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<document id="CFB145CC86BADF683A6849B4A8B6759D" ID-DOI="10.11646/zootaxa.5443.4.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064312" ID-ZooBank="F4D19D80-3772-4F85-ACB2-6140D2F3BABB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714028820295" checkinUser="plazi" docAuthor="Paz-Sedano, Sofia, Cobb, Gary, Gosliner, Terrence M. &amp; Pola, Marta" docDate="2024" docId="03B587E3FFDFEE0600F94B27FD01FE9F" docLanguage="en" docName="zootaxa.5443.4.3.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naisdoris labalsaensis Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="541" masterDocId="FF8CFF9BFFCFEE15006E4D11FFC5FFDE" masterDocTitle="Filling gaps in the knowledge of Goniodorididae taxa (Mollusca, Gastropoda, Nudibranchia) with description of seven new species" masterLastPageNumber="547" masterPageNumber="523" pageNumber="539" updateTime="1714029648312" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CFEC77E8B3728CBFECE90422DD999FA1">Filling gaps in the knowledge of Goniodorididae taxa (Mollusca, Gastropoda, Nudibranchia) with description of seven new species</mods:title>
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<mods:namePart id="A16CADDA882CE5E7249414F4D84CB68E">Paz-Sedano, Sofia</mods:namePart>
<mods:affiliation id="709407EFCA9FE96868A197B741F5D87D">Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA &amp; Department of Biology, Faculty of Sciences, Autonomous University of Madrid, Madrid, Spain</mods:affiliation>
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<mods:affiliation id="45EF2AF4E4AF72C5DB8B699FFDF9707B">Department of Biology, Faculty of Sciences, Autonomous University of Madrid, Madrid, Spain &amp; Biodiversity and Global Change Research Center (CIBC-UAM), Madrid, Spain</mods:affiliation>
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<heading id="D0EB8199FFDFEE0500F94B27FDCEF98F" bold="true" box="[151,523,1590,1617]" fontSize="11" level="1" pageId="16" pageNumber="539" reason="1">
<emphasis id="B968EAE7FFDFEE0500F94B27FDCEF98F" bold="true" box="[151,523,1590,1617]" pageId="16" pageNumber="539">
<taxonomicName id="4C1C4D76FFDFEE0500F94B27FE60F98F" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[151,421,1590,1617]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="539" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDFEE0500F94B27FE60F98F" bold="true" box="[151,421,1590,1617]" italics="true" pageId="16" pageNumber="539">Naisdoris labalsaensis</emphasis>
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<taxonomicNameLabel id="A25B579CFFDFEE0501C24B26FDCEF98F" box="[428,523,1591,1617]" pageId="16" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C306657EFFDFEE0500F94B4AFD83F8C7" pageId="16" pageNumber="539" type="description">
<paragraph id="8BA336F5FFDFEE0500F94B4AFE02F9AB" blockId="16.[151,1082,1590,1689]" box="[151,455,1627,1653]" pageId="16" pageNumber="539">
(
<figureCitation id="13272A70FFDFEE0500F14B4AFEE1F9AB" box="[159,292,1627,1653]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1917]" captionTargetBox="[227,1361,703,1870]" captionTargetId="figure-211@15.[227,1361,703,1870]" captionTargetPageId="15" captionText="FIGURE 7. Living animals. A. Naisdoris aurornata sp. nov. (CASIZ 186113) B. Naisdoris aurornata sp. nov. (CASIZ 186128). C. Naisdoris labalsaensis sp. nov. (WAM S72659). D. Naisdoris labalsaensis sp. nov. (WAM S72657). E. Naisdoris vitiligata sp. nov. (CASIZ 186126). F. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). (Photos A, B, E by T. Gosliner, C, D by G. Cobb, F by T. C. Hoon)" figureDoi="http://doi.org/10.5281/zenodo.11064326" httpUri="https://zenodo.org/record/11064326/files/figure.png" pageId="16" pageNumber="539">Figs. 7CD</figureCitation>
,
<figureCitation id="13272A70FFDFEE05015E4B4AFEAAF9AB" box="[304,367,1627,1653]" captionStart="FIGURE 6" captionStartId="13.[151,250,1823,1847]" captionTargetBox="[269,1317,193,1797]" captionTargetId="figure-19@13.[258,1328,181,1799]" captionTargetPageId="13" captionText="FIGURE 6. Buccal bulb and reproductive system. AB. Murphydoris polkadotsa sp. nov. (QM MO 86038). A. Buccal bulb. B. Reproductive system. CD. Naisdoris aurornata sp. nov. (CASIZ 186113). C. Buccal bulb. D. Reproductive system. EF. Naisdoris labalsaensis sp. nov. (WAM S72700). E. Buccal bulb. F. Reproductive system. GH. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). G. Buccal bulb. H. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064324" httpUri="https://zenodo.org/record/11064324/files/figure.png" pageId="16" pageNumber="539">6EF</figureCitation>
,
<figureCitation id="13272A70FFDFEE0501144B4AFE7BF9AB" box="[378,446,1627,1653]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="16" pageNumber="539">8CG</figureCitation>
)
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Zoobank:
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<taxonomicName id="4C1C4D76FFDFEE0500F94BD2FEC1F905" box="[151,260,1731,1755]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="539" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B968EAE7FFDFEE0500F94BD2FF25F905" box="[151,224,1731,1755]" italics="true" pageId="16" pageNumber="539">Okenia</emphasis>
sp.
</taxonomicName>
11—
<bibRefCitation id="EF8D4B04FFDFEE0501544BD2FE78F905" author="Cobb, G." box="[314,445,1731,1755]" pageId="16" pageNumber="539" refId="ref15562" refString="Cobb, G. (2012) Okenia sp. 11. Available from: https: // nudibranch. com. au / pages / G _ 3970 b. htm (accessed 25 March 2024)" type="url" year="2012">Cobb (2012)</bibRefCitation>
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<taxonomicName id="4C1C4D76FFDFEE0500F94BF3FEF2F924" baseAuthorityName="Gosliner" baseAuthorityYear="2004" box="[151,311,1762,1786]" class="Gastropoda" family="Goniodorididae" genus="Okenia" isUncertain="true" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="539" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDFEE0500F94BF3FF25F924" box="[151,224,1762,1786]" italics="true" pageId="16" pageNumber="539">Okenia</emphasis>
cf.
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<bibRefCitation id="EF8D4B04FFDFEE05013E4BF3FE2CF924" author="Mullins, D." box="[336,489,1762,1786]" pageId="16" pageNumber="539" refId="ref17152" refString="Mullins, D. (2021) Okenia cf. liklik. Available from: https: // nudibranchdomain. org / product / okenia-cf-liklik / (accessed 25 March 2024)" type="url" year="2021">Mullins (2021)</bibRefCitation>
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<taxonomicName id="4C1C4D76FFDFEE0500F94A11FEF7F8C6" box="[151,306,1792,1816]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="539" phylum="Mollusca" rank="species" species="undefined-E">
<emphasis id="B968EAE7FFDFEE0500F94A11FF3FF8C6" box="[151,250,1792,1816]" italics="true" pageId="16" pageNumber="539">Naisdoris</emphasis>
sp. E
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<bibRefCitation id="EF8D4B04FFDFEE0501224A11FD83F8C7" author="Paz-Sedano, S. &amp; Moles, J. &amp; Smirnoff, D. &amp; Gosliner, T. M. &amp; Pola, M." box="[332,582,1792,1817]" pageId="16" pageNumber="539" pagination="107990" refId="ref17463" refString="Paz-Sedano, S., Moles, J., Smirnoff, D., Gosliner, T. M. &amp; Pola, M. (2024) A combined phylogenetic strategy illuminates the evolution of Goniodorididae nudibranchs (Mollusca, Gastropoda, Heterobranchia). Molecular Phylogenetics and Evolution, 192, 107990. https: // doi. org / 10.1016 / j. ympev. 2023.107990" type="journal article" year="2024">
Paz-Sedano
<emphasis id="B968EAE7FFDFEE0501A44A10FE38F8C6" box="[458,509,1792,1816]" italics="true" pageId="16" pageNumber="539">et al.</emphasis>
(2024)
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<emphasis id="B968EAE7FFDFEE0500F94A56FE84F8BE" bold="true" box="[151,321,1862,1889]" pageId="16" pageNumber="539">Type material</emphasis>
.
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.
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,
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,
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,
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,
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depth
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,
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, col. by
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, 96% EtOH, dissected (
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: radula, labial cuticle)
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.
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<emphasis id="B968EAE7FFDFEE0504C14A7AFAE2F85B" bold="true" box="[1199,1319,1899,1925]" pageId="16" pageNumber="539">
<typeStatus id="54A78857FFDFEE0504C14A7AFAE2F85B" box="[1199,1319,1899,1925]" pageId="16" pageNumber="539" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED0DAE30FFDFEE0505564A7AFAA0F85B" box="[1336,1381,1899,1925]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="539" type="Museum">QM</collectionCode>
<specimenCode id="DBBA9E8EFFDFEE0505014A7AFF25F877" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="539" type="Museum">MO 86035</specimenCode>
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<collectorName id="26E95323FFDFEE0500824A9EFE48F877" box="[236,397,1935,1961]" pageId="16" pageNumber="539">La Balsa Park</collectorName>
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<location id="8EC3602EFFDFEE0501F44A9EFD98F877" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDFEE0501F44A9EFD98F877" box="[410,605,1935,1961]" country="Australia" name="Mooloolah River" pageId="16" pageNumber="539" stateProvince="Queensland">Mooloolah River</location>
,
<collectingRegion id="49D8F817FFDFEE0502074A9EFD35F877" box="[617,752,1935,1961]" country="Australia" name="Queensland" pageId="16" pageNumber="539">Queensland</collectingRegion>
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<collectingCountry id="F30B7665FFDFEE0502924A9EFCA3F877" box="[764,870,1935,1961]" name="Australia" pageId="16" pageNumber="539">Australia</collectingCountry>
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<quantity id="4CE49B10FFDFEE05031C4A9FFC7FF877" box="[882,954,1934,1961]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="16" pageNumber="539" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
, 2019, col. by
<collectorName id="26E95323FFDFEE0504C44A9EFAC9F877" box="[1194,1292,1935,1961]" pageId="16" pageNumber="539">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDFEE0501604AA2FE88F813" box="[270,333,1971,1997]" pageId="16" pageNumber="539">SEM</collectionCode>
: radula, labial cuticle)
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDFEE0502344AA2FAA6F82F" collectingDate="2019-12-11" collectionCode="QM, SEM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="16" pageNumber="539" specimenCode="MO 86036" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDFEE0502344AA2FD42F813" box="[602,647,1971,1997]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="539" type="Museum">QM</collectionCode>
<specimenCode id="DBBA9E8EFFDFEE0502E14AA2FCCBF813" box="[655,782,1971,1997]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="16" pageNumber="539" type="Museum">MO 86036</specimenCode>
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<location id="8EC3602EFFDFEE0503744AA2FC78F813" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDFEE0503744AA2FC78F813" box="[794,957,1971,1997]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="16" pageNumber="539" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDFEE0503A44AA2FB4BF813" box="[970,1166,1971,1997]" pageId="16" pageNumber="539">Mooloolah River</collectingMunicipality>
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<collectingRegion id="49D8F817FFDFEE0504F54AA2FAE7F813" box="[1179,1314,1971,1997]" country="Australia" name="Queensland" pageId="16" pageNumber="539">Queensland</collectingRegion>
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<collectingCountry id="F30B7665FFDFEE0505414AA2FA5CF813" box="[1327,1433,1971,1997]" name="Australia" pageId="16" pageNumber="539">Australia</collectingCountry>
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<elevation id="0031D1C6FFDFEE0500F94AC7FEE2F82F" box="[151,295,2006,2033]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="16" pageNumber="539" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDFEE0500F94AC7FF1AF82F" box="[151,223,2006,2033]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="16" pageNumber="539" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDFEE05015D4AC6FDC8F82F" box="[307,525,2007,2033]" pageId="16" pageNumber="539" value="2019-12-11">
<collectingDate id="EFE6E9DDFFDFEE05015D4AC6FDC8F82F" box="[307,525,2007,2033]" pageId="16" pageNumber="539" value="2019-12-11">11 December 2019</collectingDate>
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, col. by
<collectorName id="26E95323FFDFEE0502034AC6FD0AF82F" box="[621,719,2007,2033]" pageId="16" pageNumber="539">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDFEE0503B34AC6FBD9F82F" box="[989,1052,2007,2033]" pageId="16" pageNumber="539">SEM</collectionCode>
: radula, labial cuticle, penis)
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDFEE0705014AC6FCACFF0B" collectingDate="2019-12-31" collectionCode="AM, SEM" collectorName="G. Cobb" country="Australia" elevation="3" lastPageId="18" lastPageNumber="541" location="La Balsa Park" municipality="Mooloolah River" pageId="16" pageNumber="539" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDFEE0505014AC6FA59F82F" box="[1391,1436,2007,2033]" country="Australia" name="Australian Museum" pageId="16" pageNumber="539" type="Museum">AM</collectionCode>
C.270714,
<location id="8EC3602EFFDDEE07017B4D86FE7DFF6F" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE07017B4D86FE7DFF6F" box="[277,440,151,177]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0701AB4D86FD4FFF6F" box="[453,650,151,177]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
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<collectingRegion id="49D8F817FFDDEE0702F94D86FCDBFF6F" box="[663,798,151,177]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
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<collectingCountry id="F30B7665FFDDEE0703454D86FC50FF6F" box="[811,917,151,177]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
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<quantity id="4CE49B10FFDDEE0703CC4D87FC2EFF6F" box="[930,1003,150,177]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE07042F4D86FAD8FF6F" box="[1089,1309,151,177]" pageId="18" pageNumber="541" value="2019-12-31">
<collectingDate id="EFE6E9DDFFDDEE07042F4D86FAD8FF6F" box="[1089,1309,151,177]" pageId="18" pageNumber="541" value="2019-12-31">31 December 2019</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0705EF4D86FF13FF0B" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDDEE07018D4DAAFDE7FF0B" box="[483,546,187,213]" pageId="18" pageNumber="541">SEM</collectionCode>
: radula, labial cuticle, penis)
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE07031B4DAAFF2AFEC3" collectingDate="2018-08-31" collectionCode="WAM, SEM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72657" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE07031B4DAAFC78FF0B" box="[885,957,187,213]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE0703AA4DABFBD8FF0B" box="[964,1053,186,213]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72657</specimenCode>
,
<location id="8EC3602EFFDDEE0704464DAAFB0CFF0B" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0704464DAAFB0CFF0B" box="[1064,1225,187,213]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0704BB4DAAFA5DFF0B" box="[1237,1432,187,213]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
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<collectingRegion id="49D8F817FFDDEE0700F94DCEFEDBFF27" box="[151,286,223,249]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0701444DCEFE51FF27" box="[298,404,223,249]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE0701F14DCFFDEAFF27" box="[415,559,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE0701F14DCFFE23FF27" box="[415,486,222,249]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0702544DCEFD34FF27" box="[570,753,223,249]" pageId="18" pageNumber="541" value="2018-08-31">
<collectingDate id="EFE6E9DDFFDDEE0702544DCEFD34FF27" box="[570,753,223,249]" pageId="18" pageNumber="541" value="2018-08-31">31 August 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE07033F4DCEFC76FF27" box="[849,947,223,249]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDDEE0704AE4DCEFB3AFF27" box="[1216,1279,223,249]" pageId="18" pageNumber="541">SEM</collectionCode>
: radula, labial cuticle)
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE0700974C12FE11FE9F" collectingDate="2018-09-20" collectionCode="WAM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72658" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE0700974C12FE84FEC3" box="[249,321,259,285]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE0701294C13FE5AFEC3" box="[327,415,258,285]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72658</specimenCode>
,
<location id="8EC3602EFFDDEE0701C44C12FD8DFEC3" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0701C44C12FD8DFEC3" box="[426,584,259,285]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE07023D4C12FCD0FEC3" box="[595,789,259,285]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
,
<collectingRegion id="49D8F817FFDDEE07034E4C12FC62FEC3" box="[800,935,259,285]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0703DF4C12FBDEFEC3" box="[945,1051,259,285]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE07044B4C13FB77FEC3" box="[1061,1202,258,285]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE07044B4C13FBAEFEC3" box="[1061,1131,258,285]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0704D34C12FA5DFEC3" box="[1213,1432,259,285]" pageId="18" pageNumber="541" value="2018-09-20">
<collectingDate id="EFE6E9DDFFDDEE0704D34C12FA5DFEC3" box="[1213,1432,259,285]" pageId="18" pageNumber="541" value="2018-09-20">20 September 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0700874C36FE8FFE9F" box="[233,330,295,321]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE0701B04C36FB24FEBB" collectingDate="2018-09-06" collectionCode="WAM, SEM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72659" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE0701B04C36FDE3FE9F" box="[478,550,295,321]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE0702424C37FD40FE9F" box="[556,645,294,321]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72659</specimenCode>
,
<location id="8EC3602EFFDDEE0702E14C36FCE8FE9F" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0702E14C36FCE8FE9F" box="[655,813,295,321]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0703564C36FC3FFE9F" box="[824,1018,295,321]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
,
<collectingRegion id="49D8F817FFDDEE07046B4C36FB49FE9F" box="[1029,1164,295,321]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0704F84C36FAC5FE9F" box="[1174,1280,295,321]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE0705644C37FA5DFE9F" box="[1290,1432,294,321]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE0705644C37FA94FE9F" box="[1290,1361,294,321]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0700F94C5AFEBDFEBB" box="[151,376,331,357]" pageId="18" pageNumber="541" value="2018-09-06">
<collectingDate id="EFE6E9DDFFDDEE0700F94C5AFEBDFEBB" box="[151,376,331,357]" pageId="18" pageNumber="541" value="2018-09-06">06 September 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0701B34C5AFD84FEBB" box="[477,577,331,357]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDDEE07033B4C5AFC51FEBB" box="[853,916,331,357]" pageId="18" pageNumber="541">SEM</collectionCode>
: radula, labial cuticle, penis)
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE0704814C5AFF19FE73" collectingDate="2018-12-08" collectionCode="WAM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72698" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE0704814C5AFAF2FEBB" box="[1263,1335,331,357]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE07052E4C5AFA52FEBB" box="[1344,1431,331,357]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72698</specimenCode>
,
<location id="8EC3602EFFDDEE0700F94C7EFEFCFE57" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0700F94C7EFEFCFE57" box="[151,313,367,393]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0701284C7EFDCFFE57" box="[326,522,367,393]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
,
<collectingRegion id="49D8F817FFDDEE0702784C7EFD58FE57" box="[534,669,367,393]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0702C44C7EFCD1FE57" box="[682,788,367,393]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE07034E4C7FFC74FE57" box="[800,945,366,393]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE07034E4C7FFCADFE57" box="[800,872,366,393]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0703D34C7EFB5DFE57" box="[957,1176,367,393]" pageId="18" pageNumber="541" value="2018-12-08">
<collectingDate id="EFE6E9DDFFDDEE0703D34C7EFB5DFE57" box="[957,1176,367,393]" pageId="18" pageNumber="541" value="2018-12-08">08 December 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0704944C7EFA99FE57" box="[1274,1372,367,393]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE0700864C82FE11FE0F" collectingDate="2018-12-15" collectionCode="WAM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72699" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE0700864C82FEF5FE73" box="[232,304,403,429]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE0701564C82FE55FE73" box="[312,400,403,429]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72699</specimenCode>
,
<location id="8EC3602EFFDDEE0701F24C82FDF8FE73" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0701F24C82FDF8FE73" box="[412,573,403,429]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0702244C82FCC8FE73" box="[586,781,403,429]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
,
<collectingRegion id="49D8F817FFDDEE0703774C82FC65FE73" box="[793,928,403,429]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0703C24C82FBD3FE73" box="[940,1046,403,429]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE07044C4C83FB77FE73" box="[1058,1202,402,429]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE07044C4C83FBAFFE73" box="[1058,1130,402,429]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0704D04C83FA5DFE73" box="[1214,1432,402,429]" pageId="18" pageNumber="541" value="2018-12-15">
<collectingDate id="EFE6E9DDFFDDEE0704D04C83FA5DFE73" box="[1214,1432,402,429]" pageId="18" pageNumber="541" value="2018-12-15">15 December 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0700874CA6FE8FFE0F" box="[233,330,439,465]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH
</materialsCitation>
;
<materialsCitation id="3B743CA8FFDDEE0701B04CA6FC13FE2B" collectingDate="2018-12-15" collectionCode="WAM, SEM" collectorName="G. Cobb" country="Australia" elevation="3" location="La Balsa Park" municipality="Mooloolah River" pageId="18" pageNumber="541" specimenCode="S72700" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<collectionCode id="ED0DAE30FFDDEE0701B04CA6FDE3FE0F" box="[478,550,439,465]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">WAM</collectionCode>
<specimenCode id="DBBA9E8EFFDDEE0702424CA6FD40FE0F" box="[556,645,439,465]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="18" pageNumber="541" type="Museum">S72700</specimenCode>
,
<location id="8EC3602EFFDDEE0702E14CA6FCE8FE0F" LSID="urn:lsid:plazi:treatment:03B587E3FFDFEE0600F94B27FD01FE9F:8EC3602EFFDDEE0702E14CA6FCE8FE0F" box="[655,813,439,465]" country="Australia" municipality="Mooloolah River" name="La Balsa Park" pageId="18" pageNumber="541" stateProvince="Queensland">La Balsa Park</location>
,
<collectingMunicipality id="6BC7AC8FFFDDEE0703564CA6FC3FFE0F" box="[824,1018,439,465]" pageId="18" pageNumber="541">Mooloolah River</collectingMunicipality>
,
<collectingRegion id="49D8F817FFDDEE07046B4CA6FB49FE0F" box="[1029,1164,439,465]" country="Australia" name="Queensland" pageId="18" pageNumber="541">Queensland</collectingRegion>
,
<collectingCountry id="F30B7665FFDDEE0704F84CA6FAC5FE0F" box="[1174,1280,439,465]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
,
<elevation id="0031D1C6FFDDEE0705644CA7FA5DFE0F" box="[1290,1432,438,465]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<quantity id="4CE49B10FFDDEE0705644CA7FA94FE0F" box="[1290,1361,438,465]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="18" pageNumber="541" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">15 m</quantity>
depth
</elevation>
,
<date id="FFA21035FFDDEE0700F94CCBFEAAFE2B" box="[151,367,474,501]" pageId="18" pageNumber="541" value="2018-12-15">
<collectingDate id="EFE6E9DDFFDDEE0700F94CCBFEAAFE2B" box="[151,367,474,501]" pageId="18" pageNumber="541" value="2018-12-15">15 December 2018</collectingDate>
</date>
, col. by
<collectorName id="26E95323FFDDEE0701A34CCAFDEAFE2B" box="[461,559,475,501]" pageId="18" pageNumber="541">G. Cobb</collectorName>
, 96% EtOH, dissected (
<collectionCode id="ED0DAE30FFDDEE0703544CCAFCBCFE2B" box="[826,889,475,501]" pageId="18" pageNumber="541">SEM</collectionCode>
: radula)
</materialsCitation>
.
</paragraph>
<caption id="DF63667DFFDEEE0400F94A1DFB03F807" ID-DOI="http://doi.org/10.5281/zenodo.11064328" ID-Zenodo-Dep="11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="17" pageNumber="540" startId="17.[151,250,1804,1828]" targetBox="[341,1247,181,1780]" targetPageId="17" targetType="figure">
<paragraph id="8BA336F5FFDEEE0400F94A1DFB03F807" blockId="17.[151,1437,1804,2009]" pageId="17" pageNumber="540">
<emphasis id="B968EAE7FFDEEE0400F94A1DFED1F8FA" bold="true" box="[151,276,1804,1828]" pageId="17" pageNumber="540">FIGURE 8.</emphasis>
Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis.
<emphasis id="B968EAE7FFDEEE0404924A1DFAE9F8FA" bold="true" box="[1276,1324,1804,1828]" pageId="17" pageNumber="540">AB</emphasis>
.
<taxonomicName id="4C1C4D76FFDEEE0405544A1DFF3BF896" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="540" phylum="Mollusca" rank="species" species="aurornata" status="sp. nov.">
<emphasis id="B968EAE7FFDEEE0405544A1DFF3BF896" italics="true" pageId="17" pageNumber="540">Naisdoris aurornata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDEEE04016D4A20FE97F896" box="[259,338,1841,1864]" pageId="17" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
(CASIZ 186113).
<emphasis id="B968EAE7FFDEEE0402604A21FDE5F896" bold="true" box="[526,544,1840,1864]" pageId="17" pageNumber="540">A</emphasis>
. Detail of inner lateral teeth.
<emphasis id="B968EAE7FFDEEE04032A4A21FC90F896" bold="true" box="[836,853,1840,1864]" pageId="17" pageNumber="540">B</emphasis>
. Detail of outer lateral teeth.
<emphasis id="B968EAE7FFDEEE0404174A21FB69F896" bold="true" box="[1145,1196,1840,1864]" pageId="17" pageNumber="540">CG</emphasis>
.
<taxonomicName id="4C1C4D76FFDEEE0404D94A21FA59F896" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[1207,1436,1840,1864]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="540" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDEEE0404D94A21FA59F896" box="[1207,1436,1840,1864]" italics="true" pageId="17" pageNumber="540">Naisdoris labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDEEE0400F94A44FF2CF8B2" box="[151,233,1877,1900]" pageId="17" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B968EAE7FFDEEE04009E4A45FEC7F8B2" bold="true" box="[240,258,1876,1900]" pageId="17" pageNumber="540">C</emphasis>
. Detail of labial cuticle elements (WAM S72659).
<emphasis id="B968EAE7FFDEEE0403794A45FCECF8B2" bold="true" box="[791,809,1876,1900]" pageId="17" pageNumber="540">D</emphasis>
. Detail of inner lateral teeth with denticles (WAM S72657).
<emphasis id="B968EAE7FFDEEE0400F94A69FF6DF84E" bold="true" box="[151,168,1912,1936]" pageId="17" pageNumber="540">E</emphasis>
. Detail of inner lateral teeth without denticles (WAM S72100).
<emphasis id="B968EAE7FFDEEE0403564A69FC82F84E" bold="true" box="[824,839,1912,1936]" pageId="17" pageNumber="540">F</emphasis>
. Detail of outer lateral teeth (WAM S72659).
<emphasis id="B968EAE7FFDEEE04054F4A69FAF0F84E" bold="true" box="[1313,1333,1912,1936]" pageId="17" pageNumber="540">G</emphasis>
. Detail of penial spines (QM MO 86036).
<emphasis id="B968EAE7FFDEEE0401B24A8DFDCFF86A" bold="true" box="[476,522,1948,1972]" pageId="17" pageNumber="540">HJ</emphasis>
.
<taxonomicName id="4C1C4D76FFDEEE0402784A8DFD1EF86A" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[534,731,1948,1972]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="17" pageNumber="540" phylum="Mollusca" rank="species" species="vitiligata" status="sp. nov.">
<emphasis id="B968EAE7FFDEEE0402784A8DFD1EF86A" box="[534,731,1948,1972]" italics="true" pageId="17" pageNumber="540">Naisdoris vitiligata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDEEE04028F4A8CFCF4F86A" box="[737,817,1949,1972]" pageId="17" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
(ZCR.MOL.15410).
<emphasis id="B968EAE7FFDEEE0404644A8DFBDBF86A" bold="true" box="[1034,1054,1948,1972]" pageId="17" pageNumber="540">H</emphasis>
. Detail of inner lateral teeth.
<emphasis id="B968EAE7FFDEEE0405264A8DFA97F86A" bold="true" box="[1352,1362,1948,1972]" pageId="17" pageNumber="540">I</emphasis>
. Detail of outer lateral teeth.
<emphasis id="B968EAE7FFDEEE04011F4AD1FEBBF806" bold="true" box="[369,382,1984,2008]" pageId="17" pageNumber="540">J</emphasis>
. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm.
</paragraph>
</caption>
<paragraph id="8BA336F5FFDDEE0700A94CEEFDF3FCE7" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94CEEFE03FDC7" bold="true" box="[199,454,511,537]" pageId="18" pageNumber="541">External morphology</emphasis>
(
<figureCitation id="13272A70FFDDEE0701BD4CEEFD9DFDC7" box="[467,600,511,537]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1917]" captionTargetBox="[227,1361,703,1870]" captionTargetId="figure-211@15.[227,1361,703,1870]" captionTargetPageId="15" captionText="FIGURE 7. Living animals. A. Naisdoris aurornata sp. nov. (CASIZ 186113) B. Naisdoris aurornata sp. nov. (CASIZ 186128). C. Naisdoris labalsaensis sp. nov. (WAM S72659). D. Naisdoris labalsaensis sp. nov. (WAM S72657). E. Naisdoris vitiligata sp. nov. (CASIZ 186126). F. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). (Photos A, B, E by T. Gosliner, C, D by G. Cobb, F by T. C. Hoon)" figureDoi="http://doi.org/10.5281/zenodo.11064326" httpUri="https://zenodo.org/record/11064326/files/figure.png" pageId="18" pageNumber="541">Figs. 7CD</figureCitation>
). Preserved specimens
<quantity id="4CE49B10FFDDEE0703384CEFFC74FDC7" box="[854,945,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="18" pageNumber="541" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
in length, live up to
<quantity id="4CE49B10FFDDEE0704E04CEFFB1BFDC7" box="[1166,1246,510,537]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="541" unit="mm" value="15.0">15 mm</quantity>
. Body elongated, well developed notal border, serrated due to presence of internal spicules. One middorsal crest also supported by spicules. Two elongated, conical, and thin papillae located in front of rhinophores, one on each side. Tips of anterior papillae thinner than bases. Four elongated and thin, with homogeneous shape, papillae located behind gill, two on each side. Anterior papillae slightly larger than posterior papillae. Rhinophores non-retractile, elongated, and slender, bearing 1316 lamellae oriented posteriorly. Rhinophoral sheath absent. Four thin, simple gill branches form a semicircle around anus, two anteriormost sharing common stalk. One oral tentacle on each side of mouth, short and muscular, somewhat triangular shape. Reproductive opening located on anterior third of right lateral side of body. Mantle covered by spicules.
</paragraph>
<paragraph id="8BA336F5FFDDEE0700A94E53FCE8FBCF" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94E53FEABFC83" bold="true" box="[199,366,834,861]" pageId="18" pageNumber="541">Color pattern</emphasis>
(
<figureCitation id="13272A70FFDDEE0701EE4E52FDCCFC83" box="[384,521,835,861]" captionStart="FIGURE 7" captionStartId="15.[151,250,1893,1917]" captionTargetBox="[227,1361,703,1870]" captionTargetId="figure-211@15.[227,1361,703,1870]" captionTargetPageId="15" captionText="FIGURE 7. Living animals. A. Naisdoris aurornata sp. nov. (CASIZ 186113) B. Naisdoris aurornata sp. nov. (CASIZ 186128). C. Naisdoris labalsaensis sp. nov. (WAM S72659). D. Naisdoris labalsaensis sp. nov. (WAM S72657). E. Naisdoris vitiligata sp. nov. (CASIZ 186126). F. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). (Photos A, B, E by T. Gosliner, C, D by G. Cobb, F by T. C. Hoon)" figureDoi="http://doi.org/10.5281/zenodo.11064326" httpUri="https://zenodo.org/record/11064326/files/figure.png" pageId="18" pageNumber="541">Figs. 7CD</figureCitation>
). Body pink. Mantle edge with light yellow or white patches, interspersed with dark brown spots. Yellow or white patches on mantle edge match with small spikes of serrated border. Pink middorsal line with yellow patches more intense, surrounded with white patch, may have some dispersed small dark brown spots. Anteriormost papillae dark brown, with dark yellow base. Posteriormost papillae white at outer half of papillae, basal half same coloration as body. Gill branches translucent with light yellow tips. Rhinophores with brown tips and white basal lamellae. Foot translucent pink.
</paragraph>
<paragraph id="8BA336F5FFDDEE0700A9490AFB36FAA7" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A9490AFE5CFBEB" bold="true" box="[199,409,1051,1077]" pageId="18" pageNumber="541">Foregut anatomy</emphasis>
(
<figureCitation id="13272A70FFDDEE0701C2490AFDD4FBEB" box="[428,529,1051,1077]" captionStart="FIGURE 6" captionStartId="13.[151,250,1823,1847]" captionTargetBox="[269,1317,193,1797]" captionTargetId="figure-19@13.[258,1328,181,1799]" captionTargetPageId="13" captionText="FIGURE 6. Buccal bulb and reproductive system. AB. Murphydoris polkadotsa sp. nov. (QM MO 86038). A. Buccal bulb. B. Reproductive system. CD. Naisdoris aurornata sp. nov. (CASIZ 186113). C. Buccal bulb. D. Reproductive system. EF. Naisdoris labalsaensis sp. nov. (WAM S72700). E. Buccal bulb. F. Reproductive system. GH. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). G. Buccal bulb. H. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064324" httpUri="https://zenodo.org/record/11064324/files/figure.png" pageId="18" pageNumber="541">Figs. 6E</figureCitation>
,
<figureCitation id="13272A70FFDDEE07024E490AFDA7FBEB" box="[544,610,1051,1077]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">8CF</figureCitation>
). Buccal bulb thick and muscular (
<figureCitation id="13272A70FFDDEE070391490AFB9FFBEB" box="[1023,1114,1051,1077]" captionStart="FIGURE 6" captionStartId="13.[151,250,1823,1847]" captionTargetBox="[269,1317,193,1797]" captionTargetId="figure-19@13.[258,1328,181,1799]" captionTargetPageId="13" captionText="FIGURE 6. Buccal bulb and reproductive system. AB. Murphydoris polkadotsa sp. nov. (QM MO 86038). A. Buccal bulb. B. Reproductive system. CD. Naisdoris aurornata sp. nov. (CASIZ 186113). C. Buccal bulb. D. Reproductive system. EF. Naisdoris labalsaensis sp. nov. (WAM S72700). E. Buccal bulb. F. Reproductive system. GH. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). G. Buccal bulb. H. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064324" httpUri="https://zenodo.org/record/11064324/files/figure.png" pageId="18" pageNumber="541">Fig. 6E</figureCitation>
). Markedly large and wide, dorsal buccal pump expanding posteriorly. Small, flat radular sac located ventrally. Thin esophagus begins from buccal bulb behind buccal pump. Salivary glands rounded, located at junction of esophagus with buccal bulb. Nervous system covers this junction. Esophagus continues posteriorly and inserts into digestive-hermaphroditic gland. Labial cuticle surrounds lips and expands within buccal pump. Inside buccal bulb, thin labial cuticle bears honey-comb elements (
<figureCitation id="13272A70FFDDEE0701F149DEFE3CFB37" box="[415,505,1231,1257]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Fig. 8C</figureCitation>
). Radular formula 1820 × 1.1.0.1.1. Inner lateral tooth with single large and thin cusp, and wide, rectangular base (
<figureCitation id="13272A70FFDDEE07027D49E2FD5DFAD3" box="[531,664,1267,1293]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Figs. 8DE</figureCitation>
). Cusp large and pointed, masticatory margin may bear 2028 small, pointed denticles (
<figureCitation id="13272A70FFDDEE0701074806FE06FAEF" box="[361,451,1303,1329]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Fig. 8D</figureCitation>
) or may be smooth (
<figureCitation id="13272A70FFDDEE0702D84806FCCAFAEF" box="[694,783,1303,1329]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Fig. 8E</figureCitation>
). When present, denticles with homogenous shape, those located at extremities slightly smaller. Outer base ends in somewhat prominent wing, with rounded edge. Outer lateral tooth much smaller, rectangular with thin hook in superior part and a hooked point (
<figureCitation id="13272A70FFDDEE0704FF484EFB23FAA7" box="[1169,1254,1375,1401]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BA336F5FFDDEE0700A94893FC9AF963" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94893FE7BFA43" bold="true" box="[199,446,1410,1437]" pageId="18" pageNumber="541">Reproductive system</emphasis>
(
<figureCitation id="13272A70FFDDEE0701A44892FDE2FA43" box="[458,551,1411,1437]" captionStart="FIGURE 6" captionStartId="13.[151,250,1823,1847]" captionTargetBox="[269,1317,193,1797]" captionTargetId="figure-19@13.[258,1328,181,1799]" captionTargetPageId="13" captionText="FIGURE 6. Buccal bulb and reproductive system. AB. Murphydoris polkadotsa sp. nov. (QM MO 86038). A. Buccal bulb. B. Reproductive system. CD. Naisdoris aurornata sp. nov. (CASIZ 186113). C. Buccal bulb. D. Reproductive system. EF. Naisdoris labalsaensis sp. nov. (WAM S72700). E. Buccal bulb. F. Reproductive system. GH. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). G. Buccal bulb. H. Reproductive system. Abbreviations: am, ampulla; bc, bursa copulatrix; bp, buccal pump; fgm, female gland mass; hd, hermaphroditic duct; oe, esophagus; op, esophageal pump; p, penis; pr, prostate; ra, radular sac; rs, receptaculum seminis; sgl, salivary gland; ud, uterine duct; va, vagina; vd, vas deferens. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064324" httpUri="https://zenodo.org/record/11064324/files/figure.png" pageId="18" pageNumber="541">Figs. 6F</figureCitation>
,
<figureCitation id="13272A70FFDDEE0702414892FD90FA43" box="[559,597,1411,1437]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">8G</figureCitation>
). Reproductive system located at anterior-third of body.Thin hermaphroditic duct begins at ovotestis, located inside digestive-hermaphroditic gland. Hermaphroditic duct expands into large, kidney-shape ampulla. Postampullary duct emerges from ampulla and divides into two different, thin ducts. Short oviduct enters inside female gland mass. Second duct connects with first portion of prostate. Prostate sausage-shape, large. Prostate becomes a long and very thin vas deferens. It continues and expands to short ejaculatory duct. Penis with small, hooked penial spines (
<figureCitation id="13272A70FFDDEE07020C4B26FD78F98F" box="[610,701,1591,1617]" captionStart="FIGURE 8" captionStartId="17.[151,250,1804,1828]" captionTargetBox="[341,1247,181,1780]" captionTargetId="figure-19@17.[341,1247,181,1780]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs and light microscope photographs of cuticles, radulae and penis. AB. Naisdoris aurornata sp. nov. (CASIZ 186113). A. Detail of inner lateral teeth. B. Detail of outer lateral teeth. CG. Naisdoris labalsaensis sp. nov. C. Detail of labial cuticle elements (WAM S72659). D. Detail of inner lateral teeth with denticles (WAM S72657). E. Detail of inner lateral teeth without denticles (WAM S72100). F. Detail of outer lateral teeth (WAM S72659). G. Detail of penial spines (QM MO 86036). HJ. Naisdoris vitiligata sp. nov. (ZCR.MOL.15410). H. Detail of inner lateral teeth. I. Detail of outer lateral teeth. J. Detail penial spines. Scale bars: A, B, D, E, H: 20 μm; C, 5 μm; F, G, I, J: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.11064328" httpUri="https://zenodo.org/record/11064328/files/figure.png" pageId="18" pageNumber="541">Fig. 8G</figureCitation>
). Vagina large, similar in width to vas deferens, slightly shorter. Vagina divides in two small and thin ducts. One connects with oval bursa copulatrix, other duct connects with oval receptaculum seminis. At same point of division of ducts arises thin, large uterine duct that enters female gland mass. Receptaculum seminis similar in size to bursa copulatrix.
</paragraph>
</subSubSection>
<subSubSection id="C306657EFFDDEE0700A94BD6FD44F8DB" pageId="18" pageNumber="541" type="etymology">
<paragraph id="8BA336F5FFDDEE0700A94BD6FD44F8DB" blockId="18.[151,1437,150,2013]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94BD6FE8DF93F" bold="true" box="[199,328,1735,1761]" pageId="18" pageNumber="541">Etymology</emphasis>
. The name of the species
<taxonomicName id="4C1C4D76FFDDEE07021E4BD6FCDEF93F" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[624,795,1735,1761]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="labalsaensis">
<emphasis id="B968EAE7FFDDEE07021E4BD6FCDEF93F" box="[624,795,1735,1761]" italics="true" pageId="18" pageNumber="541">N. labalsaensis</emphasis>
</taxonomicName>
refers to the La Balsa Park, place where the species was found and where its presence was common.
</paragraph>
</subSubSection>
<subSubSection id="C306657EFFDDEE0700A94A1FFACCF893" pageId="18" pageNumber="541" type="distribution">
<paragraph id="8BA336F5FFDDEE0700A94A1FFB1CF8F7" blockId="18.[151,1437,150,2013]" box="[199,1241,1806,1833]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94A1FFE9CF8F6" bold="true" box="[199,345,1806,1832]" pageId="18" pageNumber="541">Distribution</emphasis>
.
<collectingCountry id="F30B7665FFDDEE0701094A1EFE62F8F7" box="[359,423,1807,1833]" name="Japan" pageId="18" pageNumber="541">Japan</collectingCountry>
(
<bibRefCitation id="EF8D4B04FFDDEE0701D84A1EFD95F8F7" author="Nakano, R." box="[438,592,1807,1833]" pageId="18" pageNumber="541" refId="ref17184" refString="Nakano, R. (2018) Field Guide to Sea Slugs and Nudibranchs of Japan. Bun-ichi Co., Ltd., Tokyo, 544 pp." type="book" year="2018">Nakano 2018</bibRefCitation>
) and
<collectingCountry id="F30B7665FFDDEE0702E34A1EFD31F8F7" box="[653,756,1807,1833]" name="Australia" pageId="18" pageNumber="541">Australia</collectingCountry>
(
<bibRefCitation id="EF8D4B04FFDDEE07036D4A1EFC41F8F7" author="Cobb, G." box="[771,900,1807,1833]" pageId="18" pageNumber="541" refId="ref15562" refString="Cobb, G. (2012) Okenia sp. 11. Available from: https: // nudibranch. com. au / pages / G _ 3970 b. htm (accessed 25 March 2024)" type="url" year="2012">Cobb 2012</bibRefCitation>
;
<bibRefCitation id="EF8D4B04FFDDEE0703E14A1EFBEDF8F7" author="Mullins, D." box="[911,1064,1807,1833]" pageId="18" pageNumber="541" refId="ref17152" refString="Mullins, D. (2021) Okenia cf. liklik. Available from: https: // nudibranchdomain. org / product / okenia-cf-liklik / (accessed 25 March 2024)" type="url" year="2021">Mullins 2021</bibRefCitation>
; present study).
</paragraph>
<paragraph id="8BA336F5FFDDEE0700A94A22FACCF893" blockId="18.[151,1437,150,2013]" box="[199,1289,1842,1869]" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94A22FE45F892" bold="true" box="[199,384,1842,1869]" pageId="18" pageNumber="541">Natural history</emphasis>
. The species has been found on silty, sandy substrates and rocks up to
<quantity id="4CE49B10FFDDEE0704F44A22FB00F893" box="[1178,1221,1843,1869]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="541" unit="m" value="6.0">6 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="C306657EFFDDEE0600A94A46FD01FE9F" lastPageId="19" lastPageNumber="542" pageId="18" pageNumber="541" type="discussion">
<paragraph id="8BA336F5FFDDEE0600A94A46FD01FE9F" blockId="18.[151,1437,150,2013]" lastBlockId="19.[151,1436,151,321]" lastPageId="19" lastPageNumber="542" pageId="18" pageNumber="541">
<emphasis id="B968EAE7FFDDEE0700A94A46FEF1F8AF" bold="true" box="[199,308,1879,1905]" pageId="18" pageNumber="541">Remarks</emphasis>
. The species
<taxonomicName id="4C1C4D76FFDDEE0701A64A46FDB7F8AF" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[456,626,1879,1905]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDDEE0701A64A46FDB7F8AF" box="[456,626,1879,1905]" italics="true" pageId="18" pageNumber="541">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDDEE0702164A46FD0AF8AF" box="[632,719,1879,1905]" pageId="18" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<taxonomicName id="4C1C4D76FFDDEE0703354A46FC77F8AF" baseAuthorityName="Gosliner" baseAuthorityYear="2004" box="[859,946,1879,1905]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDDEE0703354A46FC77F8AF" box="[859,946,1879,1905]" italics="true" pageId="18" pageNumber="541">N. liklik</emphasis>
</taxonomicName>
in sharing a pinkish color pattern. However,
<taxonomicName id="4C1C4D76FFDDEE0700F94A6AFE87F84B" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[151,322,1915,1941]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDDEE0700F94A6AFE87F84B" box="[151,322,1915,1941]" italics="true" pageId="18" pageNumber="541">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDDEE0701244A6AFE66F84B" box="[330,419,1915,1941]" pageId="18" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
has yellow and black patches in the dorsal part of the body while
<taxonomicName id="4C1C4D76FFDDEE0704E14A6AFB2CF84B" baseAuthorityName="Gosliner" baseAuthorityYear="2004" box="[1167,1257,1915,1941]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDDEE0704E14A6AFB2CF84B" box="[1167,1257,1915,1941]" italics="true" pageId="18" pageNumber="541">N. liklik</emphasis>
</taxonomicName>
has a chocolate brown band and white markings (
<bibRefCitation id="EF8D4B04FFDDEE07027F4A8EFD7DF867" author="Gosliner, T. M." box="[529,696,1951,1977]" pageId="18" pageNumber="541" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner 2004</bibRefCitation>
). The anteriormost papillae are dark brown with dark yellow base in
<taxonomicName id="4C1C4D76FFDDEE0700DA4AD2FE9AF803" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[180,351,1987,2013]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDDEE0700DA4AD2FE9AF803" box="[180,351,1987,2013]" italics="true" pageId="18" pageNumber="541">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDDEE0701094AD2FE05F803" box="[359,448,1987,2013]" pageId="18" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
and tipped with an orange subapical marking in
<taxonomicName id="4C1C4D76FFDDEE07038B4AD2FB3EF803" authority="(Gosliner 2004)" baseAuthorityName="Gosliner" baseAuthorityYear="2004" box="[997,1275,1987,2013]" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDDEE07038B4AD2FBFBF803" box="[997,1086,1987,2013]" italics="true" pageId="18" pageNumber="541">N. liklik</emphasis>
(
<bibRefCitation id="EF8D4B04FFDDEE0704234AD2FB36F803" author="Gosliner, T. M." box="[1101,1267,1987,2013]" pageId="18" pageNumber="541" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner 2004</bibRefCitation>
)
</taxonomicName>
. Moreover,
<taxonomicName id="4C1C4D76FFDDEE0605ED4AD2FF0AFF6F" baseAuthorityName="Gosliner" baseAuthorityYear="2004" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" lastPageId="19" lastPageNumber="542" order="Nudibranchia" pageId="18" pageNumber="541" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDDEE0605ED4AD2FF0AFF6F" italics="true" lastPageId="19" lastPageNumber="542" pageId="18" pageNumber="541">N. liklik</emphasis>
</taxonomicName>
has papillae on the margin edge, and one mid-dorsal papilla (
<bibRefCitation id="EF8D4B04FFDCEE0603FA4D86FBF9FF6F" author="Gosliner, T. M." box="[916,1084,151,177]" pageId="19" pageNumber="542" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner 2004</bibRefCitation>
). However,
<taxonomicName id="4C1C4D76FFDCEE0604A94D86FAB6FF6F" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[1223,1395,151,177]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="542" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDCEE0604A94D86FAB6FF6F" box="[1223,1395,151,177]" italics="true" pageId="19" pageNumber="542">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDCEE0605124D86FF0CFF0B" pageId="19" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
has a serrated notal border supported by internal spicules, lacking papillae. There are also some differences in the internal organs.
<taxonomicName id="4C1C4D76FFDCEE0601144DCEFDBCFF27" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[378,633,223,249]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="542" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDCEE0601144DCEFDBCFF27" box="[378,633,223,249]" italics="true" pageId="19" pageNumber="542">Naisdoris labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDCEE0602EC4DCEFD1EFF27" box="[642,731,223,249]" pageId="19" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
has much longer vagina and longer uterine duct than
<taxonomicName id="4C1C4D76FFDCEE06052C4DCEFE88FEC3" authority="(Gosliner 2004)" baseAuthorityName="Gosliner" baseAuthorityYear="2004" class="Gastropoda" family="Goniodorididae" genus="Okenia" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="542" phylum="Mollusca" rank="species" species="liklik">
<emphasis id="B968EAE7FFDCEE06052C4DCEFA59FF27" box="[1346,1436,223,249]" italics="true" pageId="19" pageNumber="542">N. liklik</emphasis>
(
<bibRefCitation id="EF8D4B04FFDCEE0600F14C12FE80FEC3" author="Gosliner, T. M." box="[159,325,259,285]" pageId="19" pageNumber="542" pagination="125" refId="ref15979" refString="Gosliner, T. M. (2004) Phylogenetic systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55 (1 / 12), 125." type="journal article" year="2004">Gosliner 2004</bibRefCitation>
)
</taxonomicName>
. The length of these ducts is characteristic of
<taxonomicName id="4C1C4D76FFDCEE0603204C12FC3CFEC3" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[846,1017,259,285]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="542" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDCEE0603204C12FC3CFEC3" box="[846,1017,259,285]" italics="true" pageId="19" pageNumber="542">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDCEE06046F4C12FB9CFEC3" box="[1025,1113,259,285]" pageId="19" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
The big buccal pump is also distinctively elongated in
<taxonomicName id="4C1C4D76FFDCEE0601D74C36FDA1FE9F" authority="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola, 2024" authorityName="Paz-Sedano &amp; Cobb &amp; Gosliner &amp; Pola" authorityYear="2024" box="[441,612,295,321]" class="Gastropoda" family="Goniodorididae" genus="Naisdoris" higherTaxonomySource="CoL" kingdom="Animalia" order="Nudibranchia" pageId="19" pageNumber="542" phylum="Mollusca" rank="species" species="labalsaensis" status="sp. nov.">
<emphasis id="B968EAE7FFDCEE0601D74C36FDA1FE9F" box="[441,612,295,321]" italics="true" pageId="19" pageNumber="542">N. labalsaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B579CFFDCEE0602054C36FD01FE9F" box="[619,708,295,321]" pageId="19" pageNumber="542" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>