353 lines
41 KiB
XML
353 lines
41 KiB
XML
<document id="4E9E4ED6AFF95D3874604004A9A48EA1" ID-CLB-Dataset="5897" ID-DOI="10.11646/zootaxa.4981.3.6" ID-GBIF-Dataset="e602c4fe-7400-4287-b3f5-62fecf81f369" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921413" ID-ZooBank="6416487B-1776-4BA5-AFB5-6D382C1940D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623282432958" checkinUser="plazi" docAuthor="Iwasa, Mitsuhiro" docDate="2021" docId="03B587C9FFB2AF29B09CE83674985977" docLanguage="en" docName="zootaxa.4981.3.6.pdf" docOrigin="Zootaxa 4981 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acerocnema incisa Iwasa 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="534" masterDocId="FF8CFFB1FFB3AF2AB00BEE1A771D5C0E" masterDocTitle="Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera" masterLastPageNumber="553" masterPageNumber="531" pageNumber="532" updateTime="1699009213171" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75524C1D3CC955AF3C80A67CDE33B204">Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera</mods:title>
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<mods:name id="4A8621B5D3CFDC64E8C42C0A78D7C583" type="personal">
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<mods:namePart id="D352B0DA1B13F8EE8987A624FBEF129A">Iwasa, Mitsuhiro</mods:namePart>
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<mods:title id="14F7F509594A8560E4A41AF48075A699">Zootaxa</mods:title>
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<mods:part id="ACE682E28B0435CAC64CCB4C923A21BB">
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<mods:date id="9CD297A8CA7866ABDD4D29F0D11B0742">2021</mods:date>
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<mods:number id="2ABEB5134534937E40CB669332918121">2021-06-08</mods:number>
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<mods:number id="D7E8CB457CD2F23FC120F659B256CE36">4981</mods:number>
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<treatment id="03B587C9FFB2AF29B09CE83674985977" ID-DOI="http://doi.org/10.5281/zenodo.4921741" ID-GBIF-Taxon="183228165" ID-Zenodo-Dep="4921741" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977" httpUri="http://treatment.plazi.org/id/03B587C9FFB2AF29B09CE83674985977" lastPageId="3" lastPageNumber="534" pageId="1" pageNumber="532">
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<subSubSection id="C3066554FFB2AF2BB09CE83676FE5A49" box="[151,483,1580,1607]" pageId="1" pageNumber="532" type="nomenclature">
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<paragraph id="8BA336DFFFB2AF2BB09CE83676FE5A49" blockId="1.[151,635,1580,1679]" box="[151,483,1580,1607]" pageId="1" pageNumber="532">
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<emphasis id="B968EACDFFB2AF2BB09CE83676FE5A49" bold="true" box="[151,483,1580,1607]" pageId="1" pageNumber="532">
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<taxonomicName id="4C1C4D5CFFB2AF2BB09CE83676635A48" ID-CoL="9HYJ8" authority="Iwasa, 2021" authorityName="Iwasa" authorityYear="2021" box="[151,382,1580,1606]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="incisa" status="sp. nov.">
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<emphasis id="B968EACDFFB2AF2BB09CE83676635A48" bold="true" box="[151,382,1580,1606]" italics="true" pageId="1" pageNumber="532">Acerocnema incisa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57B6FFB2AF2BB18EE83776FE5A49" box="[389,483,1581,1607]" pageId="1" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3066554FFB2AF2BB09CE84A75665A81" pageId="1" pageNumber="532" type="description">
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<paragraph id="8BA336DFFFB2AF2BB09CE84A76165A65" blockId="1.[151,635,1580,1679]" box="[151,267,1616,1643]" pageId="1" pageNumber="532">
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(
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<figureCitation id="13272A5AFFB2AF2BB094E84A761F5A65" box="[159,258,1616,1643]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="1" pageNumber="532">Figs 1–6</figureCitation>
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)
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</paragraph>
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<paragraph id="8BA336DFFFB2AF2BB09CE86F75665A81" blockId="1.[151,635,1580,1679]" box="[151,635,1652,1679]" pageId="1" pageNumber="532">[Japanese name: Kiashi-kuchiboso-funbae]</paragraph>
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<subSubSection id="C3066554FFB2AF2BB09CE8A775C55B69" pageId="1" pageNumber="532" type="materials_examined">
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<emphasis id="B968EACDFFB2AF2BB09CE8A776D75AD8" bold="true" box="[151,458,1724,1751]" pageId="1" pageNumber="532">Type material examined.</emphasis>
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<materialsCitation id="3B743C82FFB2AF2BB1DEE8A7769C5AF5" ID-GBIF-Occurrence="3312255304" collectingDate="2014-05-18" collectionCode="OUAVM" collectorName="H. Torikura" country="Japan" county="Inada" location="Tokachi" municipality="Obihiro City" pageId="1" pageNumber="532" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="holotype">
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<typeStatus id="54A7887DFFB2AF2BB1DEE8A7755F5AD9" box="[469,578,1725,1751]" pageId="1" pageNumber="532" type="holotype">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="9D1AFD56FFB2AF2BB246E8A775795AD9" box="[589,612,1725,1751]" pageId="1" pageNumber="532" type="male">♂</specimenCount>
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,
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<collectingCountry id="F30B764FFFB2AF2BB27FE8A775A05AD9" box="[628,701,1725,1751]" name="Japan" pageId="1" pageNumber="532">Japan</collectingCountry>
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:
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,
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<collectingMunicipality id="6BC7ACA5FFB2AF2BB32AE8A674AB5AD8" box="[801,950,1724,1750]" pageId="1" pageNumber="532">Obihiro City</collectingMunicipality>
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,
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<location id="8EC36004FFB2AF2BB3CEE8A673395AD8" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977:8EC36004FFB2AF2BB3CEE8A673395AD8" box="[965,1060,1724,1750]" country="Japan" county="Inada" municipality="Obihiro City" name="Tokachi" pageId="1" pageNumber="532" stateProvince="Hokkaido">Tokachi</location>
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,
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<collectingRegion id="49D8F83DFFB2AF2BB438E8A673B95AD8" box="[1075,1188,1724,1750]" country="Japan" name="Hokkaido" pageId="1" pageNumber="532">Hokkaido</collectingRegion>
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,
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<date id="FFA2101FFFB2AF2BB4BFE8A772205AD9" box="[1204,1341,1725,1751]" pageId="1" pageNumber="532" value="2014-05-18">
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<collectingDate id="EFE6E9F7FFB2AF2BB4BFE8A772205AD9" box="[1204,1341,1725,1751]" pageId="1" pageNumber="532" value="2014-05-18">18. v. 2014</collectingDate>
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</date>
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, leg.
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<collectorName id="26E95309FFB2AF2BB58AE8A777E45AF4" pageId="1" pageNumber="532">H. Torikura</collectorName>
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(
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<materialsCitation id="3B743C82FFB2AF2BB187E8FB760A5B11" ID-GBIF-Occurrence="3312255336" collectingDate="2014-05-15" collectingDateMax="2014-05-25" collectingDateMin="2014-05-15" collectionCode="OUAVM" collectorName="H. Torikura" country="Japan" county="Inada" location="Tokachi" municipality="Obihiro City" pageId="1" pageNumber="532" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Hokkaido" typeStatus="paratype">
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<typeStatus id="54A7887DFFB2AF2BB187E8FB75195AF5" box="[396,516,1761,1787]" pageId="1" pageNumber="532" type="paratype">Paratypes</typeStatus>
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:
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<collectingRegion id="49D8F83DFFB2AF2BB218E8FA75985AF5" box="[531,645,1760,1787]" country="Japan" name="Hokkaido" pageId="1" pageNumber="532">Hokkaido</collectingRegion>
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</emphasis>
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,
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</emphasis>
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,
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<specimenCount id="9D1AFD56FFB2AF2BB2C6E8FB75E55AF5" box="[717,760,1761,1787]" pageId="1" pageNumber="532" type="female">4 ♀</specimenCount>
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, same locality as holotype,
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</date>
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, leg.
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<collectorName id="26E95309FFB2AF2BB51CE8FB72815AF4" box="[1303,1436,1760,1787]" pageId="1" pageNumber="532">H. Torikura</collectorName>
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(
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;
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<materialsCitation id="3B743C82FFB2AF2BB128E91F73A85B11" ID-GBIF-Occurrence="3312255312" box="[291,1205,1796,1823]" collectingDate="2014-05-11" collectionCode="OUAVM" collectorName="H. Torikura" country="Japan" county="Kawanishi" location="Tokachi" municipality="Obihiro City" pageId="1" pageNumber="532" specimenCount="1" specimenCount-male="1" stateProvince="Hokkaido" typeStatus="paratype">
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<specimenCount id="9D1AFD56FFB2AF2BB128E91F76525B11" box="[291,335,1797,1823]" pageId="1" pageNumber="532" type="male">1 ♂</specimenCount>
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<collectingCounty id="62C24E53FFB2AF2BB157E91E76CA5B10" box="[348,471,1796,1822]" pageId="1" pageNumber="532">Kawanishi</collectingCounty>
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<collectingMunicipality id="6BC7ACA5FFB2AF2BB1E9E91E75695B10" box="[482,628,1796,1822]" pageId="1" pageNumber="532">Obihiro City</collectingMunicipality>
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,
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<location id="8EC36004FFB2AF2BB28BE91E75C25B10" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977:8EC36004FFB2AF2BB28BE91E75C25B10" box="[640,735,1796,1822]" country="Japan" county="Kawanishi" municipality="Obihiro City" name="Tokachi" pageId="1" pageNumber="532" stateProvince="Hokkaido">Tokachi</location>
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,
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<collectingDate id="EFE6E9F7FFB2AF2BB2E1E91F74715B11" box="[746,876,1797,1823]" pageId="1" pageNumber="532" value="2014-05-11">11. v. 2014</collectingDate>
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</date>
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, leg.
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<collectorName id="26E95309FFB2AF2BB3A3E91F73305B10" box="[936,1069,1796,1823]" pageId="1" pageNumber="532">H. Torikura</collectorName>
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(
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;
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<materialsCitation id="3B743C82FFB2AF2BB4CBE91E744E5B4D" ID-GBIF-Occurrence="3312255301" collectingDate="2017-05-18" collectingDateMax="2017-05-21" collectingDateMin="2017-05-18" collectionCode="OUAVM" collectorName="H. Torikura" country="Japan" location="Tokachi" municipality="Otofuke Town" pageId="1" pageNumber="532" specimenCount="40" specimenCount-female="34" specimenCount-male="6" stateProvince="Hokkaido" typeStatus="paratype">
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<specimenCount id="9D1AFD56FFB2AF2BB4CBE91E73F05B11" box="[1216,1261,1796,1823]" pageId="1" pageNumber="532" type="male">6 ♂</specimenCount>
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,
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<specimenCount id="9D1AFD56FFB2AF2BB4F2E91F722F5B11" box="[1273,1330,1797,1823]" pageId="1" pageNumber="532" type="female">34 ♀</specimenCount>
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,
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<collectingMunicipality id="6BC7ACA5FFB2AF2BB54BE91E77C65B4D" pageId="1" pageNumber="532">Otofuke Town</collectingMunicipality>
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,
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<location id="8EC36004FFB2AF2BB0E2E93276555B4C" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977:8EC36004FFB2AF2BB0E2E93276555B4C" box="[233,328,1832,1858]" country="Japan" municipality="Otofuke Town" name="Tokachi" pageId="1" pageNumber="532" stateProvince="Hokkaido">Tokachi</location>
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,
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</date>
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, leg.
|
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<collectorName id="26E95309FFB2AF2BB249E93375D45B4C" box="[578,713,1832,1859]" pageId="1" pageNumber="532">H. Torikura</collectorName>
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(
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.
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<emphasis id="B968EACDFFB2AF2BB354E93274A25B4D" bold="true" box="[863,959,1832,1859]" pageId="1" pageNumber="532">
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<materialsCitation id="3B743C82FFB2AF2BB354E93274A55B4D" ID-GBIF-Occurrence="3312255305" box="[863,952,1832,1859]" country="Japan" location="Kyushu" pageId="1" pageNumber="532" specimenCount="1" stateProvince="Hokkaido" typeStatus="paratype">
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<emphasis id="B968EACDFFB2AF2BB354E93274A55B4D" bold="true" box="[863,952,1832,1859]" italics="true" pageId="1" pageNumber="532">
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<location id="8EC36004FFB2AF2BB354E93274A55B4D" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977:8EC36004FFB2AF2BB354E93274A55B4D" box="[863,952,1832,1859]" country="Japan" name="Kyushu" pageId="1" pageNumber="532" stateProvince="Hokkaido">Kyushu</location>
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</emphasis>
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</materialsCitation>
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,
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</emphasis>
|
||
<materialsCitation id="3B743C82FFB2AF2BB3C2E93375C95B69" ID-GBIF-Occurrence="3312255318" collectingDate="2015-04-04" collectionCode="OUAVM" collectorName="D. Kato" country="Japan" county="Ito" location="City" municipality="Kyushu Univ." pageId="1" pageNumber="532" specimenCount="2" specimenCount-female="2" stateProvince="Fukuoka" typeStatus="paratype">
|
||
<specimenCount id="9D1AFD56FFB2AF2BB3C2E93374E85B4D" box="[969,1013,1833,1859]" pageId="1" pageNumber="532" type="female">2 ♀</specimenCount>
|
||
, south of
|
||
<collectingCounty id="62C24E53FFB2AF2BB466E93373915B4D" box="[1133,1164,1833,1859]" pageId="1" pageNumber="532">Ito</collectingCounty>
|
||
campus,
|
||
<collectingMunicipality id="6BC7ACA5FFB2AF2BB4F7E93272805B4C" box="[1276,1437,1832,1858]" pageId="1" pageNumber="532">Kyushu Univ.</collectingMunicipality>
|
||
<collectingRegion id="49D8F83DFFB2AF2BB09CE95677E45B68" box="[151,249,1868,1894]" country="Japan" name="Fukuoka" pageId="1" pageNumber="532">Fukuoka</collectingRegion>
|
||
<location id="8EC36004FFB2AF2BB10BE956762E5B68" LSID="urn:lsid:plazi:treatment:03B587C9FFB2AF29B09CE83674985977:8EC36004FFB2AF2BB10BE956762E5B68" box="[256,307,1868,1894]" country="Japan" county="Ito" municipality="Kyushu Univ." name="City" pageId="1" pageNumber="532" stateProvince="Fukuoka">City</location>
|
||
,
|
||
<date id="FFA2101FFFB2AF2BB135E95776A45B69" box="[318,441,1868,1895]" pageId="1" pageNumber="532" value="2015-04-04">
|
||
<collectingDate id="EFE6E9F7FFB2AF2BB135E95776A45B69" box="[318,441,1868,1895]" pageId="1" pageNumber="532" value="2015-04-04">4. iv. 2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26E95309FFB2AF2BB1FFE95775505B69" box="[500,589,1869,1895]" pageId="1" pageNumber="532">D. Kato</collectorName>
|
||
(
|
||
<collectionCode id="ED0DAE1AFFB2AF2BB257E95775D15B69" box="[604,716,1869,1895]" pageId="1" pageNumber="532">OUAVM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066554FFB2AF2BB0CCE96A74CA5BA0" pageId="1" pageNumber="532" type="diagnosis">
|
||
<paragraph id="8BA336DFFFB2AF2BB0CCE96A74CA5BA0" blockId="1.[151,1437,1724,1973]" pageId="1" pageNumber="532">
|
||
<emphasis id="B968EACDFFB2AF2BB0CCE96A765F5B84" bold="true" box="[199,322,1904,1930]" pageId="1" pageNumber="532">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C1C4D5CFFB2AF2BB143E96B75035B84" authority="Iwasa, 2021" authorityName="Iwasa" authorityYear="2021" box="[328,542,1905,1931]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="532" phylum="Arthropoda" rank="species" species="incisa" status="sp. nov.">
|
||
<emphasis id="B968EACDFFB2AF2BB143E96B75035B84" box="[328,542,1905,1931]" italics="true" pageId="1" pageNumber="532">Acerocnema incisa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EACDFFB2AF2BB22FE96B75625B85" bold="true" box="[548,639,1905,1931]" pageId="1" pageNumber="532">
|
||
<taxonomicNameLabel id="A25B57B6FFB2AF2BB22FE96B75625B85" box="[548,639,1905,1931]" pageId="1" pageNumber="532" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be recognized by thorax with grey microtrichose, dorsal setulae on apical portion of vein R
|
||
<subScript id="1798349AFFB2AF2BB1AAE9BD76B45BBB" attach="left" box="[417,425,1959,1973]" fontSize="6" pageId="1" pageNumber="532">1</subScript>
|
||
of wing, and male surstylus anterodistally dilated
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF636657FFB1AF28B09CE94873EF5BF4" ID-DOI="http://doi.org/10.5281/zenodo.4921415" ID-Zenodo-Dep="4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="2" pageNumber="533" startId="2.[151,264,1874,1898]" targetBox="[235,1346,195,1826]" targetPageId="2">
|
||
<paragraph id="8BA336DFFFB1AF28B09CE94873EF5BF4" blockId="2.[151,1437,1874,2043]" pageId="2" pageNumber="533">
|
||
<emphasis id="B968EACDFFB1AF28B09CE94876275B64" bold="true" box="[151,314,1874,1898]" pageId="2" pageNumber="533">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C1C4D5CFFB1AF28B14BE949751C5B64" authority="Iwasa, 2021" authorityName="Iwasa" authorityYear="2021" box="[320,513,1875,1898]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="incisa" status="sp. nov.">
|
||
<emphasis id="B968EACDFFB1AF28B14BE949751C5B64" box="[320,513,1875,1898]" italics="true" pageId="2" pageNumber="533">Acerocnema incisa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B968EACDFFB1AF28B20DE948756D5B64" bold="true" box="[518,624,1874,1898]" pageId="2" pageNumber="533">
|
||
<taxonomicNameLabel id="A25B57B6FFB1AF28B20DE948754B5B64" box="[518,598,1874,1898]" pageId="2" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1,
|
||
</emphasis>
|
||
Male head and thorax, dorsal view (holotype);
|
||
<emphasis id="B968EACDFFB1AF28B444E948737C5B64" bold="true" box="[1103,1121,1874,1898]" pageId="2" pageNumber="533">2,</emphasis>
|
||
Male head, left lateral view;
|
||
<emphasis id="B968EACDFFB1AF28B582E948728B5B64" bold="true" box="[1417,1430,1874,1898]" pageId="2" pageNumber="533">3</emphasis>
|
||
, Male sternite V, ventral view;
|
||
<emphasis id="B968EACDFFB1AF28B1C3E96C76C75B80" bold="true" box="[456,474,1910,1934]" pageId="2" pageNumber="533">4,</emphasis>
|
||
Male terminalia, left lateral view;
|
||
<emphasis id="B968EACDFFB1AF28B332E96C74505B80" bold="true" box="[825,845,1910,1934]" pageId="2" pageNumber="533">5,</emphasis>
|
||
Male terminalia, posterior view;
|
||
<emphasis id="B968EACDFFB1AF28B495E96C73AC5B80" bold="true" box="[1182,1201,1910,1934]" pageId="2" pageNumber="533">6,</emphasis>
|
||
Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3066554FFB0AF29B0CCEE8C7431581F" pageId="3" pageNumber="534" type="description">
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEE8C75AA5F53" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEE8C76F55CBF" bold="true" box="[199,488,150,177]" pageId="3" pageNumber="534">Description. Male. Head</emphasis>
|
||
(
|
||
<figureCitation id="13272A5AFFB0AF29B1FEEE8D75455CBF" box="[501,600,151,177]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Figs 1, 2</figureCitation>
|
||
): eyes reddish-brown; frons reddish; ocellar triangle black; face, parafacial, and gena yellow; occiput black and shining; postpedicel black and enlarged, approximately 2.5 times long as wide; arista basally pubescent and apically short-haired, approximately same length as postpedicel; palpi yellow and clavate; proboscis slender and sclerotized with reduced labella; vibrissal angle with a strong yellow vibrissa. Chaetotaxy (black): 3-4
|
||
<emphasis id="B968EACDFFB0AF29B152EF3D76715D4F" box="[345,364,295,321]" italics="true" pageId="3" pageNumber="534">fr</emphasis>
|
||
, 3
|
||
<emphasis id="B968EACDFFB0AF29B184EF3D76B55D4E" box="[399,424,295,320]" italics="true" pageId="3" pageNumber="534">or</emphasis>
|
||
(1 proclinate, 1 lateroclinate, 1 latero-reclinate), 1
|
||
<emphasis id="B968EACDFFB0AF29B3EEEF3D74E25D4E" box="[997,1023,295,320]" italics="true" pageId="3" pageNumber="534">oc</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B429EF3D73575D4E" box="[1058,1098,295,320]" italics="true" pageId="3" pageNumber="534">poc</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B465EF3D73975D4E" box="[1134,1162,295,320]" italics="true" pageId="3" pageNumber="534">vti</emphasis>
|
||
(strong), 1
|
||
<emphasis id="B968EACDFFB0AF29B506EF3D72335D4E" box="[1293,1326,295,320]" italics="true" pageId="3" pageNumber="534">vte</emphasis>
|
||
.
|
||
<emphasis id="B968EACDFFB0AF29B537EF3D72885D4F" bold="true" box="[1340,1429,295,321]" pageId="3" pageNumber="534">Thorax</emphasis>
|
||
: wholly black; prescutum and scutum thinly grey microtrichose, with two longitudinally shiny stripes on scutum and with inconspicuous hairs (
|
||
<figureCitation id="13272A5AFFB0AF29B1F1EF7575225D87" box="[506,575,367,393]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Fig. 1</figureCitation>
|
||
); scutellum shining; all thoracic pleura grey microtrichose; proepisternum anteriorly with hairs; anepisternum posteriorly hairy; katepisternum sparsely with long hairs; anepimeron, meron, and katatergite bare; metanotum shining in lower part. Chaetotaxy (yellow): 2
|
||
<emphasis id="B968EACDFFB0AF29B400EFAD735D5DDE" box="[1035,1088,439,464]" italics="true" pageId="3" pageNumber="534">pprn</emphasis>
|
||
, 2
|
||
<emphasis id="B968EACDFFB0AF29B468EFAD739A5DDF" box="[1123,1159,439,465]" italics="true" pageId="3" pageNumber="534">npl</emphasis>
|
||
, 1+1
|
||
<emphasis id="B968EACDFFB0AF29B4C3EFAD73FB5DDF" box="[1224,1254,439,465]" italics="true" pageId="3" pageNumber="534">ial</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B501EFAD723E5DDE" box="[1290,1315,439,464]" italics="true" pageId="3" pageNumber="534">sa</emphasis>
|
||
, 2
|
||
<emphasis id="B968EACDFFB0AF29B54DEFAD727F5DDE" box="[1350,1378,439,464]" italics="true" pageId="3" pageNumber="534">pa</emphasis>
|
||
, 1+3
|
||
<emphasis id="B968EACDFFB0AF29B09CEFC177AC5DFB" box="[151,177,475,501]" italics="true" pageId="3" pageNumber="534">dc</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B0DCEFC177E15DFB" box="[215,252,475,501]" italics="true" pageId="3" pageNumber="534">bsc</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B12AEFC176495DFA" box="[289,340,475,500]" italics="true" pageId="3" pageNumber="534">apsc</emphasis>
|
||
,1
|
||
<emphasis id="B968EACDFFB0AF29B179EFC176AA5DFA" box="[370,439,475,500]" italics="true" pageId="3" pageNumber="534">prepst</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B1D7EFC1753E5DFA" box="[476,547,475,500]" italics="true" pageId="3" pageNumber="534">prepm</emphasis>
|
||
, 6-7
|
||
<emphasis id="B968EACDFFB0AF29B254EFC175B55DFA" box="[607,680,475,500]" italics="true" pageId="3" pageNumber="534">anepst</emphasis>
|
||
, 1
|
||
<emphasis id="B968EACDFFB0AF29B2C6EFC1741A5DFB" box="[717,775,475,501]" italics="true" pageId="3" pageNumber="534">kepst</emphasis>
|
||
. Wings: hyaline, slightly tinged with brown; veins brown; R
|
||
<subScript id="1798349AFFB0AF29B0A1EC0B77AF5E11" attach="left" box="[170,178,529,543]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
with dorsal setulae on apical portion; veins R
|
||
<subScript id="1798349AFFB0AF29B2CEEC0B75C25E11" attach="left" box="[709,735,529,543]" fontSize="6" pageId="3" pageNumber="534">4+5</subScript>
|
||
and M
|
||
<subScript id="1798349AFFB0AF29B339EC0B74275E11" attach="left" box="[818,826,529,543]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
almost parallel; vein A
|
||
<subScript id="1798349AFFB0AF29B442EC0B734C5E11" attach="right" box="[1097,1105,529,543]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
+CuA
|
||
<subScript id="1798349AFFB0AF29B49EEC0B73805E11" attach="left" box="[1173,1181,529,543]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
disappear well before wing margin; calypter yellow, with hairs marginally; halteres yellow. Legs: entirely yellow, covered with hair-like setulae; fore femur with several
|
||
<emphasis id="B968EACDFFB0AF29B20BEC5D75015E6F" box="[512,540,583,609]" italics="true" pageId="3" pageNumber="534">pd</emphasis>
|
||
setae distally; mid and hind femora with preapical setae; fore tibiae with 1
|
||
<emphasis id="B968EACDFFB0AF29B56EEC5D729C5E6F" box="[1381,1409,583,609]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
at two-fifths; mid tibia 1
|
||
<emphasis id="B968EACDFFB0AF29B19CEC7176AE5E8B" box="[407,435,619,645]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
at near half and subapical setae; hind tibia with 2
|
||
<emphasis id="B968EACDFFB0AF29B3E0EC71731A5E8B" box="[1003,1031,619,645]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
at one-third and half, and subapical setae; all tarsi yellow; fifth tarsomere somewhat compressed.
|
||
<emphasis id="B968EACDFFB0AF29B358EC9474CC5EA6" bold="true" box="[851,977,654,680]" pageId="3" pageNumber="534">Abdomen:</emphasis>
|
||
tergites black and shining, covered with short hairs; sternites II to IV longitudinally rectangular, sparsely covered with hair-like setulae; sternite V (
|
||
<figureCitation id="13272A5AFFB0AF29B55BECA9728E5EC3" box="[1360,1427,691,717]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Fig. 3</figureCitation>
|
||
) deeply bilobate posteriorly; epandrium (
|
||
<figureCitation id="13272A5AFFB0AF29B256ECCD75BC5EFF" box="[605,673,727,753]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Fig. 4</figureCitation>
|
||
) developed and sparsely with hairs but without setae; surstyli (
|
||
<figureCitation id="13272A5AFFB0AF29B561ECCD77DD5F1B" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Figs 4, 5</figureCitation>
|
||
) long, slender, curved and dilated antero-apically; aedeagus (
|
||
<figureCitation id="13272A5AFFB0AF29B360ECE174B05F1B" box="[875,941,763,789]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Fig. 4</figureCitation>
|
||
) slender, apically swollen and membranous; epiphallus flattened, curved and pointed apically; parameres rod-shaped; hypandrium large, thick, and V-shaped; cerci stout and membranous, covered with hairs.
|
||
</paragraph>
|
||
<paragraph id="8BA336DFFFB0AF29B0CCED7D732D5FE3" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCED7D76385F8F" bold="true" box="[199,293,871,897]" pageId="3" pageNumber="534">Female.</emphasis>
|
||
Basal and ventral portions of postpedicel, postpronotum and proepisternum paler than those of male; abdominal tergites II to VI centrally black and laterally yellow to light brown; sternites III to VI longitudinally elongate; segments (tergites and sternites) VII to VIII (
|
||
<figureCitation id="13272A5AFFB0AF29B2DCEDB574015FC4" box="[727,796,943,970]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1874,1898]" captionTargetBox="[235,1346,195,1826]" captionTargetId="figure-19@2.[225,1363,179,1850]" captionTargetPageId="2" captionText="FIGURES 1–6. Acerocnema incisa sp. nov. 1, Male head and thorax, dorsal view (holotype); 2, Male head, left lateral view; 3, Male sternite V, ventral view; 4, Male terminalia, left lateral view; 5, Male terminalia, posterior view; 6, Female terminalia, left lateral view. Abbreviations: Head—ar, arista; fr, frontal setae; oc, ocellar setae; or, orbital setae; pal, palpus; ppd, postpedicel; prb, proboscis; vte, outer vertical setae; vti, inner vertical setae; vib, vibrissa. Abdomen - aed, aedeagus; cer, cercus; ep, epandrium; epiph, epiphallus; epipr, epiproct; hypr, hypoproct; pm, paramere; st, sternite; sur, surstylus; tg, tergite." figureDoi="http://doi.org/10.5281/zenodo.4921415" httpUri="https://zenodo.org/record/4921415/files/figure.png" pageId="3" pageNumber="534">Fig. 6</figureCitation>
|
||
) compressed and tergite VIII strongly sclerotized; cerci, epiproct, and hypoproct weakly pigmented. Other characteristics as those of male.
|
||
</paragraph>
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEDED7431581F" blockId="3.[151,1437,150,1401]" box="[199,812,1015,1041]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEDED7600581F" bold="true" box="[199,285,1015,1041]" pageId="3" pageNumber="534">Length</emphasis>
|
||
(
|
||
<specimenCount id="9D1AFD56FFB0AF29B125EDED765A581F" box="[302,327,1015,1041]" pageId="3" pageNumber="534" type="male">♂</specimenCount>
|
||
<specimenCount id="9D1AFD56FFB0AF29B14CEDED7645581F" box="[327,344,1015,1041]" pageId="3" pageNumber="534" type="female">♀</specimenCount>
|
||
). Body, 4.0–4.2 mm; wings, 4.0–4.3 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066554FFB0AF29B0CCEA0075D95838" box="[199,708,1050,1078]" pageId="3" pageNumber="534" type="distribution">
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEA0075D95838" blockId="3.[151,1437,150,1401]" box="[199,708,1050,1078]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEA007644583A" bold="true" box="[199,345,1050,1076]" pageId="3" pageNumber="534">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F30B764FFFB0AF29B16CEA0176BA583B" box="[359,423,1051,1077]" name="Japan" pageId="3" pageNumber="534">Japan</collectingCountry>
|
||
(
|
||
<collectingRegion id="49D8F83DFFB0AF29B1BDEA01753B583B" box="[438,550,1051,1077]" country="Japan" name="Hokkaido" pageId="3" pageNumber="534">Hokkaido</collectingRegion>
|
||
and Kyushu).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066554FFB0AF29B0CCEA25769B5873" pageId="3" pageNumber="534" type="etymology">
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEA25769B5873" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEA2576555857" bold="true" box="[199,328,1087,1113]" pageId="3" pageNumber="534">Etymology</emphasis>
|
||
. This new species is named after the scientific name of the flower species which has a close relationship with this fly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3066554FFB0AF29B0CCEA9D74985977" pageId="3" pageNumber="534" type="discussion">
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEA9D7521593F" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEA9D762658AF" bold="true" box="[199,315,1159,1185]" pageId="3" pageNumber="534">Remarks.</emphasis>
|
||
This new species is similar to
|
||
<taxonomicName id="4C1C4D5CFFB0AF29B284EA9D746158AF" authority="Iwasa" authorityName="Iwasa" box="[655,892,1159,1185]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="kishimotoae">
|
||
<emphasis id="B968EACDFFB0AF29B284EA9D742958AF" box="[655,820,1159,1185]" italics="true" pageId="3" pageNumber="534">A. kishimotoae</emphasis>
|
||
Iwasa
|
||
</taxonomicName>
|
||
in general appearance, but differs from it in having the following characteristics: frons reddish; scutum grey microtrichose with inconspicuous hairs; male surstyli anterodistally dilated; female postpedicel same length as that of male. In
|
||
<taxonomicName id="4C1C4D5CFFB0AF29B3C0EAD5736C58E7" box="[971,1137,1231,1257]" class="Insecta" family="Scathophagidae" genus="Acerocnema" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="kishimotoae">
|
||
<emphasis id="B968EACDFFB0AF29B3C0EAD5736C58E7" box="[971,1137,1231,1257]" italics="true" pageId="3" pageNumber="534">A. kishimotoae</emphasis>
|
||
</taxonomicName>
|
||
, frons black; scutum shining, sparsely covered with distinct hairs; male surstyli gradually dilated toward the tip, not anterodistally; female postpedicel smaller than that of male.
|
||
</paragraph>
|
||
<paragraph id="8BA336DFFFB0AF29B0CCEB2074985977" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="534">
|
||
<emphasis id="B968EACDFFB0AF29B0CCEB20765E595A" bold="true" box="[199,323,1338,1364]" pageId="3" pageNumber="534">Bionomics</emphasis>
|
||
. Adults were collected from a flower community of
|
||
<taxonomicName id="4C1C4D5CFFB0AF29B39AEB2173F0595B" authority="(Thumb.) Per." authorityName="Per." baseAuthorityName="Thumb." box="[913,1261,1339,1366]" class="Magnoliopsida" family="Papaveraceae" genus="Corydalis" kingdom="Plantae" order="Ranunculales" pageId="3" pageNumber="534" phylum="Tracheophyta" rank="species" species="incisa">
|
||
<emphasis id="B968EACDFFB0AF29B39AEB217354595A" box="[913,1097,1339,1365]" italics="true" pageId="3" pageNumber="534">Corydalis incisa</emphasis>
|
||
(Thumb.) Per.
|
||
</taxonomicName>
|
||
in early spring. The larvae probably penetrate into the seed capsules of this flower.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |