976 lines
134 KiB
XML
976 lines
134 KiB
XML
<document id="98854EC4EFAA8BFBA9B4F41F5D0B8382" ID-DOI="10.18590/euscorpius.2009.vol2009.iss90.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="12782205" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721368671199" checkinUser="felipe" docAuthor="Lowe, Graeme" docDate="2009" docId="03B787A3834CFFD2C056FAB0FD94FEF9" docLanguage="en" docName="Euscorpius.90.1-13.pdf" docOrigin="Euscorpius 90 (90)" docSource="https://mds.marshall.edu/euscorpius/vol2009/iss90/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Compsobuthus nematodactylus Lowe, 2009, sp. nov." docType="treatment" docUuid="EC96437F-2DA8-49E6-A0AB-CDF959543087" docUuidSource="ZooBank" docVersion="2" lastPageNumber="11" masterDocId="FF8EFFDB834FFFDFC14EFB40FFE7FF8E" masterDocTitle="A new lithophilic Compsobuthus Vachon, 1949 (Scorpiones: Buthidae) from northern Oman" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1721382374664" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="2DE9A1FA4C60EE01FD518905CAB2A608" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3233BBDCD69ECA4277E89FA2D3BEA70A">
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<mods:title id="1715C566819ECFDA177BE2F73A058EEB">A new lithophilic Compsobuthus Vachon, 1949 (Scorpiones: Buthidae) from northern Oman</mods:title>
|
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</mods:titleInfo>
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<mods:name id="52DB3ECF9ACDBA207517251461496249" type="personal">
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<mods:roleTerm id="661727B3C90AAEE436850C605CF9741C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="228551823C97715FB6C18B8E290BED0F">Lowe, Graeme</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3B94A8F9A72585F89A784A6EAC05BCB9">text</mods:typeOfResource>
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<mods:relatedItem id="AC534241A7F361949022ADAE3E8F24C5" type="host">
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<mods:titleInfo id="9BDC2EE4D689CD0BCC248FA042B23A29">
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<mods:title id="E3E8AE08DE74666DB22ECA0B592AD64E">Euscorpius</mods:title>
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</mods:titleInfo>
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<mods:part id="D3F4AE022264076F48AE6AA285426343">
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<mods:date id="FB96724257055D8E09275ACA04BDCE21">2009</mods:date>
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<mods:detail id="B8EDD5C8625247AE765CFC8801AD660C" type="pubDate">
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<mods:number id="4245AFA4F0567C433FA4DF9D6C491175">2009-12-31</mods:number>
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</mods:detail>
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<mods:detail id="E45354389A85924E9EE03B4C3B0C0E94" type="volume">
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<mods:number id="38B30B3A3327C6B60CCF4A4207B06735">90</mods:number>
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</mods:detail>
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<mods:detail id="2A7C97A3C27EBECA5DD1FF927FE05B4A" type="issue">
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<mods:number id="B9199C1707A117ECD74CCCD768AB48EC">90</mods:number>
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<mods:extent id="8BC3BB75DB3DCBC4D4405ACC040803B8" unit="page">
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<mods:start id="9238D289A6A74F681DC42709800B20A2">1</mods:start>
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<mods:end id="21A18D1F3DFBDB643276F2A9B78776C1">13</mods:end>
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</mods:relatedItem>
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<mods:location id="36BDE5EED034798A5614CF09F442F8AA">
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<mods:url id="C8AED6114C6C4CF268D15EDE93091549">https://mds.marshall.edu/euscorpius/vol2009/iss90/1/</mods:url>
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</mods:location>
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<mods:classification id="621D96168FBE079E341A93616227026A">journal article</mods:classification>
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<mods:identifier id="888257C6EB7AA89B10CEC640532FAE52" type="DOI">10.18590/euscorpius.2009.vol2009.iss90.1</mods:identifier>
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<mods:identifier id="D3C24D3B17956CA089339299A1059915" type="ISSN">1536-9307</mods:identifier>
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<mods:identifier id="FA7C26E671915275DF293833DB9E8196" type="Zenodo-Dep">12782205</mods:identifier>
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<mods:identifier id="81A8EF8D7C4CB3B52D4FA5AF62EAA557" type="ZooBank">BC6DC40D-96A6-4635-AC98-E11A9684AE0B</mods:identifier>
|
||
</mods:mods>
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||
<treatment id="03B787A3834CFFD2C056FAB0FD94FEF9" LSID="urn:lsid:zoobank.org:act:EC96437F-2DA8-49E6-A0AB-CDF959543087" httpUri="http://treatment.plazi.org/id/03B787A3834CFFD2C056FAB0FD94FEF9" lastPageId="13" lastPageNumber="11" pageId="3" pageNumber="2">
|
||
<subSubSection id="C304653E834CFFDCC056FAB0FD2DFD86" box="[280,714,496,520]" pageId="3" pageNumber="2" type="nomenclature">
|
||
<paragraph id="8BA136B5834CFFDCC056FAB0FD2DFD86" blockId="3.[280,714,496,551]" box="[280,714,496,520]" pageId="3" pageNumber="2">
|
||
<heading id="D0E981D9834CFFDCC056FAB0FD2DFD86" box="[280,714,496,520]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2" reason="4">
|
||
<taxonomicName id="4C1E4D36834CFFDCC056FAB0FD88FD86" ID-CoL="XHG3" authorityName="Lowe" authorityYear="2009" box="[280,623,496,520]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA7834CFFDCC056FAB0FD88FD86" bold="true" box="[280,623,496,520]" italics="true" pageId="3" pageNumber="2">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25957DC834CFFDCC338FAB1FD2DFD86" box="[630,714,497,520]" pageId="3" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E834CFFDCC0CFF950FD86FDA9" box="[385,609,528,551]" pageId="3" pageNumber="2" type="description">
|
||
<paragraph id="8BA136B5834CFFDCC0CFF950FD86FDA9" blockId="3.[280,714,496,551]" box="[385,609,528,551]" pageId="3" pageNumber="2">
|
||
(
|
||
<figureCitation id="13252A30834CFFDCC0C6F950FE1AFDA9" box="[392,509,528,551]" captionStart-0="Figures 1–9" captionStart-1="Figures 10–17" captionStart-2="Figures 18–19" captionStart-3="Figures 20–21" captionStart-4="Figures 22–23" captionStartId-0="5.[192,278,1794,1817]" captionStartId-1="6.[192,278,1625,1648]" captionStartId-2="7.[192,278,1259,1282]" captionStartId-3="8.[192,278,1193,1216]" captionStartId-4="10.[192,278,733,756]" captionTargetBox-0="[286,1362,229,1696]" captionTargetBox-1="[232,1397,207,1539]" captionTargetBox-2="[192,1438,192,1227]" captionTargetBox-3="[192,1439,192,1160]" captionTargetBox-4="[218,1411,218,671]" captionTargetId-0="figure-17@5.[192,1438,885,1062]" captionTargetId-1="figure-18@6.[192,1440,712,890]" captionTargetId-2="figure-6@7.[192,1438,192,1227]" captionTargetId-3="figure-9@8.[192,1439,192,1160]" captionTargetId-4="figure-9@10.[192,1438,192,700]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="10" captionText-0="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." captionText-1="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." captionText-2="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-3="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." captionText-4="Figures 22–23: Compsobuthus nematodactylus sp. nov. paratype adult male. 22. Pedipalp chela, left movable finger dentition. 23. Pedipalp chela, left fixed finger dentition. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782207" figureDoi-1="http://doi.org/10.5281/zenodo.12782211" figureDoi-2="http://doi.org/10.5281/zenodo.12782213" figureDoi-3="http://doi.org/10.5281/zenodo.12782215" figureDoi-4="http://doi.org/10.5281/zenodo.12782219" httpUri-0="https://zenodo.org/record/12782207/files/figure.png" httpUri-1="https://zenodo.org/record/12782211/files/figure.png" httpUri-2="https://zenodo.org/record/12782213/files/figure.png" httpUri-3="https://zenodo.org/record/12782215/files/figure.png" httpUri-4="https://zenodo.org/record/12782219/files/figure.png" pageId="3" pageNumber="2">Figs. 1–23</figureCitation>
|
||
,
|
||
<tableCitation id="C69C030E834CFFDCC349F950FDB0FDA9" box="[519,599,528,551]" captionStart="Table 1" captionStartId="9.[192,257,1266,1289]" captionText="Table 1: Variation in selected morphometric ratios for adult Compsobuthus nematodactylus sp. nov. Indicated are ranges and mean ± SD, with sample sizes in parentheses. All samples are from adults. Values of variables significantly different between the two sexes (p <0.05, Mann-Whitney test) are cited separately for males and females, and those not significantly different are pooled for both sexes." pageId="3" pageNumber="2">Table 1</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E834CFFDCC18EF911FDE5FA26" pageId="3" pageNumber="2" type="materials_examined">
|
||
<paragraph id="8BA136B5834CFFDCC18EF911FE96FDE5" blockId="3.[192,369,593,619]" box="[192,369,593,619]" pageId="3" pageNumber="2">
|
||
<emphasis id="B96AEAA7834CFFDCC18EF911FE96FDE5" bold="true" box="[192,369,593,619]" pageId="3" pageNumber="2">Type material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834CFFDCC18EF9D0FD84FC8D" blockId="3.[192,802,656,771]" pageId="3" pageNumber="2">
|
||
<materialsCitation id="3B763CE8834CFFDCC18EF9D0FD84FC8D" collectingDate="1994-09-30" collectionCode="UV, NMB" collectorName="G. Lowe" country="Oman" elevation="220" latitude="56.324" location="Khawr Najd" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Musandam" typeStatus="holotype">
|
||
<emphasis id="B96AEAA7834CFFDCC18EF9D0FEC5FD29" box="[192,290,656,679]" italics="true" pageId="3" pageNumber="2">
|
||
<typeStatus id="54A58817834CFFDCC18EF9D0FEC5FD29" box="[192,290,656,679]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D18FD3C834CFFDCC062F9D0FEA7FD29" box="[300,320,656,679]" count="1" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
,
|
||
<emphasis id="B96AEAA7834CFFDCC01FF9D0FE78FD29" bold="true" box="[337,415,656,679]" pageId="3" pageNumber="2">
|
||
<collectingCountry id="F3097625834CFFDCC01FF9D0FE7DFD29" box="[337,410,656,679]" name="Oman" pageId="3" pageNumber="2">Oman</collectingCountry>
|
||
,
|
||
</emphasis>
|
||
<collectingRegion id="49DAF857834CFFDCC0E7F9D0FDC5FD29" box="[425,546,656,679]" country="Oman" name="Musandam" pageId="3" pageNumber="2">Musandam</collectingRegion>
|
||
, mountain road SW of
|
||
<location id="8EC1606E834CFFDCC18EF9EFFEAEFD48" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC18EF9EFFEAEFD48" box="[192,329,687,710]" country="Oman" latitude="56.324" name="Khawr Najd" pageId="3" pageNumber="2" stateProvince="Musandam">Khawr Najd</location>
|
||
, west slope,
|
||
<collectionCode id="ED0FAE70834CFFDCC097F9EFFDE7FD48" box="[473,512,687,710]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
|
||
detection, rocky road cut, bare rock face, 26°05.12'
|
||
<geoCoordinate id="EE2A5072834CFFDCC09CF98EFDBFFD6B" box="[466,600,718,741]" degrees="56" direction="north" minutes="19.44" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="56.324">N 56°19.44'</geoCoordinate>
|
||
E,
|
||
<elevation id="0033D186834CFFDCC338F98EFD10FD6B" box="[630,759,718,741]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="2" unit="m" value="220.0">
|
||
<quantity id="4CE69B50834CFFDCC338F98EFD5BFD6B" box="[630,700,718,741]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="2" unit="m" value="220.0">220 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC249F98EFE96FC8D" pageId="3" pageNumber="2" value="1994-09-30">
|
||
<collectingDate id="EFE4E99D834CFFDCC249F98EFE96FC8D" pageId="3" pageNumber="2" value="1994-09-30">30 September 1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB5363834CFFDCC0E7F9ACFDE0FC8D" box="[425,519,748,771]" pageId="3" pageNumber="2">G. Lowe</collectorName>
|
||
(
|
||
<collectionCode id="ED0FAE70834CFFDCC35BF9ACFDB0FC8D" box="[533,599,748,771]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="2">NMB</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834CFFDCC18EF86CFDE5FA26" blockId="3.[192,802,812,1448]" pageId="3" pageNumber="2">
|
||
<materialsCitation id="3B763CE8834CFFDCC18EF86CFED8FCEC" collectionCode="UV" collectorName="Al Awabi" country="Oman" location="Wadi Mistal" municipality="Rte" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<emphasis id="B96AEAA7834CFFDCC18EF86CFECBFCCD" box="[192,300,812,835]" italics="true" pageId="3" pageNumber="2">
|
||
<typeStatus id="54A58817834CFFDCC18EF86CFECBFCCD" box="[192,300,812,835]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA7834CFFDCC00CF86CFE6EFCCD" bold="true" box="[322,393,812,835]" pageId="3" pageNumber="2">
|
||
<collectingCountry id="F3097625834CFFDCC00CF86CFE6EFCCD" box="[322,393,812,835]" name="Oman" pageId="3" pageNumber="2">Oman</collectingCountry>
|
||
</emphasis>
|
||
,
|
||
<specimenCount id="9D18FD3C834CFFDCC0D0F86CFE28FCCD" box="[414,463,812,835]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC5ACCF834CFFDCC0AAF86CFDEEFCCD" box="[484,521,812,835]" pageId="3" pageNumber="2">Rte</collectingMunicipality>
|
||
13,
|
||
<location id="8EC1606E834CFFDCC306F86CFD33FCCD" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC306F86CFD33FCCD" box="[584,724,812,835]" country="Oman" municipality="Rte" name="Wadi Mistal" pageId="3" pageNumber="2">Wadi Mistal</location>
|
||
to
|
||
<collectorName id="26EB5363834CFFDCC249F86CFEEDFCEC" pageId="3" pageNumber="2">Al Awabi</collectorName>
|
||
,
|
||
<collectionCode id="ED0FAE70834CFFDCC056F80BFED8FCEC" box="[280,319,843,866]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B763CE8834CFFDCC007F80BFEF5FC30" collectingDate="1994-09-26" collectionCode="NMB" collectorName="G. Lowe & M. D. Gallagher" country="Oman" elevation="400" latitude="57.594" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
detection, crevice in rocky road cut, rocky cliff, 23°19.18'
|
||
<geoCoordinate id="EE2A5072834CFFDCC022F829FE1AFC0E" box="[364,509,873,896]" degrees="57" direction="north" minutes="35.64" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="57.594">N 57°35.64'</geoCoordinate>
|
||
E, ca.
