183 lines
19 KiB
XML
183 lines
19 KiB
XML
<document id="0FA6A1A2F0828B69154E8C1642EC7061" ID-DOI="10.5281/zenodo.177278" ID-GBIF-Dataset="44e92dc8-7d91-43bd-900b-36f28549b744" ID-ISSN="1175-5326" ID-Zenodo-Dep="177278" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459857368899" checkinUser="plazi" docAuthor="Wallach, Van" docDate="2007" docId="03B587D9FF98381D90A8FB97F77FFEA7" docLanguage="en" docName="zt01515p068.pdf" docOrigin="Zootaxa 1515" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Letheobia jubana Wallach, 2007, sp. nov." docType="treatment" docVersion="11" lastPageNumber="45" masterDocId="FF8CFFA1FF953813903FFFBDF365FFA5" masterDocTitle="A review of East and Central African species of Letheobia Cope, revived from the synonymy of Rhinotyphlops Fitzinger, with descriptions of five new species (Serpentes: Typhlopidae)" masterLastPageNumber="68" masterPageNumber="31" pageNumber="44" updateTime="1698225740277" updateUser="plazi">
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<mods:mods id="6743CF37EDBD96D062D398185CE7420A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="EDE31A86E7235A879253E05F50D23C3C">
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<mods:title id="566F3339DAC62DF2D2090C79BA752A1F">A review of East and Central African species of Letheobia Cope, revived from the synonymy of Rhinotyphlops Fitzinger, with descriptions of five new species (Serpentes: Typhlopidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="AEAA66A6B988F08A03DCA29B84F27C01" type="personal">
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<mods:role id="C072C498133CDA4BCB8C2C5C1D9C54E5">
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<mods:roleTerm id="8DD7B08C4314C912CA058D38576DD3CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CBDDBF1514B20BF70D4755D0D40349E5">Wallach, Van</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2118279DBD94F015719F8B0B9E913534">text</mods:typeOfResource>
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<mods:titleInfo id="5977859E0DCA1C794BFE02CADC66E70E">
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<mods:title id="09D52BCE2BD5D45233FE3E9BC4028ED4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4AC547857BE0F9A1D5DB461543654655">
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<mods:date id="FD1E42D2239F191DE674577438F84838">2007</mods:date>
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<mods:detail id="D5CFF283BCD820D4DA470D02A772D136" type="volume">
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<mods:number id="BEF13CE604BFE2874D42EE31F711291A">1515</mods:number>
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</mods:detail>
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<mods:extent id="926AFE6071F1838F7EB92727CFFD4D42" unit="page">
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<mods:start id="A6904635920CD9D24578943F5371546C">31</mods:start>
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<mods:end id="96583E0D82F447B9A529CB2BE05F19BE">68</mods:end>
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<mods:classification id="87F40F0B774443485B8F8FA49FB4BB61">journal article</mods:classification>
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<mods:identifier id="F5F8DF86F88656B551442DE0CB68FF1F" type="DOI">10.5281/zenodo.177278</mods:identifier>
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<mods:identifier id="A85394B6852C1322D704893D02FABDD3" type="GBIF-Dataset">44e92dc8-7d91-43bd-900b-36f28549b744</mods:identifier>
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<mods:identifier id="1BD35BEE3B5F2061FDAF49D8071F0CA5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0FEC1F257D284ED8248877F7E59FF953" type="Zenodo-Dep">177278</mods:identifier>
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</mods:mods>
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<treatment id="03B587D9FF98381D90A8FB97F77FFEA7" ID-DOI="http://doi.org/10.5281/zenodo.6237798" ID-GBIF-Taxon="119363124" ID-Zenodo-Dep="6237798" LSID="urn:lsid:plazi:treatment:03B587D9FF98381D90A8FB97F77FFEA7" httpUri="http://treatment.plazi.org/id/03B587D9FF98381D90A8FB97F77FFEA7" lastPageId="14" lastPageNumber="45" pageId="13" pageNumber="44">
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<subSubSection id="C3066544FF98381E90A8FB97F39EFBC2" pageId="13" pageNumber="44" type="nomenclature">
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<paragraph id="8BA336CFFF98381E90A8FB97F2B5FBE1" blockId="13.[151,464,1066,1167]" box="[151,464,1066,1092]" pageId="13" pageNumber="44">
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<heading id="D0EB81A3FF98381E90A8FB97F2B5FBE1" bold="true" box="[151,464,1066,1092]" fontSize="11" level="1" pageId="13" pageNumber="44" reason="1">
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||
<emphasis id="B968EADDFF98381E90A8FB97F2B5FBE1" bold="true" box="[151,464,1066,1092]" pageId="13" pageNumber="44">
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<taxonomicName id="4C1C4D4CFF98381E90A8FB97F20EFBE1" box="[151,363,1066,1092]" class="Reptilia" family="Typhlopidae" genus="Letheobia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="species" species="jubana" status="sp. nov.">
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<emphasis id="B968EADDFF98381E90A8FB97F20EFBE1" bold="true" box="[151,363,1066,1092]" italics="true" pageId="13" pageNumber="44">Letheobia jubana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57A6FF98381E914CFB97F2B5FBE1" box="[371,464,1066,1092]" pageId="13" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA336CFFF98381E90A8FBF0F39EFBC2" blockId="13.[151,464,1066,1167]" box="[151,251,1101,1127]" pageId="13" pageNumber="44">
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(
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<figureCitation id="13272A4AFF98381E90A0FBF0F3B8FBC2" box="[159,221,1101,1127]" captionStart="FIGURE 3" captionStartId="12.[151,255,1740,1764]" captionTargetBox="[397,1198,579,1690]" captionTargetId="figure@12.[367,1220,554,1716]" captionTargetPageId="12" captionText="FIGURE 3. The Letheobia obtusa complex: dorsal, lateral and ventral views of head of: A. L. obtusa (BMNH 1947.1. 1.24 – Blantyre, Malawi, after Roux-Estève, 1974: Fig. 42); B. L. decorosa (MNHN 1966 - 1251 – Ngaoundéré, Cameroon, after Roux-Estève, 1974: Fig. 44); C. L. jubana sp. nov. (CAS 151201, holotype – Mareri, Juba River, Somalia, J. Boundy illustration). The lines indicate 1 mm to scale." httpUri="https://zenodo.org/record/177281/files/figure.png" pageId="13" pageNumber="44">Fig 3</figureCitation>
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C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066544FF98381E90A8FBC8F2D1FB2A" box="[151,436,1141,1167]" pageId="13" pageNumber="44" type="description">
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<paragraph id="8BA336CFFF98381E90A8FBC8F2D1FB2A" blockId="13.[151,464,1066,1167]" box="[151,436,1141,1167]" pageId="13" pageNumber="44">Juba gracile blind-snake</paragraph>
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</subSubSection>
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<subSubSection id="C3066544FF98381E90A8FB75F073FB5A" pageId="13" pageNumber="44" type="reference_group">
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<paragraph id="8BA336CFFF98381E90A8FB75F073FB5A" blockId="13.[151,1134,1224,1279]" pageId="13" pageNumber="44">
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<treatmentCitationGroup id="AB0C11E1FF98381E90A8FB75F073FB5A" pageId="13" pageNumber="44">
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<emphasis id="B968EADDFF98381E90A8FB75F28FFB7A" box="[151,490,1224,1247]" italics="true" pageId="13" pageNumber="44">
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<taxonomicName id="4C1C4D4CFF98381E90A8FB75F2B3FB7A" box="[151,470,1224,1247]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="subSpecies" species="lineolatus" subSpecies="lineolatus">Typhlops lineolatus lineolatus</taxonomicName>
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–
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</emphasis>
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(not Jan)
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<treatmentCitation id="0ABD10DEFF98381E926AFB75F0F8FB7A" author="Hoevers" box="[597,925,1224,1247]" page="182" pageId="13" pageNumber="44" year="1982">
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<bibRefCitation id="EF8D4B3EFF98381E926AFB75F002FB7A" author="Hoevers" box="[597,871,1224,1247]" lastPageId="13" lastPageNumber="66" pageId="13" pageNumber="65" refString="Hoevers, L. G. & Johnson, P. M. (1982) Notes on a collection of snakes from the Middle Juba Region, Somalia. Monitore Zoologico Italiano (Firenze), (N. S.) Supplemento 16 (8), 173 - 203." type="journal article" year="1982">Hoevers & Johnson 1982</bibRefCitation>
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: 182
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</treatmentCitation>
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;
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<treatmentCitation id="0ABD10DEFF98381E9398FB75F70FFB7A" author="Lambert" box="[935,1130,1224,1247]" page="66" pageId="13" pageNumber="44" year="1985">
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<bibRefCitation id="EF8D4B3EFF98381E9398FB75F727FB7A" author="Lambert" box="[935,1090,1224,1247]" pageId="13" pageNumber="66" refString="Lambert, M. R. K. (1985) Reptiles. In: Report on the species of vertebrates in riverine and coastal forest habitats of southern Somalia and the lower Tana in Kenya. University College London & Tropical Development and Research Institute, London, pp. 58 - 71." type="book chapter" year="1985">Lambert 1985</bibRefCitation>
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: 66
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</treatmentCitation>
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.
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<emphasis id="B968EADDFF98381E90A8FB55F21BFB5A" box="[151,382,1256,1279]" italics="true" pageId="13" pageNumber="44">
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<taxonomicName id="4C1C4D4CFF98381E90A8FB55F392FB5A" box="[151,247,1256,1279]" class="Reptilia" family="Typhlopidae" genus="Typhlops" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="genus">Typhlops</taxonomicName>
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lineolatus—
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</emphasis>
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(not Jan)
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<treatmentCitation id="0ABD10DEFF98381E91DCFB55F1FCFB5A" author="Lanza" box="[483,665,1256,1279]" page="219" pageId="13" pageNumber="44" year="1983">
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<bibRefCitation id="EF8D4B3EFF98381E91DCFB55F101FB5A" author="Lanza" box="[483,612,1256,1279]" pageId="13" pageNumber="66" refString="Lanza, B. (1983) A list of the Somali amphibians and reptiles. Monitore Zoologico Italiano, Firenze, (N. S.) Supplemento 18 (8), 193 - 247." type="journal article" year="1983">Lanza 1983</bibRefCitation>
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: 219
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</treatmentCitation>
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, 1990: 430.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3066544FF98381E90A8FA8AF71EFA57" pageId="13" pageNumber="44" type="materials_examined">
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<paragraph id="8BA336CFFF98381E90A8FA8AF2B2FA07" blockId="13.[151,1437,1335,2002]" pageId="13" pageNumber="44">
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<emphasis id="B968EADDFF98381E90A8FA8AF275FAF4" bold="true" box="[151,272,1335,1361]" pageId="13" pageNumber="44">
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<typeStatus id="54A7886DFF98381E90A8FA8AF26EFAF4" box="[151,267,1335,1361]" pageId="13" pageNumber="44" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED0DAE0AFF98381E9127FA85F234FAF7" LSID="urn:lsid:biocol.org:col:34700" box="[280,337,1336,1362]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="13" pageNumber="44">CAS</collectionCode>
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<date id="FFA2100FFF98381E9167FA85F2D6FAF7" box="[344,435,1336,1362]" pageId="13" pageNumber="44" value="01-12-15">151201</date>
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, a female from vicinity of Mareri, Juba Sugar Project, west bank of Juba River, Middle Juba region, southern
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<collectingCountry id="F30B765FFF98381E91FDFADDF147FADF" box="[450,546,1376,1402]" name="Somalia" pageId="13" pageNumber="44">Somalia</collectingCountry>
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(
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<geoCoordinate id="EE285008FF98381E920EFADDF1FFFADF" box="[561,666,1376,1402]" direction="north" orientation="latitude" pageId="13" pageNumber="44" precision="925" value="0.5">00°30’N</geoCoordinate>
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,
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<geoCoordinate id="EE285008FF98381E929AFADDF06FFADF" box="[677,778,1376,1402]" direction="east" orientation="longitude" pageId="13" pageNumber="44" precision="925" value="42.766666">42°46’E</geoCoordinate>
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, elevation ca.
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<quantity id="4CE49B2AFF98381E9390FADDF088FADF" box="[943,1005,1376,1402]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="44" unit="m" value="20.0">20 m</quantity>
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), collected by P. M. Johnson,
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<date id="FFA2100FFF98381E9578FADDF26AFA07" pageId="13" pageNumber="44" value="1979-01-25">25 January 1979</date>
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, in a termite hill.
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</paragraph>
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<paragraph id="8BA336CFFF98381E90F8FA12F71EFA57" blockId="13.[151,1437,1335,2002]" pageId="13" pageNumber="44">
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<emphasis id="B968EADDFF98381E90F8FA12F22EFA6C" bold="true" box="[199,331,1455,1481]" pageId="13" pageNumber="44">
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<typeStatus id="54A7886DFF98381E90F8FA12F222FA6C" box="[199,327,1455,1481]" pageId="13" pageNumber="44" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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<collectionCode id="ED0DAE0AFF98381E916BFA0DF2E8FA6F" LSID="urn:lsid:biocol.org:col:34700" box="[340,397,1456,1482]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="13" pageNumber="44">CAS</collectionCode>
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225258-61,
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<collectionCode id="ED0DAE0AFF98381E921AFA0DF112FA6F" box="[549,631,1456,1482]" httpUri="http://grbio.org/cool/mvk4-k796" name="Museo Zoologico La Specola, Universita di Firenze" pageId="13" pageNumber="44">MZUF</collectionCode>
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27179-84, with the same locality data as the
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<typeStatus id="54A7886DFF98381E94AAFA0DF79EFA6F" box="[1173,1275,1456,1482]" pageId="13" pageNumber="44" type="holotype">holotype</typeStatus>
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but collected by L. G. Hoevers and P. M. Johnson,
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<date id="FFA2100FFF98381E9275FA65F1B5FA57" box="[586,720,1496,1522]" pageId="13" pageNumber="44" value="1977" valueMax="1981">1977–1981</date>
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, during bush clearing by bulldozers.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066544FF98381E90F8FA42F737F934" pageId="13" pageNumber="44" type="diagnosis">
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<paragraph id="8BA336CFFF98381E90F8FA42F737F934" blockId="13.[151,1437,1335,2002]" pageId="13" pageNumber="44">
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<emphasis id="B968EADDFF98381E90F8FA42F222F9BC" bold="true" box="[199,327,1535,1561]" pageId="13" pageNumber="44">Diagnosis.</emphasis>
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Apparently most closely related to
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<taxonomicName id="4C1C4D4CFF98381E92CEF9BDF0BFF9BC" box="[753,986,1536,1561]" class="Reptilia" family="Typhlopidae" genus="Letheobia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="species" species="decorosa">
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<emphasis id="B968EADDFF98381E92CEF9BDF0BFF9BC" box="[753,986,1536,1561]" italics="true" pageId="13" pageNumber="44">Letheobia decorosa</emphasis>
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</taxonomicName>
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of
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<collectingCountry id="F30B765FFF98381E943BF9BDF7E4F9BF" box="[1028,1153,1536,1562]" name="Cameroon" pageId="13" pageNumber="44">Cameroon</collectingCountry>
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, but differing in its rostral, which tapers posteriorly to a blunt point and has an obtusely angular keratinized cutting edge (rostral rounded posteriorly and snout rounded in
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<emphasis id="B968EADDFF98381E92B8F9EDF043F9CC" box="[647,806,1616,1641]" italics="true" pageId="13" pageNumber="44">
|
||
<taxonomicName id="4C1C4D4CFF98381E92B8F9EDF070F9CC" box="[647,789,1616,1641]" class="Reptilia" family="Typhlopidae" genus="Letheobia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="species" species="decorosa">L. decorosa</taxonomicName>
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);
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</emphasis>
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its frontal being smaller than a supraocular (larger in
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<emphasis id="B968EADDFF98381E90A8F9C5F256F934" box="[151,307,1656,1681]" italics="true" pageId="13" pageNumber="44">
|
||
<taxonomicName id="4C1C4D4CFF98381E90A8F9C5F246F934" box="[151,291,1656,1681]" class="Reptilia" family="Typhlopidae" genus="Letheobia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="species" species="decorosa">L. decorosa</taxonomicName>
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||
);
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</emphasis>
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and its lower middorsal count, 391–435 vs
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||
<quantity id="4CE49B2AFF98381E930AF9C5F0DFF937" box="[821,954,1656,1682]" metricMagnitude="1" metricUnit="m" metricValue="1.2725399999999998" metricValueMax="1.37668" metricValueMin="1.1684" pageId="13" pageNumber="44" unit="in" value="501.0" valueMax="542.0" valueMin="460.0">460–542 in</quantity>
|
||
<emphasis id="B968EADDFF98381E93FEF9C5F737F934" box="[961,1106,1656,1681]" italics="true" pageId="13" pageNumber="44">
|
||
<taxonomicName id="4C1C4D4CFF98381E93FEF9C5F72BF934" box="[961,1102,1656,1681]" class="Reptilia" family="Typhlopidae" genus="Letheobia" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="44" phylum="Chordata" rank="species" species="decorosa">L. decorosa</taxonomicName>
|
||
.
|
||
</emphasis>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066544FF98381D90F8F922F77FFEA7" lastPageId="14" lastPageNumber="45" pageId="13" pageNumber="44" type="description">
|
||
<paragraph id="8BA336CFFF98381E90F8F922F203F80F" blockId="13.[151,1437,1335,2002]" pageId="13" pageNumber="44">
|
||
<emphasis id="B968EADDFF98381E90F8F922F042F91C" bold="true" box="[199,807,1695,1721]" pageId="13" pageNumber="44">
|
||
Description (
|
||
<typeStatus id="54A7886DFF98381E9159F922F2BDF91C" box="[358,472,1695,1721]" pageId="13" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
variations in parentheses).
|
||
</emphasis>
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||
Snout with an obtusely angular keratinised horizontal edge. Rostral broad and saggitate, terminating in a blunt point; ventrally broad with parallel sides; frontal, supraoculars and interparietal subequal and transverse, supraocular with its lateral apex wedged between preocular and ocular; eye faintly visible beneath preocular;
|
||
<collectingCountry id="F30B765FFF98381E9309F8A5F005F897" box="[822,864,1816,1842]" name="Cyprus" pageId="13" pageNumber="44">SIP</collectingCountry>
|
||
X (N2, P, O, O); scale rows 26-24-22; MD 435 (391–430); L/D ratio 53 (44–57). Length
|
||
<quantity id="4CE49B2AFF98381E92B7F8FDF18BF8FF" box="[648,750,1856,1882]" metricMagnitude="-1" metricUnit="m" metricValue="3.47" pageId="13" pageNumber="44" unit="mm" value="347.0">347 mm</quantity>
|
||
. Colouration – head yellow dorsally and laterally (largely brown in MZUF 27181), otherwise dorsum dark brown, each scale with a pale basal spot, larger laterally, ventrum immaculate.
|
||
</paragraph>
|
||
<paragraph id="8BA336CFFF98381E90F8F80AF756F877" blockId="13.[151,1437,1335,2002]" box="[199,1075,1975,2002]" pageId="13" pageNumber="44">
|
||
<emphasis id="B968EADDFF98381E90F8F80AF265F874" bold="true" box="[199,256,1975,2001]" pageId="13" pageNumber="44">Size.</emphasis>
|
||
Largest specimen (CAS 225258 ––
|
||
<typeStatus id="54A7886DFF98381E9294F805F077F877" box="[683,786,1976,2002]" pageId="13" pageNumber="44" type="paratype">paratype</typeStatus>
|
||
)
|
||
<quantity id="4CE49B2AFF98381E931FF805F0E4F877" box="[800,897,1976,2002]" metricMagnitude="-1" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="44" unit="mm" value="510.0">510 mm</quantity>
|
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in total length.
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</paragraph>
|
||
<paragraph id="8BA336CFFF9B381D90F8FF2AF77FFEA7" blockId="14.[151,1436,151,258]" pageId="14" pageNumber="45">
|
||
<emphasis id="B968EADDFF9B381D90F8FF2AF24BFF14" bold="true" box="[199,302,151,177]" pageId="14" pageNumber="45">Habitat.</emphasis>
|
||
The
|
||
<typeStatus id="54A7886DFF9B381D9154FF25F2FBFF17" box="[363,414,152,178]" pageId="14" pageNumber="45">type</typeStatus>
|
||
series was bulldozed out of termitaria on levees of sandy loam on the west bank of the Juba River. The original vegetation consisted of tall riverine trees with undergrowth of shrubs and creepers.
|
||
<emphasis id="B968EADDFF9B381D90F8FF5AF202FEA4" bold="true" box="[199,359,231,257]" pageId="14" pageNumber="45">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54A7886DFF9B381D924CFF55F1C3FEA7" box="[627,678,232,258]" pageId="14" pageNumber="45">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13272A4AFF9B381D932AFF55F00FFEA7" box="[789,874,232,258]" captionStart="FIGURE 11" captionStartId="30.[151,255,1128,1152]" captionTargetBox="[156,1431,199,1104]" captionTargetId="figure@30.[156,1432,194,1104]" captionTargetPageId="30" captionText="FIGURE 11. Distribution of the north-central African species of Letheobia and L. jubana in north-east Africa (inset, top left)." httpUri="https://zenodo.org/record/177289/files/figure.png" pageId="14" pageNumber="45">Fig. 11</figureCitation>
|
||
, inset top left).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |