711,1730]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="B96AEABB553D401DFCB7DD8EAF09C8BC" box="[787,873,1711,1730]" italics="true" pageId="6" pageNumber="349">T. texanus</emphasis>
</taxonomicName>
, I)
<taxonomicName id="4C1E4D2A553D401DFC2EDD8EAF9BC8BC" authorityName="Clark and Anderson" authorityYear="2019" box="[906,1019,1711,1730]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="novoleonis">
<emphasis id="B96AEABB553D401DFC2EDD8EAF9BC8BC" box="[906,1019,1711,1730]" italics="true" pageId="6" pageNumber="349">T. novoleonis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3046522553C401CFF0BD971AD68CAD0" pageId="7" pageNumber="350" type="discussion">
<paragraph id="8BA136A9553C401CFF0BD971AD68CAD0" blockId="7.[151,639,194,1198]" pageId="7" pageNumber="350">
<emphasis id="B96AEABB553C401CFF0BD971AD44CC1B" bold="true" box="[175,292,592,613]" pageId="7" pageNumber="350">Comments.</emphasis>
Within the subfamily
<taxonomicName id="4C1E4D2A553C401CFDA7D971AE1BCC1B" authorityName="Latreille" authorityYear="1802" box="[515,635,592,613]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="subFamily" subFamily="Galerucinae">Galerucinae</taxonomicName>
, the nature of the tarsal claws (simple, appendiculate, or bifid) is extensively employed to diagnose genera or even taxa above the generic level. It is therefore remarkable that some species of
<taxonomicName id="4C1E4D2A553C401CFF33D9F4AC85CC94" authorityName="Jacoby" authorityYear="1887" box="[151,229,725,746]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFF33D9F4AC85CC94" box="[151,229,725,746]" italics="true" pageId="7" pageNumber="350">Triarius</emphasis>
</taxonomicName>
have bifid claws and others have appendiculate claws. However, this situation is not unique among the New World genera in the section Scelidites, where claws are reported to be appendiculate, except in some species of
<taxonomicName id="4C1E4D2A553C401CFDA5D81EAE2DCD2A" authorityName="Jacoby" authorityYear="1887" box="[513,589,831,852]" class="Insecta" family="Chrysomelidae" genus="Triarius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFDA5D81EAE2DCD2A" box="[513,589,831,852]" italics="true" pageId="7" pageNumber="350">Triarius</emphasis>
</taxonomicName>
. Although not previously reported in the published literature, the claws are also bifid in
<taxonomicName id="4C1E4D2A553C401CFDA1D855AD7FCDDA" authority="Jacoby" authorityName="Jacoby" authorityYear="1888" class="Insecta" family="Chrysomelidae" genus="Scelida" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="B96AEABB553C401CFDA1D855ACADCDDA" italics="true" pageId="7" pageNumber="350">Scelida metallica</emphasis>
Jacoby
</taxonomicName>
, while appendiculate in all other species of
<taxonomicName id="4C1E4D2A553C401CFEACD88BAE65CDC1" authority="Chapuis. Beyond" authorityName="Chapuis. Beyond" authorityYear="1875" box="[264,517,938,959]" class="Insecta" family="Chrysomelidae" genus="Scelida" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFEACD88BAD2ECDC1" box="[264,334,938,959]" italics="true" pageId="7" pageNumber="350">Scelida</emphasis>
Chapuis. Beyond
</taxonomicName>
the section Scelidites, the lack of claw uniformity is not common, but also not unprecedented. For instance, all species of
<taxonomicName id="4C1E4D2A553C401CFEB8D8D8ADA9CA70" authority="Blake" authorityName="Blake" authorityYear="1936" box="[284,457,1017,1038]" class="Insecta" family="Chrysomelidae" genus="Erynephala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFEB8D8D8ADEBCA70" box="[284,395,1017,1038]" italics="true" pageId="7" pageNumber="350">Erynephala</emphasis>
Blake
</taxonomicName>
and some (but not all) species of
<taxonomicName id="4C1E4D2A553C401CFEB9DF35ADA9CA57" authority="LeConte" authorityName="LeConte" box="[285,457,1044,1065]" class="Insecta" family="Chrysomelidae" genus="Monoxia" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFEB9DF35AD11CA57" box="[285,369,1044,1065]" italics="true" pageId="7" pageNumber="350">Monoxia</emphasis>
LeConte
</taxonomicName>
have bifid claws in males and simple claws in females. In males of
<taxonomicName id="4C1E4D2A553C401CFF33DF68AE1ACA20" authority="Bechyne and Springlova de Bechyne" authorityName="Bechyne and Springlova de Bechyne" box="[151,634,1097,1118]" class="Insecta" family="Chrysomelidae" genus="Hyperbrotica" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEABB553C401CFF33DF68AD76CA20" box="[151,278,1097,1118]" italics="true" pageId="7" pageNumber="350">Hyperbrotica</emphasis>
Bechyné and Springlová de Bechyné
</taxonomicName>
, all claws are appendiculate in females, while only the hind claws are appendiculate in males, the others being bifid.
</paragraph>
</subSubSection>
</treatment>
</document>