plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/B5/87/03B587F4FFC12825FD99A4020645FA7B.xml

1183 lines
226 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="CC1952504A56F5BF053E3353971BC4DD" ID-DOI="10.5281/zenodo.12585387" ID-ISSN="2064-2474" ID-Zenodo-Dep="12585387" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719594345837" checkinUser="felipe" docAuthor="Norton, R. A. &amp; Behan-Pelletier, V. M." docDate="2007" docId="03B587F4FFC12825FD99A4020645FA7B" docLanguage="en" docName="ActaZoolAcadSciH.53.4.295-333.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 53 (4)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Eniochthonius mahunkai Norton &amp; Behan-Pelletier, 2007, sp. n." docType="treatment" docVersion="2" lastPageNumber="313" masterDocId="FF8CFF8CFFC32837FFC0A7030234FFC2" masterDocTitle="Eniochthonius Mahunkai Sp. N. (Acari: Oribatida: Eniochthoniidae), From North American Peatlands, With A Redescription Of Eniochthonius And A Key To North American Species" masterLastPageNumber="333" masterPageNumber="295" pageNumber="297" updateTime="1719597135592" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="33A64158724D6710233BDA5E1CF26C1A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6DC73E224F7FCA5F629A11F090918B2F">
<mods:title id="9B270B62D9FD2D44061B15BAD53D3E32">Eniochthonius Mahunkai Sp. N. (Acari: Oribatida: Eniochthoniidae), From North American Peatlands, With A Redescription Of Eniochthonius And A Key To North American Species</mods:title>
</mods:titleInfo>
<mods:name id="7DA68464B0AE98A262A30C97F4D4314B" type="personal">
<mods:role id="E41174B96F7115CBB29EC15D038E420C">
<mods:roleTerm id="A4AE5AAC1E7C92A4E71F7B2A69F3E76C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BCBCF39877D80E1C726B4ACA97B77DC">Norton, R. A.</mods:namePart>
</mods:name>
<mods:name id="CC1908B67641892FA70E012F0A9D28DF" type="personal">
<mods:role id="99199AB02ECF3A468060AF92EB5BE6D2">
<mods:roleTerm id="B5E1AAB0EF26D33E3897E47B75AA8EC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECE34EE3270D27A4738AF6655D42A7B2">Behan-Pelletier, V. M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="97A0B0E12CAC231D6489CF63E49D7320">text</mods:typeOfResource>
<mods:relatedItem id="975FDC1298E8A1AAAF66F9A9D14D116B" type="host">
<mods:titleInfo id="624DD45D229E28C190F8C54727E88017">
<mods:title id="12A8AF28C4448CE134C9A69B80256C99">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="B7DDEDDB3399B6535403866881F930FD">
<mods:date id="83AE46693756872EF7D03D98A4CCD2FE">2007</mods:date>
<mods:detail id="CFCE5B8E33115BEB7FBAE0199A0A898F" type="volume">
<mods:number id="6291A3C355CDBF07DABFCB58E4B31FF0">53</mods:number>
</mods:detail>
<mods:detail id="CB8EE32233D9AF87382DA644FFAF88C1" type="issue">
<mods:number id="2B5E29AE390049655379A1F51D0416BB">4</mods:number>
</mods:detail>
<mods:extent id="3BCD3CC34C31B62FE499533DF8301BF8" unit="page">
<mods:start id="F830AF9D32180B61ACA66ED46E4823F6">295</mods:start>
<mods:end id="D7AFB43AB856234D0CBE8AEE8EA497BC">333</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="25B6DB08774A8C881377CA529F365ECD">journal article</mods:classification>
<mods:identifier id="A6E38D20049D633E7356FE571FCCFAD6" type="DOI">10.5281/zenodo.12585387</mods:identifier>
<mods:identifier id="631A0A06C50A46E4E3145E522FDFAF3A" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="1F9D17242806A617FEB36C60DABA80ED" type="Zenodo-Dep">12585387</mods:identifier>
</mods:mods>
<treatment id="03B587F4FFC12825FD99A4020645FA7B" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B" httpUri="http://treatment.plazi.org/id/03B587F4FFC12825FD99A4020645FA7B" lastPageId="18" lastPageNumber="313" pageId="2" pageNumber="297">
<subSubSection id="C3066569FFC12835FD99A40201EEFCD9" box="[601,986,769,795]" pageId="2" pageNumber="297" type="nomenclature">
<paragraph id="8BA336E2FFC12835FD99A40201EEFCD9" blockId="2.[601,986,769,795]" box="[601,986,769,795]" pageId="2" pageNumber="297">
<heading id="D0EB818EFFC12835FD99A40201EEFCD9" box="[601,986,769,795]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="297" reason="2">
<taxonomicName id="4C1C4D61FFC12835FD99A40201A0FCD9" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[601,916,769,795]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC12835FD99A40201A0FCD9" bold="true" box="[601,916,769,795]" pageId="2" pageNumber="297">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC12835FC5BA40201EEFCD9" box="[923,986,769,795]" pageId="2" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3066569FFC12825FEBEA4440645FA7B" lastPageId="18" lastPageNumber="313" pageId="2" pageNumber="297" type="diagnosis">
<paragraph id="8BA336E2FFC12835FEBEA4440385FBF7" blockId="2.[317,1270,839,1077]" pageId="2" pageNumber="297">
Diagnosis. Adult relatively small (length 309–343 µm); notogaster moderately arched in lateral aspect. Bothridial seta slightly broadened and flattened distally, with 2–4 long tines and sparse small barbs. Notogaster obovate in dorsal aspect, with relatively long setae (
<emphasis id="B968EAF0FFC12835FD77A4B200FCFC0C" box="[695,712,945,974]" italics="true" pageId="2" pageNumber="297">
f
<subScript id="179834A7FFC12835FD00A4C300FCFC0C" attach="left" box="[704,712,960,974]" fontSize="6" pageId="2" pageNumber="297">1</subScript>
</emphasis>
reaches insertion of
<emphasis id="B968EAF0FFC12835FC7EA4B201E0FC0C" box="[958,980,945,974]" italics="true" pageId="2" pageNumber="297">
h
<subScript id="179834A7FFC12835FC0CA4C301E0FC0C" attach="left" box="[972,980,960,974]" fontSize="6" pageId="2" pageNumber="297">
<quantity id="4CE49B07FFC12835FC0CA4C301E0FC0C" box="[972,980,960,974]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="297" unit="in" value="1.0">1</quantity>
</subScript>
</emphasis>
in dorsal aspect). Aggenital region with three separate plates; seta
<emphasis id="B968EAF0FFC12835FCFDA4D7016EFC2C" box="[829,858,980,1006]" italics="true" pageId="2" pageNumber="297">ag</emphasis>
inserted on smallest, middle plate. Leg tarsus setation (I–IV, famulus included) 18–15–13–13; each with
<emphasis id="B968EAF0FFC12835FBB2A4FB06A7FBD0" box="[1138,1171,1016,1042]" italics="true" pageId="2" pageNumber="297">it”</emphasis>
present,
<emphasis id="B968EAF0FFC12835FEFDA3180363FBF7" box="[317,343,1051,1077]" italics="true" pageId="2" pageNumber="297">it’</emphasis>
absent.
</paragraph>
<paragraph id="8BA336E2FFC12835FEFDA3700078FB4F" blockId="2.[317,588,1139,1165]" box="[317,588,1139,1165]" pageId="2" pageNumber="297">
<heading id="D0EB818EFFC12835FEFDA3700078FB4F" box="[317,588,1139,1165]" fontSize="11" level="3" pageId="2" pageNumber="297" reason="4">
<emphasis id="B968EAF0FFC12835FEFDA3700078FB4F" box="[317,588,1139,1165]" italics="true" pageId="2" pageNumber="297">
Adult (
<figureCitation id="13272A67FFC12835FE4DA3700032FB4F" box="[397,518,1139,1165]" captionStart-0="Figs 1–3" captionStart-1="Figs 4–8" captionStart-2="Figs 9–11" captionStart-3="Figs 12–18" captionStart-4="Figs 19–22" captionStart-5="Figs 23–28" captionStart-6="Figs 29–35" captionStartId-0="4.[317,360,1748,1769]" captionStartId-1="5.[317,360,1644,1665]" captionStartId-2="6.[317,360,1749,1770]" captionStartId-3="8.[317,360,1631,1652]" captionStartId-4="10.[317,360,1636,1657]" captionStartId-5="11.[317,360,1522,1543]" captionStartId-6="12.[317,360,1439,1460]" captionTargetBox-0="[362,1222,429,1712]" captionTargetBox-1="[325,1264,428,1595]" captionTargetBox-2="[323,1268,1174,1717]" captionTargetBox-3="[364,1222,423,1603]" captionTargetBox-4="[379,1208,421,1608]" captionTargetBox-5="[332,1256,419,1493]" captionTargetBox-6="[378,1209,418,1411]" captionTargetId-0="figure-19@4.[361,1225,424,1720]" captionTargetId-1="figure-15@5.[317,1269,428,1610]" captionTargetId-2="figure-384@6.[319,1269,1174,1721]" captionTargetId-3="figure-19@8.[364,1222,423,1603]" captionTargetId-4="figure-19@10.[379,1208,421,1608]" captionTargetId-5="figure-15@11.[332,1256,419,1493]" captionTargetId-6="figure-19@12.[378,1209,418,1411]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" captionText-1="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" captionText-2="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" captionText-3="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" captionText-4="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" captionText-5="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," captionText-6="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi-0="http://doi.org/10.5281/zenodo.12585389" figureDoi-1="http://doi.org/10.5281/zenodo.12585391" figureDoi-2="http://doi.org/10.5281/zenodo.12585393" figureDoi-3="http://doi.org/10.5281/zenodo.12585395" figureDoi-4="http://doi.org/10.5281/zenodo.12585397" figureDoi-5="http://doi.org/10.5281/zenodo.12585399" figureDoi-6="http://doi.org/10.5281/zenodo.12585401" httpUri-0="https://zenodo.org/record/12585389/files/figure.png" httpUri-1="https://zenodo.org/record/12585391/files/figure.png" httpUri-2="https://zenodo.org/record/12585393/files/figure.png" httpUri-3="https://zenodo.org/record/12585395/files/figure.png" httpUri-4="https://zenodo.org/record/12585397/files/figure.png" httpUri-5="https://zenodo.org/record/12585399/files/figure.png" httpUri-6="https://zenodo.org/record/12585401/files/figure.png" pageId="2" pageNumber="297">Figs 1–35</figureCitation>
,
<figureCitation id="13272A67FFC12835FDD3A3700077FB4F" box="[531,579,1139,1165]" captionStart="Figs 51–56" captionStartId="17.[317,360,1548,1569]" captionTargetBox="[319,1268,689,1520]" captionTargetId="figure-141@17.[319,1268,689,1520]" captionTargetPageId="17" captionText="Figs 51–56. Eniochthonius spp., adults. 51A = lateral aspect of E. mahunkai n. sp. (top, e = egg); 51B = E. minutissimus (bottom) from same sample at Cicero, New York; 52 = E. minutissimus, anterolateral aspect of bothridial seta (bo) and nearby region, specimen from Germany; 53 = E. crosbyi, dorsal aspect of bothridial seta, with insert showing dorsolateral aspect of famulus of E. crosbyi (A) andE.minutissimus (B), each ca.10 µm; 54 = E.mahunkai sp.n., ventral aspect of genital region (cf. Fig. 3; arrow points to oblique articulation between middle and anterior aggenital plates; 55 = E. crosbyi, same, but from dissected specimen; 56 = E. minutissimus, same (intact specimen from Poland). Scale bars: 100 µm (51), 10 µm (all others); ag = alveolus of aggenital seta; e = egg; pp = plicature plate. Fig. 52 SEM micrograph (from ALBERTI et al. 1994); others from light microscope" figureDoi="http://doi.org/10.5281/zenodo.12585407" httpUri="https://zenodo.org/record/12585407/files/figure.png" pageId="2" pageNumber="297">51A</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336E2FFC12835FEBEA3A6062DFAEF" blockId="2.[317,1271,1188,1814]" pageId="2" pageNumber="297">
<emphasis id="B968EAF0FFC12835FEBEA3A603DBFB78" box="[382,495,1189,1210]" italics="true" pageId="2" pageNumber="297">Dimensions</emphasis>
(n = 20). Mean total length 325 µm (range 309–343 µm); mean maximum width 154 µm (range 147–162 µm). Mean length/width ratio of individual specimens 2.11 (varies with degree of distension); proportions of total length of fully distended specimen contributed by prodorsum 0.35; by soft, pleated sejugal band 0.04; and by hysterosoma 0.61. When fully contracted or distended, alignment of protero- and opisthosoma nearly linear (
<figureCitation id="13272A67FFC12835FC56A21B01E2FAEF" box="[918,982,1304,1325]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="2" pageNumber="297">Figs 2</figureCitation>
,
<figureCitation id="13272A67FFC12835FC21A21B063AFAEF" box="[993,1038,1304,1325]" captionStart="Figs 51–56" captionStartId="17.[317,360,1548,1569]" captionTargetBox="[319,1268,689,1520]" captionTargetId="figure-141@17.[319,1268,689,1520]" captionTargetPageId="17" captionText="Figs 51–56. Eniochthonius spp., adults. 51A = lateral aspect of E. mahunkai n. sp. (top, e = egg); 51B = E. minutissimus (bottom) from same sample at Cicero, New York; 52 = E. minutissimus, anterolateral aspect of bothridial seta (bo) and nearby region, specimen from Germany; 53 = E. crosbyi, dorsal aspect of bothridial seta, with insert showing dorsolateral aspect of famulus of E. crosbyi (A) andE.minutissimus (B), each ca.10 µm; 54 = E.mahunkai sp.n., ventral aspect of genital region (cf. Fig. 3; arrow points to oblique articulation between middle and anterior aggenital plates; 55 = E. crosbyi, same, but from dissected specimen; 56 = E. minutissimus, same (intact specimen from Poland). Scale bars: 100 µm (51), 10 µm (all others); ag = alveolus of aggenital seta; e = egg; pp = plicature plate. Fig. 52 SEM micrograph (from ALBERTI et al. 1994); others from light microscope" figureDoi="http://doi.org/10.5281/zenodo.12585407" httpUri="https://zenodo.org/record/12585407/files/figure.png" pageId="2" pageNumber="297">51A</figureCitation>
).
</paragraph>
<paragraph id="8BA336E2FFC12835FEBEA236067AF944" blockId="2.[317,1271,1188,1814]" pageId="2" pageNumber="297">
<emphasis id="B968EAF0FFC12835FEBEA23603C5FA88" box="[382,497,1333,1354]" italics="true" pageId="2" pageNumber="297">Integument.</emphasis>
In reflected light mostly smooth, shiny, pale brownish-yellow; epimeral borders, apodemes, grooves and overlapping edges of sclerites appear darker. Notogaster with two conspicuous dark transverse bands: anterior derives from sulcus (see below), posterior band derives from thickened anterior border of pygidium, showing through tectal limb of scissure by transparency. Cerotegument apparent only as small patches of granular excrescences, inconspicuous except in SEM (
<figureCitation id="13272A67FFC12835FEBFA2C703C5FA1B" box="[383,497,1476,1497]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Figs 12–13</figureCitation>
,
<figureCitation id="13272A67FFC12835FE3FA2C7000BFA1B" box="[511,575,1476,1497]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="2" pageNumber="297">20–21</figureCitation>
), distributed as indicated below. Most regions with irregular shallow grooves or wrinkles, hardly noticeable except in SEM (
<figureCitation id="13272A67FFC12835FC83A2E201BDFA34" box="[835,905,1505,1526]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 13</figureCitation>
). Except as noted, epicuticle of cuticular plates with continuous pattern of small, highly refractive chambers, giving irregular, falselyporose appearance in bright-field illumination, both before and after demineralization (
<figureCitation id="13272A67FFC12835FB44A11806F8F9F2" box="[1156,1228,1563,1584]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 18</figureCitation>
; see Note 1). Chambers on legs 2–3 times as large as those on body (
<figureCitation id="13272A67FFC12835FC66A13B01D9F98F" box="[934,1005,1592,1613]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 17</figureCitation>
); most chambers on genital plates elongated, directed obliquely (
<figureCitation id="13272A67FFC12835FD59A15700D4F9AB" box="[665,736,1620,1641]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 16</figureCitation>
). General cuticle pale brown after demineralization, giving evidence of light sclerotization. Setae evenly birefringent throughout length.
</paragraph>
<paragraph id="8BA336E2FFC12834FEBEA18D0089FCE1" blockId="2.[317,1271,1188,1814]" lastBlockId="3.[317,1270,436,1811]" lastPageId="3" lastPageNumber="298" pageId="2" pageNumber="297">
<emphasis id="B968EAF0FFC12835FEBEA18D03DEF961" box="[382,490,1678,1699]" italics="true" pageId="2" pageNumber="297">Prodorsum</emphasis>
. Roughly triangular in dorsal aspect, with slight constriction at level of lamellar setae (
<figureCitation id="13272A67FFC12835FEBAA1A80382F902" box="[378,438,1707,1728]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="2" pageNumber="297">Figs 1</figureCitation>
,
<figureCitation id="13272A67FFC12835FE02A1A803E9F902" box="[450,477,1707,1728]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">12</figureCitation>
). Lateral prodorsal margins undulating; two concavities form niches accommodating legs I and II, respectively, with small flange between them (
<figureCitation id="13272A67FFC12835FC4EA1CB01F3F91F" box="[910,967,1736,1757]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="2" pageNumber="297">Fig. 2</figureCitation>
); more distal concavity delimits rostral lobe, best seen in anterior aspect (
<figureCitation id="13272A67FFC12835FD10A1E7012CF93B" box="[720,792,1764,1785]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 15</figureCitation>
). Rostrum with smooth, slightly reflexed margin (
<figureCitation id="13272A67FFC12835FE84A00203BFF8D4" box="[324,395,1793,1814]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="2" pageNumber="297">Fig. 14</figureCitation>
). Cuticle thicker in vague groove between bothridia (
<figureCitation id="13272A67FFC12835FC41A00201F3F8D4" box="[897,967,1793,1814]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="2" pageNumber="297">Fig. 24</figureCitation>
, arrowhead), giving appearance of darker band connecting bothridia in transmitted light (
<figureCitation id="13272A67FFC02834FC9AA6B701A5FE0B" box="[858,913,436,457]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Fig. 1</figureCitation>
), bearing sigilla for cheliceral retractor muscles. Cuticle thinner posterior to groove, where prodorsum narrows slightly, telescoping into hysterosoma, but posterior margin strongly thickened where it attaches to soft sejugal cuticle (
<figureCitation id="13272A67FFC02834FB0BA6ED034EFDE2" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="3" pageNumber="298">Figs 23, 24</figureCitation>
). Interlamellar (
<emphasis id="B968EAF0FFC02834FDD2A5080011FDE2" box="[530,549,523,544]" italics="true" pageId="3" pageNumber="298">in</emphasis>
), lamellar (
<emphasis id="B968EAF0FFC02834FD52A5080097FDE2" box="[658,675,523,544]" italics="true" pageId="3" pageNumber="298">le</emphasis>
) and rostral setae (
<emphasis id="B968EAF0FFC02834FC94A508015DFDE2" box="[852,873,523,544]" italics="true" pageId="3" pageNumber="298">ro</emphasis>
) setae attenuate, with sparse minute barbs on outer curvature; respective lengths ca. 40, 35 and 30 µm; respective mutual distances of pairs 44, 30 and 20 µm. Setae
<emphasis id="B968EAF0FFC02834FDC5A546002CFD98" box="[517,536,581,602]" italics="true" pageId="3" pageNumber="298">in</emphasis>
and
<emphasis id="B968EAF0FFC02834FD85A5460062FD98" box="[581,598,581,602]" italics="true" pageId="3" pageNumber="298">le</emphasis>
usually erect, slightly curved posteromedially and anteromedially, respectively; seta
<emphasis id="B968EAF0FFC02834FE13A56203DCFDB4" box="[467,488,609,630]" italics="true" pageId="3" pageNumber="298">ro</emphasis>
strongly curved at base, directed posterodorsally (all can be deformed during preparation). Two exobothridial setae finely attenuate;
<emphasis id="B968EAF0FFC02834FCFCA57D0167FD51" box="[828,851,638,659]" italics="true" pageId="3" pageNumber="298">xa</emphasis>
closely lateral to bothridium, ca. 33 µm, directed anterodorsally;
<emphasis id="B968EAF0FFC02834FDE8A598000BFD72" box="[552,575,667,688]" italics="true" pageId="3" pageNumber="298">xp</emphasis>
posteroventral to
<emphasis id="B968EAF0FFC02834FD30A5980133FD72" box="[752,775,667,688]" italics="true" pageId="3" pageNumber="298">xa</emphasis>
, ca. 24 µm, directed ventrally (
<figureCitation id="13272A67FFC02834FBF8A5980640FD72" box="[1080,1140,667,688]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Fig. 2</figureCitation>
). Bothridium opening on distinct swelling (
<figureCitation id="13272A67FFC02834FD99A5BB00ABFD0F" box="[601,671,696,717]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="3" pageNumber="298">Fig. 13</figureCitation>
); with three chambers, outer two with smooth walls, inner one densely spiculate (
<figureCitation id="13272A67FFC02834FE2DA5D70000FD2B" box="[493,564,724,745]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="3" pageNumber="298">Fig. 29</figureCitation>
; see Note 3). Bothridial seta (
<emphasis id="B968EAF0FFC02834FC8DA5D60151FD28" box="[845,869,725,746]" italics="true" pageId="3" pageNumber="298">bo</emphasis>
) slightly broadened and flattened distally, with 2–4 long tines (ca. 10 µm) and sparse small barbs (
<figureCitation id="13272A67FFC02834FC90A5F201B8FCC4" box="[848,908,753,774]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 1</figureCitation>
,
<figureCitation id="13272A67FFC02834FC58A5F20186FCC4" box="[920,946,753,774]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="3" pageNumber="298">13</figureCitation>
); length ca. 65 µm, about equal to mutual distance of bothridial apertures.
</paragraph>
<paragraph id="8BA336E2FFC02834FEBEA4280153F97F" blockId="3.[317,1270,436,1811]" pageId="3" pageNumber="298">
<emphasis id="B968EAF0FFC02834FEBEA42803DDFC82" box="[382,489,811,832]" italics="true" pageId="3" pageNumber="298">Notogaster</emphasis>
(
<figureCitation id="13272A67FFC02834FE36A4280078FC82" box="[502,588,811,832]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 1, 2</figureCitation>
,
<figureCitation id="13272A67FFC02834FD96A4280045FC82" box="[598,625,811,832]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="3" pageNumber="298">12</figureCitation>
). Roughly obovate in dorsal aspect, anteriorly about same width as prodorsum; 1.3–1.5 times longer than maximum width (slightly posterior to mid-length), depending on distension; moderately arched in lateral aspect, confluent with curvature of prodorsum (
<figureCitation id="13272A67FFC02834FB53A46706DFFCBB" box="[1171,1259,868,889]" captionStart="Figs 51–56" captionStartId="17.[317,360,1548,1569]" captionTargetBox="[319,1268,689,1520]" captionTargetId="figure-141@17.[319,1268,689,1520]" captionTargetPageId="17" captionText="Figs 51–56. Eniochthonius spp., adults. 51A = lateral aspect of E. mahunkai n. sp. (top, e = egg); 51B = E. minutissimus (bottom) from same sample at Cicero, New York; 52 = E. minutissimus, anterolateral aspect of bothridial seta (bo) and nearby region, specimen from Germany; 53 = E. crosbyi, dorsal aspect of bothridial seta, with insert showing dorsolateral aspect of famulus of E. crosbyi (A) andE.minutissimus (B), each ca.10 µm; 54 = E.mahunkai sp.n., ventral aspect of genital region (cf. Fig. 3; arrow points to oblique articulation between middle and anterior aggenital plates; 55 = E. crosbyi, same, but from dissected specimen; 56 = E. minutissimus, same (intact specimen from Poland). Scale bars: 100 µm (51), 10 µm (all others); ag = alveolus of aggenital seta; e = egg; pp = plicature plate. Fig. 52 SEM micrograph (from ALBERTI et al. 1994); others from light microscope" figureDoi="http://doi.org/10.5281/zenodo.12585407" httpUri="https://zenodo.org/record/12585407/files/figure.png" pageId="3" pageNumber="298">Fig. 51A</figureCitation>
). With anterior plate (pronotaspis, NA) bearing setal rows
<emphasis id="B968EAF0FFC02834FCACA4820143FC54" box="[876,887,897,918]" italics="true" pageId="3" pageNumber="298">c</emphasis>
,
<emphasis id="B968EAF0FFC02834FC44A48201A4FC54" box="[900,912,897,918]" italics="true" pageId="3" pageNumber="298">d</emphasis>
and
<emphasis id="B968EAF0FFC02834FC01A48201F8FC54" box="[961,972,897,918]" italics="true" pageId="3" pageNumber="298">e</emphasis>
; isolated from pygidial region (PY) by single transverse tectiform (type-L) scissure; tectum ca. 20 µm wide (of which solid limb ca. 15 µm) in sagittal plane, decreasing laterally to end at suprapleural plate (
<emphasis id="B968EAF0FFC02834FBC1A4B80622FC12" box="[1025,1046,955,976]" italics="true" pageId="3" pageNumber="298">sp</emphasis>
); difficult to discern in dorsal aspect of whole specimens as posterior border extremely thin (
<figureCitation id="13272A67FFC02834FC20A4DB0629FC2F" box="[992,1053,984,1005]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 1</figureCitation>
,
<figureCitation id="13272A67FFC02834FBEAA4DB0670FC2F" box="[1066,1092,984,1005]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="3" pageNumber="298">26</figureCitation>
,
<emphasis id="B968EAF0FFC02834FB8FA4DB0656FC2F" box="[1103,1122,984,1005]" italics="true" pageId="3" pageNumber="298">bt</emphasis>
). In contracted specimens, anterior margin of pygidium (
<emphasis id="B968EAF0FFC02834FD09A4F600D5FBC8" box="[713,737,1013,1034]" italics="true" pageId="3" pageNumber="298">bp</emphasis>
) telescoping underneath tectum to level ca. 20 µm anterior to seta
<emphasis id="B968EAF0FFC02834FE65A3120384FBE4" box="[421,432,1041,1062]" italics="true" pageId="3" pageNumber="298">e</emphasis>
<subScript id="179834A7FFC02834FE6FA31D0382FBE8" attach="left" box="[431,438,1054,1066]" fontSize="5" pageId="3" pageNumber="298">1</subScript>
. Transverse sulcus of pronotaspis emphasized in transmitted light by slightly thicker cuticle (
<figureCitation id="13272A67FFC02834FEB4A32D0385FB81" box="[372,433,1070,1091]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 1</figureCitation>
,
<figureCitation id="13272A67FFC02834FE7DA32D03E3FB81" box="[445,471,1070,1091]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="3" pageNumber="298">25</figureCitation>
), weakest medially, between setal pair
<emphasis id="B968EAF0FFC02834FC93A32D016BFB81" box="[851,863,1070,1091]" italics="true" pageId="3" pageNumber="298">d</emphasis>
<subScript id="179834A7FFC02834FC9FA3390152FB84" attach="left" box="[863,870,1082,1094]" fontSize="5" pageId="3" pageNumber="298">1</subScript>
. Pronotaspis isolated from pleural region (PL) by paired elongated suprapleural plate (ca. 15 times longer than wide), extending anteriorly from about level of setal row
<emphasis id="B968EAF0FFC02834FD91A36B006CFBBF" box="[593,600,1128,1149]" italics="true" pageId="3" pageNumber="298">f</emphasis>
to within about 10 µm of sejugal articulation; plate ca. 125 µm long, 10 µm at widest point, bearing setae
<emphasis id="B968EAF0FFC02834FDBBA38600BAFB5F" box="[635,654,1157,1181]" italics="true" pageId="3" pageNumber="298">
h
<subScript id="179834A7FFC02834FD47A39200BAFB5F" attach="left" box="[647,654,1169,1181]" fontSize="5" pageId="3" pageNumber="298">2</subScript>
</emphasis>
,
<emphasis id="B968EAF0FFC02834FD59A3860098FB5F" box="[665,684,1157,1181]" italics="true" pageId="3" pageNumber="298">
h
<subScript id="179834A7FFC02834FD65A3920098FB5F" attach="left" box="[677,684,1169,1181]" fontSize="5" pageId="3" pageNumber="298">
<quantity id="4CE49B07FFC02834FD65A3920098FB5F" box="[677,684,1169,1181]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="298" unit="in" value="3.0">3</quantity>
</subScript>
</emphasis>
in posterior quarter (
<figureCitation id="13272A67FFC02834FCB4A387019AFB5B" box="[884,942,1156,1177]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Fig. 2</figureCitation>
);
<emphasis id="B968EAF0FFC02834FC00A38601E7FB5F" box="[960,979,1157,1181]" italics="true" pageId="3" pageNumber="298">
h
<subScript id="179834A7FFC02834FC0CA39201E7FB5F" attach="left" box="[972,979,1169,1181]" fontSize="5" pageId="3" pageNumber="298">3</subScript>
</emphasis>
reaches posterior end of plate,
<emphasis id="B968EAF0FFC02834FEFDA3A20364FB78" box="[317,336,1185,1210]" italics="true" pageId="3" pageNumber="298">
h
<subScript id="179834A7FFC02834FE89A3AD0364FB78" attach="left" box="[329,336,1198,1210]" fontSize="5" pageId="3" pageNumber="298">2</subScript>
</emphasis>
extends well beyond it. Pleural region with narrow, laterally projecting carina originating near level of seta
<emphasis id="B968EAF0FFC02834FE76A3BD03F3FB15" box="[438,455,1214,1239]" italics="true" pageId="3" pageNumber="298">
c
<subScript id="179834A7FFC02834FE00A3C803F3FB15" attach="left" box="[448,455,1227,1239]" fontSize="5" pageId="3" pageNumber="298">3</subScript>
</emphasis>
and merging posteriorly with notogastral margin, just posterior to level of legs IV; in anterior half, ventral surface of carina invaginated as taenidium (
<figureCitation id="13272A67FFC02834FC7AA3D80618FB32" box="[954,1068,1243,1264]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="3" pageNumber="298">Figs 30–32</figureCitation>
). Posterior to leg IV taenidium curves ventrad, away from pleural carina; continuing as short, shallow groove onto aggenital plate (see below). Hidden arched wall of taenidium bearing inconspicuous elongated porose area (ca. 25–30 µm long; see Note 4) in region dorsal to legs III and IV (
<figureCitation id="13272A67FFC02834FBE8A2320651FA84" box="[1064,1125,1329,1350]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 2</figureCitation>
,
<emphasis id="B968EAF0FFC02834FBAEA23206B2FA84" box="[1134,1158,1329,1350]" italics="true" pageId="3" pageNumber="298">pa</emphasis>
; 33); rarely two widely separated smaller porose areas present instead. Lateral margin of notogaster sharply folded dorsomediad in posterior two-thirds to form plicature plate (
<figureCitation id="13272A67FFC02834FC01A26801CFFA42" box="[961,1019,1387,1408]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Fig. 3</figureCitation>
,
<emphasis id="B968EAF0FFC02834FBC6A268062AFA42" box="[1030,1054,1387,1408]" italics="true" pageId="3" pageNumber="298">pp</emphasis>
), hidden in contracted specimens (see Note 5); plicature plate not mineralized, without epicuticular chambers. Five pairs of lyrifissures present (see Note 6);
<emphasis id="B968EAF0FFC02834FDBBA2A600BAFA78" box="[635,654,1445,1466]" italics="true" pageId="3" pageNumber="298">ia</emphasis>
slit-like, inconspicuous, aligned with edge of pleural carina and lying between it and porose area (
<figureCitation id="13272A67FFC02834FD51A2C200EEFA14" box="[657,730,1473,1494]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="3" pageNumber="298">Fig. 33</figureCitation>
, arrowhead). Others with cupular form (
<figureCitation id="13272A67FFC02834FBADA2C206FCFA14" box="[1133,1224,1473,1494]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 2, 3</figureCitation>
):
<emphasis id="B968EAF0FFC02834FB1DA2C206C1FA14" box="[1245,1269,1473,1494]" italics="true" pageId="3" pageNumber="298">im</emphasis>
posterolateral on pronotaspis, about 10 µm from its lateral edge;
<emphasis id="B968EAF0FFC02834FC62A2DD0181FA31" box="[930,949,1502,1523]" italics="true" pageId="3" pageNumber="298">ip</emphasis>
far anterior in pleural region near posterior end of pleural carina;
<emphasis id="B968EAF0FFC02834FDABA2F8004AF9D2" box="[619,638,1531,1552]" italics="true" pageId="3" pageNumber="298">ih</emphasis>
approximately aligned between
<emphasis id="B968EAF0FFC02834FC78A2F801E4F9D2" box="[952,976,1531,1552]" italics="true" pageId="3" pageNumber="298">im</emphasis>
and seta
<emphasis id="B968EAF0FFC02834FBEAA2F80609F9D1" box="[1066,1085,1531,1555]" italics="true" pageId="3" pageNumber="298">
p
<subScript id="179834A7FFC02834FBF6A1040609F9D1" attach="left" box="[1078,1085,1543,1555]" fontSize="5" pageId="3" pageNumber="298">3</subScript>
</emphasis>
, about 30 µm from the latter;
<emphasis id="B968EAF0FFC02834FE5DA11B038DF9EF" box="[413,441,1560,1581]" italics="true" pageId="3" pageNumber="298">ips</emphasis>
on plicature plate about at level of seta
<emphasis id="B968EAF0FFC02834FCF0A11B017CF9EF" box="[816,840,1560,1581]" italics="true" pageId="3" pageNumber="298">ad</emphasis>
<subScript id="179834A7FFC02834FC88A127017BF9F2" attach="left" box="[840,847,1572,1584]" fontSize="5" pageId="3" pageNumber="298">3</subScript>
. Notogastral setae mostly attenuate, posteriorly directed, slightly curved, with minute serration of barbs dorsally, difficult to see except in profile (
<figureCitation id="13272A67FFC02834FE84A15203B8F9A4" box="[324,396,1617,1638]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="3" pageNumber="298">Fig. 35</figureCitation>
). Setae mostly 40–55 µm long; when flattened by coverglass many reaching level of insertions for next most posterior row;
<emphasis id="B968EAF0FFC02834FD40A16D00A5F945" box="[640,657,1646,1671]" italics="true" pageId="3" pageNumber="298">
e
<subScript id="179834A7FFC02834FD4AA17800A5F945" attach="left" box="[650,657,1659,1671]" fontSize="5" pageId="3" pageNumber="298">2</subScript>
</emphasis>
and
<emphasis id="B968EAF0FFC02834FD7CA16D00FBF945" box="[700,719,1646,1671]" italics="true" pageId="3" pageNumber="298">
p
<subScript id="179834A7FFC02834FD08A17800FBF945" attach="left" box="[712,719,1659,1671]" fontSize="5" pageId="3" pageNumber="298">1</subScript>
</emphasis>
longest, latter extending slightly beyond posterior margin in dorsal aspect; seta
<emphasis id="B968EAF0FFC02834FDC9A188002FF961" box="[521,539,1675,1699]" italics="true" pageId="3" pageNumber="298">
c
<subScript id="179834A7FFC02834FDD4A194002FF961" attach="left" box="[532,539,1687,1699]" fontSize="5" pageId="3" pageNumber="298">3</subScript>
</emphasis>
much smaller (ca. 15 µm), finely attenuate and flexible, inconspicuously inserted in anterolateral corner of pronotaspis (
<figureCitation id="13272A67FFC02834FD35A1AB0107F97F" box="[757,819,1704,1725]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="3" pageNumber="298">Figs 1</figureCitation>
,
<figureCitation id="13272A67FFC02834FC80A1AB016EF97F" box="[832,858,1704,1725]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="3" pageNumber="298">13</figureCitation>
).
</paragraph>
<paragraph id="8BA336E2FFC02833FEBEA1C60174F8E1" blockId="3.[317,1270,436,1811]" lastBlockId="4.[317,1270,1748,1827]" lastPageId="4" lastPageNumber="299" pageId="3" pageNumber="298">
<emphasis id="B968EAF0FFC02834FEBEA1C603CDF918" box="[382,505,1733,1754]" italics="true" pageId="3" pageNumber="298">Coxisternum</emphasis>
. Epimeres I and II fully fused, ca. 80 µm wide; most of surface with granular cerotegument (
<figureCitation id="13272A67FFC02834FE0DA1E20027F934" box="[461,531,1761,1782]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="3" pageNumber="298">Fig. 21</figureCitation>
) inconspicuous in transmitted light due to epicuticular chambers. Epimere II bending abruptly dorsad just posterior to seta
<emphasis id="B968EAF0FFC02834FD29A1FD0135F8D1" box="[745,769,1790,1811]" italics="true" pageId="3" pageNumber="298">2a</emphasis>
, then again posteriorly, prior to joining soft articulat- (50 µm)
</paragraph>
<caption id="DF63666AFFC72833FEFDA1D706C2F8C4" ID-DOI="http://doi.org/10.5281/zenodo.12585389" ID-Zenodo-Dep="12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="4" pageNumber="299" startId="4.[317,360,1748,1769]" targetBox="[362,1222,429,1712]" targetPageId="4" targetType="figure">
<paragraph id="8BA336E2FFC72833FEFDA1D706C2F8C4" blockId="4.[317,1270,1748,1827]" pageId="4" pageNumber="299">
<emphasis id="B968EAF0FFC72833FEFDA1D703ACF92B" bold="true" box="[317,408,1748,1769]" pageId="4" pageNumber="299">Figs 1–3.</emphasis>
<taxonomicName id="4C1C4D61FFC72833FE5DA1D700B9F92B" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[413,653,1748,1769]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC72833FE5DA1D700B9F92B" box="[413,653,1748,1769]" italics="true" pageId="4" pageNumber="299">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC72833FD52A1D700F3F92B" box="[658,711,1748,1769]" pageId="4" pageNumber="299" rank="species">sp. n.</taxonomicNameLabel>
, adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale
</paragraph>
</caption>
<caption id="DF63666AFFC62832FEFDA16F06C1F915" ID-DOI="http://doi.org/10.5281/zenodo.12585391" ID-Zenodo-Dep="12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="5" pageNumber="300" startId="5.[317,360,1644,1665]" targetBox="[325,1264,428,1595]" targetPageId="5" targetType="figure">
<paragraph id="8BA336E2FFC62832FEFDA16F06C1F915" blockId="5.[317,1270,1644,1780]" pageId="5" pageNumber="300">
<emphasis id="B968EAF0FFC62832FEFDA16F03ACF943" bold="true" box="[317,408,1644,1665]" pageId="5" pageNumber="300">Figs 4–8.</emphasis>
<taxonomicName id="4C1C4D61FFC62832FE5EA16F00B9F943" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[414,653,1644,1665]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC62832FE5EA16F00B9F943" box="[414,653,1644,1665]" italics="true" pageId="5" pageNumber="300">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC62832FD53A16F00FCF943" box="[659,712,1644,1665]" pageId="5" pageNumber="300" rank="species">sp. n.</taxonomicNameLabel>
, adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (
<emphasis id="B968EAF0FFC62832FD5DA1A5009DF979" box="[669,681,1702,1723]" italics="true" pageId="5" pageNumber="300">p</emphasis>
) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig. 5, indicated only by dotted circles:
<emphasis id="B968EAF0FFC62832FCEEA1C10176F915" box="[814,834,1730,1751]" italics="true" pageId="5" pageNumber="300">a’</emphasis>
and
<emphasis id="B968EAF0FFC62832FCAEA1C101B3F915" box="[878,903,1730,1751]" italics="true" pageId="5" pageNumber="300">tc’</emphasis>
(respectively below and above seta a”
</paragraph>
</caption>
<paragraph id="8BA336E2FFC62832FE87A1DC06D9F936" blockId="5.[317,1270,1644,1780]" box="[327,1261,1759,1780]" pageId="5" pageNumber="300">
in figure); and several others on
<figureCitation id="13272A67FFC62832FDB4A1DC00DFF936" box="[628,747,1759,1780]" captionStart="Figs 4–8" captionStartId="5.[317,360,1644,1665]" captionTargetBox="[325,1264,428,1595]" captionTargetId="figure-15@5.[317,1269,428,1610]" captionTargetPageId="5" captionText="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" figureDoi="http://doi.org/10.5281/zenodo.12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="5" pageNumber="300">Figs. 6 and 8</figureCitation>
that can be interpreted from
<figureCitation id="13272A67FFC62832FC37A1DC065DF936" box="[1015,1129,1759,1780]" captionStart="Figs 4–8" captionStartId="5.[317,360,1644,1665]" captionTargetBox="[325,1264,428,1595]" captionTargetId="figure-15@5.[317,1269,428,1610]" captionTargetPageId="5" captionText="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" figureDoi="http://doi.org/10.5281/zenodo.12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="5" pageNumber="300">Figs 4 and 7</figureCitation>
, respectively.
</paragraph>
<paragraph id="8BA336E2FFC52831FEFDA6B7003DFC82" blockId="6.[317,1271,436,1149]" pageId="6" pageNumber="301">
ing sejugal cuticle, forming “neck” in proterosoma that inserts into hysterosoma during contracted state (
<figureCitation id="13272A67FFC52831FEB6A6D20381FE24" box="[374,437,465,486]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="6" pageNumber="301">Figs 2</figureCitation>
,
<figureCitation id="13272A67FFC52831FE00A6D20034FE24" box="[448,512,465,486]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="6" pageNumber="301">23, 27</figureCitation>
). Sub-cruciform pattern of narrow grooves on surface (
<figureCitation id="13272A67FFC52831FBD6A6D20668FE24" box="[1046,1116,465,486]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="6" pageNumber="301">Fig. 21</figureCitation>
) overlying well formed apodeme 2 and anterior sternal apodeme; latter ending posteriorly at change in contour. Supracoxal gland opening at dorsal margin of epimere I, at level of epimeral seta
<emphasis id="B968EAF0FFC52831FBF7A508067AFDE2" box="[1079,1102,523,544]" italics="true" pageId="6" pageNumber="301">1a</emphasis>
; cuticle along canal border projecting internally in row of small teeth for about 15 µm anterior to aperture. Supracoxal seta
<emphasis id="B968EAF0FFC52831FEA8A546034FFD98" box="[360,379,581,602]" italics="true" pageId="6" pageNumber="301">eI</emphasis>
ca. 5 µm, spiniform, nearly isodiametric but with minute terminal fork; inserted immediately dorsal to trochanter I, below podocephalic canal. Epimeres III and IV fully fused with each other, ca. 90 µm wide; surface with weak grooves and cerotegument granules marking positions of underlying apodemes (
<figureCitation id="13272A67FFC52831FE6BA59803C1FD72" box="[427,501,667,688]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="6" pageNumber="301">Fig. 21</figureCitation>
). Epimere IV also fully fused to anterior aggenital plate. Posterior sternal apodeme conspicuous, about 30 µm long, ending posteriorly about at level of seta
<emphasis id="B968EAF0FFC52831FBE8A5BB0674FD0F" box="[1064,1088,696,717]" italics="true" pageId="6" pageNumber="301">4b</emphasis>
. Apodeme 3 short, reaching at most halfway to midline; apodeme 4 absent (
<figureCitation id="13272A67FFC52831FCA3A5D701AAFD2B" box="[867,926,724,745]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="6" pageNumber="301">Fig. 3</figureCitation>
). Articulations with leg trochanters sunken, no soft cuticle exposed, approaching ball-socket formation. Epimeral setal formula 3–1–3–4; setae finely attenuate, apparently smooth, mostly 10–15 µm; seta
<emphasis id="B968EAF0FFC52831FC68A40D01F4FCE1" box="[936,960,782,803]" italics="true" pageId="6" pageNumber="301">1a</emphasis>
slightly longer (ca. 20 µm), with inconspicuous barbs.
</paragraph>
<paragraph id="8BA336E2FFC52831FEBEA44B01E9F8E6" blockId="6.[317,1271,436,1149]" lastBlockId="6.[317,1270,1749,1828]" pageId="6" pageNumber="301">
<emphasis id="B968EAF0FFC52831FEBEA44B0000FC9F" box="[382,564,840,861]" italics="true" pageId="6" pageNumber="301">Anogenital region.</emphasis>
With three pairs of mutually articulated aggenital plates that collectively bracket the genital and preanal plates (
<figureCitation id="13272A67FFC52831FD73A46700D9FCBB" box="[691,749,868,889]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="6" pageNumber="301">Fig. 3</figureCitation>
). Anterior plate largest, roughly triangular, ca. 40 µm long, imperceptibly fused to epimere IV, with two well marked impressions; medial one (close to genital plate) broad, shallow; lateral one (postpedal fossa
<emphasis id="B968EAF0FFC52831FCB0A49D01B7FC71" box="[880,899,926,947]" italics="true" pageId="6" pageNumber="301">fo</emphasis>
) behind leg IV insertion, narrow with well-defined aperture (
<figureCitation id="13272A67FFC52831FDDBA4B8006FFC12" box="[539,603,955,976]" captionStart="Figs 1–3" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[362,1222,429,1712]" captionTargetId="figure-19@4.[361,1225,424,1720]" captionTargetPageId="4" captionText="Figs 1–3. Eniochthonius mahunkai sp. n., adult (contracted specimen): 1 = dorsal aspect; 2 = lateral aspect; 3 = ventral aspect. Legs and gnathosoma partly shown (1, 3) or omitted (2). All to same scale" figureDoi="http://doi.org/10.5281/zenodo.12585389" httpUri="https://zenodo.org/record/12585389/files/figure.png" pageId="6" pageNumber="301">Figs 2</figureCitation>
,
<figureCitation id="13272A67FFC52831FDA7A4B800B6FC12" box="[615,642,955,976]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="6" pageNumber="301">21</figureCitation>
). Vague, shallow groove extending laterally from fossa towards pleural region of notogaster, aligned with pleural taenidium (
<figureCitation id="13272A67FFC52831FC45A4DB01F9FC2F" box="[901,973,984,1005]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="6" pageNumber="301">Fig. 31</figureCitation>
). Middle aggenital plate small, 4-sided polygon, ca. 22 µm in longest dimension; bearing seta
<emphasis id="B968EAF0FFC52831FC6FA4F601F3FBC8" box="[943,967,1013,1034]" italics="true" pageId="6" pageNumber="301">ag</emphasis>
(simple, ca. 12 µm). Posterior aggenital plate narrowly sub-rectangular, slightly wider posteriorly, ca. 35 µm long. Genital plates roughly pentagonal as pair, each truncated posteriorly; large, ca. 90 µm long, 55 µm at widest point; narrow band of unsclerotized cuticle cutting transversely (slightly obliquely) across plate at widest point and meeting articulation between second and third aggenital plates laterally; band evident by bars: 20 µm (9), 10 µm (10), 25 µm (11).
</paragraph>
<caption id="DF63666AFFC52831FEFDA1D606C2F8C5" ID-DOI="http://doi.org/10.5281/zenodo.12585393" ID-Zenodo-Dep="12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="6" pageNumber="301" startId="6.[317,360,1749,1770]" targetBox="[323,1268,1174,1717]" targetPageId="6" targetType="figure">
<paragraph id="8BA336E2FFC52831FEFDA1D606C2F8C5" blockId="6.[317,1270,1749,1828]" pageId="6" pageNumber="301">
<emphasis id="B968EAF0FFC52831FEFDA1D60397F928" bold="true" box="[317,419,1749,1770]" pageId="6" pageNumber="301">Figs 9–11.</emphasis>
<taxonomicName id="4C1C4D61FFC52831FE68A1D500A3F929" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[424,663,1750,1771]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="301" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC52831FE68A1D500A3F929" box="[424,663,1750,1771]" italics="true" pageId="6" pageNumber="301">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC52831FD5CA1D500E4F929" box="[668,720,1750,1771]" pageId="6" pageNumber="301" rank="species">sp. n.</taxonomicNameLabel>
, adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale
</paragraph>
</caption>
<paragraph id="8BA336E2FFC42830FEFDA6B706C2FD51" blockId="7.[317,1270,436,662]" pageId="7" pageNumber="302">
lightness in transmitted light, chambered epicuticle above it remaining mineralized, but in pattern different from that of surrounding epicuticle (
<figureCitation id="13272A67FFC42830FD2FA6D20102FE24" box="[751,822,465,486]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="7" pageNumber="302">Fig. 16</figureCitation>
); with sparse cerotegument granules anteriorly (
<figureCitation id="13272A67FFC42830FE84A6ED03B8FDC1" box="[324,396,494,515]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="7" pageNumber="302">Fig. 21</figureCitation>
). Ten pairs of genital setae forming two longitudinal rows, seven in medial and three in lateral; four medial and one lateral seta anterior to transverse band; spacing of medial row setae slightly variable; setae attenuate, without noticeable barbs, 12–15 µm. Ovipositor strongly pleated, short, only ca. 30 µm long when folded (just proximal to terminal lobes); directed posteroventrad when in body (
<figureCitation id="13272A67FFC42830FEB8A562038AFDB4" box="[376,446,609,630]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="7" pageNumber="302">Fig. 23</figureCitation>
,
<emphasis id="B968EAF0FFC42830FE06A56203E9FDB4" box="[454,477,609,630]" italics="true" pageId="7" pageNumber="302">ov</emphasis>
), so that paired dorsal lobes in ventral position; with nine pairs of setae: τ
<subScript id="179834A7FFC42830FB4AA56D06A5FDB8" attach="left" box="[1162,1169,622,634]" fontSize="5" pageId="7" pageNumber="302">1</subScript>
and Ψ
<subScript id="179834A7FFC42830FB0FA56D06E2FDB8" attach="left" box="[1231,1238,622,634]" fontSize="5" pageId="7" pageNumber="302">1</subScript>
ca. 35 µm; τ
<subScript id="179834A7FFC42830FE4FA5890390FD54" attach="left" box="[399,420,650,662]" fontSize="5" pageId="7" pageNumber="302">2–4</subScript>
, Ψ
<subScript id="179834A7FFC42830FE01A58903FCFD54" attach="left" box="[449,456,650,662]" fontSize="5" pageId="7" pageNumber="302">2</subScript>
and three pairs of coronal setae 13–15 µm; all setae eupathidial. Genital papillae rela-
</paragraph>
<caption id="DF63666AFFC42830FE82A5CB06C4FD38" pageId="7" pageNumber="302" startId="7.[322,381,712,733]" targetType="table">
<paragraph id="8BA336E2FFC42830FE82A5CB06C4FD38" blockId="7.[322,1264,712,864]" pageId="7" pageNumber="302">
<emphasis id="B968EAF0FFC42830FE82A5CB03A0FD1F" bold="true" box="[322,404,712,733]" pageId="7" pageNumber="302">Table 1.</emphasis>
Ontogeny of setae and solenidia in
<taxonomicName id="4C1C4D61FFC42830FD21A5CB01FBFD1F" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[737,975,712,733]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="302" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC42830FD21A5CB01FBFD1F" box="[737,975,712,733]" italics="true" pageId="7" pageNumber="302">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC42830FC13A5CB0631FD1F" box="[979,1029,712,733]" pageId="7" pageNumber="302" rank="species">sp. n.</taxonomicNameLabel>
(Structures are indicated where they are first added and are present through the rest of ontogeny; setae in parentheses repre-
</paragraph>
</caption>
<paragraph id="8BA336E2FFC42830FE35A401060AFCD5" blockId="7.[322,1264,712,864]" box="[501,1086,770,791]" pageId="7" pageNumber="302">sent pairs; dash indicates no additions. See text for variation)</paragraph>
<caption id="DF63666AFFCB283FFEFDA15C06C2F8C7" ID-DOI="http://doi.org/10.5281/zenodo.12585395" ID-Zenodo-Dep="12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="8" pageNumber="303" startId="8.[317,360,1631,1652]" targetBox="[364,1222,423,1603]" targetPageId="8" targetType="figure">
<paragraph id="8BA336E2FFCB283FFEFDA15C06C2F8C7" blockId="8.[317,1270,1631,1825]" pageId="8" pageNumber="303">
<emphasis id="B968EAF0FFCB283FFEFDA15C039BF9B6" bold="true" box="[317,431,1631,1652]" pageId="8" pageNumber="303">Figs 12–18.</emphasis>
<taxonomicName id="4C1C4D61FFCB283FFE75A1630090F9B7" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[437,676,1632,1653]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="303" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFCB283FFE75A1630090F9B7" box="[437,676,1632,1653]" italics="true" pageId="8" pageNumber="303">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFCB283FFD69A16300EAF9B7" box="[681,734,1632,1653]" pageId="8" pageNumber="303" rank="species">sp. n.</taxonomicNameLabel>
, adult: 12 = dorsal habitus; 13 = bothridial seta (
<emphasis id="B968EAF0FFCB283FFB6DA16306F1F9B7" box="[1197,1221,1632,1653]" italics="true" pageId="8" pageNumber="303">bo</emphasis>
) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-
</paragraph>
</caption>
<paragraph id="8BA336E2FFCB283FFEADA00F06F3F8E3" blockId="8.[317,1270,1631,1825]" box="[365,1223,1804,1825]" pageId="8" pageNumber="303">graphs; 16–18 light micrographs. Scale bars: 50 µm (12), 10 µm (13–15), 20 µm (16–18).</paragraph>
<paragraph id="8BA336E2FFCA283EFEFDA6B7068BFD9B" blockId="9.[317,1270,436,1811]" pageId="9" pageNumber="304">
tively large (10–12 µm diameter), all three similar. Preanal plate (
<figureCitation id="13272A67FFCA283EFC6EA6B701CDFE0B" box="[942,1017,436,457]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="9" pageNumber="304">Figs 22</figureCitation>
,
<figureCitation id="13272A67FFCA283EFBC2A6B70628FE0B" box="[1026,1052,436,457]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="9" pageNumber="304">28</figureCitation>
,
<emphasis id="B968EAF0FFCA283EFBE6A6B6060FFE08" box="[1062,1083,437,458]" italics="true" pageId="9" pageNumber="304">pr</emphasis>
) narrow, ca. 45 µm across, may be inconspicuous in contracted specimens. Collectively, anal and adanal plates ca. 60 µm long, 50 µm wide anteriorly, tapering posteriorly to 20–25 µm wide at level of seta
<emphasis id="B968EAF0FFCA283EFB9BA6ED0647FDC1" box="[1115,1139,494,515]" italics="true" pageId="9" pageNumber="304">ad</emphasis>
<subScript id="179834A7FFCA283EFBB3A6F9064EFDC4" attach="left" box="[1139,1146,506,518]" fontSize="5" pageId="9" pageNumber="304">1</subScript>
; without noticeable cerotegument. Three pairs of adanal setae 15 µm, two pairs of anal setae 15 µm; all attenuate, smooth. Anteromedial margin of each anal plate projected internally as contiguous pair of solid, horn-like, heavily sclerotized anal apodemes (
<figureCitation id="13272A67FFCA283EFD3EA5470173FD9B" box="[766,839,580,601]" captionStart="Figs 23–28" captionStartId="11.[317,360,1522,1543]" captionTargetBox="[332,1256,419,1493]" captionTargetId="figure-15@11.[332,1256,419,1493]" captionTargetPageId="11" captionText="Figs 23–28. Eniochthonius mahunkai sp. n., adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla).Scale bar: 50 µm (23)," figureDoi="http://doi.org/10.5281/zenodo.12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="9" pageNumber="304">Fig. 28</figureCitation>
,
<emphasis id="B968EAF0FFCA283EFC92A546015EFD98" box="[850,874,581,602]" italics="true" pageId="9" pageNumber="304">aa</emphasis>
), conspicuous in transmitted light.
</paragraph>
<paragraph id="8BA336E2FFCA283EFEBEA5620032FAEB" blockId="9.[317,1270,436,1811]" pageId="9" pageNumber="304">
<emphasis id="B968EAF0FFCA283EFEBEA562039FFDB4" box="[382,427,609,630]" italics="true" pageId="9" pageNumber="304">Legs</emphasis>
(
<figureCitation id="13272A67FFCA283EFE77A5620039FDB4" box="[439,525,609,630]" captionStart="Figs 4–8" captionStartId="5.[317,360,1644,1665]" captionTargetBox="[325,1264,428,1595]" captionTargetId="figure-15@5.[317,1269,428,1610]" captionTargetPageId="5" captionText="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" figureDoi="http://doi.org/10.5281/zenodo.12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="9" pageNumber="304">Figs 4–8</figureCitation>
). Legs all relatively short, similar in size, ca. 0.35 to 0.40 times body length, IV longest; simple in structure, without bosses, tubercles or surface sculpturing other than fine, wrinkle-like grooves running obliquely to transversely across segments. Basally, segments with constriction near articulation; constriction strongest dorsally on femora, ventrally on more distal segments, and only weakly defined on tarsi III–IV. Articulations tight, almost ball-socket
<typeStatus id="54A78840FFCA283EFBF1A5D70669FD2B" box="[1073,1117,724,745]" pageId="9" pageNumber="304">type</typeStatus>
, without clearly defined protecta, but soft articulating cuticle not exposed, even ventrally, regardless of degree of flexing. All trochanters, and femora I and II, with cerotegument granules. All tarsi with normal, proximodorsal lyrifissure. Single claw simple in structure, without barbs or dentition. Leg setae attenuate to sub-flagellate, mostly smooth or with sparse, minute barbs; seta
<emphasis id="B968EAF0FFCA283EFBD3A44B0613FC9F" box="[1043,1063,840,861]" italics="true" pageId="9" pageNumber="304">a’</emphasis>
on tarsi I and II with several strong barbs (not illustrated). Setation (legs I-IV, including famulus but not solenidia) of trochanters (0–0–2–2), femora (3–5–3–3), genua (5–3–3–3), tibiae (5–4–3–3), tarsi (18–15–13–13); no variation noted (but slight variations in ontogeny, described below); setal homologies given in
<figureCitation id="13272A67FFCA283EFB0CA49D0355FC12" captionStart="Figs 4–8" captionStartId="5.[317,360,1644,1665]" captionTargetBox="[325,1264,428,1595]" captionTargetId="figure-15@5.[317,1269,428,1610]" captionTargetPageId="5" captionText="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" figureDoi="http://doi.org/10.5281/zenodo.12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="9" pageNumber="304">Figs 4–8</figureCitation>
and in
<tableCitation id="C69E0359FFCA283EFE63A4B803DEFC12" box="[419,490,955,976]" captionStart="Table 1" captionStartId="7.[322,381,712,733]" captionText="Table 1. Ontogeny of setae and solenidia in Eniochthonius mahunkai sp.n. (Structures are indicated where they are first added and are present through the rest of ontogeny; setae in parentheses repre-" pageId="9" pageNumber="304">Table 1</tableCitation>
(see also Note 11). Seta
<emphasis id="B968EAF0FFCA283EFD0FA4B800EFFC12" box="[719,731,955,976]" italics="true" pageId="9" pageNumber="304">d</emphasis>
of tibiae I and II minute, coupled to respective solenidion ϕ, in adjacent but separate alveoli; seta
<emphasis id="B968EAF0FFCA283EFD0BA4DB00E3FC2F" box="[715,727,984,1005]" italics="true" pageId="9" pageNumber="304">d</emphasis>
of tibia III and of all genua independent of respective solenidia. Solenidial formula (legs I–IV) of genua (2–1–1–1), tibiae (1–1–1–0), tarsi (3–1–0–0). Genual solenidia all piliform; σ”I proximal to seta
<emphasis id="B968EAF0FFCA283EFCE6A3120106FBE4" box="[806,818,1041,1062]" italics="true" pageId="9" pageNumber="304">d</emphasis>
, σ’ I approximately lateral to it; σII placed like σ”I, slightly on posterior face; σ of genua III and IV also slightly on posterior face, but well distal to seta
<emphasis id="B968EAF0FFCA283EFEA8A3480340FBA2" box="[360,372,1099,1120]" italics="true" pageId="9" pageNumber="304">d</emphasis>
. Tibial solenidion ϕI tactile (flagellate, ca. 70 µm), directed dorsodistally, about half length of leg I; solenidion ϕII ceratiform, slightly shorter than segment length, strongly curved toward posterior face; ϕIII piliform, with form and position similar to σIII, distal to
<emphasis id="B968EAF0FFCA283EFC1DA38601DDFB58" box="[989,1001,1157,1178]" italics="true" pageId="9" pageNumber="304">d</emphasis>
. Tarsal solenidion ω1 large, ceratiform, curved toward segment, and ending directed toward base of claw; ω2 and ω3 piliform, closely aligned together, lateral to famulus on posterior face; ω of tarsus II similar to ω1 of tarsus I, but slightly smaller. Famulus (
<emphasis id="B968EAF0FFCA283EFDACA3D80043FB32" box="[620,631,1243,1264]" italics="true" pageId="9" pageNumber="304">e</emphasis>
) aligned between seta
<emphasis id="B968EAF0FFCA283EFC9CA3D80145FB32" box="[860,881,1243,1264]" italics="true" pageId="9" pageNumber="304">ft’</emphasis>
and solenidion ω2; forked: one branch thick, terminating bluntly or with slight knob-like swelling, other attenuate, bract-like, about onethird longer (
<figureCitation id="13272A67FFCA283EFE7EA21703CEFAEB" box="[446,506,1300,1321]" captionStart="Figs 4–8" captionStartId="5.[317,360,1644,1665]" captionTargetBox="[325,1264,428,1595]" captionTargetId="figure-15@5.[317,1269,428,1610]" captionTargetPageId="5" captionText="Figs 4–8. Eniochthonius mahunkai sp. n., adult: 4 = leg I, femur, genu and tibia; 5 = leg I, tibia and tarsus; 6 = leg II, except trochanter; 7 = leg III; 8 = leg IV. All abaxial view; all to same scale (50 µm). Unlabeled setae include: proral pair (p) inserted dorsally at base of claws on all legs; two setae on reverse side of Fig.5, indicated only by dotted circles: a’ and tc’ (respectively below and above seta a”" figureDoi="http://doi.org/10.5281/zenodo.12585391" httpUri="https://zenodo.org/record/12585391/files/figure.png" pageId="9" pageNumber="304">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BA336E2FFCA283CFEBEA23201B4F8E5" blockId="9.[317,1270,436,1811]" lastBlockId="11.[317,1270,1522,1831]" lastPageId="11" lastPageNumber="306" pageId="9" pageNumber="304">
<taxonomicName id="4C1C4D61FFCA283EFEBEA23203CDFA84" authorityName="Krefft" authorityYear="1871" box="[382,505,1329,1350]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="304" phylum="Nematoda" rank="genus">
<emphasis id="B968EAF0FFCA283EFEBEA23203CDFA84" box="[382,505,1329,1350]" italics="true" pageId="9" pageNumber="304">Gnathosoma</emphasis>
</taxonomicName>
. Subcapitulum anarthric but with paired pattern of wrinkles and grooves leading posterolaterally from inferior commissure of mouth (in normal position of labiogenal articulation; see Note 14), delineating region of triangular mentum from paired genae (
<figureCitation id="13272A67FFCA283EFC3FA2680609FA42" box="[1023,1085,1387,1408]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="9" pageNumber="304">Figs 9</figureCitation>
,
<figureCitation id="13272A67FFCA283EFB87A2680656FA42" box="[1095,1122,1387,1408]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="9" pageNumber="304">20</figureCitation>
); cerotegument granules small, vague, restricted to periphery. Dorsal surface of genae with extensive, paired, rasp-like region. No dentition observed on labrum. With small, rectangular capitular apodeme, ca. 10 µm wide, projecting 15 µm posteriorly from subcapitular cervix. Rutellum (
<figureCitation id="13272A67FFCA283EFBEFA2C20659FA14" box="[1071,1133,1473,1494]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="9" pageNumber="304">Fig. 9</figureCitation>
, RU) narrow, seemingly inserted in genu like large seta, slightly broadened distally with three obliquely aligned terminal teeth (see Note 7). Four setae on subcapitular face, positioned as in
<figureCitation id="13272A67FFCA283EFBCBA2F80670F9D2" box="[1035,1092,1531,1552]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="9" pageNumber="304">Fig. 9</figureCitation>
; all attenuate, with several minute barbs; hypostomal seta (
<emphasis id="B968EAF0FFCA283EFD75A11B00F5F9EF" box="[693,705,1560,1581]" italics="true" pageId="9" pageNumber="304">h</emphasis>
) 15 µm, median setae (
<emphasis id="B968EAF0FFCA283EFC5EA11B0182F9F2" box="[926,950,1560,1584]" italics="true" pageId="9" pageNumber="304">
m
<subScript id="179834A7FFCA283EFC6FA1270182F9F2" attach="left" box="[943,950,1572,1584]" fontSize="5" pageId="9" pageNumber="304">1</subScript>
</emphasis>
,
<emphasis id="B968EAF0FFCA283EFC01A11B01EDF9F2" box="[961,985,1560,1584]" italics="true" pageId="9" pageNumber="304">
m
<subScript id="179834A7FFCA283EFC12A12701EDF9F2" attach="left" box="[978,985,1572,1584]" fontSize="5" pageId="9" pageNumber="304">2</subScript>
</emphasis>
) 17 µm, anterior seta (
<emphasis id="B968EAF0FFCA283EFB6FA11B068FF9EF" box="[1199,1211,1560,1581]" italics="true" pageId="9" pageNumber="304">a</emphasis>
) largest (23 µm) and less rapidly tapered than others. Three heterogeneous adoral setae on exposed lateral lips:
<emphasis id="B968EAF0FFCA283EFEABA15203B4F9A4" box="[363,384,1617,1638]" italics="true" pageId="9" pageNumber="304">or</emphasis>
<subScript id="179834A7FFCA283EFE41A15D03BCF9A8" attach="left" box="[385,392,1630,1642]" fontSize="5" pageId="9" pageNumber="304">1</subScript>
and
<emphasis id="B968EAF0FFCA283EFE73A15203E4F9A8" box="[435,464,1617,1642]" italics="true" pageId="9" pageNumber="304">
or
<subScript id="179834A7FFCA283EFE09A15D03E4F9A8" attach="left" box="[457,464,1630,1642]" fontSize="5" pageId="9" pageNumber="304">3</subScript>
</emphasis>
attenuate, without ornamentation, ca. 8 µm and 20 µm respectively. Seta
<emphasis id="B968EAF0FFCA283EFB42A15206AAF9A8" box="[1154,1182,1617,1642]" italics="true" pageId="9" pageNumber="304">
or
<subScript id="179834A7FFCA283EFB57A15D06AAF9A8" attach="left" box="[1175,1182,1630,1642]" fontSize="5" pageId="9" pageNumber="304">2</subScript>
</emphasis>
complex (
<figureCitation id="13272A67FFCA283EFE84A16D03B6F941" box="[324,386,1646,1667]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="9" pageNumber="304">Figs 9</figureCitation>
,
<figureCitation id="13272A67FFCA283EFE4BA16D0391F941" box="[395,421,1646,1667]" captionStart="Figs 12–18" captionStartId="8.[317,360,1631,1652]" captionTargetBox="[364,1222,423,1603]" captionTargetId="figure-19@8.[364,1222,423,1603]" captionTargetPageId="8" captionText="Figs 12–18. Eniochthonius mahunkai sp. n., adult: 12 = dorsal habitus; 13 = bothridial seta (bo) and sejugal region, dorsolateral aspect; 14 = rostrum and distal region of subcapitulum, lateral aspect; 15 = proterosoma, partial anterior view; 16 = genital region in polarized transmitted light (* marks location of different mineralization pattern covering narrow transverse band of unsclerotized procuticle that divides genital plate); 17 = mineralization pattern on leg III; 18 = mineralization pattern in central notogaster, polarized light on left, unpolarized on right. Figs 12–15 scanning electron micro-" figureDoi="http://doi.org/10.5281/zenodo.12585395" httpUri="https://zenodo.org/record/12585395/files/figure.png" pageId="9" pageNumber="304">14</figureCitation>
,
<figureCitation id="13272A67FFCA283EFE6FA16D03FFF941" box="[431,459,1646,1667]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="9" pageNumber="304">20</figureCitation>
): main axis (ca. 24 µm) finely attenuate, with small ventral tooth at mid-length, distal half bent strongly dorsad (slightly bowed like parenthesis) then again anteriorly for short distance; vertical portion of
<emphasis id="B968EAF0FFCA283EFE2BA1AB003CF902" box="[491,520,1704,1728]" italics="true" pageId="9" pageNumber="304">
or
<subScript id="179834A7FFCA283EFDC1A1B7003CF902" attach="left" box="[513,520,1716,1728]" fontSize="5" pageId="9" pageNumber="304">2</subScript>
</emphasis>
pectinate with more than 10 long, fine, closely adjacent, medially directed cilia, which (as setal pair) interdigitate to form baleen-like sieve. Postpalpal seta 7–8 µm, little tapered, terminating in distal fork; attached without alveolus, and at near right-angle, to vertical post-like tubercle about as long as and little wider than seta (
<figureCitation id="13272A67FFCA283EFD26A1FD0118F8D1" box="[742,812,1790,1811]" captionStart="Figs 29–35" captionStartId="12.[317,360,1439,1460]" captionTargetBox="[378,1209,418,1411]" captionTargetId="figure-19@12.[378,1209,418,1411]" captionTargetPageId="12" captionText="Figs 29–35. Eniochthonius mahunkai sp. n., adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section inFig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure ia); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta e1, showing fine barbs.All but Fig.31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="9" pageNumber="304">Fig. 34</figureCitation>
). Palp simple, all five segments fully articulated; others by proportion.
</paragraph>
<caption id="DF63666AFFC9283DFEFDA16706C2F92F" ID-DOI="http://doi.org/10.5281/zenodo.12585397" ID-Zenodo-Dep="12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="10" pageNumber="305" startId="10.[317,360,1636,1657]" targetBox="[379,1208,421,1608]" targetPageId="10" targetType="figure">
<paragraph id="8BA336E2FFC9283DFEFDA16706C2F92F" blockId="10.[317,1270,1636,1802]" pageId="10" pageNumber="305">
<emphasis id="B968EAF0FFC9283DFEFDA167039AF9BB" bold="true" box="[317,430,1636,1657]" pageId="10" pageNumber="305">Figs 19–22.</emphasis>
<taxonomicName id="4C1C4D61FFC9283DFE73A1660095F9B8" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[435,673,1637,1658]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="305" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC9283DFE73A1660095F9B8" box="[435,673,1637,1658]" italics="true" pageId="10" pageNumber="305">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC9283DFD65A16600ECF9B8" box="[677,728,1637,1658]" pageId="10" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
, adult: 19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm
</paragraph>
</caption>
<footnote id="E8072AECFFC9283DFD75A1F6014BF8C8" box="[693,895,1781,1802]" pageId="10" pageNumber="305">
<paragraph id="8BA336E2FFC9283DFD75A1F6014BF8C8" blockId="10.[317,1270,1636,1802]" box="[693,895,1781,1802]" pageId="10" pageNumber="305">(20); 20 µm (21, 22).</paragraph>
</footnote>
<caption id="DF63666AFFC8283CFEFDA2F106C1F8C8" ID-DOI="http://doi.org/10.5281/zenodo.12585399" ID-Zenodo-Dep="12585399" httpUri="https://zenodo.org/record/12585399/files/figure.png" pageId="11" pageNumber="306" startId="11.[317,360,1522,1543]" targetBox="[332,1256,419,1493]" targetPageId="11" targetType="figure">
<paragraph id="8BA336E2FFC8283CFEFDA2F106C1F8C8" blockId="11.[317,1270,1522,1831]" pageId="11" pageNumber="306">
<emphasis id="B968EAF0FFC8283CFEFDA2F10384F9C5" bold="true" box="[317,432,1522,1543]" pageId="11" pageNumber="306">Figs 23–28.</emphasis>
<taxonomicName id="4C1C4D61FFC8283CFE76A2F10092F9C5" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[438,678,1522,1543]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="11" pageNumber="306" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFC8283CFE76A2F10092F9C5" box="[438,678,1522,1543]" italics="true" pageId="11" pageNumber="306">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFC8283CFD6BA2F100D5F9C5" box="[683,737,1522,1543]" pageId="11" pageNumber="306" rank="species">sp. n.</taxonomicNameLabel>
, adult, lateral aspect of near-sagittal section, anterior to left: 23 = overview of section, with position of enlarged images marked by respective figure number in similar relative position (ch = chelicera, fb = food bolus, ov = ovipositor); 24 = dorsal sejugal region (sej = soft articulating cuticle, arrowhead indicates thickened cuticle behind which proterosoma narrows slightly to proximal “neck”); 25 = region of transverse sulcus, showing thickened cuticle and overlying chambered epicuticle (demineralized, marked with arrowhead); 26 = telescoping transverse scissure (type-L), showing tectum on pronotaspis (asterisks mark limits of sclerotized cuticle on respective plates; voluminous connecting soft cuticle not shown); 27 = ventral sejugal region (* marks cuticle of proximal “neck”, exposed only in distended individuals); 28 = juncture of genital, preanal (pr) and anal plates (aa = anal apodeme; gp = posterior genital papilla). Scale bar: 50 µm (23),
</paragraph>
</caption>
<caption id="DF63666AFFCF283BFEFDA29C06C2F8CC" ID-DOI="http://doi.org/10.5281/zenodo.12585401" ID-Zenodo-Dep="12585401" httpUri="https://zenodo.org/record/12585401/files/figure.png" pageId="12" pageNumber="307" startId="12.[317,360,1439,1460]" targetBox="[378,1209,418,1411]" targetPageId="12" targetType="figure">
<paragraph id="8BA336E2FFCF283BFEFDA29C06C2F8CC" blockId="12.[317,1270,1439,1835]" pageId="12" pageNumber="307">
<emphasis id="B968EAF0FFCF283BFEFDA29C039BFA76" bold="true" box="[317,431,1439,1460]" pageId="12" pageNumber="307">Figs 29–35.</emphasis>
<taxonomicName id="4C1C4D61FFCF283BFE74A2A30097FA77" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[436,675,1440,1461]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="12" pageNumber="307" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFCF283BFE74A2A30097FA77" box="[436,675,1440,1461]" italics="true" pageId="12" pageNumber="307">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFCF283BFD67A2A300E8FA77" box="[679,732,1440,1461]" pageId="12" pageNumber="307" rank="species">sp. n.</taxonomicNameLabel>
, adult: 29 = bothridium, optical section (arrowhead indicates spiculated inner chamber); 30 = anterior region of isolated suprapleural plate (sp) and pleural region of notogaster (pl), lateral aspect (cf. Fig. 2), showing taenidium (t) curving ventrad (* and IV mark approximate positions of postpedal fossa on aggenital plate and leg IV insertion, respectively); 31 = taenidium (t) and fossa (fo) on left side, showing curved alignment seen by transparency on dorsally positioned, slightly flattened specimen (III = leg III, * marks approximate level of section in Fig. 32); 32 = frontal section through notogastral taenidium and porose area (t = taenidium, sa = sternal apodeme of epimere III, tr III = trochanter III, arrow marks open slot of taenidium); 33 = notogastral taenidium and porose area, lateral aspect (cf. Fig. 30), at three progressively deeper focal planes (arrowhead indicates lyrifissure
<emphasis id="B968EAF0FFCF283BFD96A1A0005DF97A" box="[598,617,1699,1720]" italics="true" pageId="12" pageNumber="307">ia</emphasis>
); 34 = posterodorsal corner of subcapitulum, lateral aspect, showing forked postpalpal seta (P = femur of palp, arrowhead indicates cylindrical stalk on which seta inserts at strong angle); 35 = notogastral seta
<emphasis id="B968EAF0FFCF283BFD63A1DE009AF930" box="[675,686,1757,1778]" italics="true" pageId="12" pageNumber="307">e</emphasis>
<subScript id="179834A7FFCF283BFD6DA1EA0080F937" attach="left" box="[685,692,1769,1781]" fontSize="5" pageId="12" pageNumber="307">1</subScript>
, showing fine barbs. All but Fig. 31 with differential contrast transmitted illumination; 30 and 34 layered images. Scale bars: 50 µm (30), 20 µm (31, 32), 10 µm
</paragraph>
</caption>
<paragraph id="8BA336E2FFCF283BFD1EA0150161F8E9" blockId="12.[317,1270,1439,1835]" box="[734,853,1814,1835]" pageId="12" pageNumber="307">(29, 33–35).</paragraph>
<paragraph id="8BA336E2FFCE283AFEFDA6B70653FCE1" blockId="13.[317,1270,436,803]" pageId="13" pageNumber="308">
femur and tarsus approximately of equal length, about twice that of genu and tibia (
<figureCitation id="13272A67FFCE283AFB90A6B706A2FE0B" box="[1104,1174,436,457]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="13" pageNumber="308">Fig. 11</figureCitation>
). Setation (trochanter to tarsus) 0–2–1–3–11, plus solenidion ω. All setae on femur to tibia and most setae on tarsus attenuate, with sparse minute barbs; unpaired anteroculminal,
<emphasis id="B968EAF0FFCE283AFC06A6ED01DAFDC1" box="[966,1006,494,515]" italics="true" pageId="13" pageNumber="308">acm</emphasis>
, and ultimal pair (
<emphasis id="B968EAF0FFCE283AFB5CA6ED069BFDC1" box="[1180,1199,494,515]" italics="true" pageId="13" pageNumber="308">ul</emphasis>
) of tarsus eupathidial, little tapered; ultimal pair fused near base to form strongly diverging fork;
<emphasis id="B968EAF0FFCE283AFB17A50806D6FDE2" box="[1239,1250,523,544]" italics="true" pageId="13" pageNumber="308">v</emphasis>
<subScript id="179834A7FFCE283AFB21A51406DCFDE1" attach="left" box="[1249,1256,535,547]" fontSize="5" pageId="13" pageNumber="308">1</subScript>
<emphasis id="B968EAF0FFCE283AFB28A50806C1FDE2" box="[1256,1269,523,544]" italics="true" pageId="13" pageNumber="308">”</emphasis>
spiniform. Solenidion ω about half length of tarsus, narrowly ceratiform, curved toward tip of segment to end near ultimal fork (
<figureCitation id="13272A67FFCE283AFD9EA5470093FD9B" box="[606,679,580,601]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="13" pageNumber="308">Figs 11</figureCitation>
,
<figureCitation id="13272A67FFCE283AFD71A54700F8FD9B" box="[689,716,580,601]" captionStart="Figs 19–22" captionStartId="10.[317,360,1636,1657]" captionTargetBox="[379,1208,421,1608]" captionTargetId="figure-19@10.[379,1208,421,1608]" captionTargetPageId="10" captionText="Figs 19–22. Eniochthonius mahunkai sp. n., adult:19 = ventral habitus; 20 = subcapitulum and vicinity, ventral aspect (white arrowhead indicates fused pair of ultimal setae, * marks groove separating gena and mentum); 21 = coxisternum and vicinity, ventral aspect; 22 = anogenital region and adjacent notogaster (AD, AN indicate adanal and anal valves, respectively; G marks position of narrow transverse band of unsclerotized procuticle that divides genital plate). Scale bars: 50 µm (19), 10 µm" figureDoi="http://doi.org/10.5281/zenodo.12585397" httpUri="https://zenodo.org/record/12585397/files/figure.png" pageId="13" pageNumber="308">20</figureCitation>
). Chelicera (
<figureCitation id="13272A67FFCE283AFC85A54701BEFD9B" box="[837,906,580,601]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="13" pageNumber="308">Fig. 10</figureCitation>
) relatively narrow, ca. 65 µm long, 22 µm greatest width, gradually tapered distally; small proximal portion (less than one-tenth of length) internalized through cheliceral frame (body wall). Not mineralized. Ventral trochanter remnant relatively large, extending about half length of chelicera. Main cheliceral body with fine longitudinal striae dorsally (not illustrated), proximal to seta
<emphasis id="B968EAF0FFCE283AFCE9A5BB0178FD0F" box="[809,844,696,717]" italics="true" pageId="13" pageNumber="308">cha</emphasis>
; adaxial face with several scattered sharp spicules. Digits ca. 21 µm long, tapered to fine tips, each with four teeth; abaxial face of fixed digit with small, seemingly porose region of unknown nature; small, sharp mid-ventral tooth near base of movable digit. Setae inserted as in
<figureCitation id="13272A67FFCE283AFD55A40D00E9FCE1" box="[661,733,782,803]" captionStart="Figs 9–11" captionStartId="6.[317,360,1749,1770]" captionTargetBox="[323,1268,1174,1717]" captionTargetId="figure-384@6.[319,1269,1174,1721]" captionTargetPageId="6" captionText="Figs 9–11. Eniochthonius mahunkai sp. n., adult: 9 = subcapitulum, ventral aspect (* marks position of postpalpal setae on dorsal face); 10 = chelicera, adaxial aspect; 11 = palp, abaxial aspect. Scale" figureDoi="http://doi.org/10.5281/zenodo.12585393" httpUri="https://zenodo.org/record/12585393/files/figure.png" pageId="13" pageNumber="308">Fig. 10</figureCitation>
;
<emphasis id="B968EAF0FFCE283AFD29A40D0138FCE1" box="[745,780,782,803]" italics="true" pageId="13" pageNumber="308">cha</emphasis>
,
<emphasis id="B968EAF0FFCE283AFCD9A40D0108FCE1" box="[793,828,782,803]" italics="true" pageId="13" pageNumber="308">chb</emphasis>
ca. 7 µm, 16 µm respectively.
</paragraph>
<paragraph id="8BA336E2FFCE283AFEFDA460007BFCBF" blockId="13.[317,591,867,893]" box="[317,591,867,893]" pageId="13" pageNumber="308">
<heading id="D0EB818EFFCE283AFEFDA460007BFCBF" box="[317,591,867,893]" fontSize="11" level="3" pageId="13" pageNumber="308" reason="4">
<emphasis id="B968EAF0FFCE283AFEFDA460007BFCBF" box="[317,591,867,893]" italics="true" pageId="13" pageNumber="308">
Ontogeny (
<figureCitation id="13272A67FFCE283AFE7FA4600072FCBF" box="[447,582,867,893]" captionStart-0="Figs 36–41" captionStart-1="Figs 42–50" captionStartId-0="14.[317,360,1600,1621]" captionStartId-1="16.[317,360,1551,1572]" captionTargetBox-0="[371,1212,422,1571]" captionTargetBox-1="[346,1241,417,1523]" captionTargetId-0="figure-19@14.[370,1216,418,1571]" captionTargetId-1="figure-19@16.[346,1241,417,1523]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." captionText-1="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi-0="http://doi.org/10.5281/zenodo.12585403" figureDoi-1="http://doi.org/10.5281/zenodo.12585405" httpUri-0="https://zenodo.org/record/12585403/files/figure.png" httpUri-1="https://zenodo.org/record/12585405/files/figure.png" pageId="13" pageNumber="308">Figs 36–50</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA336E2FFCE283AFEBEA49603C4FBFB" blockId="13.[317,1271,916,1801]" pageId="13" pageNumber="308">
<emphasis id="B968EAF0FFCE283AFEBEA49603DBFC68" box="[382,495,917,938]" italics="true" pageId="13" pageNumber="308">Dimensions</emphasis>
. Mean (range) total length and maximum width measured from
<specimenCount id="9D1AFD6BFFCE283AFB9AA49706E8FC6B" box="[1114,1244,916,937]" count="10" pageId="13" pageNumber="308" type="generic">10 specimens</specimenCount>
of each instar: larva (La): 218 µm (208–230 µm) by 97 µm (93–108 µm); protonymph (Pn) 247 µm (230–260 µm) by 113 µm (105–118 µm); deutonymph (Dn) 279 µm (259–289 µm) by 126 µm (118–132 µm); tritonymph (Tn) 310 µm (294–323 µm) by 146 µm (132–157 µm). As in adult, length varies widely depending on degree of distention; width varies also, being slightly narrower in distended specimens.
</paragraph>
<paragraph id="8BA336E2FFCE283AFEBEA3420096FB6F" blockId="13.[317,1271,916,1801]" pageId="13" pageNumber="308">
<emphasis id="B968EAF0FFCE283AFEBEA34203DEFB94" box="[382,490,1089,1110]" italics="true" pageId="13" pageNumber="308">Integument</emphasis>
. Shiny in reflected light; nearly white in early instars, may have slight yellow tint in nymphs, but always paler than adult. Except at articulations, with strongly chambered, mineralized cuticle in all immature instars, essentially like that of adult; when demineralized, chambers more discernable in nymphs than in larva.
</paragraph>
<paragraph id="8BA336E2FFCE283AFEBEA3B60698FAB4" blockId="13.[317,1271,916,1801]" pageId="13" pageNumber="308">
<emphasis id="B968EAF0FFCE283AFEBEA3B603DEFB08" box="[382,490,1205,1226]" italics="true" pageId="13" pageNumber="308">Prodorsum</emphasis>
. General structure similar to that of adult, except lateral margin rather simple in La; distinct rostral lobe and concavities for legs appear in Pn. Posterior margin of prodorsum not noticeably thickened in any immature instar. Setation generally similar to that of adult, but setae
<emphasis id="B968EAF0FFCE283AFB7BA3ED06FAFAC1" box="[1211,1230,1262,1283]" italics="true" pageId="13" pageNumber="308">in</emphasis>
and
<emphasis id="B968EAF0FFCE283AFEFDA208037AFAE2" box="[317,334,1291,1312]" italics="true" pageId="13" pageNumber="308">le</emphasis>
acicular and more clearly barbed in La, becoming attenuate in nymphs. Posterior exobothridial seta (
<emphasis id="B968EAF0FFCE283AFE85A22B0368FAFF" box="[325,348,1320,1341]" italics="true" pageId="13" pageNumber="308">xp</emphasis>
) proportionally smaller than in adult; about one-quarter length of anterior (
<emphasis id="B968EAF0FFCE283AFBFCA22B0667FAFF" box="[1084,1107,1320,1341]" italics="true" pageId="13" pageNumber="308">xa</emphasis>
). Bothridial seta distally broadened and barbed; number of long tines
<quantity id="4CE49B07FFCE283AFC9AA24701AFFA9B" box="[858,923,1348,1369]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" metricValueMax="5.08" metricValueMin="0.0" pageId="13" pageNumber="308" unit="in" value="1.0" valueMax="2.0" valueMin="0.0">0–2 in</quantity>
La (
<figureCitation id="13272A67FFCE283AFC10A247062EFA9B" box="[976,1050,1348,1369]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="13" pageNumber="308">Fig. 43</figureCitation>
) and
<quantity id="4CE49B07FFCE283AFB98A24706AEFA9B" box="[1112,1178,1348,1369]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="10.16" metricValueMin="0.0" pageId="13" pageNumber="308" unit="in" value="2.0" valueMax="4.0" valueMin="0.0">0–4 in</quantity>
nymphs. Bothridium two-chambered in all immature instars, with inner chamber densely spiculate.
</paragraph>
<paragraph id="8BA336E2FFCE283AFEBEA27D0065F8CB" blockId="13.[317,1271,916,1801]" pageId="13" pageNumber="308">
<emphasis id="B968EAF0FFCE283AFEBEA27D000FFA51" box="[382,571,1406,1427]" italics="true" pageId="13" pageNumber="308">Gastronotic region</emphasis>
. Larva with notogaster divided into three parts – pronotaspis (NA), mesonotaspis (NM) and pygidium (PY) – by two simple, type-E transverse scissures (
<figureCitation id="13272A67FFCE283AFBA8A298069BFA72" box="[1128,1199,1435,1456]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="13" pageNumber="308">Fig. 36</figureCitation>
). Anterior scissure, between setal rows
<emphasis id="B968EAF0FFCE283AFDB2A2BB004AFA0F" box="[626,638,1464,1485]" italics="true" pageId="13" pageNumber="308">d</emphasis>
and
<emphasis id="B968EAF0FFCE283AFD6AA2BB0081FA0F" box="[682,693,1464,1485]" italics="true" pageId="13" pageNumber="308">e</emphasis>
(scissure
<emphasis id="B968EAF0FFCE283AFCD1A2BB011AFA12" box="[785,814,1464,1488]" italics="true" pageId="13" pageNumber="308">
ar
<subScript id="179834A7FFCE283AFCE7A2C7011AFA12" attach="left" box="[807,814,1476,1488]" fontSize="5" pageId="13" pageNumber="308">2</subScript>
</emphasis>
of
<bibRefCitation id="EF8D4B13FFCE283AFC8AA2BB01C0FA0F" author="GRANDJEAN, F." box="[842,1012,1464,1485]" pageId="13" pageNumber="308" pagination="213 - 248" refId="ref17987" refString="GRANDJEAN, F. (1947) Les Enarthronota (Acariens). Premiere serie. Annales des Sciences Naturelles Zoologie 11 e serie 8: 213 - 248." type="journal article" year="1947">
<smallCapsWord id="8D45A03EFFCE283AFC8AA2BB018AFA09" baselines="1480,1479" box="[842,958,1464,1485]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Grandjean" pageId="13" pageNumber="308">GRANDJEAN</smallCapsWord>
1947
</bibRefCitation>
) with relatively broader articulating cuticle; posterior scissure, between rows
<emphasis id="B968EAF0FFCE283AFCF3A2D6010AFA28" box="[819,830,1493,1514]" italics="true" pageId="13" pageNumber="308">e</emphasis>
and
<emphasis id="B968EAF0FFCE283AFCB0A2D60143FA28" box="[880,887,1493,1514]" italics="true" pageId="13" pageNumber="308">f</emphasis>
(
<emphasis id="B968EAF0FFCE283AFC46A2D60193FA28" box="[902,935,1493,1514]" italics="true" pageId="13" pageNumber="308">ar3</emphasis>
), very narrow. Scissures reaching laterally to broad, soft, striated pleural band bordering notogaster. Neither suprapleural plate nor plicature plate developed; without distinct pleural carina. Single small, oval porose area (ca. 6 µm long) on vague pleural extension of pronotaspis, posteroventral to seta
<emphasis id="B968EAF0FFCE283AFC23A12801C1F981" box="[995,1013,1579,1603]" italics="true" pageId="13" pageNumber="308">
c
<subScript id="179834A7FFCE283AFC2EA13401C1F981" attach="left" box="[1006,1013,1591,1603]" fontSize="5" pageId="13" pageNumber="308">3</subScript>
</emphasis>
(
<figureCitation id="13272A67FFCE283AFBC1A128067EF982" box="[1025,1098,1579,1600]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="13" pageNumber="308">Fig. 44</figureCitation>
). With 13 pairs of setae representing typical holotrichous setation (
<figureCitation id="13272A67FFCE283AFCCAA14B0165F99F" box="[778,849,1608,1629]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="13" pageNumber="308">Fig. 36</figureCitation>
). Setae heterotrichous:
<emphasis id="B968EAF0FFCE283AFBF0A14B0676F9A2" box="[1072,1090,1608,1632]" italics="true" pageId="13" pageNumber="308">
c
<subScript id="179834A7FFCE283AFBFBA1570676F9A2" attach="left" box="[1083,1090,1620,1632]" fontSize="5" pageId="13" pageNumber="308">1</subScript>
</emphasis>
,
<emphasis id="B968EAF0FFCE283AFB8DA14B066AF9A2" box="[1101,1118,1608,1632]" italics="true" pageId="13" pageNumber="308">
c
<subScript id="179834A7FFCE283AFB97A157066AF9A2" attach="left" box="[1111,1118,1620,1632]" fontSize="5" pageId="13" pageNumber="308">2</subScript>
</emphasis>
, and
<emphasis id="B968EAF0FFCE283AFB51A14B06A9F99F" box="[1169,1181,1608,1629]" italics="true" pageId="13" pageNumber="308">d</emphasis>
<subScript id="179834A7FFCE283AFB5DA1570690F9A2" attach="left" box="[1181,1188,1620,1632]" fontSize="5" pageId="13" pageNumber="308">1</subScript>
acicular and more clearly barbed than in other instars (
<figureCitation id="13272A67FFCE283AFD34A167010FF9BB" box="[756,827,1636,1657]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="13" pageNumber="308">Fig. 42</figureCitation>
);
<emphasis id="B968EAF0FFCE283AFC8DA1660150F9B8" box="[845,868,1637,1658]" italics="true" pageId="13" pageNumber="308">cp</emphasis>
variable (acicular, attenuate or intermediate);
<emphasis id="B968EAF0FFCE283AFEADA182034EF958" box="[365,378,1665,1690]" italics="true" pageId="13" pageNumber="308">
f
<subScript id="179834A7FFCE283AFEB3A18D034EF958" attach="left" box="[371,378,1678,1690]" fontSize="5" pageId="13" pageNumber="308">1</subScript>
</emphasis>
thickest of all notogastral setae, acicular with dorsal row of inconspicuous fine barbs;
<emphasis id="B968EAF0FFCE283AFB69A182068EF958" box="[1193,1210,1665,1690]" italics="true" pageId="13" pageNumber="308">
c
<subScript id="179834A7FFCE283AFB73A18D068EF958" attach="left" box="[1203,1210,1678,1690]" fontSize="5" pageId="13" pageNumber="308">3</subScript>
</emphasis>
attenuate but relatively longer than in nymphs and adult (longer than acicular setae) (
<figureCitation id="13272A67FFCE283AFBEDA19D0647F971" box="[1069,1139,1694,1715]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="13" pageNumber="308">Fig. 37</figureCitation>
). Lyrifissures cupular:
<emphasis id="B968EAF0FFCE283AFE51A1B80390F912" box="[401,420,1723,1744]" italics="true" pageId="13" pageNumber="308">ia</emphasis>
on lateral margin of pronotaspis, about 15 µm anterior to scissure;
<emphasis id="B968EAF0FFCE283AFBE3A1B8060FF912" box="[1059,1083,1723,1744]" italics="true" pageId="13" pageNumber="308">im</emphasis>
near lateral margin of mesonotaspis;
<emphasis id="B968EAF0FFCE283AFE22A1DB03C1F92F" box="[482,501,1752,1773]" italics="true" pageId="13" pageNumber="308">ip</emphasis>
far anterior from usual position (see Note 6), ca. 15 µm posteroventral to
<emphasis id="B968EAF0FFCE283AFB66A1DB068AF92F" box="[1190,1214,1752,1773]" italics="true" pageId="13" pageNumber="308">im</emphasis>
and 5 µm from notogastral border.