|
||
<elevation id="0033D186834CFFDCC319F829FD0AFC0E" box="[599,749,873,896]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="2" unit="m" value="400.0">
|
||
<quantity id="4CE69B50834CFFDCC319F829FD4FFC0E" box="[599,680,873,896]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="2" unit="m" value="400.0">400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC249F829FE9DFC11" pageId="3" pageNumber="2" value="1994-09-26">
|
||
<collectingDate id="EFE4E99D834CFFDCC249F829FE9DFC11" pageId="3" pageNumber="2" value="1994-09-26">26 September 1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB5363834CFFDCC08BF8C8FDCBFC11" box="[453,556,904,927]" pageId="3" pageNumber="2">G. Lowe</collectorName>
|
||
&
|
||
<collectorName id="26EB5363834CFFDCC32EF8C8FCC6FC11" box="[608,801,904,927]" pageId="3" pageNumber="2">M. D. Gallagher</collectorName>
|
||
(
|
||
<collectionCode id="ED0FAE70834CFFDCC186F8E7FEEDFC30" box="[200,266,935,958]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="2">NMB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B763CE8834CFFDCC06FF8E7FD37FB94" collectingDate="1994-09-28" collectionCode="ONHM" collectorName="G. Lowe" country="Oman" elevation="750" latitude="56.2115" location="S of Khasab" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Musandam" typeStatus="paratype">
|
||
<specimenCount id="9D18FD3C834CFFDCC06FF8E7FEAAFC30" box="[289,333,935,958]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49DAF857834CFFDCC010F8E7FE30FC30" box="[350,471,935,958]" country="Oman" name="Musandam" pageId="3" pageNumber="2">Musandam</collectingRegion>
|
||
, mountain road
|
||
<location id="8EC1606E834CFFDCC3DEF8E7FCF9FC30" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC3DEF8E7FCF9FC30" box="[656,798,935,958]" country="Oman" latitude="56.2115" name="S of Khasab" pageId="3" pageNumber="2" stateProvince="Musandam">S of Khasab</location>
|
||
, ultraviolet detection, in crack under rock at base of rocky cliff, road cut, 26°01.81'
|
||
<geoCoordinate id="EE2A5072834CFFDCC369F8A4FD55FC75" box="[551,690,996,1019]" degrees="56" direction="north" minutes="12.69" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="56.2115">N 56°12.69'</geoCoordinate>
|
||
E,
|
||
<elevation id="0033D186834CFFDCC398F8A4FF15FB94" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="2" unit="m" value="750.0">
|
||
<quantity id="4CE69B50834CFFDCC398F8A4FCC5FC75" box="[726,802,996,1019]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="2" unit="m" value="750.0">750 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC045FF43FE12FB94" box="[267,501,1027,1050]" pageId="3" pageNumber="2" value="1994-09-28">
|
||
<collectingDate id="EFE4E99D834CFFDCC045FF43FE12FB94" box="[267,501,1027,1050]" pageId="3" pageNumber="2" value="1994-09-28">28 September 1994</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EB5363834CFFDCC345FF43FD82FB94" box="[523,613,1027,1050]" pageId="3" pageNumber="2">G.Lowe</collectorName>
|
||
,
|
||
<collectionCode id="ED0FAE70834CFFDCC335FF43FD37FB94" box="[635,720,1027,1050]" pageId="3" pageNumber="2">ONHM</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B763CE8834CFFDCC3A9FF43FD57FBB6" collectionCode="UV" country="Oman" location="S of Khasab" pageId="3" pageNumber="2" specimenCount="2" specimenCount-male="2" stateProvince="Musandam" typeStatus="paratype">
|
||
<specimenCount id="9D18FD3C834CFFDCC3A9FF43FCFCFB94" box="[743,795,1027,1050]" count="2" pageId="3" pageNumber="2" type="male">2 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49DAF857834CFFDCC18EFF61FEDEFBB6" box="[192,313,1057,1080]" country="Oman" name="Musandam" pageId="3" pageNumber="2">Musandam</collectingRegion>
|
||
, mountain road
|
||
<location id="8EC1606E834CFFDCC0A1FF61FD9CFBB6" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC0A1FF61FD9CFBB6" box="[495,635,1057,1080]" country="Oman" name="S of Khasab" pageId="3" pageNumber="2" stateProvince="Musandam">S of Khasab</location>
|
||
,
|
||
<collectionCode id="ED0FAE70834CFFDCC3C7FF61FD57FBB6" box="[649,688,1057,1080]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B763CE8834CFFDCC3F7FF61FD59FBF8" collectingDate="1994-09-28" collectionCode="NMB" collectorName="G. Lowe" country="Oman" elevation="980" latitude="56.2115" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
detection, rocky road cuts, 26°00.32'
|
||
<geoCoordinate id="EE2A5072834CFFDCC095FF00FDB8FBD9" box="[475,607,1088,1111]" degrees="56" direction="north" minutes="12.69" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="56.2115">N 56°12.69'</geoCoordinate>
|
||
E,
|
||
<elevation id="0033D186834CFFDCC335FF00FD1EFBD9" box="[635,761,1088,1111]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="2" unit="m" value="980.0">
|
||
<quantity id="4CE69B50834CFFDCC335FF00FD27FBD9" box="[635,704,1088,1111]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="3" pageNumber="2" unit="m" value="980.0">980 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC249FF00FE9FFBF8" pageId="3" pageNumber="2" value="1994-09-28">
|
||
<collectingDate id="EFE4E99D834CFFDCC249FF00FE9FFBF8" pageId="3" pageNumber="2" value="1994-09-28">28 September 1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB5363834CFFDCC08EFF1FFDC2FBF8" box="[448,549,1119,1142]" pageId="3" pageNumber="2">G. Lowe</collectorName>
|
||
(
|
||
<collectionCode id="ED0FAE70834CFFDCC375FF1FFD9BFBF8" box="[571,636,1119,1142]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="2">NMB</collectionCode>
|
||
, GL)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B763CE8834CFFDCC39EFF1FFE6BFB3D" collectionCode="UV" collectorName="Bukha" country="Oman" location="Hanah" municipality="Khasab" pageId="3" pageNumber="2" specimenCount="3" specimenCount-female="1" specimenCount-immature="1" specimenCount-male="1" stateProvince="Musandam" typeStatus="paratype">
|
||
<specimenCount id="9D18FD3C834CFFDCC39EFF1FFCE7FBF8" box="[720,768,1119,1142]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D18FD3C834CFFDCC25AFF1FFEC1FB1A" count="1" pageId="3" pageNumber="2" type="immature">1 immature</specimenCount>
|
||
<specimenCount id="9D18FD3C834CFFDCC07CFF3DFEA1FB1A" box="[306,326,1149,1172]" count="1" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
,
|
||
<collectingRegion id="49DAF857834CFFDCC019FF3DFE37FB1A" box="[343,464,1149,1172]" country="Oman" name="Musandam" pageId="3" pageNumber="2">Musandam</collectingRegion>
|
||
, road from
|
||
<collectingMunicipality id="6BC5ACCF834CFFDCC314FF3DFD4DFB1A" box="[602,682,1149,1172]" pageId="3" pageNumber="2">Khasab</collectingMunicipality>
|
||
to
|
||
<collectorName id="26EB5363834CFFDCC398FF3DFCF9FB1A" box="[726,798,1149,1172]" pageId="3" pageNumber="2">Bukha</collectorName>
|
||
, east of
|
||
<location id="8EC1606E834CFFDCC05FFFDCFEBEFB3D" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC05FFFDCFEBEFB3D" box="[273,345,1180,1203]" country="Oman" municipality="Khasab" name="Hanah" pageId="3" pageNumber="2" stateProvince="Musandam">Hanah</location>
|
||
,
|
||
<collectionCode id="ED0FAE70834CFFDCC02BFFDCFE6BFB3D" box="[357,396,1180,1203]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B763CE8834CFFDCC0DAFFDCFEB1FAA0" collectingDate="1994-09-29" collectionCode="NMB" collectorName="G. Lowe" country="Oman" elevation="100" latitude="56.20933" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
detection, female in crevice in rocky road cut, damaged during extraction from rock, male on rock surface, rocky road cut, mountain pass, 26°13.98'
|
||
<geoCoordinate id="EE2A5072834CFFDCC243FF99FECEFA81" degrees="56" direction="north" minutes="12.56" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="56.20933">N 56°12.56'</geoCoordinate>
|
||
E,
|
||
<elevation id="0033D186834CFFDCC004FFB8FE35FA81" box="[330,466,1272,1295]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="2" unit="m" value="100.0">
|
||
<quantity id="4CE69B50834CFFDCC004FFB8FE75FA81" box="[330,402,1272,1295]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="2" unit="m" value="100.0">100 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC0ABFFB8FD24FA81" box="[485,707,1272,1295]" pageId="3" pageNumber="2" value="1994-09-29">
|
||
<collectingDate id="EFE4E99D834CFFDCC0ABFFB8FD24FA81" box="[485,707,1272,1295]" pageId="3" pageNumber="2" value="1994-09-29">29 September 1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB5363834CFFDCC246FFB8FF1AFAA0" pageId="3" pageNumber="2">G. Lowe</collectorName>
|
||
(
|
||
<collectionCode id="ED0FAE70834CFFDCC042FE57FEA9FAA0" box="[268,334,1303,1326]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="2">NMB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B763CE8834CFFDCC02FFE57FECEFAC2" collectionCode="UV" country="Oman" location="S of Khawr Najd" pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Musandam" typeStatus="paratype">
|
||
<specimenCount id="9D18FD3C834CFFDCC02FFE57FE6DFAA0" box="[353,394,1303,1326]" count="1" pageId="3" pageNumber="2" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49DAF857834CFFDCC0D6FE57FDF5FAA0" box="[408,530,1303,1326]" country="Oman" name="Musandam" pageId="3" pageNumber="2">Musandam</collectingRegion>
|
||
; mountain rd
|
||
<location id="8EC1606E834CFFDCC3E8FE57FF10FAC2" LSID="urn:lsid:plazi:treatment:03B787A3834CFFD2C056FAB0FD94FEF9:8EC1606E834CFFDCC3E8FE57FF10FAC2" country="Oman" name="S of Khawr Najd" pageId="3" pageNumber="2" stateProvince="Musandam">S of Khawr Najd</location>
|
||
,
|
||
<collectionCode id="ED0FAE70834CFFDCC04CFE75FECEFAC2" box="[258,297,1333,1356]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="3" pageNumber="2">UV</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B763CE8834CFFDCC061FE75FD69FA04" collectingDate="1994-09-30" collectionCode="NMB" collectorName="G. Lowe" country="Oman" elevation="5" latitude="56.3235" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
|
||
detection, rocky road cut, split rock face, ca.
|
||
<quantity id="4CE69B50834CFFDCC25AFE75FF32FAE5" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="2" unit="m" value="5.0">
|
||
<elevation id="0033D186834CFFDCC25AFE75FF32FAE5" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="2" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
above road, 26°05.39'
|
||
<geoCoordinate id="EE2A5072834CFFDCC080FE14FDB3FAE5" box="[462,596,1364,1387]" degrees="56" direction="north" minutes="19.41" orientation="latitude" pageId="3" pageNumber="2" precision="9" value="56.3235">N 56°19.41'</geoCoordinate>
|
||
E,
|
||
<elevation id="0033D186834CFFDCC33DFE14FD10FAE5" box="[627,759,1364,1387]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="2" unit="m" value="275.0">
|
||
<quantity id="4CE69B50834CFFDCC33DFE14FD5DFAE5" box="[627,698,1364,1387]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="3" pageNumber="2" unit="m" value="275.0">275 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="FFA01075834CFFDCC249FE14FE9DFA04" pageId="3" pageNumber="2" value="1994-09-30">
|
||
<collectingDate id="EFE4E99D834CFFDCC249FE14FE9DFA04" pageId="3" pageNumber="2" value="1994-09-30">30 September 1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB5363834CFFDCC08BFE33FDCAFA04" box="[453,557,1395,1418]" pageId="3" pageNumber="2">G. Lowe</collectorName>
|
||
(
|
||
<collectionCode id="ED0FAE70834CFFDCC30AFE33FD61FA04" box="[580,646,1395,1418]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="3" pageNumber="2">NMB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B763CE8834CFFDCC3ECFE33FE19FA26" collectingDate="1994-09-30" collectionCode="ONHM" collectorName="G. Lowe" country="Oman" elevation="220" latitude="56.324" location="Khawr Najd" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Musandam" typeStatus="paratype">
|
||
<specimenCount id="9D18FD3C834CFFDCC3ECFE33FD33FA04" box="[674,724,1395,1418]" count="1" pageId="3" pageNumber="2" type="male">1 ♂</specimenCount>
|
||
, same locality as holotype (
|
||
<collectionCode id="ED0FAE70834CFFDCC0D1FED1FE10FA26" box="[415,503,1425,1448]" pageId="3" pageNumber="2">ONHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E834CFFDCC18EFE8FFE93F9CC" pageId="3" pageNumber="2" type="etymology">
|
||
<paragraph id="8BA136B5834CFFDCC18EFE8FFE93F9CC" blockId="3.[192,802,1487,1602]" pageId="3" pageNumber="2">
|
||
<emphasis id="B96AEAA7834CFFDCC18EFE8FFEA5FA68" bold="true" box="[192,322,1487,1510]" pageId="3" pageNumber="2">Etymology.</emphasis>
|
||
The specific epithet is derived by combining the Greek nouns,
|
||
<emphasis id="B96AEAA7834CFFDCC369FEADFD67F98A" box="[551,640,1517,1540]" italics="true" pageId="3" pageNumber="2">nematos</emphasis>
|
||
(thread) and
|
||
<emphasis id="B96AEAA7834CFFDCC18EFD4CFEFEF9AD" box="[192,281,1548,1571]" italics="true" pageId="3" pageNumber="2">daktylos</emphasis>
|
||
(finger), a reference to the long, tenuous pedipalp fingers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E834CFFDBC18EFD28FD54FC88" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="8BA136B5834CFFDCC18EFD28FB9AFC75" blockId="3.[192,802,1640,1908]" lastBlockId="3.[830,1441,199,1019]" pageId="3" pageNumber="2">
|
||
<emphasis id="B96AEAA7834CFFDCC18EFD28FED2F9F1" bold="true" box="[192,309,1640,1663]" pageId="3" pageNumber="2">Diagnosis.</emphasis>
|
||
Medium-sized
|
||
<taxonomicName id="4C1E4D36834CFFDCC0AAFD28FD64F9F1" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[484,643,1640,1663]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA7834CFFDCC0AAFD28FD64F9F1" box="[484,643,1640,1663]" italics="true" pageId="3" pageNumber="2">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
, adults
|
||
<quantity id="4CE69B50834CFFDCC391FD28FF0AF913" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="4.3" metricValueMin="3.1" pageId="3" pageNumber="2" unit="mm" value="37.0" valueMax="43.0" valueMin="31.0">31–43 mm</quantity>
|
||
; body strongly compressed dorsoventrally; color pale yellow with or without faint fuscous markings on carapace, tergites and legs; carapace nearly parallelsided; carapace and tergites with weak or moderate carinae; carapace with central median carinae curved, partially fused or separated from posterior median carinae, anterior median and central lateral carinae weak or obsolete; pedipalps very slender, femur L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC3E4FC1DFD0AF8FA" box="[682,749,1885,1908]" degrees="3.0" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-3.0">W 3.0</geoCoordinate>
|
||
–4.4, patella L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC2EBFB87FC0FFF50" box="[933,1000,199,222]" degrees="3.5" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-3.5">W 3.5</geoCoordinate>
|
||
–3.9, chela L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC533FB87FB26FF50" box="[1149,1217,199,222]" degrees="7.4" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-7.4">W 7.4</geoCoordinate>
|
||
–8.4; pedipalp femur and patella with smooth intercarinal surfaces, patella with dorsomedian and dorsoexternal carinae very weak, nearly obsolete; chela manus not swollen, smooth with all carinae obsolete, fingers very long and tenuous, movable finger L/manus ventral L 3.2–3.8; chela fingers with primary denticles on fixed and movable fingers divided into 11–12 subrows, lacking external accessory denticles, with full compliment of internal accessory denticles (i.e. a member of the ‘acutecarinatus’ group of
|
||
<taxonomicName id="4C1E4D36834CFFDCC270FABAFB0EFD9F" authority="(Levy & Amitai, 1980))" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[830,1257,506,529]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA7834CFFDCC270FABAFC3AFD9F" box="[830,989,506,529]" italics="true" pageId="3" pageNumber="2">Compsobuthus</emphasis>
|
||
(Levy & Amitai, 1980))
|
||
</taxonomicName>
|
||
; trichobothria
|
||
<emphasis id="B96AEAA7834CFFDCC4CBFABAFA47FD9F" box="[1413,1440,506,529]" italics="true" pageId="3" pageNumber="2">db</emphasis>
|
||
and
|
||
<emphasis id="B96AEAA7834CFFDCC23CF958FC60FDA1" box="[882,903,536,559]" italics="true" pageId="3" pageNumber="2">dt</emphasis>
|
||
located in distal half of fixed finger; pedipalp fingers without scalloping on proximal margins; legs long, slender, laterigrade; tergites I–VI with several macrosetae on posterior margins; sternite VI smooth with carinae weak or obsolete; sternite VII smooth, tetracarinate, with weak median carinae and moderate, granulated lateral carinae; metasomal carinae moderately developed, smooth to granulate; median lateral carinae present on metasoma I, absent on metasoma II–IV; metasomal slender, elongate, metasoma IV L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC464F86CFA8CFCCD" box="[1322,1387,812,835]" degrees="2.3" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-2.3">W 2.3</geoCoordinate>
|
||
–2.7, metasoma V L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC2B4F80BFBAFFCEC" box="[1018,1096,843,866]" degrees="2.7" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-2.7">W 2.7</geoCoordinate>
|
||
–3.3; carinae and intercarinal surfaces of metasoma and sternite VII bearing numerous long, reddish macrosetae; telson moderately elongate (L/D 2.9–3.3), with very weak subaculear tubercle, bearing several long reddish macrosetae; pectine teeth: males, 31–33, females 28–34.
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834CFFDBC270FF62FD54FC88" blockId="3.[830,1441,1058,1909]" lastBlockId="4.[192,802,199,774]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
|
||
<emphasis id="B96AEAA7834CFFDCC270FF62FC3AFBB7" bold="true" box="[830,989,1058,1081]" pageId="3" pageNumber="2">Comparisons.</emphasis>
|
||
<taxonomicName id="4C1E4D36834CFFDCC2A4FF62FB5FFBB7" authorityName="Lowe" authorityYear="2009" box="[1002,1208,1058,1081]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA7834CFFDCC2A4FF62FC1BFBB7" box="[1002,1020,1058,1081]" italics="true" pageId="3" pageNumber="2">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA7834CFFDCC55EFF62FB5FFBB7" box="[1040,1208,1058,1081]" italics="true" pageId="3" pageNumber="2">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
is clearly differentiated from other
|
||
<taxonomicName id="4C1E4D36834CFFDCC2BAFF00FB74FBD9" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[1012,1171,1088,1111]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA7834CFFDCC2BAFF00FB74FBD9" box="[1012,1171,1088,1111]" italics="true" pageId="3" pageNumber="2">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
by having (i) the central median carinae on the carapace curved and partially or weakly fused with the linear posterior median carinae, and (ii) very slender pedipalp chelae with extremely elongated pedipalp fingers greatly exceeding the length of the manus. The only other known member of the genus with similar pedipalp proportions is
|
||
<taxonomicName id="4C1E4D36834CFFDCC447FFB8FC92FAA0" authority="Levy" authorityName="Levy" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="longipalpis">
|
||
<emphasis id="B96AEAA7834CFFDCC447FFB8FAFCFA81" box="[1289,1307,1272,1295]" italics="true" pageId="3" pageNumber="2">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA7834CFFDCC466FFB8FA47FA81" box="[1320,1440,1272,1295]" italics="true" pageId="3" pageNumber="2">longipalpis</emphasis>
|
||
Levy
|
||
</taxonomicName>
|
||
et. al. 1973, which differs in having dark pigment on metasoma V, less elongated pedipalp chelae (L/
|
||
<geoCoordinate id="EE2A5072834CFFDCC4C8FE76FC96FAE5" degrees="6.86" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="555" value="-6.86">W 6.86</geoCoordinate>
|
||
), chela fingers with external accessory denticles present, 14 subrows of denticles on the fingers, and fixed finger trichobothrium
|
||
<emphasis id="B96AEAA7834CFFDCC575FED2FBBEFA27" box="[1083,1113,1426,1449]" italics="true" pageId="3" pageNumber="2">est</emphasis>
|
||
closer to
|
||
<emphasis id="B96AEAA7834CFFDCC597FED2FB13FA27" box="[1241,1268,1426,1449]" italics="true" pageId="3" pageNumber="2">db</emphasis>
|
||
than
|
||
<emphasis id="B96AEAA7834CFFDCC471FED2FAB3FA27" box="[1343,1364,1426,1449]" italics="true" pageId="3" pageNumber="2">dt</emphasis>
|
||
(Hendrixson, 2006). Other
|
||
<taxonomicName id="4C1E4D36834CFFDCC579FEF0FB31FA49" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[1079,1238,1456,1479]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA7834CFFDCC579FEF0FB31FA49" box="[1079,1238,1456,1479]" italics="true" pageId="3" pageNumber="2">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
from the Arabian Peninsula without external accessory denticles but with internal accessory denticles on the pedipalp fingers (Kovařík, 2003) are differentiated as follows:
|
||
<taxonomicName id="4C1E4D36834CFFDCC416FD4CFB14F9CC" authority="(Simon, 1882)" baseAuthorityName="Simon" baseAuthorityYear="1882" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="acutecarinatus">
|
||
<emphasis id="B96AEAA7834CFFDCC416FD4CFBA0F9CC" italics="true" pageId="3" pageNumber="2">Compsobuthus acutecarinatus</emphasis>
|
||
(Simon, 1882)
|
||
</taxonomicName>
|
||
: pedipalp chela with manus swollen, with smooth carinae, fingers much shorter, movable finger with 10 subrows of denticles, proximal margin of fingers weakly scalloped, sternite VI with 4 well developed carinae, carapace and tergites with strong carinae, telson vesicle bulbous, and fewer pectine teeth (males 23–29, females 20–27);
|
||
<taxonomicName id="4C1E4D36834CFFDCC416FDA3FB6EF897" authority="Lowe, 2001" authorityName="Lowe" authorityYear="2001" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="polisi">
|
||
<emphasis id="B96AEAA7834CFFDCC416FDA3FC0DF897" italics="true" pageId="3" pageNumber="2">Compsobuthus polisi</emphasis>
|
||
Lowe, 2001
|
||
</taxonomicName>
|
||
: smaller species, fewer pectine teeth (males 18–20, females 16–20), metasoma finely granulated, median lateral carinae well developed on metasoma II, telson more slender with prominent subaculear tubercle;
|
||
<taxonomicName id="4C1E4D36834BFFDBC0E1FB87FEB6FF73" authority="(Kraepelin, 1900)" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="maindroni">
|
||
<emphasis id="B96AEAA7834BFFDBC0E1FB87FD2BFF50" box="[431,716,199,222]" italics="true" pageId="4" pageNumber="3">Compsobuthus maindroni</emphasis>
|
||
(Kraepelin, 1900)
|
||
</taxonomicName>
|
||
: smaller, narrower body, color darker, orange-brown with fuscous metasoma V, pedipalp chela with granulated carinae, movable finger denticles divided into 9 subrows, sternite VI with 4 carinae, metasoma with surfaces finely granulose, macrosetae sparse or absent, median lateral carinae well developed and complete on metasoma II–III, fewer pectine teeth (males 21–22, females 19–21);
|
||
<taxonomicName id="4C1E4D36834BFFDBC352FAFCFDC6FE7C" authority="(Levy, Amitai et Shulov, 1973)" baseAuthorityName="Levy, Amitai et Shulov" baseAuthorityYear="1973" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="arabicus">
|
||
<emphasis id="B96AEAA7834BFFDBC352FAFCFCC6FE5D" box="[540,801,444,467]" italics="true" pageId="4" pageNumber="3">Compsobuthus arabicus</emphasis>
|
||
(Levy, Amitai et Shulov, 1973)
|
||
</taxonomicName>
|
||
: much smaller species, carapace with fuscous interocular markings, pedipalp chelae with distinct carinae, fewer pectine teeth (9–15), legs short and stout, sternite VI with distinct lateral carinae, sternite VII finely granular; metasoma densely, finely granular without numerous intercarinal macrosetae;
|
||
<taxonomicName id="4C1E4D36834BFFDBC043F9D3FD58FD24" authority="Sissom, 1994" authorityName="Sissom" authorityYear="1994" box="[269,703,659,682]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="vachoni">
|
||
<emphasis id="B96AEAA7834BFFDBC043F9D3FDF6FD24" box="[269,529,659,682]" italics="true" pageId="4" pageNumber="3">Compsobuthus vachoni</emphasis>
|
||
Sissom, 1994
|
||
</taxonomicName>
|
||
: smaller species, fewer pectine teeth (female 15), pedipalp movable finger with 9 subrows of primary denticles, trichobothrium
|
||
<emphasis id="B96AEAA7834BFFDBC029F9AFFE65FC88" box="[359,386,751,774]" italics="true" pageId="4" pageNumber="3">db</emphasis>
|
||
in basal half of fixed finger.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E834BFFD5C18EF86DFC4FF929" lastPageId="10" lastPageNumber="9" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="8BA136B5834BFFDBC18EF86DFDC0FCC9" blockId="4.[192,551,813,839]" box="[192,551,813,839]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC18EF86DFDC0FCC9" bold="true" box="[192,551,813,839]" pageId="4" pageNumber="3">
|
||
Description of
|
||
<typeStatus id="54A58817834BFFDBC036F86DFE04FCC9" box="[376,483,813,839]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
male
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFDBC18EF82DFD6FFB38" blockId="4.[192,802,877,1206]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC18EF82DFEA5FC0A" bold="true" box="[192,322,877,900]" pageId="4" pageNumber="3">Coloration.</emphasis>
|
||
Base colour pale translucent yellow; with faint fuscosity on lateral areas of carapace and tergites and anterior margin of carapace, darker pigmentation concentrated on posterior margins of carapace and tergites I–VI, and on dorsal retrolateral and ventral prolateral surfaces of femora and patellae of legs I–IV; median and lateral eyes with underlying melanic pigment; denticles of chelicerae and pedipalp fingers, articular condyles of pedipalp movable fingers, tarsal ungues and aculeus of telson castaneous; macrosetae on body and appendages a dark reddish color.
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFDBC18EFF9DFBA1FF73" blockId="4.[192,802,1245,1912]" lastBlockId="4.[830,1440,199,253]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC18EFF9DFE7AFB7A" bold="true" box="[192,413,1245,1268]" pageId="4" pageNumber="3">
|
||
Carapace (
|
||
<figureCitation id="13252A30834BFFDBC00AFF9DFE69FB7A" box="[324,398,1245,1268]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Weakly quadrate, nearly parallel sided, convergent on anterolateral margins; posterior width nearly equal to length; surface strongly compressed dorsoventrally, planar; median ocular tubercle depressed, level with surrounding surfaces; lateral eyes with 5 ocelli (3 large, 2 small); anterior margin concave, smooth; anterior median carinae granular, weak, confined to posterior part of interocular triangle; superciliary carinae distinct, granular; central median carinae weak, irregularly granulose, curved outward posteriorly; posterior median carinae strong, linear, with coarse regular granulation, terminating posteriorly in spiniform granules projecting beyond posterior margin of carapace; central median and central posterior carinae joined on the left, separated on the right; central lateral and lateral ocular carinae weak, less distinct, marked by irregular series of granules; other carinae indistinct; anterior margin bordered with series of weak, coarse granules; interocular triangle smooth medially, fine granulate laterally; medial and mediolateral surfaces smooth to finely shagreened, lateral flanks with scattered coarse and fine granulation; anterior margin of carapace with 3 pairs of macrosetae, carapace otherwise bare.
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFDBC270FA63FC42FDA1" blockId="4.[830,1440,291,559]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC270FA63FBD4FEB4" bold="true" box="[830,1075,291,314]" pageId="4" pageNumber="3">
|
||
Chelicera (
|
||
<figureCitation id="13252A30834BFFDBC2F2FA63FBC1FEB4" box="[956,1062,291,314]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="4" pageNumber="3">Figs. 2–3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Dorsal surface of manus smooth, glabrous, with pair of short pale microsetae on apical border, long reddish subapical macroseta on dorsointernal carina, and short pale microseta located anteromedially; dorsal surface of movable finger with 2 short pale microsetae; cheliceral dentition with characteristic buthid pattern (Sissom, 1990; Soleglad & Fet, 2003), ventral surface of fixed finger with two enlarged denticles.
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFDBC270F916FA99FCAB" blockId="4.[830,1440,598,805]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC270F916FB47FDE3" bold="true" box="[830,1184,598,621]" pageId="4" pageNumber="3">
|
||
Coxosternal area (
|
||
<figureCitation id="13252A30834BFFDBC55EF916FB73FDE3" box="[1040,1172,598,621]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="4" pageNumber="3">Figs. 19–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Coxal surfaces smooth, anterior margins of coxae II–III and anterior and proximal posterior margins of coxa IV finely shagreened; setation sparse, fewer than 4 macrosetae per coxal segment; sternum triangular, smooth with shallow posterior depression; genital opercula smooth, with 2–4 lateral macrosetae; genital papillae present (in males).
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFDBC270F80DFC47FB18" blockId="4.[830,1441,845,1174]" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC270F80DFB75FCEA" bold="true" box="[830,1170,845,868]" pageId="4" pageNumber="3">
|
||
Pectines (
|
||
<figureCitation id="13252A30834BFFDBC2E2F80DFBD6FCEA" box="[940,1073,845,868]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="4" pageNumber="3">Figs. 10–11</figureCitation>
|
||
,
|
||
<figureCitation id="13252A30834BFFDBC50EF80DFB61FCEA" box="[1088,1158,845,868]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="4" pageNumber="3">19–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Basal piece rectangular, smooth, with deep anteromedian incision, distal tips of pectines extending to apical limit of trochanter IV; pectines with 3 marginal lamellae, 10–11 middle lamellae, and small intermediate lamella at distal end of basal marginal lamella, 32–32 teeth; all lamellae bearing numerous short reddish macrosetae, including 2–5 on fulcra; when anterior margins of left and right pectines are aligned with posterior margins of coxae IV, basal 2 pectine teeth overlap, with a gap between basal middle lamellae.
|
||
</paragraph>
|
||
<paragraph id="8BA136B5834BFFD9C270FFFEFF0BF8F6" blockId="4.[830,1441,1214,1911]" lastBlockId="6.[192,801,1797,1912]" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3">
|
||
<emphasis id="B96AEAA7834BFFDBC270FFFEFBBBFB5B" bold="true" box="[830,1116,1214,1237]" pageId="4" pageNumber="3">
|
||
Mesosoma (
|
||
<figureCitation id="13252A30834BFFDBC286FFFEFBB7FB5B" box="[968,1104,1214,1237]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="4" pageNumber="3">Figs. 18–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<emphasis id="B96AEAA7834BFFDBC528FFFEFB5AFB5B" box="[1126,1213,1214,1237]" italics="true" pageId="4" pageNumber="3">Tergites</emphasis>
|
||
: pretergites smooth; tergite I with single pair of short granulose, lateral carinae, median carina obsolete, marked by single median granule; tergites II–VI with 3 granulose, moderately strong, longitudinal carinae; lateral carinae of all tergites terminating posteriorly in spiniform granules projecting beyond posterior margins of sclerite; median carinae of II–VI slightly projecting beyond posterior margins; lateral carinae of all tergites curved outwards anteriorly; tergite VII with two pairs of strong granular lateral carinae, and weak granular median carina; lateral flanks of tergites I–VI weakly sloped, finely shagreened with scattered coarse granulation; median intercarinal surfaces finely shagreened, without granules; tergite VII finely shagreened with scattered fine granulation on lateral flanks; tergites I–VI with 4 posterior marginal macrosetae, one medial pair on inner side of lateral carinae, one lateral pair (some setae lost, marked by insertion sockets); tergite VII lacking macrosetae.
|
||
<emphasis id="B96AEAA7834BFFDBC284FC44FBC0F895" box="[970,1063,1796,1819]" italics="true" pageId="4" pageNumber="3">Sternites</emphasis>
|
||
: all sternites smooth, sternite III without carinae, carinae on sternites IV–V obsolete; sternite VI with median carinae obsolete, lateral carinae smooth and nearly obsolete; sternite VII with median carinae weak, smooth, lateral carinae moderate, granular; all sternites with scattered, sparse short to long reddish macrosetae, 9 non-marginal setae on sternite VII.
|
||
</paragraph>
|
||
<caption id="DF61663D834AFFDAC18EFC42FEAAF8F2" ID-DOI="http://doi.org/10.5281/zenodo.12782207" ID-Zenodo-Dep="12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,278,1794,1817]" targetBox="[286,1362,229,1696]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BA136B5834AFFDAC18EFC42FEAAF8F2" blockId="5.[192,1441,1794,1916]" pageId="5" pageNumber="4">
|
||
<emphasis id="B96AEAA7834AFFDAC18EFC42FEA9F897" bold="true" box="[192,334,1794,1817]" pageId="5" pageNumber="4">Figures 1–9:</emphasis>
|
||
<taxonomicName id="4C1E4D36834AFFDAC01AFC43FD98F896" authorityName="Lowe" authorityYear="2009" box="[340,639,1795,1816]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA7834AFFDAC01AFC43FD98F896" box="[340,639,1795,1816]" italics="true" pageId="5" pageNumber="4">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA7834AFFDAC3CBFC43FCF2F896" bold="true" box="[645,789,1795,1816]" pageId="5" pageNumber="4">
|
||
<taxonomicNameLabel id="A25957DC834AFFDAC3CBFC43FD34F896" box="[645,723,1795,1816]" pageId="5" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1, 6–9
|
||
</emphasis>
|
||
, holotype male,
|
||
<emphasis id="B96AEAA7834AFFDAC2F8FC43FC3DF896" bold="true" box="[950,986,1795,1816]" pageId="5" pageNumber="4">2–5</emphasis>
|
||
, paratype adult males.
|
||
<emphasis id="B96AEAA7834AFFDAC5F7FC43FB2BF896" bold="true" box="[1209,1228,1795,1816]" pageId="5" pageNumber="4">1.</emphasis>
|
||
Carapace and tergites I–II.
|
||
<emphasis id="B96AEAA7834AFFDAC1BFFC5CFEFDF8BF" bold="true" box="[241,282,1820,1841]" pageId="5" pageNumber="4">2–3.</emphasis>
|
||
Right chelicera, dorsal (2) and ventral (3) aspect.
|
||
<emphasis id="B96AEAA7834AFFDAC24BFC5CFCC9F8BF" bold="true" box="[773,814,1820,1841]" pageId="5" pageNumber="4">4–5.</emphasis>
|
||
Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects.
|
||
<emphasis id="B96AEAA7834AFFDAC0DDFC75FE42F8C4" bold="true" box="[403,421,1845,1866]" pageId="5" pageNumber="4">6.</emphasis>
|
||
Metasoma and telson, right lateral aspect.
|
||
<emphasis id="B96AEAA7834AFFDAC20AFC75FCB0F8C4" bold="true" box="[836,855,1845,1866]" pageId="5" pageNumber="4">7.</emphasis>
|
||
Metasoma and telson, ventral aspect.
|
||
<emphasis id="B96AEAA7834AFFDAC587FC75FB3CF8C4" bold="true" box="[1225,1243,1845,1866]" pageId="5" pageNumber="4">8.</emphasis>
|
||
Right telotarsus III, ventral aspect.
|
||
<emphasis id="B96AEAA7834AFFDAC01AFC0EFE81F8ED" bold="true" box="[340,358,1870,1891]" pageId="5" pageNumber="4">9.</emphasis>
|
||
Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2
|
||
<emphasis id="B96AEAA7834AFFDAC505FC0EFBB0F8ED" bold="true" box="[1099,1111,1870,1891]" pageId="5" pageNumber="4">–</emphasis>
|
||
3, 8
|
||
<emphasis id="B96AEAA7834AFFDAC532FC0EFB6FF8ED" bold="true" box="[1148,1160,1870,1891]" pageId="5" pageNumber="4">–</emphasis>
|
||
9: 1.45 mm; 4
|
||
<emphasis id="B96AEAA7834AFFDAC45FFC0EFAFAF8ED" bold="true" box="[1297,1309,1870,1891]" pageId="5" pageNumber="4">–</emphasis>
|
||
5: 0.725 mm; 6
|
||
<emphasis id="B96AEAA7834AFFDAC182FC27FF3FF8F2" bold="true" box="[204,216,1895,1916]" pageId="5" pageNumber="4">–</emphasis>
|
||
7: 4.17 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF61663D8349FFD9C18EFD19FDE7F950" ID-DOI="http://doi.org/10.5281/zenodo.12782211" ID-Zenodo-Dep="12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,278,1625,1648]" targetBox="[232,1397,207,1539]" targetPageId="6" targetType="figure">
|
||
<paragraph id="8BA136B58349FFD9C18EFD19FDE7F950" blockId="6.[192,1441,1625,1758]" pageId="6" pageNumber="5">
|
||
<emphasis id="B96AEAA78349FFD9C18EFD19FE8DF9FE" bold="true" box="[192,362,1625,1648]" pageId="6" pageNumber="5">Figures 10–17:</emphasis>
|
||
<taxonomicName id="4C1E4D368349FFD9C03DFD1BFD47F9FE" authorityName="Lowe" authorityYear="2009" box="[371,672,1627,1648]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78349FFD9C03DFD1BFD47F9FE" box="[371,672,1627,1648]" italics="true" pageId="6" pageNumber="5">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78349FFD9C3E6FD1BFC84F9FE" bold="true" box="[680,867,1627,1648]" pageId="6" pageNumber="5">
|
||
<taxonomicNameLabel id="A25957DC8349FFD9C3E6FD1BFD1FF9FE" box="[680,760,1627,1648]" pageId="6" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
11, 12–16
|
||
</emphasis>
|
||
, holotype male,
|
||
<emphasis id="B96AEAA78349FFD9C544FD1BFBAFF9FE" bold="true" box="[1034,1096,1627,1648]" pageId="6" pageNumber="5">10, 17</emphasis>
|
||
, paratype adult female (near Wadi Mistal).
|
||
<emphasis id="B96AEAA78349FFD9C05BFD37FED4F902" bold="true" box="[277,307,1655,1676]" pageId="6" pageNumber="5">10.</emphasis>
|
||
Left pectine of female, ventral aspect.
|
||
<emphasis id="B96AEAA78349FFD9C38FFD37FD38F902" bold="true" box="[705,735,1655,1676]" pageId="6" pageNumber="5">11.</emphasis>
|
||
Left pectine of male, ventral aspect.
|
||
<emphasis id="B96AEAA78349FFD9C515FD37FB9EF902" bold="true" box="[1115,1145,1655,1676]" pageId="6" pageNumber="5">12.</emphasis>
|
||
Right pedipalp femur, dorsal aspect.
|
||
<emphasis id="B96AEAA78349FFD9C044FDD2FECEF929" bold="true" box="[266,297,1682,1703]" pageId="6" pageNumber="5">13.</emphasis>
|
||
Right pedipalp patella, dorsal aspect.
|
||
<emphasis id="B96AEAA78349FFD9C3EBFDD2FD24F929" bold="true" box="[677,707,1682,1703]" pageId="6" pageNumber="5">14.</emphasis>
|
||
Right pedipalp patella, external aspect.
|
||
<emphasis id="B96AEAA78349FFD9C51DFDD2FB96F929" bold="true" box="[1107,1137,1682,1703]" pageId="6" pageNumber="5">15.</emphasis>
|
||
Right pedipalp chela, external aspect.
|
||
<emphasis id="B96AEAA78349FFD9C047FDEEFECFF94D" bold="true" box="[265,296,1710,1731]" pageId="6" pageNumber="5">16.</emphasis>
|
||
Right pedipalp chela, ventral aspect.
|
||
<emphasis id="B96AEAA78349FFD9C3D7FDEEFD5FF94D" bold="true" box="[665,696,1710,1731]" pageId="6" pageNumber="5">17.</emphasis>
|
||
Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12
|
||
<emphasis id="B96AEAA78349FFD9C044FD89FEF1F950" bold="true" box="[266,278,1737,1758]" pageId="6" pageNumber="5">–</emphasis>
|
||
16: 2 mm; 17: 4.17 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58349FFD9C270FC45FA79F8F6" blockId="6.[830,1441,1797,1912]" pageId="6" pageNumber="5">
|
||
<emphasis id="B96AEAA78349FFD9C270FC45FA46F892" bold="true" box="[830,1441,1797,1820]" pageId="6" pageNumber="5">
|
||
Hemispermatophore lobes (
|
||
<typeStatus id="54A588178349FFD9C5DCFC45FB1DF892" box="[1170,1274,1797,1820]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
|
||
) (
|
||
<figureCitation id="13252A308349FFD9C450FC45FA76F892" box="[1310,1425,1797,1820]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="6" pageNumber="5">Figs. 4–5</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Inner lobe broad, triangular; median lobe broad, triangular, 0.6 times length of inner lobe; outer lobe narrow, tapering apically, almost as long as inner lobe;
|
||
</paragraph>
|
||
<caption id="DF61663D8348FFD8C18EFFABFECDFA90" ID-DOI="http://doi.org/10.5281/zenodo.12782213" ID-Zenodo-Dep="12782213" httpUri="https://zenodo.org/record/12782213/files/figure.png" pageId="7" pageNumber="6" startId="7.[192,278,1259,1282]" targetBox="[192,1438,192,1227]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8BA136B58348FFD8C18EFFABFECDFA90" blockId="7.[192,1402,1259,1310]" pageId="7" pageNumber="6">
|
||
<emphasis id="B96AEAA78348FFD8C18EFFABFE8FFA8C" bold="true" box="[192,360,1259,1282]" pageId="7" pageNumber="6">Figures 18–19:</emphasis>
|
||
<taxonomicName id="4C1E4D368348FFD8C020FFADFD7DFA8C" authorityName="Lowe" authorityYear="2009" box="[366,666,1261,1282]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78348FFD8C020FFADFD7DFA8C" box="[366,666,1261,1282]" italics="true" pageId="7" pageNumber="6">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78348FFD8C3D1FFADFD0AFA8C" bold="true" box="[671,749,1261,1282]" pageId="7" pageNumber="6">
|
||
<taxonomicNameLabel id="A25957DC8348FFD8C3D1FFADFD0AFA8C" box="[671,749,1261,1282]" pageId="7" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male, habitus.
|
||
<emphasis id="B96AEAA78348FFD8C294FFADFC1FFA8C" bold="true" box="[986,1016,1261,1282]" pageId="7" pageNumber="6">18.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96AEAA78348FFD8C5C2FFADFB4EFA8C" bold="true" box="[1164,1193,1261,1282]" pageId="7" pageNumber="6">19.</emphasis>
|
||
Ventral aspect. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58348FFD8C18EFE04FF1DFAF4" blockId="7.[192,801,1348,1402]" pageId="7" pageNumber="6">basal lobe with broad longitudinal base, and hook-like apex.</paragraph>
|
||
<paragraph id="8BA136B58348FFD8C18EFEE0FBC3FA7A" blockId="7.[192,803,1440,1892]" lastBlockId="7.[830,1440,1348,1524]" pageId="7" pageNumber="6">
|
||
<emphasis id="B96AEAA78348FFD8C18EFEE0FDA9FA39" bold="true" box="[192,590,1440,1463]" pageId="7" pageNumber="6">
|
||
Metasoma (
|
||
<figureCitation id="13252A308348FFD8C003FEE0FE5FFA39" box="[333,440,1440,1463]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="7" pageNumber="6">Figs. 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308348FFD8C083FEE0FE0FFA39" box="[461,488,1440,1463]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="7" pageNumber="6">17</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308348FFD8C0B5FEE0FDD8FA39" box="[507,575,1440,1463]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="7" pageNumber="6">18–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Slender, elongate, exceeding length of prosoma and mesosoma, segments narrowing posteriorly; segment I with 10 carinae, segments II–IV with 8 carinae (median lateral carinae absent), segment V with 5 carinae; dorsosubmedian and dorsolateral carinae strong, crenulate-granulate on all segments; median lateral carinae moderate, crenulate-granulate on I, absent on other segments; ventrolateral carinae strong to moderate, crenulate-granulate all segments; ventromedian carinae weak on I–II, moderate, on III–IV, weakly crenulate; metasoma V with ventromedian carina strong, granular, ventrosubmedian carinae absent, intercarinal surfaces smooth on all segments; lateral anal lobes not dissected; ventral anal arc with fine granules.
|
||
<emphasis id="B96AEAA78348FFD8C07FFC0DFE6FF8EA" box="[305,392,1869,1892]" italics="true" pageId="7" pageNumber="6">Setation</emphasis>
|
||
: dorsosubmedian carinae on metasoma I–VI with irregular (1–5) macrosetae; dorsolateral carinae on metasoma I–VI with irregular macrosetae (1– 3); dorsal surface of metasoma V with linear series of 3– 4 macrosetae flanking dorsolateral carinae; lateral and ventral surfaces, and carinae of all segments with sparse, scattered macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8BA136B58348FFD8C270FD5BFC8BF886" blockId="7.[830,1441,1563,1800]" pageId="7" pageNumber="6">
|
||
<emphasis id="B96AEAA78348FFD8C270FD5BFB68F9BC" bold="true" box="[830,1167,1563,1586]" pageId="7" pageNumber="6">
|
||
Telson (
|
||
<figureCitation id="13252A308348FFD8C2D2FD5BFBE5F9BC" box="[924,1026,1563,1586]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="7" pageNumber="6">Figs. 6–7</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308348FFD8C55AFD5BFBCAF9BC" box="[1044,1069,1563,1586]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="7" pageNumber="6">17</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308348FFD8C571FD5BFB67F9BC" box="[1087,1152,1563,1586]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="7" pageNumber="6">18–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Vesicle pyriform, dorsal and dorsolateral surfaces smooth, without setae; lateral and ventral surfaces with median and lateral longitudinal series of weak bumps corresponding to insertions of fine microsetae, and scattered macrosetae; subaculear tubercle weak, flanked by pair of long curved macrosetae; aculeus long, slender, strongly curved, as long as vesicle.
|
||
</paragraph>
|
||
<paragraph id="8BA136B58348FFD7C270FC6EFBB9F929" blockId="7.[830,1440,1838,1892]" lastBlockId="8.[830,1441,1281,1703]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
|
||
<emphasis id="B96AEAA78348FFD8C270FC6EFB5BF8CB" bold="true" box="[830,1212,1838,1861]" pageId="7" pageNumber="6">
|
||
Pedipalp (
|
||
<figureCitation id="13252A308348FFD8C2F1FC6EFBA8F8CB" box="[959,1103,1838,1861]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="7" pageNumber="6">Figs. 12–16</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308348FFD8C524FC6EFB4BF8CB" box="[1130,1196,1838,1861]" captionStart="Figures 22–23" captionStartId="10.[192,278,733,756]" captionTargetBox="[218,1411,218,671]" captionTargetId="figure-9@10.[192,1438,192,700]" captionTargetPageId="10" captionText="Figures 22–23: Compsobuthus nematodactylus sp. nov. paratype adult male. 22. Pedipalp chela, left movable finger dentition. 23. Pedipalp chela, left fixed finger dentition. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12782219" httpUri="https://zenodo.org/record/12782219/files/figure.png" pageId="7" pageNumber="6">22–23</figureCitation>
|
||
).
|
||
</emphasis>
|
||
<emphasis id="B96AEAA78348FFD8C59FFC6EFAFFF8CB" box="[1233,1304,1838,1861]" italics="true" pageId="7" pageNumber="6">Femur</emphasis>
|
||
(
|
||
<figureCitation id="13252A308348FFD8C47AFC6EFA74F8CB" box="[1332,1427,1838,1861]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="7" pageNumber="6">Fig. 12</figureCitation>
|
||
): slender, 4.18 times longer than wide; dorsoexternal and dorsointernal carinae moderate, finely granular; ventrointernal carina moderate, with larger granulation; other carinae obsolete; dorsal surface flat smooth, external surface convex with dispersed fine granules, ventral surface weakly concave proximally, smooth; internal surface convex with scattered enlarged granules; 9–10 macrosetae scattered on external surface, distal cluster not well defined.
|
||
<emphasis id="B96AEAA78347FFD7C0C0FE98FE3CFA61" box="[398,475,1496,1519]" italics="true" pageId="8" pageNumber="7">Patella</emphasis>
|
||
(
|
||
<figureCitation id="13252A308347FFD7C0BEFE98FD92FA61" box="[496,629,1496,1519]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="8" pageNumber="7">Figs. 13–14</figureCitation>
|
||
): slender, 3.52 times longer than wide; dorsointernal, dorsomedian, ventroexternal and ventrointernal carinae weak with very fine granules; dorsoexternal, external and ventromedian carinae weak, smooth; internal carina moderate, finely crenulate-granulate, with fewer larger granules; all intercarinal surfaces smooth; mostly bare, with two large proximal internal macrosetae and a few short macrosetae.
|
||
<emphasis id="B96AEAA78347FFD7C045FD8DFEACF96A" box="[267,331,1741,1764]" italics="true" pageId="8" pageNumber="7">Chela</emphasis>
|
||
(
|
||
<figureCitation id="13252A308347FFD7C02CFD8DFE0DF96A" box="[354,490,1741,1764]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="8" pageNumber="7">Figs. 15–16</figureCitation>
|
||
): very slender, 7.45 times longer than wide, with long, tenuous fingers, movable finger 3.29 times manus ventral length; surface smooth with all carinae obsolete, proximal margins of fingers not scalloped; sparse short macrosetae scattered on external and ventral surfaces of manus and ventral aspect of movable finger; apex of movable finger with ventral and internal clusters of macrosetae, apex of fixed finger with dorsal and internal clusters of macrosetae; both fixed and movable fingers terminate in enlarged apical tooth capped by smooth patches of thickened cuticle; a pair of short, laminate, reddish setae present sub-distally under enlarged terminal teeth of fixed and movable fingers; movable finger with 4 subdistal denticles; 11–12 linear, non-imbricated primary denticle subrows on fixed and movable fingers (11 subrows correspond to proximal subrow fusions) (
|
||
<figureCitation id="13252A308347FFD7C443FD74FA76F9C5" box="[1293,1425,1588,1611]" captionStart="Figures 22–23" captionStartId="10.[192,278,733,756]" captionTargetBox="[218,1411,218,671]" captionTargetId="figure-9@10.[192,1438,192,700]" captionTargetPageId="10" captionText="Figures 22–23: Compsobuthus nematodactylus sp. nov. paratype adult male. 22. Pedipalp chela, left movable finger dentition. 23. Pedipalp chela, left fixed finger dentition. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12782219" httpUri="https://zenodo.org/record/12782219/files/figure.png" pageId="8" pageNumber="7">Figs. 22–23</figureCitation>
|
||
), separated by enlarged proximal denticles; all denticle subrows flanked by internal accessory denticles, external accessory denticles absent.
|
||
</paragraph>
|
||
<caption id="DF61663D8347FFD7C18EFFE9FDFAFB55" ID-DOI="http://doi.org/10.5281/zenodo.12782215" ID-Zenodo-Dep="12782215" httpUri="https://zenodo.org/record/12782215/files/figure.png" pageId="8" pageNumber="7" startId="8.[192,278,1193,1216]" targetBox="[192,1439,192,1160]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BA136B58347FFD7C18EFFE9FDFAFB55" blockId="8.[192,1440,1193,1243]" pageId="8" pageNumber="7">
|
||
<emphasis id="B96AEAA78347FFD7C18EFFE9FE8DFB4E" bold="true" box="[192,362,1193,1216]" pageId="8" pageNumber="7">Figures 20–21:</emphasis>
|
||
<taxonomicName id="4C1E4D368347FFD7C03FFFEAFD79FB31" authorityName="Lowe" authorityYear="2009" box="[369,670,1194,1215]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78347FFD7C03FFFEAFD79FB31" box="[369,670,1194,1215]" italics="true" pageId="8" pageNumber="7">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78347FFD7C3EBFFEAFD12FB31" bold="true" box="[677,757,1194,1215]" pageId="8" pageNumber="7">
|
||
<taxonomicNameLabel id="A25957DC8347FFD7C3EBFFEAFD12FB31" box="[677,757,1194,1215]" pageId="8" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype adult female (near Wadi Mistal), habitus.
|
||
<emphasis id="B96AEAA78347FFD7C5BCFFEAFAF7FB31" bold="true" box="[1266,1296,1194,1215]" pageId="8" pageNumber="7">20.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96AEAA78347FFD7C18FFF86FF07FB55" bold="true" box="[193,224,1222,1243]" pageId="8" pageNumber="7">21.</emphasis>
|
||
Ventral aspect. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58347FFD7C270FD8DFC59F8F3" blockId="8.[830,1440,1741,1917]" pageId="8" pageNumber="7">
|
||
<emphasis id="B96AEAA78347FFD7C270FD8DFAF4F96A" bold="true" box="[830,1299,1741,1764]" pageId="8" pageNumber="7">
|
||
Trichobothrial pattern (
|
||
<figureCitation id="13252A308347FFD7C53DFD8DFAE4F96A" box="[1139,1283,1741,1764]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="8" pageNumber="7">Figs. 12–16</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Orthobothriotaxic,
|
||
<typeStatus id="54A588178347FFD7C2E6FDACFC31F88D" box="[936,982,1772,1795]" pageId="8" pageNumber="7">type</typeStatus>
|
||
Aβ (Vachon, 1974, 1975); femur
|
||
<emphasis id="B96AEAA78347FFD7C4CDFDACFA77F88D" box="[1411,1424,1772,1795]" italics="true" pageId="8" pageNumber="7">d</emphasis>
|
||
<subScript id="179A34F08347FFD7C4DFFDB6FA7DF88B" attach="right" box="[1425,1434,1782,1797]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
|
||
, patella
|
||
<emphasis id="B96AEAA78347FFD7C2DDFC4AFC47F8AF" box="[915,928,1802,1825]" italics="true" pageId="8" pageNumber="7">d</emphasis>
|
||
<subScript id="179A34F08347FFD7C2EFFC55FC4DF8AA" attach="right" box="[929,938,1813,1828]" fontSize="6" pageId="8" pageNumber="7">2</subScript>
|
||
, chela
|
||
<emphasis id="B96AEAA78347FFD7C54AFC4AFBC5F8AF" box="[1028,1058,1802,1825]" italics="true" pageId="8" pageNumber="7">Eb</emphasis>
|
||
<subScript id="179A34F08347FFD7C56CFC55FBCCF8AA" attach="left" box="[1058,1067,1813,1828]" fontSize="6" pageId="8" pageNumber="7">3</subScript>
|
||
,
|
||
<emphasis id="B96AEAA78347FFD7C50EFC4AFB8FF8AF" box="[1088,1128,1802,1825]" italics="true" pageId="8" pageNumber="7">Esb</emphasis>
|
||
and
|
||
<emphasis id="B96AEAA78347FFD7C5E4FC4AFB29F8AF" box="[1194,1230,1802,1825]" italics="true" pageId="8" pageNumber="7">esb</emphasis>
|
||
petite;
|
||
<emphasis id="B96AEAA78347FFD7C462FC4AFAADF8AF" box="[1324,1354,1802,1825]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
and
|
||
<emphasis id="B96AEAA78347FFD7C4C3FC4AFA47F8AF" box="[1421,1440,1802,1825]" italics="true" pageId="8" pageNumber="7">et</emphasis>
|
||
located on fixed finger between
|
||
<emphasis id="B96AEAA78347FFD7C5F3FC69FB3FF8CE" box="[1213,1240,1833,1856]" italics="true" pageId="8" pageNumber="7">db</emphasis>
|
||
and
|
||
<emphasis id="B96AEAA78347FFD7C455FC69FAD7F8CE" box="[1307,1328,1833,1856]" italics="true" pageId="8" pageNumber="7">dt</emphasis>
|
||
, with
|
||
<emphasis id="B96AEAA78347FFD7C4CCFC69FA47F8CE" box="[1410,1440,1833,1856]" italics="true" pageId="8" pageNumber="7">est</emphasis>
|
||
approximately midway between
|
||
<emphasis id="B96AEAA78347FFD7C5EDFC08FB59F8D1" box="[1187,1214,1864,1887]" italics="true" pageId="8" pageNumber="7">db</emphasis>
|
||
and
|
||
<emphasis id="B96AEAA78347FFD7C5B8FC08FAEEF8D1" box="[1270,1289,1864,1887]" italics="true" pageId="8" pageNumber="7">dt</emphasis>
|
||
, but closer to
|
||
<emphasis id="B96AEAA78347FFD7C270FC26FCB4F8F3" box="[830,851,1894,1917]" italics="true" pageId="8" pageNumber="7">dt</emphasis>
|
||
(
|
||
<figureCitation id="13252A308347FFD7C22FFC26FC48F8F3" box="[865,943,1894,1917]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="8" pageNumber="7">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C209FB89FA88FEAB" blockId="9.[212,1398,201,1234]" pageId="9" pageNumber="8">
|
||
<emphasis id="B96AEAA78346FFD6C209FB89FAF7FF6D" bold="true" box="[839,1296,201,227]" pageId="9" pageNumber="8">Males Females</emphasis>
|
||
Carapace posterior W/ L 0.94 – 0.9 9, 0.96 ± 0.02 (4) 0.99 – 1.01, 1.00 ± 0.01 (3) Pedipalp movable finger L/manus ventral L 3.19 – 3.29, 3.22 ± 0.04 (4) 3.61 – 3.83, 3.73 ± 0.09 (4)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFA7AFA88FEDF" blockId="9.[212,1398,201,1234]" box="[213,1391,314,337]" pageId="9" pageNumber="8">
|
||
Telson vesicle D/
|
||
<geoCoordinate id="EE2A50728346FFD6C0D8FA7AFCEEFEDF" box="[406,777,314,337]" degrees="0.98" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.98">W 0.98</geoCoordinate>
|
||
– 1.00, 0.99 ± 0.01 (4) 0.92 – 0.97, 0.94 ± 0.02 (4)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFA25FA91FEF2" blockId="9.[212,1398,201,1234]" box="[213,1398,357,380]" pageId="9" pageNumber="8">Pedipalp chela manus D/ carapace L 0.28 – 0.29, 0.28 ± 0.004 (4) 0.24 – 0.26, 0.25 ± 0.005 (4)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFAD0FA88FE29" blockId="9.[212,1398,201,1234]" box="[213,1391,400,423]" pageId="9" pageNumber="8">Pectine L/ carapace L 1.21 – 1.25, 1.22 ± 0.02 (4) 1.10 – 1.16, 1.13 ± 0.03 (4)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFAFBFB5CFE5C" blockId="9.[212,1398,201,1234]" box="[213,1211,443,466]" pageId="9" pageNumber="8">
|
||
Pedipalp femur L/
|
||
<geoCoordinate id="EE2A50728346FFD6C0D9FAFBFC21FE5C" box="[407,966,443,466]" degrees="3.00" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-3.0">W 3.00</geoCoordinate>
|
||
– 4.33, 4.05 ± 0.40 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFA9BFB5CFE7C" blockId="9.[212,1398,201,1234]" box="[213,1211,475,498]" pageId="9" pageNumber="8">
|
||
Pedipalp patella L/
|
||
<geoCoordinate id="EE2A50728346FFD6C0D0FA9BFC21FE7C" box="[414,966,475,498]" degrees="3.45" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-3.45">W 3.45</geoCoordinate>
|
||
– 3.91, 3.61 ± 0.14 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFABBFB5CFD9C" blockId="9.[212,1398,201,1234]" box="[213,1211,507,530]" pageId="9" pageNumber="8">
|
||
Pedipalp chela L/manus
|
||
<geoCoordinate id="EE2A50728346FFD6C092FABBFC21FD9C" box="[476,966,507,530]" degrees="7.43" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-7.43">W 7.43</geoCoordinate>
|
||
– 8.86, 7.97 ± 0.50 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF95BFB5CFDBC" blockId="9.[212,1398,201,1234]" box="[213,1211,539,562]" pageId="9" pageNumber="8">Pedipalp femur L/ carapace L 1.02 – 1.11, 1.06 ± 0.03 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF97BFB5CFDDC" blockId="9.[212,1398,201,1234]" box="[213,1211,571,594]" pageId="9" pageNumber="8">Pedipalp patella L/ carapace L 1.22 – 1.36, 1.29 ± 0.05 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF91BFB5CFDFC" blockId="9.[212,1398,201,1234]" box="[213,1211,603,626]" pageId="9" pageNumber="8">Pedipalp chela L/ carapace L 1.92 – 2.17, 2.03 ± 0.07 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF93BFB5CFD1C" blockId="9.[212,1398,201,1234]" box="[213,1211,635,658]" pageId="9" pageNumber="8">Pedipalp movable finger L/ carapace L 1.53 – 1.72, 1.60 ± 0.06 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF9DBFB5CFD3C" blockId="9.[212,1398,201,1234]" box="[213,1211,667,690]" pageId="9" pageNumber="8">Pedipalp chela manus W/ carapace L 0.24 – 0.28, 0.26 ± 0.02 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF9FBFB5CFD5C" blockId="9.[212,1398,201,1234]" box="[213,1211,699,722]" pageId="9" pageNumber="8">Leg III patella L/D 4.35 – 4.81, 4.58 ± 0.15 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF99BFB5CFD7C" blockId="9.[212,1398,201,1234]" box="[213,1211,731,754]" pageId="9" pageNumber="8">Leg III patella L/ carapace L 0.86 – 0.92, 0.89 ± 0.02 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF9BBFB5CFC9C" blockId="9.[212,1398,201,1234]" box="[213,1211,763,786]" pageId="9" pageNumber="8">
|
||
Metasoma I L/
|
||
<geoCoordinate id="EE2A50728346FFD6C03CF9BBFC21FC9C" box="[370,966,763,786]" degrees="1.23" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.23">W 1.23</geoCoordinate>
|
||
– 1.50, 1.32 ± 0.08 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF85BFB5CFCBC" blockId="9.[212,1398,201,1234]" box="[213,1211,795,818]" pageId="9" pageNumber="8">
|
||
Metasoma II L/
|
||
<geoCoordinate id="EE2A50728346FFD6C035F85BFC21FCBC" box="[379,966,795,818]" degrees="1.67" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.67">W 1.67</geoCoordinate>
|
||
– 2.01, 1.78 ± 0.10 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF87BFB5CFCDC" blockId="9.[212,1398,201,1234]" box="[213,1211,827,850]" pageId="9" pageNumber="8">
|
||
Metasoma III L/
|
||
<geoCoordinate id="EE2A50728346FFD6C0CAF87BFC21FCDC" box="[388,966,827,850]" degrees="1.85" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.85">W 1.85</geoCoordinate>
|
||
– 2.15, 1.97 ± 0.09 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF81BFB5CFCFC" blockId="9.[212,1398,201,1234]" box="[213,1211,859,882]" pageId="9" pageNumber="8">
|
||
Metasoma IV L/
|
||
<geoCoordinate id="EE2A50728346FFD6C0CBF81BFC21FCFC" box="[389,966,859,882]" degrees="2.33" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.33">W 2.33</geoCoordinate>
|
||
– 2.69, 2.47 ± 0.11 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF83BFB5CFC1C" blockId="9.[212,1398,201,1234]" box="[213,1211,891,914]" pageId="9" pageNumber="8">
|
||
<materialsCitation id="3B763CE88346FFD6C19BF83BFC21FC1C" box="[213,966,891,914]" collectionCode="V" longitude="-2.71" pageId="9" pageNumber="8" specimenCount="1">
|
||
Metasoma
|
||
<collectionCode id="ED0FAE708346FFD6C004F83BFEBAFC1C" box="[330,349,891,914]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="8" type="Museum">V</collectionCode>
|
||
L/
|
||
<geoCoordinate id="EE2A50728346FFD6C032F83BFC21FC1C" box="[380,966,891,914]" degrees="2.71" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-2.71">W 2.71</geoCoordinate>
|
||
</materialsCitation>
|
||
– 3.25, 2.89 ± 0.16 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF8DBFB5BFC3C" blockId="9.[212,1398,201,1234]" box="[213,1212,923,946]" pageId="9" pageNumber="8">Vesicle L/ telson L 0.44 – 0.51, 0.47 ± 0.02 (8)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF8FBFB5BFC5C" blockId="9.[212,1398,201,1234]" box="[213,1212,955,978]" pageId="9" pageNumber="8">Metasoma I L/ carapace L 0.66 – 0.73, 0.69 ± 0.03 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF89BFB5CFC7C" blockId="9.[212,1398,201,1234]" box="[213,1211,987,1010]" pageId="9" pageNumber="8">Metasoma II L/ carapace L 0.76 – 0.85, 0.80 ± 0.03 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BF8BBFB5CFB9C" blockId="9.[212,1398,201,1234]" box="[213,1211,1019,1042]" pageId="9" pageNumber="8">Metasoma III L/ carapace L 0.79 – 0.89, 0.83 ± 0.03 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFF5BFB5CFBBC" blockId="9.[212,1398,201,1234]" box="[213,1211,1051,1074]" pageId="9" pageNumber="8">Metasoma IV L/ carapace L 0.91 – 1.01, 0.95 ± 0.03 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFF7BFB5BFBDC" blockId="9.[212,1398,201,1234]" box="[213,1212,1083,1106]" pageId="9" pageNumber="8">Metasoma V L/ carapace L 1.04 – 1.09, 1.06 ± 0.02 (9)</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFF1BFB5BFBFC" blockId="9.[212,1398,201,1234]" box="[213,1212,1115,1138]" pageId="9" pageNumber="8">
|
||
Metasoma II W/ Metasoma I
|
||
<geoCoordinate id="EE2A50728346FFD6C341FF1BFC21FBFC" box="[527,966,1115,1138]" degrees="0.83" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.83">W 0.83</geoCoordinate>
|
||
– 0.88, 0.86 ± 0.02 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFF3BFB5CFB1C" blockId="9.[212,1398,201,1234]" box="[213,1211,1147,1170]" pageId="9" pageNumber="8">
|
||
Metasoma III W/ Metasoma I
|
||
<geoCoordinate id="EE2A50728346FFD6C356FF3BFC21FB1C" box="[536,966,1147,1170]" degrees="0.79" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.79">W 0.79</geoCoordinate>
|
||
– 0.83, 0.81 ± 0.01 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFFDBFB5CFB3C" blockId="9.[212,1398,201,1234]" box="[213,1211,1179,1202]" pageId="9" pageNumber="8">
|
||
Metasoma IV W/ Metasoma I
|
||
<geoCoordinate id="EE2A50728346FFD6C357FFDBFC21FB3C" box="[537,966,1179,1202]" degrees="0.72" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.72">W 0.72</geoCoordinate>
|
||
– 0.77, 0.74 ± 0.01 (9)
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C19BFFFBFB5BFB5C" blockId="9.[212,1398,201,1234]" box="[213,1212,1211,1234]" pageId="9" pageNumber="8">
|
||
<materialsCitation id="3B763CE88346FFD6C19BFFFBFC21FB5C" box="[213,966,1211,1234]" collectionCode="V" collectorName="Metasoma I" longitude="-0.68" pageId="9" pageNumber="8" specimenCount="1">
|
||
Metasoma
|
||
<collectionCode id="ED0FAE708346FFD6C004FFFBFEBAFB5C" box="[330,349,1211,1234]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="9" pageNumber="8" type="Museum">V</collectionCode>
|
||
W/
|
||
<collectorName id="26EB53638346FFD6C0C2FFFBFDEDFB5C" box="[396,522,1211,1234]" pageId="9" pageNumber="8">Metasoma I</collectorName>
|
||
<geoCoordinate id="EE2A50728346FFD6C35FFFFBFC21FB5C" box="[529,966,1211,1234]" degrees="0.68" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.68">W 0.68</geoCoordinate>
|
||
</materialsCitation>
|
||
– 0.74, 0.71 ± 0.02 (9)
|
||
</paragraph>
|
||
<caption id="DF61663D8346FFD6C18EFFB2FE72FADD" pageId="9" pageNumber="8" startId="9.[192,257,1266,1289]" targetType="table">
|
||
<paragraph id="8BA136B58346FFD6C18EFFB2FE72FADD" blockId="9.[192,1440,1266,1363]" pageId="9" pageNumber="8">
|
||
<emphasis id="B96AEAA78346FFD6C18EFFB2FEC7FA87" bold="true" box="[192,288,1266,1289]" pageId="9" pageNumber="8">Table 1:</emphasis>
|
||
Variation in selected morphometric ratios for adult
|
||
<taxonomicName id="4C1E4D368346FFD6C26AFFB4FBB6FA87" authorityName="Lowe" authorityYear="2009" box="[804,1105,1268,1289]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78346FFD6C26AFFB4FBB6FA87" box="[804,1105,1268,1289]" italics="true" pageId="9" pageNumber="8">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78346FFD6C517FFB4FB4FFA87" bold="true" box="[1113,1192,1268,1289]" pageId="9" pageNumber="8">
|
||
<taxonomicNameLabel id="A25957DC8346FFD6C517FFB4FB4FFA87" box="[1113,1192,1268,1289]" pageId="9" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Indicated are ranges and mean ± SD, with sample sizes in parentheses. All samples are from adults. Values of variables significantly different between the two sexes (p <0.05, Mann-Whitney test) are cited separately for males and females, and those not significantly different are pooled for both sexes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58346FFD6C18EFE39FE11F9E8" blockId="9.[192,805,1401,1638]" pageId="9" pageNumber="8">
|
||
<emphasis id="B96AEAA78346FFD6C18EFE39FE22FA1E" bold="true" box="[192,453,1401,1424]" pageId="9" pageNumber="8">
|
||
Legs (
|
||
<figureCitation id="13252A308346FFD6C048FE39FE81FA1E" box="[262,358,1401,1424]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="9" pageNumber="8">Figs 8–9</figureCitation>
|
||
,
|
||
<figureCitation id="13252A308346FFD6C03DFE39FE5EFA1E" box="[371,441,1401,1424]" captionStart-0="Figures 18–19" captionStart-1="Figures 20–21" captionStartId-0="7.[192,278,1259,1282]" captionStartId-1="8.[192,278,1193,1216]" captionTargetBox-0="[192,1438,192,1227]" captionTargetBox-1="[192,1439,192,1160]" captionTargetId-0="figure-6@7.[192,1438,192,1227]" captionTargetId-1="figure-9@8.[192,1439,192,1160]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 18–19: Compsobuthus nematodactylus sp. nov. holotype male, habitus. 18. Dorsal aspect. 19. Ventral aspect. Scale bar: 5 mm." captionText-1="Figures 20–21: Compsobuthus nematodactylus sp. nov. paratype adult female (near Wadi Mistal), habitus. 20. Dorsal aspect. 21. Ventral aspect. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12782213" figureDoi-1="http://doi.org/10.5281/zenodo.12782215" httpUri-0="https://zenodo.org/record/12782213/files/figure.png" httpUri-1="https://zenodo.org/record/12782215/files/figure.png" pageId="9" pageNumber="8">18–21</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Very slender, laterigrade; leg III patella L/D 4.55; legs III–IV with tibial spurs; retrolateral pedal spurs simple, without setae; prolateral pedal spurs basally bifurcate, with 0–1 setae; telotarsi elongate, with two rows of short macrosetae on ventral surfaces (
|
||
<figureCitation id="13252A308346FFD6C065FD52FE96F9A7" box="[299,369,1554,1577]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="9" pageNumber="8">Fig. 8</figureCitation>
|
||
); basitarsi elongate, with two rows of stout macrosetae on ventral surface; bristle combs absent; ungues stout (
|
||
<figureCitation id="13252A308346FFD6C0E9FD0FFE0EF9E8" box="[423,489,1615,1638]" captionStart="Figures 1–9" captionStartId="5.[192,278,1794,1817]" captionTargetBox="[286,1362,229,1696]" captionTargetId="figure-17@5.[192,1438,885,1062]" captionTargetPageId="5" captionText="Figures 1–9: Compsobuthus nematodactylus sp. nov. 1, 6–9, holotype male, 2–5, paratype adult males. 1. Carapace and tergites I–II. 2–3. Right chelicera, dorsal (2) and ventral (3) aspect. 4–5. Right hemispermatophore, lobes at base of flagellum, dorsal (4) and ectal (5) aspects. 6. Metasoma and telson, right lateral aspect. 7. Metasoma and telson, ventral aspect. 8. Right telotarsus III, ventral aspect. 9. Right basitarsus and telotarsus III, retrolateral aspect. Scale bar: 1: 2 mm; 2–3, 8–9: 1.45 mm; 4–5: 0.725 mm; 6–7: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782207" httpUri="https://zenodo.org/record/12782207/files/figure.png" pageId="9" pageNumber="8">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD6C18EFDCDFBC8FA43" blockId="9.[192,802,1677,1914]" lastBlockId="9.[830,1440,1401,1485]" pageId="9" pageNumber="8">
|
||
<emphasis id="B96AEAA78346FFD6C18EFDCDFD9AF92A" bold="true" box="[192,637,1677,1700]" pageId="9" pageNumber="8">
|
||
Measurements of
|
||
<typeStatus id="54A588178346FFD6C0C0FDCDFE17F92A" box="[398,496,1677,1700]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
male (mm).
|
||
</emphasis>
|
||
Total L 32.50; metasoma + telson L 20.00; carapace L 3.69,
|
||
<geoCoordinate id="EE2A50728346FFD6C380FDEBFCC5F94C" box="[718,802,1707,1730]" degrees="1.54" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.54">W 1.54</geoCoordinate>
|
||
(anterior interocular), 3.48 (posterior), carapace preocular L 1.31; metasomal segments (L/D/W) I 2.46/1.63/2.00, II 2.92/1.59/1.75, III 3.08/1.58/1.67, IV 3.54/1.54/1.50, V 3.92/1.50/1.35; telson L 3.54; vesicle L 1.67, D 1.17,
|
||
<geoCoordinate id="EE2A50728346FFD6C039FC05FE2BF8D2" box="[375,460,1861,1884]" degrees="1.17" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.17">W 1.17</geoCoordinate>
|
||
; pedipalp chela L 7.52, manus ventral L 1.75, chela
|
||
<geoCoordinate id="EE2A50728346FFD6C0F7FC23FDF7F8F4" box="[441,528,1891,1914]" degrees="1.01" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.01">W 1.01</geoCoordinate>
|
||
, D 1.03, fixed finger L 5.21, movable finger L 5.75; pedipalp femur L 3.83,
|
||
<geoCoordinate id="EE2A50728346FFD6C4C9FE39FC88FA20" degrees="0.92" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-0.92">W 0.92</geoCoordinate>
|
||
, patella L 4.69,
|
||
<geoCoordinate id="EE2A50728346FFD6C57AFED7FB6AFA20" box="[1076,1165,1431,1454]" degrees="1.33" direction="west" orientation="longitude" pageId="9" pageNumber="8" precision="555" value="-1.33">W 1.33</geoCoordinate>
|
||
; pectine L 4.46; leg III patella L 3.25, D 0.71.
|
||
</paragraph>
|
||
<paragraph id="8BA136B58346FFD5C270FEB3FF32FBAD" blockId="9.[830,1441,1523,1914]" lastBlockId="10.[192,802,821,1059]" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
|
||
<emphasis id="B96AEAA78346FFD6C270FEB3FBC2F984" bold="true" box="[830,1061,1523,1546]" pageId="9" pageNumber="8">Sexual dimorphism.</emphasis>
|
||
Adult females differed as follows: larger size, adult female total L 40.00 – 43.00 (41.25 ± 1.26,
|
||
<emphasis id="B96AEAA78346FFD6C232FD71FC6EF9C6" box="[892,905,1585,1608]" italics="true" pageId="9" pageNumber="8">n</emphasis>
|
||
= 4), carapace L 4.67–4.87 (4.78 ± 0.08,
|
||
<emphasis id="B96AEAA78346FFD6C418FD71FA84F9C6" box="[1366,1379,1585,1608]" italics="true" pageId="9" pageNumber="8">n</emphasis>
|
||
= 4), adult male total L 26.00 – 32.50 (30.70 ± 2.68,
|
||
<emphasis id="B96AEAA78346FFD6C41AFD0FFA86F9E8" box="[1364,1377,1615,1638]" italics="true" pageId="9" pageNumber="8">n</emphasis>
|
||
= 5), carapace L 3.00 – 3.69 (3.53 ± 0.30,
|
||
<emphasis id="B96AEAA78346FFD6C5B4FD2EFAE0F90B" box="[1274,1287,1646,1669]" italics="true" pageId="9" pageNumber="8">n</emphasis>
|
||
= 5); denser setation on sternite VII and ventral surfaces of metasoma (
|
||
<figureCitation id="13252A308346FFD6C208FDEBFC70F94C" box="[838,919,1707,1730]" captionStart="Figures 10–17" captionStartId="6.[192,278,1625,1648]" captionTargetBox="[232,1397,207,1539]" captionTargetId="figure-18@6.[192,1440,712,890]" captionTargetPageId="6" captionText="Figures 10–17: Compsobuthus nematodactylus sp. nov. 11, 12–16, holotype male, 10, 17, paratype adult female (near Wadi Mistal). 10. Left pectine of female, ventral aspect. 11. Left pectine of male, ventral aspect. 12. Right pedipalp femur, dorsal aspect. 13. Right pedipalp patella, dorsal aspect. 14. Right pedipalp patella, external aspect. 15. Right pedipalp chela, external aspect. 16. Right pedipalp chela, ventral aspect. 17. Metasoma IV, V and telson of female, ventral aspect. Scale bar: 10, 11: 2.8 mm; 12–16: 2 mm; 17: 4.17 mm." figureDoi="http://doi.org/10.5281/zenodo.12782211" httpUri="https://zenodo.org/record/12782211/files/figure.png" pageId="9" pageNumber="8">Fig. 17</figureCitation>
|
||
); genital papillae absent; pectines shorter, distal tips not extending to distal limits of trochanter IV, smaller ratio of pectine L/carapace L (p = 0.014), pectine teeth shorter, such that when anterior margins of left and right pectines are aligned with posterior margins of coxae IV, basal pectine teeth do not overlap; lateral carinae obsolete on sternite VI, more weakly developed on sternite VII; wider prosoma and mesosoma, larger ratio of carapace posterior W/ carapace L (p = 0.025), slightly longer pedipalp movable finger relative to manus (p = 0.05), telson vesicle depth smaller relative to width (p = 0.014), and less deep chela manus relative to carapace L (p = 0.014). Both dimorphic and nondimorphic morphometric ratios are summarized in
|
||
<tableCitation id="C69C030E8345FFD5C3ABF8ADFF28FBAD" captionStart="Table 1" captionStartId="9.[192,257,1266,1289]" captionText="Table 1: Variation in selected morphometric ratios for adult Compsobuthus nematodactylus sp. nov. Indicated are ranges and mean ± SD, with sample sizes in parentheses. All samples are from adults. Values of variables significantly different between the two sexes (p <0.05, Mann-Whitney test) are cited separately for males and females, and those not significantly different are pooled for both sexes." pageId="10" pageNumber="9">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF61663D8345FFD5C18EF99DFCF9FC81" ID-DOI="http://doi.org/10.5281/zenodo.12782219" ID-Zenodo-Dep="12782219" httpUri="https://zenodo.org/record/12782219/files/figure.png" pageId="10" pageNumber="9" startId="10.[192,278,733,756]" targetBox="[218,1411,218,671]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BA136B58345FFD5C18EF99DFCF9FC81" blockId="10.[192,1440,733,783]" pageId="10" pageNumber="9">
|
||
<emphasis id="B96AEAA78345FFD5C18EF99DFE8EFD7A" bold="true" box="[192,361,733,756]" pageId="10" pageNumber="9">Figures 22–23:</emphasis>
|
||
<taxonomicName id="4C1E4D368345FFD5C03EF99EFD7AFD7D" authorityName="Lowe" authorityYear="2009" box="[368,669,734,755]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78345FFD5C03EF99EFD7AFD7D" box="[368,669,734,755]" italics="true" pageId="10" pageNumber="9">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78345FFD5C3EDF99EFD15FD7D" bold="true" box="[675,754,734,755]" pageId="10" pageNumber="9">
|
||
<taxonomicNameLabel id="A25957DC8345FFD5C3EDF99EFD15FD7D" box="[675,754,734,755]" pageId="10" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
paratype adult male.
|
||
<emphasis id="B96AEAA78345FFD5C288F99EFC04FD7D" bold="true" box="[966,995,734,755]" pageId="10" pageNumber="9">22.</emphasis>
|
||
Pedipalp chela, left movable finger dentition.
|
||
<emphasis id="B96AEAA78345FFD5C18EF9BAFF39FC81" bold="true" box="[192,222,762,783]" pageId="10" pageNumber="9">23.</emphasis>
|
||
Pedipalp chela, left fixed finger dentition. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58345FFD5C18EFF0AFD47FA1D" blockId="10.[192,802,1098,1427]" pageId="10" pageNumber="9">
|
||
<emphasis id="B96AEAA78345FFD5C18EFF0AFE82FBF1" bold="true" pageId="10" pageNumber="9">
|
||
Measurements of
|
||
<typeStatus id="54A588178345FFD5C0E3FF0AFDF4FBEF" box="[429,531,1098,1121]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
female (near Wadi Mistal) (mm).
|
||
</emphasis>
|
||
Total L 43.00; metasoma + telson L 26.00; carapace L 4.81,
|
||
<geoCoordinate id="EE2A50728345FFD5C099FFC7FDCCFB10" box="[471,555,1159,1182]" degrees="2.10" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-2.1">W 2.10</geoCoordinate>
|
||
(anterior interocular), 4.83 (posterior), carapace preocular L 1.71; metasomal segments (L/D/W) I 3.33/2.08/2.46, II 3.83/2.00/2.05, III 4.08/2.00/1.99, IV 4.67/1.88/1.81, V 5.15/1.88/1.75; telson L 4.58; vesicle L 2.00, D 1.40,
|
||
<geoCoordinate id="EE2A50728345FFD5C329FE41FD5EFA96" box="[615,697,1281,1304]" degrees="1.50" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-1.5">W 1.50</geoCoordinate>
|
||
; pedipalp chela L 9.80, manus ventral L 2.10, chela
|
||
<geoCoordinate id="EE2A50728345FFD5C3E7FE60FD1AFAB9" box="[681,765,1312,1335]" degrees="1.18" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-1.18">W 1.18</geoCoordinate>
|
||
, D 1.20, fixed finger L 7.08, movable finger L 7.91; pedipalp femur L 5.08,
|
||
<geoCoordinate id="EE2A50728345FFD5C08AFE1DFDF3FAFA" box="[452,532,1373,1396]" degrees="1.18" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-1.18">W 1.18</geoCoordinate>
|
||
, patella L 6.21,
|
||
<geoCoordinate id="EE2A50728345FFD5C384FE1DFCFCFAFA" box="[714,795,1373,1396]" degrees="1.75" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-1.75">W 1.75</geoCoordinate>
|
||
; pectine L 5.29; leg III patella L 4.21, D 0.94.
|
||
</paragraph>
|
||
<paragraph id="8BA136B58345FFD5C18EFEF9FB1EFB55" blockId="10.[192,804,1465,1917]" lastBlockId="10.[830,1441,821,1243]" pageId="10" pageNumber="9">
|
||
<emphasis id="B96AEAA78345FFD5C18EFEF9FE44FA5E" bold="true" box="[192,419,1465,1488]" pageId="10" pageNumber="9">Meristic variation.</emphasis>
|
||
Pectinal tooth count was not sexually dimorphic in the small sample size available (p = 0.49): males 31–33 (mode 32; 1/10 combs with 31, 5/10 with 32, and 4/10 with 33 teeth), females 28–33 (mode 31; 1/8 combs with 28, 29, 32 and 33 teeth, 2/8 combs with 31 and 34 teeth). The number of denticle subrows on the pedipalp fingers was either 11 or 12, and the lower counts were associated with fusion, often of the proximal subrows. Among intact fixed fingers, there were 5/18 with 11, and 13/18 with 12 subrows; among intact movable fingers there were 8/17 with 11 subrows, and 9/17 with 12 subrows. Thus, the fixed fingers had the higher mean subrow count. Males had a significantly higher count of movable finger subrows than females (p = 0.01): among male intact movable fingers, there were 2/10 with 11, and 8/10 with 12 subrows; among intact female movable fingers, there were 6/7 with 11 and 1/7 with 12 subrows. The numbers of posterior marginal macrosetae on tergites I–VI was variable. In specimens of both sexes from the
|
||
<collectingRegion id="49DAF8578345FFD5C50BF8F0FB5CFC49" box="[1093,1211,944,967]" country="Oman" name="Musandam" pageId="10" pageNumber="9">Musandam</collectingRegion>
|
||
, the medial margins (inside the lateral carinae) normally had 2, and occasionally 3 macrosetae, and the left or right lateral margins (outside the lateral carinae) normally had 1, and occasionally 2 macrosetae, i.e. the setation formula (left lateral, medial, right lateral) was 0–1, 2–3, 0 –1. The female sample from near Wadi Mistal was distinctive in having higher numbers of marginal macrosetae, with setation formula for each tergite: I = 1,3,2, II = 3,3,2, III = 2,5,4, IV = 2,5,3, V = 3,7,3, VI = 1,6,2.
|
||
</paragraph>
|
||
<paragraph id="8BA136B58345FFD5C270FE41FC4FF929" blockId="10.[830,1441,1281,1703]" pageId="10" pageNumber="9">
|
||
<emphasis id="B96AEAA78345FFD5C270FE41FBD1FA96" bold="true" box="[830,1078,1281,1304]" pageId="10" pageNumber="9">Variation with age.</emphasis>
|
||
Study of a single available intermediate instar male (total L
|
||
<quantity id="4CE69B508345FFD5C5FEFE60FB19FAB9" box="[1200,1278,1312,1335]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="9" unit="mm" value="26.0">26 mm</quantity>
|
||
, carapace L
|
||
<quantity id="4CE69B508345FFD5C4DDFE60FC80FADB" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="9" unit="mm" value="3.0">3 mm</quantity>
|
||
) revealed significant (p <0.05) morphometric differences from adult males: more slender pedipalp chelae (chela L/
|
||
<geoCoordinate id="EE2A50728345FFD5C2B5FE3CFBB3FA1D" box="[1019,1108,1404,1427]" degrees="8.86" direction="west" orientation="longitude" pageId="10" pageNumber="9" precision="555" value="-8.86">W 8.86</geoCoordinate>
|
||
), longer pedipalp fixed and movable fingers (fixed finger L/ manus ventral L 3.42, movable finger L/manus ventral L 3.72), more slender metasomal segments (segment L/W: I 1.5, II 2.0, III 2.15, IV 3.25), less bulbous telson vesicle (vesicle L/D 1.61). Intercarinal granulation was sparser and weaker on the carapace and tergites, and carinal granulation was weaker or finer on all sclerites. On the metasomal segments, ventromedian carinae were smooth, not crenulate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E8345FFD4C270FD8DFEC4FA09" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9" type="distribution">
|
||
<paragraph id="8BA136B58345FFD4C270FD8DFEC4FA09" blockId="10.[830,1441,1741,1917]" lastBlockId="11.[192,802,1178,1415]" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
|
||
<emphasis id="B96AEAA78345FFD5C270FD8DFC36F96A" bold="true" box="[830,977,1741,1764]" pageId="10" pageNumber="9">Distribution.</emphasis>
|
||
<taxonomicName id="4C1E4D368345FFD5C296FD8DFB46F96A" authorityName="Lowe" authorityYear="2009" box="[984,1185,1741,1764]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA78345FFD5C296FD8DFC0DF96A" box="[984,1002,1741,1764]" italics="true" pageId="10" pageNumber="9">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78345FFD5C2B7FD8DFB46F96A" box="[1017,1185,1741,1764]" italics="true" pageId="10" pageNumber="9">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
is probably endemic to Al Hajar Al Gharbi, the western section of northern
|
||
<collectingCountry id="F30976258345FFD5C270FC4AFC98F8AF" box="[830,895,1802,1825]" name="Oman" pageId="10" pageNumber="9">Oman</collectingCountry>
|
||
mountain ranges (
|
||
<figureCitation id="13252A308345FFD5C502FC4AFB7BF8AF" box="[1100,1180,1802,1825]" captionStart="Figure 25" captionStartId="12.[1092,1168,1069,1092]" captionTargetBox="[177,1061,256,1316]" captionTargetId="figure-9@12.[169,1068,240,1321]" captionTargetPageId="12" captionText="Figure 25: Topographic map of Oman in the southeastern Arabian Peninsula, with collection localities of C. nematodactylus plotted as blue circles (modified from a public domain topographic map available at http://en.wikipedia.org/wiki/File: Oman_Topography.png)." figureDoi="http://doi.org/10.5281/zenodo.12782223" httpUri="https://zenodo.org/record/12782223/files/figure.png" pageId="10" pageNumber="9">Fig. 25</figureCitation>
|
||
). Although eight of the
|
||
<specimenCount id="9D18FD3C8345FFD5C270FC69FC45F8CE" box="[830,930,1833,1856]" count="9" pageId="10" pageNumber="9" type="generic" typeStatus="type">nine type</specimenCount>
|
||
specimens were collected from the
|
||
<collectingRegion id="49DAF8578345FFD5C464FC69FA46F8CE" box="[1322,1441,1833,1856]" country="Oman" name="Musandam" pageId="10" pageNumber="9">Musandam</collectingRegion>
|
||
Peninsula, the record from near Wadi Mistal indicates a wider and perhaps continuous distribution. It is likely to occupy the intervening mountains including parts within the borders of the
|
||
<collectingCountry id="F30976258344FFD4C0DEFFF8FD62FB41" box="[400,645,1208,1231]" name="United Arab Emirates" pageId="11" pageNumber="10">United Arab Emirates</collectingCountry>
|
||
. Although the lack of records from Al Hajar Ash Sharqi (eastern part of the ranges) might indicate that the species does not bridge the Sumail Gap (the large wadi separating eastern and western ranges), this might just be a reflection of insufficient sampling of this relatively small, secretive scorpion.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF61663D8344FFD4C18EFF66FBF8FBFD" ID-DOI="http://doi.org/10.5281/zenodo.12782221" ID-Zenodo-Dep="12782221" httpUri="https://zenodo.org/record/12782221/files/figure.png" pageId="11" pageNumber="10" startId="11.[192,268,1062,1085]" targetBox="[191,1441,191,1029]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8BA136B58344FFD4C18EFF66FBF8FBFD" blockId="11.[192,1441,1062,1139]" pageId="11" pageNumber="10">
|
||
<emphasis id="B96AEAA78344FFD4C18EFF66FED1FBB3" bold="true" box="[192,310,1062,1085]" pageId="11" pageNumber="10">Figure 24:</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="4C1E4D368344FFD4C093FF67FCEEFBB2" authorityName="Lowe" authorityYear="2009" box="[477,777,1063,1084]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="nematodactylus" status="sp. nov.">
|
||
<emphasis id="B96AEAA78344FFD4C093FF67FCEEFBB2" box="[477,777,1063,1084]" italics="true" pageId="11" pageNumber="10">Compsobuthus nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEAA78344FFD4C241FF67FCBAFBB2" bold="true" box="[783,861,1063,1084]" pageId="11" pageNumber="10">
|
||
<taxonomicNameLabel id="A25957DC8344FFD4C241FF67FCBAFBB2" box="[783,861,1063,1084]" pageId="11" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Southward view of the western slope of the mountains and road SW of Khawr Najd, Musandam Peninsula, showing extensive layers of shelf carbonate with an abundance of rock fissures providing microhabitat for lithophilic scorpions. Date of photograph: 30 September 1994.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C304653E8344FFD4C18EFEEEFBF5FAA5" pageId="11" pageNumber="10" type="biology_ecology">
|
||
<paragraph id="8BA136B58344FFD4C18EFEEEFBF5FAA5" blockId="11.[192,802,1454,1906]" lastBlockId="11.[830,1440,1178,1323]" pageId="11" pageNumber="10">
|
||
<emphasis id="B96AEAA78344FFD4C18EFEEEFEC6FA4B" bold="true" box="[192,289,1454,1477]" pageId="11" pageNumber="10">Ecology.</emphasis>
|
||
All samples of the new species were collected at lower elevations (
|
||
<quantity id="4CE69B508344FFD4C0FBFE8CFDDDFA6D" box="[437,570,1484,1507]" metricMagnitude="2" metricUnit="m" metricValue="5.4" metricValueMax="9.8" metricValueMin="1.0" pageId="11" pageNumber="10" unit="m" value="540.0" valueMax="980.0" valueMin="100.0">100–980 m</quantity>
|
||
a.s.l.) at night, by ultraviolet detection on vertical rock cliffs and road cuts. This species is apparently an obligate ‘ultralithophile’, a term describing scorpions that exclusively inhabit solid rock substrates, sheltering in fissures by day and emerging at night to forage on bare rock faces. The strongly flattened body, slender metasoma, long laterigrade legs armed with stout ungues, and elongated pedipalps with extremely tenuous fingers, are structural adaptations to an existence in narrow rock crevices (Polis, 1990; Prendini, 2001a). A similar morphology is observed in many members the lithophilic North American vaejovid genera
|
||
<taxonomicName id="4C1E4D368344FFD4C0BCFC7CFD91F8DD" authorityName="Stahnke" authorityYear="1974" box="[498,630,1852,1875]" class="Arachnida" family="Vaejovidae" genus="Serradigitus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78344FFD4C0BCFC7CFD91F8DD" box="[498,630,1852,1875]" italics="true" pageId="11" pageNumber="10">Serradigitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E4D368344FFD4C3F8FC7CFCFCF8DD" authorityName="Kraepelin" authorityYear="1900" box="[694,795,1852,1875]" class="Arachnida" family="Vaejovidae" genus="Syntropis" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78344FFD4C3F8FC7CFCFCF8DD" box="[694,795,1852,1875]" italics="true" pageId="11" pageNumber="10">Syntropis</emphasis>
|
||
</taxonomicName>
|
||
, which also developed very slender pedipalp fingers with linear series of denticles, terminating in an enlarged tooth with smooth cap and apical clusters of sensory setae (Stahnke, 1974; Williams & Berke, 1986; Sissom & Stockwell, 1991; Soleglad & Fet, 2006; Soleglad, Lowe & Fet, 2007).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304653E8344FFD2C270FE12FD94FEF9" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10" type="discussion">
|
||
<paragraph id="8BA136B58344FFD3C270FE12FE5CF8F1" blockId="11.[830,1441,1362,1906]" lastBlockId="12.[192,802,1375,1919]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
|
||
<emphasis id="B96AEAA78344FFD4C270FE12FC59FAE7" bold="true" box="[830,958,1362,1385]" pageId="11" pageNumber="10">Discussion.</emphasis>
|
||
The new species displays some significant differences from other known species of
|
||
<taxonomicName id="4C1E4D368344FFD4C5B2FE30FA7EFA09" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[1276,1433,1392,1415]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78344FFD4C5B2FE30FA7EFA09" box="[1276,1433,1392,1415]" italics="true" pageId="11" pageNumber="10">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
. An original diagnostic character for the genus was the fusion of the central median and posterior median carinae of the carapace into a single continuous linear keel (Vachon, 1949, 1952). In the new species, these two carinae were frequently observed to be separated by gaps of one or more granule diameters. Although the posterior median carinae were linear with regular granulation, the central median carinae were curved and often exhibited irregular granulation. The carinae were fused in 10/18 cases, and separated in 8/18 cases. Fusion was not always left-right symmetric: carinae of 3/9 carapaces were bilaterally fused, 4/9 unilaterally fused (3 left, 1 right), and 2/9 bilaterally separated. In other respects, the new species conforms to the current definition of
|
||
<taxonomicName id="4C1E4D368344FFD4C29FFC7CFB97F8DD" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[977,1136,1852,1875]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78344FFD4C29FFC7CFB97F8DD" box="[977,1136,1852,1875]" italics="true" pageId="11" pageNumber="10">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
, so it is placed within that genus. Sissom & Fet (1998) arrived at a similar taxonomic decision for
|
||
<taxonomicName id="4C1E4D368343FFD3C0FDFE1FFDB8FAF8" baseAuthorityName="Birula" baseAuthorityYear="1905" box="[435,607,1375,1398]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="matthiesseni">
|
||
<emphasis id="B96AEAA78343FFD3C0FDFE1FFE22FAF8" box="[435,453,1375,1398]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C094FE1FFDB8FAF8" box="[474,607,1375,1398]" italics="true" pageId="12" pageNumber="11">matthiesseni</emphasis>
|
||
</taxonomicName>
|
||
, which can also have these two carinae incompletely fused.
|
||
<taxonomicName id="4C1E4D368343FFD3C39BFE3EFEA9FA3D" authorityName="Lowe" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA78343FFD3C39BFE3EFD00FA1B" box="[725,743,1406,1429]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C24EFE3EFEA9FA3D" italics="true" pageId="12" pageNumber="11">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
is further distinguished by having the carinae on the carapace and tergites more weakly developed than many other
|
||
<taxonomicName id="4C1E4D368343FFD3C368FE9AFD23FA7F" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[550,708,1498,1521]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78343FFD3C368FE9AFD23FA7F" box="[550,708,1498,1521]" italics="true" pageId="12" pageNumber="11">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
. These differences in carinal fusion and development can be interpreted as synapomorphic modifications linked to flattening and streamlining of the body. Robust, raised carinae are likely to serve a biomechanical function in arthropods, acting as reinforcing ribs for the rigid exoskeleton, while maintaining a relatively lightweight structure (Vincent & Wegst, 2004). In ultralithophilic scorpions, reduction of carinae has enabled them to squeeze tightly into very narrow fissures. It seems that any structural weakening of the integument incurred by carinal attenuation is amply compensated by protection gained from being able to retreat effortlessly into fortresses of solid rock.
|
||
</paragraph>
|
||
<caption id="DF61663D8343FFD3C50AFF6DFADEFA8B" ID-DOI="http://doi.org/10.5281/zenodo.12782223" ID-Zenodo-Dep="12782223" httpUri="https://zenodo.org/record/12782223/files/figure.png" pageId="12" pageNumber="11" startId="12.[1092,1168,1069,1092]" targetBox="[177,1061,256,1316]" targetPageId="12" targetType="figure">
|
||
<paragraph id="8BA136B58343FFD3C50AFF6DFADEFA8B" blockId="12.[1092,1441,1069,1285]" pageId="12" pageNumber="11">
|
||
<emphasis id="B96AEAA78343FFD3C50AFF6DFB26FBCA" bold="true" box="[1092,1217,1069,1092]" pageId="12" pageNumber="11">Figure 25:</emphasis>
|
||
Topographic map of Oman in the southeastern Arabian Peninsula, with collection localities of
|
||
<taxonomicName id="4C1E4D368343FFD3C527FFC1FAC0FB18" authorityName="Lowe" authorityYear="2009" box="[1129,1319,1153,1174]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA78343FFD3C527FFC1FB9EFB18" box="[1129,1145,1153,1174]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C5DEFFC1FAC0FB18" box="[1168,1319,1153,1174]" italics="true" pageId="12" pageNumber="11">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
plotted as blue circles (modified from a public domain topographic map available at http://en.wikipedia.org/wiki/File: Oman_Topography.png).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA136B58343FFD2C220FE1FFD94FEF9" blockId="12.[830,1441,1375,1919]" lastBlockId="13.[192,802,199,375]" lastPageId="13" lastPageNumber="12" pageId="12" pageNumber="11">
|
||
The species closest to
|
||
<taxonomicName id="4C1E4D368343FFD3C5C6FE1FFABFFAF8" authorityName="Lowe" authorityYear="2009" box="[1160,1368,1375,1398]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA78343FFD3C5C6FE1FFB7DFAF8" box="[1160,1178,1375,1398]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C5FFFE1FFABFFAF8" box="[1201,1368,1375,1398]" italics="true" pageId="12" pageNumber="11">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
, both systematically and biogeographically, appear to be
|
||
<taxonomicName id="4C1E4D368343FFD3C4C6FE3EFC3AFA3D" baseAuthorityName="Simon" baseAuthorityYear="1882" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="acutecarinatus">
|
||
<emphasis id="B96AEAA78343FFD3C4C6FE3EFA7DFA1B" box="[1416,1434,1406,1429]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C270FEDCFC3AFA3D" box="[830,989,1436,1459]" italics="true" pageId="12" pageNumber="11">acutecarinatus</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F30976258343FFD3C5E1FEDCFB17FA3D" box="[1199,1264,1436,1459]" name="Oman" pageId="12" pageNumber="11">Oman</collectingCountry>
|
||
(
|
||
<collectingRegion id="49DAF8578343FFD3C441FEDCFABBFA3D" box="[1295,1372,1436,1459]" country="Oman" name="Zufar" pageId="12" pageNumber="11">Dhofar</collectingRegion>
|
||
) and
|
||
<collectingCountry id="F30976258343FFD3C270FEFBFC69FA5C" box="[830,910,1467,1490]" name="Yemen" pageId="12" pageNumber="11">Yemen</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C1E4D368343FFD3C29EFEFBFBC8FA5C" authorityName="Lowe" authorityYear="2001" box="[976,1071,1467,1490]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="polisi">
|
||
<emphasis id="B96AEAA78343FFD3C29EFEFBFC05FA5C" box="[976,994,1467,1490]" italics="true" pageId="12" pageNumber="11">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78343FFD3C2BAFEFBFBC8FA5C" box="[1012,1071,1467,1490]" italics="true" pageId="12" pageNumber="11">polisi</emphasis>
|
||
</taxonomicName>
|
||
from the central coast of
|
||
<collectingCountry id="F30976258343FFD3C411FEFBFA47FA5C" box="[1375,1440,1467,1490]" name="Oman" pageId="12" pageNumber="11">Oman</collectingCountry>
|
||
and Masirah Island. All three species have lost the external accessory denticles of the pedipalp fixed fingers, presumably a derived condition of the ‘acutecarinatus’ group of
|
||
<taxonomicName id="4C1E4D368343FFD3C562FD76FB2DF9C3" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[1068,1226,1590,1613]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78343FFD3C562FD76FB2DF9C3" box="[1068,1226,1590,1613]" italics="true" pageId="12" pageNumber="11">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
(Levy & Amitai, 19 8 0). In addition, the sternites and ventrolateral and ventral aspects of the metasoma of these species bear numerous straight, reddish macrosetae on the carinae and intercarinal surfaces. This heavy setation is absent in most other known
|
||
<taxonomicName id="4C1E4D368343FFD3C57DFD8FFB35F968" authorityName="Vachon" authorityYear="1949" baseAuthorityName="Levy & Amitai" baseAuthorityYear="1980" box="[1075,1234,1743,1766]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AEAA78343FFD3C57DFD8FFB35F968" box="[1075,1234,1743,1766]" italics="true" pageId="12" pageNumber="11">Compsobuthus</emphasis>
|
||
</taxonomicName>
|
||
(Kovařík, 2003; Kovařík & Ahmed, 2007), so it may be synapomorphy indicating shared ancestry within the ‘acutecarinatus’ group. The three species comprise an allopatric series of populations deployed along the south eastern coast of the Arabian Peninsula, extending from Aden to the
|
||
<collectingRegion id="49DAF8578343FFD2C426FC28FEE0FF50" country="Oman" lastPageId="13" lastPageNumber="12" name="Musandam" pageId="12" pageNumber="11">Musandam</collectingRegion>
|
||
Peninsula. In the northern mountains, competition between an ‘acutecarinatus’ ancestor and other scorpions species not present in the south (for example,
|
||
<taxonomicName id="4C1E4D368342FFD2C18EFA63FE38FEB4" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[192,479,291,314]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="maindroni">
|
||
<emphasis id="B96AEAA78342FFD2C18EFA63FE38FEB4" box="[192,479,291,314]" italics="true" pageId="13" pageNumber="12">Compsobuthus maindroni</emphasis>
|
||
</taxonomicName>
|
||
), may have been a factor leading to speciation of
|
||
<taxonomicName id="4C1E4D368342FFD2C09DFA02FD7AFED7" authorityName="Lowe" authorityYear="2009" box="[467,669,322,345]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="nematodactylus">
|
||
<emphasis id="B96AEAA78342FFD2C09DFA02FE02FED7" box="[467,485,322,345]" italics="true" pageId="13" pageNumber="12">C</emphasis>
|
||
.
|
||
<emphasis id="B96AEAA78342FFD2C0B8FA02FD7AFED7" box="[502,669,322,345]" italics="true" pageId="13" pageNumber="12">nematodactylus</emphasis>
|
||
</taxonomicName>
|
||
by specialization for ultralithophilic microhabitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |