treatments-xml/data/03/B7/87/03B787B74D20324CFF7747D7FEDF34D6.xml
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<document id="54DF35F5873E28940F5A788A9D3066A0" ID-CLB-Dataset="53621" ID-DOI="10.11646/zootaxa.5124.5.6" ID-GBIF-Dataset="2309e531-0c74-42e7-9d0a-bb669dd7a66c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6417067" ID-ZooBank="CB42D636-9ED7-4BCA-AF3B-20F4412F0020" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649229388743" checkinUser="plazi" docAuthor="Nascimento, Elynton Alves Do &amp; Bocakova, Milada" docDate="2022" docId="03B787B74D20324CFF7747D7FEDF34D6" docLanguage="en" docName="zootaxa.5124.5.6.pdf" docOrigin="Zootaxa 5124 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladocalon bugabense Nascimento &amp; Bocakova 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="580" masterDocId="FF8EFFCF4D22324FFFE04255FFDE3507" masterDocTitle="Cladocalon, a new genus of net-winged beetles from Central America and Mexico (Coleoptera: Lycidae)" masterLastPageNumber="584" masterPageNumber="577" pageNumber="579" updateTime="1699319478996" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="70650FBE1E25047CFA1FD94120DECFF9">Cladocalon, a new genus of net-winged beetles from Central America and Mexico (Coleoptera: Lycidae)</mods:title>
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<mods:namePart id="4C0199E73FB472A03886F32B0E23DA77">Nascimento, Elynton Alves Do</mods:namePart>
<mods:affiliation id="9D68CDE338B752F1094E15C9185B2BED">Departamento de Engenharia Ambiental, Universidade Estadual do Centro-Oeste, Rua Professora Maria Roza Zanon de Almeida, s / n, Engenheiro Gutierrez, Irati-PR, Brazil, CEP 84505 - 677.</mods:affiliation>
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<mods:namePart id="E98E5BE91DCDDCE84F91D39F0B3F8E5E">Bocakova, Milada</mods:namePart>
<mods:nameIdentifier id="4C568FF12DC2ADA772173CB5ABE29EA2" type="ORCID">0000-0002-2507-0887</mods:nameIdentifier>
<mods:affiliation id="FB99DF5727AC1C0A7BC4B016F2FFC9B8">Department of Biology, Faculty of Education, Palacky University, Purkrabska 2, CZ- 77140 Olomouc, Czech Republic. milada. bocakova @ upol. cz; https: // orcid. org / 0000 - 0002 - 2507 - 0887</mods:affiliation>
<mods:nameIdentifier id="3A35BE636CB2EACA9565C66463214C16" type="email">milada.bocakova@upol.cz</mods:nameIdentifier>
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<mods:date id="850FFFB869EFC21036A8FF4777A491C7">2022</mods:date>
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<mods:number id="59384E5A050BED0351FCF4B4832FD7EA">2022-04-06</mods:number>
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<treatment id="03B787B74D20324CFF7747D7FEDF34D6" ID-DOI="http://doi.org/10.5281/zenodo.6416871" ID-GBIF-Taxon="194469468" ID-Zenodo-Dep="6416871" LSID="urn:lsid:plazi:treatment:03B787B74D20324CFF7747D7FEDF34D6" httpUri="http://treatment.plazi.org/id/03B787B74D20324CFF7747D7FEDF34D6" lastPageId="3" lastPageNumber="580" pageId="2" pageNumber="579">
<subSubSection id="C304652A4D20324DFF7747D7FDD1309A" box="[151,527,1410,1437]" pageId="2" pageNumber="579" type="nomenclature">
<paragraph id="8BA136A14D20324DFF7747D7FDD1309A" blockId="2.[151,527,1410,1473]" box="[151,527,1410,1437]" pageId="2" pageNumber="579">
<heading id="D0E981CD4D20324DFF7747D7FDD1309A" bold="true" box="[151,527,1410,1437]" fontSize="11" level="1" pageId="2" pageNumber="579" reason="1">
<emphasis id="B96AEAB34D20324DFF7747D7FDD1309A" bold="true" box="[151,527,1410,1437]" pageId="2" pageNumber="579">
<taxonomicName id="4C1E4D224D20324DFF7747D7FE74309A" authority="Nascimento &amp; Bocakova, 2022" authorityName="Nascimento &amp; Bocakova" authorityYear="2022" box="[151,426,1410,1437]" class="Insecta" family="Lycidae" genus="Cladocalon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="bugabense" status="sp. nov.">
<emphasis id="B96AEAB34D20324DFF7747D7FE74309A" bold="true" box="[151,426,1410,1437]" italics="true" pageId="2" pageNumber="579">Cladocalon bugabense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957C84D20324DFE5147D6FDD1309A" box="[433,527,1411,1437]" pageId="2" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C304652A4D20324DFF7747F2FEF730C6" box="[151,297,1447,1473]" pageId="2" pageNumber="579" type="description">
<paragraph id="8BA136A14D20324DFF7747F2FEF730C6" blockId="2.[151,527,1410,1473]" box="[151,297,1447,1473]" pageId="2" pageNumber="579">
<figureCitation id="13252A244D20324DFF7747F2FEDF30C6" box="[151,257,1447,1473]" captionStart="FIGURES 115" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[162,1422,199,1796]" captionTargetId="figure-25@4.[151,1436,181,1807]" captionTargetPageId="4" captionText="FIGURES 115. 1. male antennae of Cladocalon mexicanus sp. nov. 2. Terminal maxillary palpi of C. histrionicum comb. nov. 34: Pronotum. 3. C. chiriquense comb. nov., 4. C. histrionicum. 5. Male abdominal sternum 9 of C. chiriquense. 6-15: Male genitalia (lateral and ventral views, respectively). 67. C. chiriquense. 89. C. bugabense sp. nov.1011. C. histrionicum.1213. C. mexicanus sp. nov. 1415. C. guatemalense sp. nov. Scales: 0.25 mm (Fig. 2), 0.5 mm (Figs. 1, 3-15)." figureDoi="http://doi.org/10.5281/zenodo.6417069" httpUri="https://zenodo.org/record/6417069/files/figure.png" pageId="2" pageNumber="579">
Figs. 8
<emphasis id="B96AEAB34D20324DFF0347F2FF2F30C6" bold="true" box="[227,241,1447,1473]" pageId="2" pageNumber="579"></emphasis>
9
</figureCitation>
,
<figureCitation id="13252A244D20324DFEED47F2FEF730C6" box="[269,297,1447,1473]" captionStart="FIGURES 1620" captionStartId="5.[151,264,1754,1778]" captionTargetBox="[175,1420,326,1716]" captionTargetId="figure-69@5.[151,1436,309,1731]" captionTargetPageId="5" captionText="FIGURES 1620. Habitus: 16. Cladocalon chiriquense comb. nov. 17. C. bugabense sp. nov. 18. C. histrionicum. comb. nov.19. C. mexicanus sp. nov. 20. C. guatemalense sp. nov. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6417071" httpUri="https://zenodo.org/record/6417071/files/figure.png" pageId="2" pageNumber="579">17</figureCitation>
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<paragraph id="8BA136A14D20324DFF7747BAFA553372" blockId="2.[151,1437,1518,2013]" pageId="2" pageNumber="579">
<emphasis id="B96AEAB34D20324DFF7747BAFE9A330F" bold="true" box="[151,324,1518,1545]" pageId="2" pageNumber="579">Type material:</emphasis>
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<typeStatus id="54A588034D20324DFEA747BAFE6A330E" box="[327,436,1519,1545]" pageId="2" pageNumber="579" type="holotype">Holotype</typeStatus>
</emphasis>
, male,
<collectingCountry id="F30976314D20324DFDE047BAFDA5330E" box="[512,635,1519,1545]" name="Panama" pageId="2" pageNumber="579">PANAMA</collectingCountry>
:“
<location id="8EC1607A4D20324DFD6C47BAFD39330E" LSID="urn:lsid:plazi:treatment:03B787B74D20324CFF7747D7FEDF34D6:8EC1607A4D20324DFD6C47BAFD39330E" box="[652,743,1519,1545]" country="Panama" municipality="Champion" name="Bugaba" pageId="2" pageNumber="579">Bugaba</location>
,
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800
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1500ft.
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,
<collectingMunicipality id="6BC5ACDB4D20324DFC6547BAFBDD330E" box="[901,1027,1519,1545]" pageId="2" pageNumber="579">Champion</collectingMunicipality>
.”.Secondary label:
<collectorName id="26EB53774D20324DFB3447BAFA8D330D" box="[1236,1363,1519,1546]" pageId="2" pageNumber="579">B.C.A.Col.</collectorName>
III.(2).
<taxonomicName id="4C1E4D224D20324DFF774446FE7C332A" authorityName="Gorham" authorityYear="1884" box="[151,418,1555,1581]" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="chiriquense">Calocladon chiriquense</taxonomicName>
. [
<typeStatus id="54A588034D20324DFE514447FDD0332A" box="[433,526,1554,1581]" pageId="2" pageNumber="579" type="syntype">syntype</typeStatus>
] (
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)
</materialsCitation>
.
<materialsCitation id="3B763CFC4D20324DFD434446FB883356" ID-GBIF-Occurrence="3747946311" collectionCode="NHMUK" collectorName="B. C. A. Col." country="Panama" elevation="351" location="Champion" municipality="Bugaba" pageId="2" pageNumber="579" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B96AEAB34D20324DFD434446FCC5332A" bold="true" box="[675,795,1555,1581]" pageId="2" pageNumber="579">
<typeStatus id="54A588034D20324DFD434446FCC5332A" box="[675,795,1555,1581]" pageId="2" pageNumber="579" type="paratype">Paratypes</typeStatus>
</emphasis>
,
<specimenCount id="9D18FD284D20324DFCC84446FCAB332A" box="[808,885,1555,1581]" pageId="2" pageNumber="579" type="male">1 male</specimenCount>
,
<collectingCountry id="F30976314D20324DFC9F4446FC24332A" box="[895,1018,1555,1581]" name="Panama" pageId="2" pageNumber="579">PANAMA</collectingCountry>
: “
<collectingMunicipality id="6BC5ACDB4D20324DFBED4446FBB6332A" box="[1037,1128,1555,1581]" pageId="2" pageNumber="579">Bugaba</collectingMunicipality>
,
<quantity id="4CE69B444D20324DFB924446FADD332A" box="[1138,1283,1554,1581]" metricMagnitude="2" metricUnit="m" metricValue="3.5052" metricValueMax="4.572" metricValueMin="2.4384" pageId="2" pageNumber="579" unit="ft" value="1150.0" valueMax="1500.0" valueMin="800.0">
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800
<emphasis id="B96AEAB34D20324DFB7C4446FB74332A" bold="true" box="[1180,1194,1555,1581]" pageId="2" pageNumber="579"></emphasis>
1500 ft.
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,
<location id="8EC1607A4D20324DFAED4446FA55332A" LSID="urn:lsid:plazi:treatment:03B787B74D20324CFF7747D7FEDF34D6:8EC1607A4D20324DFAED4446FA55332A" box="[1293,1419,1555,1581]" country="Panama" municipality="Bugaba" name="Champion" pageId="2" pageNumber="579">Champion</location>
.”. Secondary label:
<collectorName id="26EB53774D20324DFE834462FE313355" box="[355,495,1591,1618]" pageId="2" pageNumber="579">B.C.A. Col.</collectorName>
III.(2).
<taxonomicName id="4C1E4D224D20324DFDA24462FC8C3356" authorityName="Gorham" authorityYear="1884" box="[578,850,1591,1617]" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="chiriquense">Calocladon chiriquense</taxonomicName>
. [
<typeStatus id="54A588034D20324DFC874463FC1A3356" box="[871,964,1590,1617]" pageId="2" pageNumber="579" type="syntype">syntype</typeStatus>
] (
<collectionCode id="ED0FAE644D20324DFC3E4462FB913356" box="[990,1103,1591,1617]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="579" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
;
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<specimenCount id="9D18FD284D20324DFB864462FB663356" box="[1126,1208,1591,1617]" pageId="2" pageNumber="579" type="male">1 male</specimenCount>
,
<collectingCountry id="F30976314D20324DFB284462FA9D3356" box="[1224,1347,1591,1617]" name="Panama" pageId="2" pageNumber="579">PANAMA</collectingCountry>
: “V. de
<collectingRegion id="49DAF8434D20324DFF77440EFF293372" box="[151,247,1627,1653]" country="Panama" name="Chiriqui" pageId="2" pageNumber="579">Chiriqui</collectingRegion>
,
<quantity id="4CE69B444D20324DFEE3440FFE5A3372" box="[259,388,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" pageId="2" pageNumber="579" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">
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25
<emphasis id="B96AEAB34D20324DFEFF440EFEF33372" bold="true" box="[287,301,1627,1653]" pageId="2" pageNumber="579"></emphasis>
4000 ft.
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</quantity>
<location id="8EC1607A4D20324DFE6B440EFDD33372" LSID="urn:lsid:plazi:treatment:03B787B74D20324CFF7747D7FEDF34D6:8EC1607A4D20324DFE6B440EFDD33372" box="[395,525,1627,1653]" country="Panama" name="Champion." pageId="2" pageNumber="579" stateProvince="Chiriqui">Champion.</location>
”. Secondary labels: “
<collectorName id="26EB53774D20324DFD19440EFCA73371" box="[761,889,1627,1654]" pageId="2" pageNumber="579">B.C.A.Col.</collectorName>
III.(2).
<taxonomicName id="4C1E4D224D20324DFC2A440EFB073372" authorityName="Gorham" authorityYear="1884" box="[970,1241,1627,1653]" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="chiriquense">Calocladon chiriquense</taxonomicName>
.” “
<typeStatus id="54A588034D20324DFB16440EFAA33372" box="[1270,1405,1627,1653]" pageId="2" pageNumber="579" type="syntype">SYNTYPE</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C304652A4D20324DFF27442BFC8A3202" pageId="2" pageNumber="579" type="diagnosis">
<paragraph id="8BA136A14D20324DFF27442BFC8A3202" blockId="2.[151,1437,1518,2013]" pageId="2" pageNumber="579">
<emphasis id="B96AEAB34D20324DFF27442BFEE4339F" bold="true" box="[199,314,1662,1688]" pageId="2" pageNumber="579">Diagnosis</emphasis>
. The species can be differentiated by body coloration similar to
<taxonomicName id="4C1E4D224D20324DFBDE442AFB38339E" authorityName="Gorham" authorityYear="1884" box="[1086,1254,1663,1689]" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="579" phylum="Arthropoda" rank="species" species="chiriquense">
<emphasis id="B96AEAB34D20324DFBDE442AFB38339E" box="[1086,1254,1663,1689]" italics="true" pageId="2" pageNumber="579">C. chiriquense</emphasis>
</taxonomicName>
, but presenting maxillary palpomere 2 2.4× longer than palpomere 3, and branch of third antennomere 2.3× longer than antennomere, pronotum almost as long as wide, and male genitalia with shorter, V-shaped and apically single-hooked laterally projected parameres, and phallobase as long as half of phallus.
</paragraph>
</subSubSection>
<subSubSection id="C304652A4D20324CFF27455BFB8A3446" lastPageId="3" lastPageNumber="580" pageId="2" pageNumber="579" type="description">
<paragraph id="8BA136A14D20324DFF27455BFC19324A" blockId="2.[151,1437,1518,2013]" pageId="2" pageNumber="579">
<emphasis id="B96AEAB34D20324DFF27455BFE8F322F" bold="true" box="[199,337,1806,1832]" pageId="2" pageNumber="579">Description</emphasis>
. Pronotum yellow. Elytra yellow, elytral suture light brown at basal ¼ (
<figureCitation id="13252A244D20324DFB96455AFB17322E" box="[1142,1225,1807,1833]" captionStart="FIGURES 1620" captionStartId="5.[151,264,1754,1778]" captionTargetBox="[175,1420,326,1716]" captionTargetId="figure-69@5.[151,1436,309,1731]" captionTargetPageId="5" captionText="FIGURES 1620. Habitus: 16. Cladocalon chiriquense comb. nov. 17. C. bugabense sp. nov. 18. C. histrionicum. comb. nov.19. C. mexicanus sp. nov. 20. C. guatemalense sp. nov. Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6417071" httpUri="https://zenodo.org/record/6417071/files/figure.png" pageId="2" pageNumber="579">Fig. 17</figureCitation>
). Trochanters light brown, femur dark brown with basal 1/3 light brown. Tibiae dark brown.
</paragraph>
<paragraph id="8BA136A14D20324CFF274502FB8A3446" blockId="2.[151,1437,1518,2013]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="580" pageId="2" pageNumber="579">
Eyes large, eye diameter 1.7× longer interocular distance. Antennomere 4 as long as 5, 1.4× longer than 3. Branch of antennomere 3 2.3× longer than antennomere. Mandibles hooked. Labial palpi short, terminal palpomere rounded apically. Maxillary palpi elongate, palpomere 2 about 2.4× longer than palpomere 3 and 1.1× longer than 4, palpomere 4 rounded distally. Pronotum almost as long as wide, posterior margin bisinuate. Elytra 3.6× longer than humeral width. Trochanters as long as third of femur length, tibiae as long as femur. Phallus broad medially being as wide as phallobase in ventral view, forming phallic folding projected ventrally in lateral view. Phallus distally narrow and rounded. Internal sac with many conspicuous spiculae distally. Parameres almost V-shaped, basal half projected dorsally, apical half with distal hook laterally projected, parameres as long as phallus half. Phallobase as long as half of phallus (
<figureCitation id="13252A244D21324CFE7F4372FDD23446" box="[415,524,295,321]" captionStart="FIGURES 115" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[162,1422,199,1796]" captionTargetId="figure-25@4.[151,1436,181,1807]" captionTargetPageId="4" captionText="FIGURES 115. 1. male antennae of Cladocalon mexicanus sp. nov. 2. Terminal maxillary palpi of C. histrionicum comb. nov. 34: Pronotum. 3. C. chiriquense comb. nov., 4. C. histrionicum. 5. Male abdominal sternum 9 of C. chiriquense. 6-15: Male genitalia (lateral and ventral views, respectively). 67. C. chiriquense. 89. C. bugabense sp. nov.1011. C. histrionicum.1213. C. mexicanus sp. nov. 1415. C. guatemalense sp. nov. Scales: 0.25 mm (Fig. 2), 0.5 mm (Figs. 1, 3-15)." figureDoi="http://doi.org/10.5281/zenodo.6417069" httpUri="https://zenodo.org/record/6417069/files/figure.png" pageId="3" pageNumber="580">Figs. 89</figureCitation>
). Body length:
<quantity id="4CE69B444D21324CFD584373FCE53446" box="[696,827,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.4" metricValueMin="5.8" pageId="3" pageNumber="580" unit="mm" value="6.1" valueMax="6.4" valueMin="5.8">
5.8
<emphasis id="B96AEAB34D21324CFD3B4372FD373446" bold="true" box="[731,745,295,321]" pageId="3" pageNumber="580"></emphasis>
6.4mm
</quantity>
, humeral width:
<quantity id="4CE69B444D21324CFC174372FB8E3446" box="[1015,1104,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="580" unit="mm" value="1.3">1.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C304652A4D21324CFF27431FFE173462" box="[199,457,330,357]" pageId="3" pageNumber="580" type="distribution">
<paragraph id="8BA136A14D21324CFF27431FFE173462" blockId="3.[151,1437,151,465]" box="[199,457,330,357]" pageId="3" pageNumber="580">
<emphasis id="B96AEAB34D21324CFF27431FFE873463" bold="true" box="[199,345,330,356]" pageId="3" pageNumber="580">Distribution</emphasis>
:
<collectingCountry id="F30976314D21324CFE88431EFE1A3462" box="[360,452,331,357]" name="Panama" pageId="3" pageNumber="580">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C304652A4D21324CFF27433AFD6E348E" box="[199,688,367,393]" pageId="3" pageNumber="580" type="etymology">
<paragraph id="8BA136A14D21324CFF27433AFD6E348E" blockId="3.[151,1437,151,465]" box="[199,688,367,393]" pageId="3" pageNumber="580">
<emphasis id="B96AEAB34D21324CFF27433AFE96348E" bold="true" box="[199,328,367,393]" pageId="3" pageNumber="580">Etymology</emphasis>
: named after the collection site.
</paragraph>
</subSubSection>
<subSubSection id="C304652A4D21324CFF2743C7FEDF34D6" pageId="3" pageNumber="580" type="discussion">
<paragraph id="8BA136A14D21324CFF2743C7FEDF34D6" blockId="3.[151,1437,151,465]" pageId="3" pageNumber="580">
<emphasis id="B96AEAB34D21324CFF2743C7FE8C34AB" bold="true" box="[199,338,402,428]" pageId="3" pageNumber="580">Comments:</emphasis>
The type series of this new species is comprising three individuals from
<taxonomicName id="4C1E4D224D21324CFB6F43C6FA4334AA" authorityName="Gorham" authorityYear="1884" box="[1167,1437,403,429]" class="Insecta" family="Lycidae" genus="Calocladon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="580" phylum="Arthropoda" rank="species" species="chiriquense">
<emphasis id="B96AEAB34D21324CFB6F43C6FA4334AA" box="[1167,1437,403,429]" italics="true" pageId="3" pageNumber="580">Calocladon chiriquense</emphasis>
</taxonomicName>
<typeStatus id="54A588034D21324CFF7743E2FF2334D6" box="[151,253,439,465]" pageId="3" pageNumber="580" type="syntype">syntypes</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>