</paragraph>
<caption id="DF63666AFFCD2839FEFDA14306C1F8DD" ID-DOI="http://doi.org/10.5281/zenodo.12585403" ID-Zenodo-Dep="12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="14" pageNumber="309" startId="14.[317,360,1600,1621]" targetBox="[371,1212,422,1571]" targetPageId="14" targetType="figure">
<paragraph id="8BA336E2FFCD2839FEFDA14306C1F8DD" blockId="14.[317,1270,1600,1823]" pageId="14" pageNumber="309">
<emphasis id="B968EAF0FFCD2839FEFDA143039AF997" bold="true" box="[317,430,1600,1621]" pageId="14" pageNumber="309">Figs 36–41.</emphasis>
<taxonomicName id="4C1C4D61FFCD2839FE74A1430096F997" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[436,674,1600,1621]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="309" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFCD2839FE74A1430096F997" box="[436,674,1600,1621]" italics="true" pageId="14" pageNumber="309">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFCD2839FD67A14300EFF997" box="[679,731,1600,1621]" pageId="14" pageNumber="309" rank="species">sp. n.</taxonomicNameLabel>
, immatures: 36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from
<bibRefCitation id="EF8D4B13FFCD2839FEB2A1790012F94D" author="GRANDJEAN, F." box="[370,550,1658,1679]" pageId="14" pageNumber="309" pagination="30 - 61" refId="ref17742" refString="GRANDJEAN, F. (1933) Etude sur le developpement des Oribates. Bulletin de la Societe Zoologique de France 58: 30 - 61." type="journal article" year="1933">
<smallCapsWord id="8D45A03EFFCD2839FEB2A17903D9F94C" baselines="1674,1673" box="[370,493,1658,1679]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Grandjean" pageId="14" pageNumber="309">GRANDJEAN</smallCapsWord>
1933
</bibRefCitation>
), contrasting presence of inguinal seta
<emphasis id="B968EAF0FFCD2839FC5BA1790193F94D" box="[923,935,1658,1679]" italics="true" pageId="14" pageNumber="309">h</emphasis>
<subScript id="179834A7FFCD2839FC67A185019AF950" attach="left" box="[935,942,1670,1682]" fontSize="5" pageId="14" pageNumber="309">4</subScript>
in
<taxonomicName id="4C1C4D61FFCD2839FC0CA1790672F94D" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[972,1094,1658,1679]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="309" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="B968EAF0FFCD2839FC0CA1790672F94D" box="[972,1094,1658,1679]" italics="true" pageId="14" pageNumber="309">E. mahunkai</emphasis>
</taxonomicName>
(left side) with its absence in
<taxonomicName id="4C1C4D61FFCD2839FE64A1940000F96E" baseAuthorityName="Berlese" baseAuthorityYear="1903" box="[420,564,1687,1708]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="309" phylum="Arthropoda" rank="species" species="minutissmus">
<emphasis id="B968EAF0FFCD2839FE64A1940000F96E" box="[420,564,1687,1708]" italics="true" pageId="14" pageNumber="309">E. minutissmus</emphasis>
</taxonomicName>
(right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38).
</paragraph>
</caption>
<paragraph id="8BA336E2FFCC2838FEBEA6B706DCFC71" blockId="15.[317,1271,436,1811]" pageId="15" pageNumber="310">
Nymphs with notogastral structure similar to that of adult in having single transverse scissure immediately posterior to setal row
<emphasis id="B968EAF0FFCC2838FD47A6D200A6FE24" box="[647,658,465,486]" italics="true" pageId="15" pageNumber="310">e</emphasis>
(i.e. pronotaspis carries setal rows
<emphasis id="B968EAF0FFCC2838FC1EA6D201CFFE24" box="[990,1019,465,486]" italics="true" pageId="15" pageNumber="310">c-e</emphasis>
;
<figureCitation id="13272A67FFCC2838FBC7A6D2067FFE24" box="[1031,1099,465,486]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 39</figureCitation>
) and suprapleural plate, bearing setae
<emphasis id="B968EAF0FFCC2838FDCAA6ED0029FDC5" box="[522,541,494,519]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FDD6A6F80029FDC5" attach="left" box="[534,541,507,519]" fontSize="5" pageId="15" pageNumber="310">2</subScript>
</emphasis>
,
<emphasis id="B968EAF0FFCC2838FDEEA6ED0075FDC5" box="[558,577,494,519]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FDFAA6F80075FDC5" attach="left" box="[570,577,507,519]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
, but with soft, striated cuticle between notogaster and coxisternum. Suprapleural plate shorter and broader (6–7 times longer than broad) than in adult, reaching anteriorly only to level of leg IV. Transverse scissure without tectal limb, but pronotaspis cuticle bends sharply ventrad at level of row
<emphasis id="B968EAF0FFCC2838FDA7A5460046FD98" box="[615,626,581,602]" italics="true" pageId="15" pageNumber="310">e</emphasis>
, forming narrow vertical wall ending at very narrow, hinge-like articulation with pygidial region (
<figureCitation id="13272A67FFCC2838FDACA5620087FDB4" box="[620,691,609,630]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="15" pageNumber="310">Fig. 46</figureCitation>
, arrowhead); anterior sclerite therefore higher than posterior one in lateral aspect, but without ability to telescope. Indistinct groove present posterior to setal row
<emphasis id="B968EAF0FFCC2838FB29A57D06C1FD51" box="[1257,1269,638,659]" italics="true" pageId="15" pageNumber="310">d</emphasis>
(remnant of
<emphasis id="B968EAF0FFCC2838FE75A59803E5FD71" box="[437,465,667,691]" italics="true" pageId="15" pageNumber="310">
ar
<subScript id="179834A7FFCC2838FE0AA5A403E5FD71" attach="left" box="[458,465,679,691]" fontSize="5" pageId="15" pageNumber="310">2</subScript>
</emphasis>
), less conspicuous than homologous transverse sulcus of adult. Pleural region with porose area similar to that of La, lying between seta
<emphasis id="B968EAF0FFCC2838FCE5A5BB0102FD12" box="[805,822,696,720]" italics="true" pageId="15" pageNumber="310">
c
<subScript id="179834A7FFCC2838FCEFA5C70102FD12" attach="left" box="[815,822,708,720]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
and anterolateral corner of pronotaspis. Pleural carina present but weakly developed; with second oval porose area (
<emphasis id="B968EAF0FFCC2838FC0CA5D601C4FD28" box="[972,1008,725,746]" italics="true" pageId="15" pageNumber="310">app</emphasis>
) on underside (
<figureCitation id="13272A67FFCC2838FB46A5D706F9FD2B" box="[1158,1229,724,745]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="15" pageNumber="310">Fig. 45</figureCitation>
; see Note 4); in dorsoventral aspect, porose area appears between levels of legs III and IV. Cupule
<emphasis id="B968EAF0FFCC2838FB7FA5F206E6FCC4" box="[1215,1234,753,774]" italics="true" pageId="15" pageNumber="310">ia</emphasis>
between porose area and anterior corner of suprapleural plate;
<emphasis id="B968EAF0FFCC2838FCBDA40D01A1FCE1" box="[893,917,782,803]" italics="true" pageId="15" pageNumber="310">im</emphasis>
dorsal to latter plate;
<emphasis id="B968EAF0FFCC2838FBA6A40D064DFCE1" box="[1126,1145,782,803]" italics="true" pageId="15" pageNumber="310">ip</emphasis>
further anterior in pleural region than in La, approximately as in adult;
<emphasis id="B968EAF0FFCC2838FC4AA42801A9FC82" box="[906,925,811,832]" italics="true" pageId="15" pageNumber="310">ih</emphasis>
anterodorsal to seta
<emphasis id="B968EAF0FFCC2838FBA9A4280648FC81" box="[1129,1148,811,835]" italics="true" pageId="15" pageNumber="310">
p
<subScript id="179834A7FFCC2838FBB5A4340648FC81" attach="left" box="[1141,1148,823,835]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
, as in adult. Ventrolateral plicature plate develops posteriorly in Pn, fully formed (as in adult) in Dn and Tn; contains cupule
<emphasis id="B968EAF0FFCC2838FE76A46603E6FCB8" box="[438,466,869,890]" italics="true" pageId="15" pageNumber="310">ips</emphasis>
as in adult. All nymphs with 16 pairs of setae, representing larval complement plus pseudanal setae
<emphasis id="B968EAF0FFCC2838FE16A4820030FC58" box="[470,516,897,922]" italics="true" pageId="15" pageNumber="310">
p
<subScript id="179834A7FFCC2838FE22A48D03DDFC58" attach="left" box="[482,489,910,922]" fontSize="5" pageId="15" pageNumber="310">1</subScript>
-p
<subScript id="179834A7FFCC2838FE3DA48D0030FC58" attach="left" box="[509,516,910,922]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
(
<emphasis id="B968EAF0FFCC2838FDD0A4820017FC58" box="[528,547,897,922]" italics="true" pageId="15" pageNumber="310">
p
<subScript id="179834A7FFCC2838FDDCA48D0017FC58" attach="left" box="[540,547,910,922]" fontSize="5" pageId="15" pageNumber="310">4</subScript>
</emphasis>
absent from nymphs and adult; see Note 8). All notogastral setae with dorsal barbs, usually inconspicuous. Setae of nymphs all attenuate as in adult and with similar relative size.
</paragraph>
<paragraph id="8BA336E2FFCC2838FEBEA4B80679FACF" blockId="15.[317,1271,436,1811]" pageId="15" pageNumber="310">
<emphasis id="B968EAF0FFCC2838FEBEA4B803CDFC12" box="[382,505,955,976]" italics="true" pageId="15" pageNumber="310">Coxisternum</emphasis>
. Soft cuticle between epimere I and subcapitulum granular. All epimeres with medial band of soft, longitudinally striated cuticle; more apparent in La (
<figureCitation id="13272A67FFCC2838FBC3A4DB067EFC2F" box="[1027,1098,984,1005]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 37</figureCitation>
) since band folds more deeply in nymphs (
<figureCitation id="13272A67FFCC2838FDEEA4F70043FBCB" box="[558,631,1012,1033]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 40</figureCitation>
). Sejugal articulation with transverse striae, blending into those of epimera II and III, respectively, in dactylographic pattern. In all immatures, each half of epimere I at least weakly fused to respective half of epimere II; separated by conspicuous groove, but without apparent articulation. In nymphs, halves of epimeres III and IV fully fused, with no external or internal delimitation. Postpedal fossa absent from all immatures. Larval Claparède’s organ of typical form, without indication of annulations; seta
<emphasis id="B968EAF0FFCC2838FD01A38600E3FB58" box="[705,727,1157,1178]" italics="true" pageId="15" pageNumber="310">1c</emphasis>
scale-like, reverting to normal form in Pn. Otherwise, epimeral setae relatively short, attenuate, smooth or with sparse, minute barbs; generally similar in form and relative size to those of adult. Setal ontogeny as follows: La (3–1–2), Pn (3–1–2–1), Dn (3–1–3 –3), Tn (3–1–3–4); in
<figureCitation id="13272A67FFCC2838FD97A3D800AEFB32" box="[599,666,1243,1264]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 40</figureCitation>
setae of epimere IV lettered in order of appearance (a in Pn, b, c in Dn, d in Tn); in two of 11 tritonymphs examined, seta
<emphasis id="B968EAF0FFCC2838FCB4A3FB01B8FACF" box="[884,908,1272,1293]" italics="true" pageId="15" pageNumber="310">4d</emphasis>
absent unilaterally.
</paragraph>
<paragraph id="8BA336E2FFCC2838FEBEA21600C7F97F" blockId="15.[317,1271,436,1811]" pageId="15" pageNumber="310">
<emphasis id="B968EAF0FFCC2838FEBEA2160001FAE8" box="[382,565,1301,1322]" italics="true" pageId="15" pageNumber="310">Anogenital region.</emphasis>
Genital region of larva represented by broad expanse of soft, longitudinally striated cuticle (
<figureCitation id="13272A67FFCC2838FDC9A232007AFA84" box="[521,590,1329,1350]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 37</figureCitation>
), except for pair of conspicuous thorn-shaped, sclerotized, laterally flattened preanal spines projecting posteroventrad (
<figureCitation id="13272A67FFCC2838FCD1A24D01B0FAA1" box="[785,900,1358,1379]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="15" pageNumber="310">Figs 49, 50</figureCitation>
; see Note 9); nymphs without spines. Genital valves relatively larger in successive nymphs; setation (Pn-Tn) 1–3–7, with Tn having 3 setae in lateral row,
<quantity id="4CE49B07FFCC2838FE06A28B03DDFA5F" box="[454,489,1416,1437]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="15" pageNumber="310" unit="in" value="4.0">4 in</quantity>
medial row (
<figureCitation id="13272A67FFCC2838FDA5A28B0099FA5F" box="[613,685,1416,1437]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 40</figureCitation>
). Aggenital plates absent. Aggenital seta first forms in Tn (often unilaterally; see Note 10) or adult; if in Tn, it inserts at extreme lateral edge of genital plate (
<figureCitation id="13272A67FFCC2838FB0FA2A7036DFA14" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 40</figureCitation>
). Preanal plate absent from all immatures. All immatures with apodeme extending internally from anterior walls of paraproctal valves (
<figureCitation id="13272A67FFCC2838FD58A2DD0132FA31" box="[664,774,1502,1523]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="15" pageNumber="310">Figs 49, 50</figureCitation>
;
<emphasis id="B968EAF0FFCC2838FCCEA2DD0112FA31" box="[782,806,1502,1523]" italics="true" pageId="15" pageNumber="310">aa</emphasis>
): with dark V-shaped cross section in ventral aspect (“V” pointed anteriorly), subtriangular in lateral aspect. Paraproctal valves of La (segment PS) narrow, well defined, consistently with 4 pairs of setae arranged in uniformly spaced row (
<figureCitation id="13272A67FFCC2838FB6AA11B06C6F9EF" box="[1194,1266,1560,1581]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Fig. 38</figureCitation>
, left side). Cupule
<emphasis id="B968EAF0FFCC2838FE24A1360034F988" box="[484,512,1589,1610]" italics="true" pageId="15" pageNumber="310">ips</emphasis>
not present on paraprocts;
<emphasis id="B968EAF0FFCC2838FD3CA136013BF988" box="[764,783,1589,1610]" italics="true" pageId="15" pageNumber="310">ih</emphasis>
in soft cuticle between most anterior pseudanal seta (inguinal seta
<emphasis id="B968EAF0FFCC2838FE05A15203ECF9A8" box="[453,472,1617,1642]" italics="true" pageId="15" pageNumber="310">
p
<subScript id="179834A7FFCC2838FE11A15D03ECF9A8" attach="left" box="[465,472,1630,1642]" fontSize="5" pageId="15" pageNumber="310">4</subScript>
</emphasis>
) and notogastral seta
<emphasis id="B968EAF0FFCC2838FD6BA152008AF9A8" box="[683,702,1617,1642]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FD77A15D008AF9A8" attach="left" box="[695,702,1630,1642]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
. Inguinal seta
<emphasis id="B968EAF0FFCC2838FC8CA152016BF9A8" box="[844,863,1617,1642]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FC98A15D016BF9A8" attach="left" box="[856,863,1630,1642]" fontSize="5" pageId="15" pageNumber="310">4</subScript>
</emphasis>
present near anterior end of paraprocts on opposite side of cupule
<emphasis id="B968EAF0FFCC2838FDDBA16D001AF941" box="[539,558,1646,1667]" italics="true" pageId="15" pageNumber="310">ih</emphasis>
from
<emphasis id="B968EAF0FFCC2838FDA5A16D004CF945" box="[613,632,1646,1671]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FDB1A178004CF945" attach="left" box="[625,632,1659,1671]" fontSize="5" pageId="15" pageNumber="310">3</subScript>
</emphasis>
(transcupular) (
<figureCitation id="13272A67FFCC2838FCCDA16D014EF941" box="[781,890,1646,1667]" captionStart="Figs 36–41" captionStartId="14.[317,360,1600,1621]" captionTargetBox="[371,1212,422,1571]" captionTargetId="figure-19@14.[370,1216,418,1571]" captionTargetPageId="14" captionText="Figs 36–41. Eniochthonius mahunkai sp. n., immatures:36 = larva, dorsal aspect; 37 = same, ventral aspect (specimen contracted, legs incomplete); 38 = opisthosomal venter of distended larva (adapted from GRANDJEAN 1933), contrasting presence of inguinal seta h4 in E. mahunkai (left side) with its absence in E.minutissmus (right side); 39 = tritonymph, dorsal aspect; 40 = same, ventral aspect (legs incomplete; apa = anterior porose area); 41 = schematic dorsal view of left tarsi III (top) and IV (bottom) at instars indicated, anterior (‘) face to right, distal at top (La = larva, Pn = protonymph, Ad = adult); setal insertions indicated by circles: black = dorsal position, visible directly (half-circles are lateral); dotted = ventral position, by transparency). Scale bars: 50 µm (36–37, 39–40), 10 µm (38)." figureDoi="http://doi.org/10.5281/zenodo.12585403" httpUri="https://zenodo.org/record/12585403/files/figure.png" pageId="15" pageNumber="310">Figs 37, 38</figureCitation>
). Both
<emphasis id="B968EAF0FFCC2838FC7CA16D01FBF945" box="[956,975,1646,1671]" italics="true" pageId="15" pageNumber="310">
p
<subScript id="179834A7FFCC2838FC08A17801FBF945" attach="left" box="[968,975,1659,1671]" fontSize="5" pageId="15" pageNumber="310">4</subScript>
</emphasis>
and
<emphasis id="B968EAF0FFCC2838FC3AA16D0639F945" box="[1018,1037,1646,1671]" italics="true" pageId="15" pageNumber="310">
h
<subScript id="179834A7FFCC2838FBC6A1780639F945" attach="left" box="[1030,1037,1659,1671]" fontSize="5" pageId="15" pageNumber="310">4</subScript>
</emphasis>
lost from all later instars (see Note 8). Paraproctal setation 4–3–2–2 (La-Tn); without paraproctal atrichosy. Adanal and anal plates acquire adult form and setation in Dn.
</paragraph>
<paragraph id="8BA336E2FFCC2827FEBEA1C606F0F8E5" blockId="15.[317,1271,436,1811]" lastBlockId="16.[317,1271,1551,1831]" lastPageId="16" lastPageNumber="311" pageId="15" pageNumber="310">
<emphasis id="B968EAF0FFCC2838FEBEA1C6039FF918" box="[382,427,1733,1754]" italics="true" pageId="15" pageNumber="310">Legs</emphasis>
. General form and proportions similar to those of adult, but articulations with soft cuticle exposed, clearly broader ventrally, as typical for oribatid mites. Ontogeny of setae and solenidia given in
<tableCitation id="C69E0359FFCC2838FE4FA1FD03ECF8D1" box="[399,472,1790,1811]" captionStart="Table 1" captionStartId="7.[322,381,712,733]" captionText="Table 1. Ontogeny of setae and solenidia in Eniochthonius mahunkai sp.n. (Structures are indicated where they are first added and are present through the rest of ontogeny; setae in parentheses repre-" pageId="15" pageNumber="310">Table 1</tableCitation>
; their forms and positions generally as in adult, except as noted below. No variation Scale bars: 10 µm (43, 45, 47–49), 5 µm (42, 44, 46, 50).
<figureCitation id="13272A67FFD32827FC58A0110612F8E5" box="[920,1062,1810,1831]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="16" pageNumber="311">Figs 42 and 44</figureCitation>
layered images.
</paragraph>
<caption id="DF63666AFFD32827FEFDA10C06C3F8C9" ID-DOI="http://doi.org/10.5281/zenodo.12585405" ID-Zenodo-Dep="12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="16" pageNumber="311" startId="16.[317,360,1551,1572]" targetBox="[346,1241,417,1523]" targetPageId="16" targetType="figure">
<paragraph id="8BA336E2FFD32827FEFDA10C06C3F8C9" blockId="16.[317,1271,1551,1831]" pageId="16" pageNumber="311">
<emphasis id="B968EAF0FFD32827FEFDA10C039AF9E6" bold="true" box="[317,430,1551,1572]" pageId="16" pageNumber="311">Figs 42–50.</emphasis>
<taxonomicName id="4C1C4D61FFD32827FE73A10C0095F9E6" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[435,673,1551,1572]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="16" pageNumber="311" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. nov.">
<emphasis id="B968EAF0FFD32827FE73A10C0095F9E6" box="[435,673,1551,1572]" italics="true" pageId="16" pageNumber="311">Eniochthonius mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFD32827FD66A10C00EDF9E6" box="[678,729,1551,1572]" pageId="16" pageNumber="311" rank="species">sp. n.</taxonomicNameLabel>
, immatures: 42 = notogastral seta
<emphasis id="B968EAF0FFD32827FBDBA10C0612F9E6" box="[1051,1062,1551,1572]" italics="true" pageId="16" pageNumber="311">c</emphasis>
<subScript id="179834A7FFD32827FBE6A11F0619F9EA" attach="left" box="[1062,1069,1564,1576]" fontSize="5" pageId="16" pageNumber="311">1</subScript>
of larva; 43 = bothridial seta and interlamellar seta (
<emphasis id="B968EAF0FFD32827FDB0A12F00B7F983" box="[624,643,1580,1601]" italics="true" pageId="16" pageNumber="311">in</emphasis>
) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta
<emphasis id="B968EAF0FFD32827FC68A14A0187F99C" box="[936,947,1609,1630]" italics="true" pageId="16" pageNumber="311">c</emphasis>
<subScript id="179834A7FFD32827FC73A156018EF9A3" attach="left" box="[947,954,1621,1633]" fontSize="5" pageId="16" pageNumber="311">3</subScript>
; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule
<emphasis id="B968EAF0FFD32827FC50A1650197F9B9" box="[912,931,1638,1659]" italics="true" pageId="16" pageNumber="311">ia</emphasis>
on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged.
</paragraph>
</caption>
<paragraph id="8BA336E2FFD22826FEFDA6B706A3F8E7" blockId="17.[317,1270,436,630]" lastBlockId="17.[317,1270,1548,1829]" pageId="17" pageNumber="312">
seen in La, Pn or Dn, but slight asymmetrical variations in Tn: of five individuals studied, one lacked seta
<emphasis id="B968EAF0FFD22826FEA7A6D20342FE24" box="[359,374,465,486]" italics="true" pageId="17" pageNumber="312">l’</emphasis>
from one trochanter IV, another lacked
<emphasis id="B968EAF0FFD22826FD2EA6D20135FE24" box="[750,769,465,486]" italics="true" pageId="17" pageNumber="312">v’</emphasis>
from one genu I and one genu III, and another lacked
<emphasis id="B968EAF0FFD22826FEFDA6ED036CFDC1" box="[317,344,494,515]" italics="true" pageId="17" pageNumber="312">it”</emphasis>
from one tarsus III. Seta
<emphasis id="B968EAF0FFD22826FD87A6ED0067FDC1" box="[583,595,494,515]" italics="true" pageId="17" pageNumber="312">d</emphasis>
of genua II and III minute (2–3 µm), blunt, subspiniform in La, despite not being coupled to respective solenidia; both attaining adult shape and form in Pn. Seta
<emphasis id="B968EAF0FFD22826FB4FA50806AFFDE2" box="[1167,1179,523,544]" italics="true" pageId="17" pageNumber="312">d</emphasis>
of genu I with similar change in form, but position also changes: one alveolar diameter distal to solenidion σ’ in La, but 2–3 diameters distal in nymphs. Leg IV of Pn with typical numerical setation (0–0–0–0–7), but single fastigial seta slightly on anterior, rather than posterior face; position maintained to adult with brightfield (51) or differential interference contrast;
<figureCitation id="13272A67FFD22826FC03A0130633F8E7" box="[963,1031,1808,1829]" captionStart="Figs 51–56" captionStartId="17.[317,360,1548,1569]" captionTargetBox="[319,1268,689,1520]" captionTargetId="figure-141@17.[319,1268,689,1520]" captionTargetPageId="17" captionText="Figs 51–56. Eniochthonius spp., adults. 51A = lateral aspect of E. mahunkai n. sp. (top, e = egg); 51B = E. minutissimus (bottom) from same sample at Cicero, New York; 52 = E. minutissimus, anterolateral aspect of bothridial seta (bo) and nearby region, specimen from Germany; 53 = E. crosbyi, dorsal aspect of bothridial seta, with insert showing dorsolateral aspect of famulus of E. crosbyi (A) andE.minutissimus (B), each ca.10 µm; 54 = E.mahunkai sp.n., ventral aspect of genital region (cf. Fig. 3; arrow points to oblique articulation between middle and anterior aggenital plates; 55 = E. crosbyi, same, but from dissected specimen; 56 = E. minutissimus, same (intact specimen from Poland). Scale bars: 100 µm (51), 10 µm (all others); ag = alveolus of aggenital seta; e = egg; pp = plicature plate. Fig. 52 SEM micrograph (from ALBERTI et al. 1994); others from light microscope" figureDoi="http://doi.org/10.5281/zenodo.12585407" httpUri="https://zenodo.org/record/12585407/files/figure.png" pageId="17" pageNumber="312">Fig. 53</figureCitation>
insert layered.
</paragraph>
<caption id="DF63666AFFD22826FEFDA10F06C2F8CA" ID-DOI="http://doi.org/10.5281/zenodo.12585407" ID-Zenodo-Dep="12585407" httpUri="https://zenodo.org/record/12585407/files/figure.png" pageId="17" pageNumber="312" startId="17.[317,360,1548,1569]" targetBox="[319,1268,689,1520]" targetPageId="17" targetType="figure">
<paragraph id="8BA336E2FFD22826FEFDA10F06C2F8CA" blockId="17.[317,1270,1548,1829]" pageId="17" pageNumber="312">
<emphasis id="B968EAF0FFD22826FEFDA10F0385F9E3" bold="true" box="[317,433,1548,1569]" pageId="17" pageNumber="312">Figs 51–56.</emphasis>
<taxonomicName id="4C1C4D61FFD22826FE78A10E0040F9E0" box="[440,628,1549,1570]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B968EAF0FFD22826FE78A10E0077F9E0" box="[440,579,1549,1570]" italics="true" pageId="17" pageNumber="312">Eniochthonius</emphasis>
spp.
</taxonomicName>
, adults. 51A = lateral aspect of
<taxonomicName id="4C1C4D61FFD22826FC6CA10E0613F9E0" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[940,1063,1549,1570]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="mahunkai" status="n. sp.">
<emphasis id="B968EAF0FFD22826FC6CA10E0613F9E0" box="[940,1063,1549,1570]" italics="true" pageId="17" pageNumber="312">E. mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFD22826FBEEA10E0657F9E0" box="[1070,1123,1549,1570]" pageId="17" pageNumber="312" rank="species">n. sp.</taxonomicNameLabel>
(top, e = egg); 51B =
<taxonomicName id="4C1C4D61FFD22826FE43A129002AF9FD" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[387,542,1578,1599]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD22826FE43A129002AF9FD" box="[387,542,1578,1599]" italics="true" pageId="17" pageNumber="312">E. minutissimus</emphasis>
</taxonomicName>
(bottom) from same sample at Cicero, New York; 52 =
<taxonomicName id="4C1C4D61FFD22826FB95A12906DBF9FD" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[1109,1263,1578,1599]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD22826FB95A12906DBF9FD" box="[1109,1263,1578,1599]" italics="true" pageId="17" pageNumber="312">E. minutissimus</emphasis>
</taxonomicName>
, anterolateral aspect of bothridial seta (
<emphasis id="B968EAF0FFD22826FD7CA14500E0F999" box="[700,724,1606,1627]" italics="true" pageId="17" pageNumber="312">bo</emphasis>
) and nearby region, specimen from Germany; 53 =
<taxonomicName id="4C1C4D61FFD22826FB21A14503B0F9BA" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" class="Arachnida" family="Hypochthoniidae" genus="Hypochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="crosbyi">
<emphasis id="B968EAF0FFD22826FB21A14503B0F9BA" italics="true" pageId="17" pageNumber="312">E. crosbyi</emphasis>
</taxonomicName>
, dorsal aspect of bothridial seta, with insert showing dorsolateral aspect of famulus of
<taxonomicName id="4C1C4D61FFD22826FB21A16003B0F957" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" class="Arachnida" family="Hypochthoniidae" genus="Hypochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="crosbyi">
<emphasis id="B968EAF0FFD22826FB21A16003B0F957" italics="true" pageId="17" pageNumber="312">E. crosbyi</emphasis>
</taxonomicName>
(A) and
<taxonomicName id="4C1C4D61FFD22826FE14A183005EF957" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[468,618,1664,1685]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD22826FE14A183005EF957" box="[468,618,1664,1685]" italics="true" pageId="17" pageNumber="312">E. minutissimus</emphasis>
</taxonomicName>
(B), each ca. 10 µm; 54 =
<taxonomicName id="4C1C4D61FFD22826FC9DA18301E1F957" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[861,981,1664,1685]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. n.">
<emphasis id="B968EAF0FFD22826FC9DA18301E1F957" box="[861,981,1664,1685]" italics="true" pageId="17" pageNumber="312">E. mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFD22826FC19A1830638F957" box="[985,1036,1664,1685]" pageId="17" pageNumber="312" rank="species">sp. n.</taxonomicNameLabel>
, ventral aspect of genital region (cf. Fig. 3; arrow points to oblique articulation between middle and anterior aggenital plates; 55 =
<taxonomicName id="4C1C4D61FFD22826FEB1A1B903E0F90D" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[369,468,1722,1743]" class="Arachnida" family="Hypochthoniidae" genus="Hypochthonius" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="crosbyi">
<emphasis id="B968EAF0FFD22826FEB1A1B903E0F90D" box="[369,468,1722,1743]" italics="true" pageId="17" pageNumber="312">E. crosbyi</emphasis>
</taxonomicName>
, same, but from dissected specimen; 56 =
<taxonomicName id="4C1C4D61FFD22826FCB7A1B90624F90D" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[887,1040,1722,1743]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="17" pageNumber="312" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD22826FCB7A1B90624F90D" box="[887,1040,1722,1743]" italics="true" pageId="17" pageNumber="312">E. minutissimus</emphasis>
</taxonomicName>
, same (intact specimen from Poland). Scale bars: 100 µm (51), 10 µm (all others); ag = alveolus of aggenital seta; e = egg; pp = plicature plate. Fig. 52 SEM micrograph (from
<bibRefCitation id="EF8D4B13FFD22826FCCFA1F001E7F8CA" author="ALBERTI, G. &amp; MORENO, A. I. &amp; KRATZMANN, M." box="[783,979,1779,1800]" pageId="17" pageNumber="312" pagination="57 - 74" refId="ref17235" refString="ALBERTI, G., MORENO, A. I. &amp; KRATZMANN, M. (1994) The fine structure of trichobothria in moss mites with special emphasis on Acrogalumna longipluma (Berlese, 1904) (Oribatida, Acari, Arachnida). Acta Zoologica, Stockholm 75: 57 - 74." type="journal article" year="1994">
<smallCapsWord id="8D45A03EFFD22826FCCFA1F00152F8C4" baselines="1795,1794" box="[783,870,1779,1800]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Alberti" pageId="17" pageNumber="312">ALBERTI</smallCapsWord>
<emphasis id="B968EAF0FFD22826FCABA1F001AFF8CA" box="[875,923,1779,1800]" italics="true" pageId="17" pageNumber="312">et al.</emphasis>
1994
</bibRefCitation>
); others from light microscope
</paragraph>
</caption>
<paragraph id="8BA336E2FFD12825FEFDA6B703D6FDE2" blockId="18.[317,1270,436,630]" pageId="18" pageNumber="313">
(see Note 11). Solenidia on genua II and III and on tibia III less attenuated (ceratiform) in La, becoming attenuate (piliform) in Pn; other solenidia relatively unchanged through ontogeny. Immatures often have thinner branch of famulus not bract-like as in adult, but blunt and same length as knobbed branch (
<figureCitation id="13272A67FFD12825FE4DA50803E2FDE2" box="[397,470,523,544]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="18" pageNumber="313">Fig. 48</figureCitation>
).
</paragraph>
<paragraph id="8BA336E2FFD12825FEBEA52B06B5FDB4" blockId="18.[317,1270,436,630]" pageId="18" pageNumber="313">
<taxonomicName id="4C1C4D61FFD12825FEBEA52B03CDFDFF" authorityName="Krefft" authorityYear="1871" box="[382,505,552,573]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="18" pageNumber="313" phylum="Nematoda" rank="genus">
<emphasis id="B968EAF0FFD12825FEBEA52B03CDFDFF" box="[382,505,552,573]" italics="true" pageId="18" pageNumber="313">Gnathosoma</emphasis>
</taxonomicName>
. General structure as in adult except as noted. Subcapitulum with labiogenal groove slightly less conspicuous (see Note 12,
<figureCitation id="13272A67FFD12825FCC0A5470173FD9B" box="[768,839,580,601]" captionStart="Figs 42–50" captionStartId="16.[317,360,1551,1572]" captionTargetBox="[346,1241,417,1523]" captionTargetId="figure-19@16.[346,1241,417,1523]" captionTargetPageId="16" captionText="Figs 42–50. Eniochthonius mahunkai sp. n., immatures:42 = notogastral seta c1 of larva; 43 = bothridial seta and interlamellar seta (in) of larva, near-dorsal aspect; 44 = humeral region of larval notogaster, lateral aspect, showing porose area (arrowhead) and seta c3; 45 = same, but tritonymph with pleural carina bearing posterior porose area (app) and cupule ia on underside seen by transparency (apa = anterior porose area); 46 = sagittal section of tritonymph notogaster at transverse scissure (anterior to left; arrowhead indicates extremely narrow articulating cuticle); 47 = subcapitulum of tritonymph, ventral aspect in polarized light, showing lack of demineralization in mentum; 48 = partial tarsus I of tritonymph, lateral aspect, showing famulus (e); 49 = posteroventral region of larval opisthosoma, lateral aspect, showing preanal spine (sp) and anal apodeme (aa); 50 = same, enlarged." figureDoi="http://doi.org/10.5281/zenodo.12585405" httpUri="https://zenodo.org/record/12585405/files/figure.png" pageId="18" pageNumber="313">Fig. 47</figureCitation>
). Adoral seta
<emphasis id="B968EAF0FFD12825FC0CA54601DCFD9F" box="[972,1000,581,605]" italics="true" pageId="18" pageNumber="313">
or
<subScript id="179834A7FFD12825FC21A55201DCFD9F" attach="left" box="[993,1000,593,605]" fontSize="5" pageId="18" pageNumber="313">3</subScript>
</emphasis>
formed in Pn; genal seta
<emphasis id="B968EAF0FFD12825FB1DA54606C2FD9F" box="[1245,1270,581,605]" italics="true" pageId="18" pageNumber="313">
m
<subScript id="179834A7FFD12825FB2FA55206C2FD9F" attach="right" box="[1263,1270,593,605]" fontSize="5" pageId="18" pageNumber="313">2</subScript>
</emphasis>
formed in Dn. Palp femoral seta
<emphasis id="B968EAF0FFD12825FD40A56200F5FDB4" box="[640,705,609,630]" italics="true" pageId="18" pageNumber="313">v” (inf</emphasis>
) formed in Dn, tarsal seta
<emphasis id="B968EAF0FFD12825FC0AA56201C6FDB4" box="[970,1010,609,630]" italics="true" pageId="18" pageNumber="313">cmp</emphasis>
formed in Pn.
</paragraph>
<paragraph id="8BA336E2FFD12825FEBEA598039CFCC4" blockId="18.[317,1270,667,774]" pageId="18" pageNumber="313">
<emphasis id="B968EAF0FFD12825FEBEA59803D1FD72" box="[382,485,667,688]" italics="true" pageId="18" pageNumber="313">Etymology</emphasis>
. This species is named in honor of Dr. Sándor
<smallCapsWord id="8D45A03EFFD12825FC61A5980633FD6C" baselines="683,682" box="[929,1031,667,688]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Mahunka" pageId="18" pageNumber="313">MAHUNKA</smallCapsWord>
(Hungarian Natural History Museum, Budapest), who has studied the systematics of acariform mites for more than four decades. Simply put, his explorations of the global diversity of oribatid mites are unmatched and invaluable.
</paragraph>
<paragraph id="8BA336E2FFD12825FEBEA4280128FBCB" blockId="18.[317,1270,811,1466]" pageId="18" pageNumber="313">
<emphasis id="B968EAF0FFD12825FEBEA4280000FC82" box="[382,564,811,832]" italics="true" pageId="18" pageNumber="313">Material examined</emphasis>
.
<materialsCitation id="3B743CBFFFD12825FD80A428006EFCBB" collectingDate="2005-10-05" collectionCode="R" country="United States of America" county="Onondaga Co." latitude="43.178165" location="Beaver Lake Nature Center" longLatPrecision="11" longitude="-76.40483" municipality="Baldwinsville" pageId="18" pageNumber="313" specimenCount="1" stateProvince="New York" typeStatus="holotype">
<emphasis id="B968EAF0FFD12825FD80A42800ACFC82" box="[576,664,811,832]" italics="true" pageId="18" pageNumber="313">
<typeStatus id="54A78840FFD12825FD80A42800ACFC82" box="[576,664,811,832]" pageId="18" pageNumber="313" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F30B7672FFD12825FD65A42800ECFC82" box="[677,728,811,832]" name="United States of America" pageId="18" pageNumber="313">USA</collectingCountry>
;
<collectingRegion id="49D8F800FFD12825FD21A428017DFC82" box="[737,841,811,832]" country="United States of America" name="New York" pageId="18" pageNumber="313">New York</collectingRegion>
;
<collectingCounty id="62C24E6EFFD12825FC93A42801D4FC82" box="[851,992,811,832]" pageId="18" pageNumber="313">Onondaga Co.</collectingCounty>
,
<collectingMunicipality id="6BC7AC98FFD12825FC29A4280645FC82" box="[1001,1137,811,832]" pageId="18" pageNumber="313">Baldwinsville</collectingMunicipality>
;
<location id="8EC36039FFD12825FBBCA42803F2FC9F" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FBBCA42803F2FC9F" country="United States of America" county="Onondaga Co." latitude="43.178165" longLatPrecision="11" longitude="-76.40483" municipality="Baldwinsville" name="Beaver Lake Nature Center" pageId="18" pageNumber="313" stateProvince="New York">
<location id="8EC36039FFD12825FBBCA42806C2FC82" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FBBCA42806C2FC82" box="[1148,1270,811,832]" country="United States of America" county="Onondaga Co." latitude="43.178165" longLatPrecision="11" longitude="-76.40483" municipality="Baldwinsville" name="Beaver Lake" pageId="18" pageNumber="313" stateProvince="New York">
Beaver
<location id="8EC36039FFD12825FB06A42806C2FC82" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FB06A42806C2FC82" box="[1222,1270,811,832]" country="United States of America" county="Onondaga Co." latitude="43.178165" longLatPrecision="11" longitude="-76.40483" municipality="Baldwinsville" name="Lake Nature Center" pageId="18" pageNumber="313" stateProvince="New York">Lake</location>
</location>
Nature Center
</location>
,
<geoCoordinate id="EE285025FFD12825FE0FA44B0076FC9F" box="[463,578,840,861]" degrees="43" direction="north" minutes="10.69" orientation="latitude" pageId="18" pageNumber="313" precision="9" value="43.178165">43°10.69’N</geoCoordinate>
,
<geoCoordinate id="EE285025FFD12825FD8CA44B00F0FC9F" box="[588,708,840,861]" degrees="76" direction="west" minutes="24.29" orientation="longitude" pageId="18" pageNumber="313" precision="9" value="-76.40483">76°24.29’W</geoCoordinate>
, col.
<collectionCode id="ED0DAE27FFD12825FD37A44B013EFC9F" box="[759,778,840,861]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="313">R</collectionCode>
.A.
<smallCapsWord id="8D45A03EFFD12825FCE9A44B014FFC99" baselines="856,855" box="[809,891,840,861]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norton" pageId="18" pageNumber="313">NORTON</smallCapsWord>
,
<date id="FFA21022FFD12825FC47A44B01D2FC9F" box="[903,998,840,861]" pageId="18" pageNumber="313" value="2005-10-05">
<collectingDate id="EFE6E9CAFFD12825FC47A44B01D2FC9F" box="[903,998,840,861]" pageId="18" pageNumber="313" value="2005-10-05">5-X-2005</collectingDate>
</date>
, ex: sphagnum moss on side of hummocks in lakeside fen
</materialsCitation>
.
<materialsCitation id="3B743CBFFFD12825FDA4A466067DFCBB" box="[612,1097,868,890]" collectingDate="2005-10-05" collectionCode="R" country="United States of America" county="Onondaga Co." latitude="43.178165" location="Beaver Lake Nature Center" longLatPrecision="11" longitude="-76.40483" municipality="Baldwinsville" pageId="18" pageNumber="313" specimenCount="130" specimenCount-adult="130" stateProvince="New York" typeStatus="paratype">
<emphasis id="B968EAF0FFD12825FDA4A46600F1FCB8" box="[612,709,869,890]" italics="true" pageId="18" pageNumber="313">
<typeStatus id="54A78840FFD12825FDA4A46600F1FCB8" box="[612,709,869,890]" pageId="18" pageNumber="313" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D1AFD6BFFD12825FD13A4670103FCBB" box="[723,823,868,889]" count="130" pageId="18" pageNumber="313" type="adult">130 adults</specimenCount>
with same data as holotype
</materialsCitation>
.
<materialsCitation id="3B743CBFFFD12825FB93A467012CFBCB" collectionCode="FM, R" country="United States of America" county="Division of Insects" location="Chicago" municipality="Field Museum" pageId="18" pageNumber="313" specimenCount="10" stateProvince="Illinois" typeStatus="holotype">
<typeStatus id="54A78840FFD12825FB93A4670698FCBB" box="[1107,1196,868,889]" pageId="18" pageNumber="313" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D1AFD6BFFD12825FB1DA46703ADFC54" count="10" pageId="18" pageNumber="313" type="generic" typeStatus="paratype">10 paratypes</specimenCount>
deposited in the collection of the
<collectingCounty id="62C24E6EFFD12825FD22A48201AEFC54" box="[738,922,897,918]" pageId="18" pageNumber="313">Division of Insects</collectingCounty>
,
<collectingMunicipality id="6BC7AC98FFD12825FC64A4820604FC54" box="[932,1072,897,918]" pageId="18" pageNumber="313">Field Museum</collectingMunicipality>
(
<collectionCode id="ED0DAE27FFD12825FBFDA4820650FC54" box="[1085,1124,897,918]" country="China" name="Department of Nature, Fujian Province Museum" pageId="18" pageNumber="313">FM</collectionCode>
),
<location id="8EC36039FFD12825FBB5A48206F3FC54" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FBB5A48206F3FC54" box="[1141,1223,897,918]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Chicago" pageId="18" pageNumber="313" stateProvince="Illinois">Chicago</location>
,
<collectingRegion id="49D8F800FFD12825FB12A482035CFC71" country="United States of America" name="Illinois" pageId="18" pageNumber="313">Illinois</collectingRegion>
.
<location id="8EC36039FFD12825FEB0A49D0393FC71" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FEB0A49D0393FC71" box="[368,423,926,947]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Other" pageId="18" pageNumber="313" stateProvince="Illinois">Other</location>
<typeStatus id="54A78840FFD12825FE6CA49D003CFC71" box="[428,520,926,947]" pageId="18" pageNumber="313" type="paratype">paratypes</typeStatus>
deposited as follows:
<quantity id="4CE49B07FFD12825FD1BA49D013FFC71" box="[731,779,926,947]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="18" pageNumber="313" unit="in" value="10.0">10 in</quantity>
the
<location id="8EC36039FFD12825FCF2A49D06B9FC71" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FCF2A49D06B9FC71" box="[818,1165,926,947]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Hungarian Natural History Museum" pageId="18" pageNumber="313" stateProvince="Illinois">Hungarian Natural History Museum</location>
,
<location id="8EC36039FFD12825FB56A49D06C6FC71" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FB56A49D06C6FC71" box="[1174,1266,926,947]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Budapest" pageId="18" pageNumber="313" stateProvince="Illinois">Budapest</location>
;
<quantity id="4CE49B07FFD12825FEFDA4B80359FC12" box="[317,365,955,976]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="18" pageNumber="313" unit="in" value="10.0">10 in</quantity>
the
<location id="8EC36039FFD12825FE53A4B8005DFC12" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FE53A4B8005DFC12" box="[403,617,955,976]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Acarology Laboratory" pageId="18" pageNumber="313" stateProvince="Illinois">Acarology Laboratory</location>
,
<location id="8EC36039FFD12825FDB2A4B800A3FC12" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FDB2A4B800A3FC12" box="[626,663,955,976]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="The" pageId="18" pageNumber="313" stateProvince="Illinois">The</location>
<collectingRegion id="49D8F800FFD12825FD5BA4B80134FC12" box="[667,768,955,976]" country="United States of America" name="Ohio" pageId="18" pageNumber="313">Ohio State</collectingRegion>
<location id="8EC36039FFD12825FCC5A4B80159FC12" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FCC5A4B80159FC12" box="[773,877,955,976]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="University" pageId="18" pageNumber="313" stateProvince="Illinois">University</location>
,
<location id="8EC36039FFD12825FCB4A4B801EDFC12" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FCB4A4B801EDFC12" box="[884,985,955,976]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Columbus" pageId="18" pageNumber="313" stateProvince="Illinois">Columbus</location>
,
<collectingRegion id="49D8F800FFD12825FC22A4B80621FC12" box="[994,1045,955,976]" country="United States of America" name="Ohio" pageId="18" pageNumber="313">Ohio</collectingRegion>
;
<quantity id="4CE49B07FFD12825FBDDA4B80678FC12" box="[1053,1100,955,976]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="18" pageNumber="313" unit="in" value="14.0">14 in</quantity>
the
<location id="8EC36039FFD12825FBB2A4B803D4FC2F" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FBB2A4B803D4FC2F" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Canadian National Collection" pageId="18" pageNumber="313" stateProvince="Illinois">Canadian National Collection</location>
,
<location id="8EC36039FFD12825FE29A4DB0005FC2F" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FE29A4DB0005FC2F" box="[489,561,984,1005]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="Ottawa" pageId="18" pageNumber="313" stateProvince="Illinois">Ottawa</location>
; and
<quantity id="4CE49B07FFD12825FDA2A4DB00A7FC2F" box="[610,659,984,1005]" metricMagnitude="0" metricUnit="m" metricValue="2.1844" pageId="18" pageNumber="313" unit="in" value="86.0">86 in</quantity>
the collection of
<collectionCode id="ED0DAE27FFD12825FCF8A4DB017FFC2F" box="[824,843,984,1005]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="18" pageNumber="313">R</collectionCode>
. A.
<smallCapsWord id="8D45A03EFFD12825FCB0A4DB01F5FC29" baselines="1000,999" box="[880,961,984,1005]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norton" pageId="18" pageNumber="313">NORTON</smallCapsWord>
.
<location id="8EC36039FFD12825FC0DA4DB01D8FC2F" LSID="urn:lsid:plazi:treatment:03B587F4FFC12825FD99A4020645FA7B:8EC36039FFD12825FC0DA4DB01D8FC2F" box="[973,1004,984,1005]" country="United States of America" county="Division of Insects" municipality="Field Museum" name="All" pageId="18" pageNumber="313" stateProvince="Illinois">All</location>
<typeStatus id="54A78840FFD12825FC31A4DB0679FC2F" box="[1009,1101,984,1005]" pageId="18" pageNumber="313" type="paratype">paratypes</typeStatus>
in alcohol except for several on slides in the latter two collections
</materialsCitation>
.
</paragraph>
<paragraph id="8BA336E2FFD12825FEBEA3120128FB11" blockId="18.[317,1270,811,1466]" pageId="18" pageNumber="313">
<emphasis id="B968EAF0FFD12825FEBEA3120038FBE4" box="[382,524,1041,1062]" italics="true" pageId="18" pageNumber="313">Other records.</emphasis>
USA. New York. Onondaga Co., Cicero Game Management Area (
<geoCoordinate id="EE285025FFD12825FB55A3120364FB81" degrees="43" direction="north" minutes="08.87" orientation="latitude" pageId="18" pageNumber="313" precision="9" value="43.147835">43°08.87’ N</geoCoordinate>
,
<geoCoordinate id="EE285025FFD12825FE9CA32D03E1FB81" box="[348,469,1070,1091]" degrees="76" direction="west" minutes="02.87" orientation="longitude" pageId="18" pageNumber="313" precision="9" value="-76.04784">76°02.87’W</geoCoordinate>
),
<date id="FFA21022FFD12825FE27A32D0073FB81" box="[487,583,1070,1091]" pageId="18" pageNumber="313" value="2005-05-03">3-V-2005</date>
, R. A.
<smallCapsWord id="8D45A03EFFD12825FD4EA32D00D4FB80" baselines="1086,1086" box="[654,736,1070,1091]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Norton" pageId="18" pageNumber="313">NORTON</smallCapsWord>
col., ex:
<emphasis id="B968EAF0FFD12825FCFDA32D0196FB81" box="[829,930,1070,1091]" italics="true" pageId="18" pageNumber="313">Sphagnum</emphasis>
and
<taxonomicName id="4C1C4D61FFD12825FC14A32D067FFB81" box="[980,1099,1070,1091]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="18" pageNumber="313" phylum="Bryophyta" rank="genus">
<emphasis id="B968EAF0FFD12825FC14A32D067FFB81" box="[980,1099,1070,1091]" italics="true" pageId="18" pageNumber="313">Polytrichum</emphasis>
</taxonomicName>
moss on sides of hummocks in blueberry bog.
<emphasis id="B968EAF0FFD12825FD9AA3480089FBA2" box="[602,701,1099,1120]" italics="true" pageId="18" pageNumber="313">Wisconsin</emphasis>
. Kenosha Co., Salem (W), Van Halter Bog,
<date id="FFA21022FFD12825FBAEA34806C6FBA2" box="[1134,1266,1099,1120]" pageId="18" pageNumber="313" value="1972-08-15">15-VIII-1972</date>
, W. Suter, col., ex: sphagnum moss.
<emphasis id="B968EAF0FFD12825FD5EA36400DAFBBE" bold="true" box="[670,750,1127,1148]" pageId="18" pageNumber="313">
<collectingCountry id="F30B7672FFD12825FD5EA36400DAFBBE" box="[670,750,1127,1148]" name="Canada" pageId="18" pageNumber="313">Canada</collectingCountry>
</emphasis>
.
<emphasis id="B968EAF0FFD12825FD3BA36B01A5FBBF" box="[763,913,1128,1149]" italics="true" pageId="18" pageNumber="313">
<collectingRegion id="49D8F800FFD12825FD3BA36B01A5FBBF" box="[763,913,1128,1149]" country="Canada" name="New Brunswick" pageId="18" pageNumber="313">New Brunswick</collectingRegion>
</emphasis>
. Kent Co., Kouchibouguac National Park,
<date id="FFA21022FFD12825FEB4A38703DFFB5B" box="[372,491,1156,1177]" pageId="18" pageNumber="313" value="1978-06-16">16-VI–1978</date>
, E. Rickey, col., ex: sphagnum and rhododendron litter.
<emphasis id="B968EAF0FFD12825FBC0A386064EFB58" box="[1024,1146,1157,1178]" italics="true" pageId="18" pageNumber="313">
<collectingRegion id="49D8F800FFD12825FBC0A3860642FB58" box="[1024,1142,1157,1178]" country="Canada" name="Nova Scotia" pageId="18" pageNumber="313">Nova Scotia</collectingRegion>
.
</emphasis>
Cape Breton Highlands National Park, North Mt. spruce forest and bog,
<date id="FFA21022FFD12825FCBCA3A201D3FB74" box="[892,999,1185,1206]" pageId="18" pageNumber="313" value="1983-09-08">8-IX–1983</date>
, V.
<smallCapsWord id="8D45A03EFFD12825FBCFA3A2068BFB77" baselines="1201,1201,1201,1201,1201" box="[1039,1215,1185,1206]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Behan-Pelletier" pageId="18" pageNumber="313">BEHAN-PELLETIER</smallCapsWord>
, col., ex:
<taxonomicName id="4C1C4D61FFD12825FEA2A3BD000BFB11" authority=", Sphagnum" authorityName="Sphagnum" box="[354,575,1214,1235]" class="Magnoliopsida" family="Sarraceniaceae" genus="Sarracenia" kingdom="Plantae" order="Ericales" pageId="18" pageNumber="313" phylum="Tracheophyta" rank="genus">
<emphasis id="B968EAF0FFD12825FEA2A3BD03F9FB11" box="[354,461,1214,1235]" italics="true" pageId="18" pageNumber="313">Sarracenia</emphasis>
,
<emphasis id="B968EAF0FFD12825FE1AA3BD000BFB11" box="[474,575,1214,1235]" italics="true" pageId="18" pageNumber="313">Sphagnum</emphasis>
</taxonomicName>
and dwarf larch litter.
</paragraph>
<paragraph id="8BA336E2FFD12825FEBEA3D80645FA7B" blockId="18.[317,1270,811,1466]" pageId="18" pageNumber="313">
All known sites inhabited by
<taxonomicName id="4C1C4D61FFD12825FD57A3D80124FB32" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[663,784,1243,1264]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="mahunkai" status="sp. n.">
<emphasis id="B968EAF0FFD12825FD57A3D80124FB32" box="[663,784,1243,1264]" italics="true" pageId="18" pageNumber="313">E. mahunkai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B578BFFD12825FCD6A3D8017DFB32" box="[790,841,1243,1264]" pageId="18" pageNumber="313" rank="species">sp. n.</taxonomicNameLabel>
are peatlands (fens or bogs), with sphagnum moss being a consistent component of the microhabitat. It is also consistently collected with
<taxonomicName id="4C1C4D61FFD12825FB21A3FB038EFAE8" baseAuthorityName="BERLESE" baseAuthorityYear="1910" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD12825FB21A3FB038EFAE8" italics="true" pageId="18" pageNumber="313">E. minutissimus</emphasis>
</taxonomicName>
, although the microhabitat preferences of these species may differ. Based on unstructured sampling at the
<typeStatus id="54A78840FFD12825FDC9A2320006FA84" box="[521,562,1329,1350]" pageId="18" pageNumber="313">type</typeStatus>
locality and in a blueberry bog (technically a fen) at Cicero,
<collectingRegion id="49D8F800FFD12825FBB2A23206ECFA84" box="[1138,1240,1329,1350]" country="United States of America" name="New York" pageId="18" pageNumber="313">New York</collectingRegion>
,
<taxonomicName id="4C1C4D61FFD12825FB21A23203A8FAA1" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="B968EAF0FFD12825FB21A23203A8FAA1" italics="true" pageId="18" pageNumber="313">E. mahunkai</emphasis>
</taxonomicName>
is a dominant mite species in sphagnum moss on hummocks, while
<taxonomicName id="4C1C4D61FFD12825FBDAA24D0684FAA1" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[1050,1200,1358,1379]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD12825FBDAA24D0684FAA1" box="[1050,1200,1358,1379]" italics="true" pageId="18" pageNumber="313">E. minutissimus</emphasis>
</taxonomicName>
is relatively infrequent. By contrast, in adjacent clumps of
<taxonomicName id="4C1C4D61FFD12825FCF0A2680193FA42" box="[816,935,1387,1408]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="18" pageNumber="313" phylum="Bryophyta" rank="genus">
<emphasis id="B968EAF0FFD12825FCF0A2680193FA42" box="[816,935,1387,1408]" italics="true" pageId="18" pageNumber="313">Polytrichum</emphasis>
</taxonomicName>
moss at each location
<taxonomicName id="4C1C4D61FFD12825FBBDA26806C2FA42" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[1149,1270,1387,1408]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="B968EAF0FFD12825FBBDA26806C2FA42" box="[1149,1270,1387,1408]" italics="true" pageId="18" pageNumber="313">E. mahunkai</emphasis>
</taxonomicName>
is much less abundant, about as frequent as
<taxonomicName id="4C1C4D61FFD12825FD21A28B014DFA5F" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[737,889,1416,1437]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD12825FD21A28B014DFA5F" box="[737,889,1416,1437]" italics="true" pageId="18" pageNumber="313">E. minutissimus</emphasis>
</taxonomicName>
. In forest litter and humus about
<quantity id="4CE49B07FFD12825FB79A28B06C2FA5F" box="[1209,1270,1416,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="313" unit="m" value="100.0">100 m</quantity>
from the
<typeStatus id="54A78840FFD12825FE58A2A703F5FA7B" box="[408,449,1444,1465]" pageId="18" pageNumber="313">type</typeStatus>
locality
<taxonomicName id="4C1C4D61FFD12825FDD8A2A600A0FA78" authorityName="Norton &amp; Behan-Pelletier" authorityYear="2007" box="[536,660,1445,1466]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="B968EAF0FFD12825FDD8A2A600A0FA78" box="[536,660,1445,1466]" italics="true" pageId="18" pageNumber="313">E. mahunkai</emphasis>
</taxonomicName>
does not occur, but
<taxonomicName id="4C1C4D61FFD12825FC9DA2A601C2FA78" baseAuthorityName="BERLESE" baseAuthorityYear="1910" box="[861,1014,1445,1466]" class="Arachnida" family="Eniochthoniidae" genus="Eniochthonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="18" pageNumber="313" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="B968EAF0FFD12825FC9DA2A601C2FA78" box="[861,1014,1445,1466]" italics="true" pageId="18" pageNumber="313">E. minutissimus</emphasis>
</taxonomicName>
is common.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink